treatments-xml/data/32/73/E8/3273E86FFFD7FFA17F9EFB3C86080DA6.xml
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<document id="BEA6ADFDD5C933C009BD91D1C094C2CF" ID-CLB-Dataset="2602" ID-DOI="10.11646/zootaxa.5091.2.8" ID-GBIF-Dataset="30952597-d5c2-4b06-8f7c-005070536131" ID-ISSN="1175-5326" ID-Zenodo-Dep="5843827" ID-ZooBank="E7B06F66-AB66-43DB-9F95-AFBA71088446" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642061910504" checkinUser="plazi" docAuthor="Liu, Tian-Tian, Nozaki, Tsubasa, Yamamoto, Shûhei &amp; Maruyama, Munetoshi" docDate="2022" docId="3273E86FFFD7FFA17F9EFB3C86080DA6" docLanguage="en" docName="zootaxa.5091.2.8.pdf" docOrigin="Zootaxa 5091 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Myllaena japonica &quot; Sharp 1888" docType="treatment" docVersion="8" lastPageNumber="375" masterDocId="CE4A9017FFD6FFA37F09FF9285270F57" masterDocTitle="Synopsis of the Japanese species of Aleocharinae (Coleoptera: Staphylinidae), with review of the type specimens II. Genus Myllaena Erichson of tribe Myllaenini with redescription of four Japanese species" masterLastPageNumber="382" masterPageNumber="373" pageNumber="374" updateTime="1699302085872" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B4E9E466507AC8F23CE60B2B1D212CB7">Synopsis of the Japanese species of Aleocharinae (Coleoptera: Staphylinidae), with review of the type specimens II. Genus Myllaena Erichson of tribe Myllaenini with redescription of four Japanese species</mods:title>
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<mods:namePart id="1BFBF250E51A23C29180482DEE050539">Liu, Tian-Tian</mods:namePart>
<mods:nameIdentifier id="38C47B414BA7CC0FD461AF696A6B609D" type="ORCID">0000-0002-1971-7883</mods:nameIdentifier>
<mods:affiliation id="7A10E0EED4AE8E965BC549A931E9A6E1">The Kyushu University Museum, Fukuoka 812 - 8581, Japan &amp; 13131203225 @ 163. com; https: // orcid. org / 0000 - 0002 - 1971 - 7883</mods:affiliation>
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<mods:namePart id="257A5D8341D167896FDFD33B327A7170">Nozaki, Tsubasa</mods:namePart>
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<mods:affiliation id="06FA5258C7BB94B72D1388FB281B6DE1">The Kyushu University Museum, Fukuoka 812 - 8581, Japan &amp; 1 nozakki. 4 ori. 27 arthropoda. 256 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2419 - 4968</mods:affiliation>
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<mods:namePart id="BD83FACD1C20A66E377D9C026A3CF6F5">Yamamoto, Shûhei</mods:namePart>
<mods:affiliation id="B6AFEE06B28833A375C7C94400A51E84">The Hokkaido University Museum, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="754FA5271C6037529D7A18D2584F151C">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="E694D2BF2751B860F4C648725BAEA4C4">The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
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<mods:date id="24D2C852CDEF36485FFB5AAAD747D968">2022</mods:date>
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<treatment id="3273E86FFFD7FFA17F9EFB3C86080DA6" ID-DOI="http://doi.org/10.5281/zenodo.5844057" ID-GBIF-Taxon="192084985" ID-Zenodo-Dep="5844057" LSID="urn:lsid:plazi:treatment:3273E86FFFD7FFA17F9EFB3C86080DA6" httpUri="http://treatment.plazi.org/id/3273E86FFFD7FFA17F9EFB3C86080DA6" lastPageId="2" lastPageNumber="375" pageId="1" pageNumber="374">
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<paragraph id="BA655979FFD7FFA27F9EFB3C87300B9F" blockId="1.[151,588,1198,1297]" box="[151,535,1198,1225]" pageId="1" pageNumber="374">
<heading id="E12DEE15FFD7FFA27F9EFB3C87300B9F" bold="true" box="[151,535,1198,1225]" fontSize="11" level="1" pageId="1" pageNumber="374" reason="1">
<taxonomicName id="7DDA22FAFFD7FFA27F9EFB3C87300B9F" ID-CoL="743Q8" authority="Sharp, 1888" authorityName="&quot; Sharp" authorityYear="1888" box="[151,535,1198,1225]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="88AE856BFFD7FFA27F9EFB3C87300B9F" bold="true" box="[151,535,1198,1225]" pageId="1" pageNumber="374">
<emphasis id="88AE856BFFD7FFA27F9EFB3C845E0B9E" bold="true" box="[151,377,1198,1225]" italics="true" pageId="1" pageNumber="374">Myllaena japonica</emphasis>
<bibRefCitation id="DE4B2488FFD7FFA27E89FB3C87300B9F" author="Sharp, D." box="[384,535,1198,1224]" pageId="1" pageNumber="374" pagination="369 - 387" refId="ref4775" refString="Sharp, D. (1888) The Staphylinidae of Japan. Annals and Magazine of Natural History, 2 (6), 277 - 295, 369 - 387. https: // doi. org / 10.1080 / 00222938809460928" type="journal article" year="1888">Sharp, 1888</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="BA655979FFD7FFA27F9EFB41876B0BBA" blockId="1.[151,588,1198,1297]" box="[151,588,1234,1261]" pageId="1" pageNumber="374">[Japanese name: Migiwa-hanekakushi]</paragraph>
</subSubSection>
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<paragraph id="BA655979FFD7FFA27F9EFB65847F0A46" blockId="1.[151,588,1198,1297]" box="[151,344,1270,1297]" pageId="1" pageNumber="374">(Figs. 1, 58, 18)</paragraph>
</subSubSection>
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<treatmentCitation id="3B7B7F68FFD7FFA27F9EFAA9872F0A04" author="Sharp, D." box="[151,520,1339,1363]" page="377" pageId="1" pageNumber="374" year="1888">
<taxonomicName id="7DDA22FAFFD7FFA27F9EFAA9872F0A04" ID-CoL="743Q8" authority="Sharp, 1888: 377" authorityName="&quot; Sharp" authorityPageNumber="377" authorityYear="1888" box="[151,520,1339,1363]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="88AE856BFFD7FFA27F9EFAA984710A04" box="[151,342,1339,1363]" italics="true" pageId="1" pageNumber="374">Myllaena japonica</emphasis>
<bibRefCitation id="DE4B2488FFD7FFA27E52FAA9872F0A04" author="Sharp, D." box="[347,520,1339,1363]" pageId="1" pageNumber="374" pagination="369 - 387" refId="ref4775" refString="Sharp, D. (1888) The Staphylinidae of Japan. Annals and Magazine of Natural History, 2 (6), 277 - 295, 369 - 387. https: // doi. org / 10.1080 / 00222938809460928" type="journal article" year="1888">Sharp, 1888: 377</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="6561E7DBFFD7FFA27DE0FAAE86320A03" box="[745,789,1340,1364]" pageId="1" pageNumber="374">type</typeStatus>
locality:
<collectingCountry id="C2CD19E9FFD7FFA27C7AFAAE86880A04" box="[883,943,1340,1363]" name="Japan" pageId="1" pageNumber="374">Japan</collectingCountry>
, “Miyanoshita” [Hakone-machi, Kanagawa-ken, eastern Honshû]); Bernhauer &amp; Scheerpeltz 1926: 507 (catalogue);
<treatmentCitation id="3B7B7F68FFD7FFA27C8CFAC886D80A25" author="Li, J." box="[901,1023,1370,1394]" page="55" pageId="1" pageNumber="374" year="1992">
<bibRefCitation id="DE4B2488FFD7FFA27C8CFAC886D80A25" author="Li, J." box="[901,1023,1370,1394]" pageId="1" pageNumber="374" refId="ref4394" refString="Li, J. (1992) The Coleoptera fauna of Northeast China. Jilin Education Publishing House, Jinlin, 208 pp. (In Chinese, with English title.)" type="book" year="1992">Li 1992: 55</bibRefCitation>
</treatmentCitation>
(record from
<collectingCountry id="C2CD19E9FFD7FFA27B9DFAC881F20A25" box="[1172,1237,1370,1394]" name="China" pageId="1" pageNumber="374">China</collectingCountry>
);
<treatmentCitation id="3B7B7F68FFD7FFA27BE1FAC880BB0A25" author="Pasnik, G." box="[1256,1436,1370,1394]" page="189" pageId="1" pageNumber="374" year="2001">
<bibRefCitation id="DE4B2488FFD7FFA27BE1FAC880BB0A25" author="Pasnik, G." box="[1256,1436,1370,1394]" pageId="1" pageNumber="374" pagination="185 - 234" refId="ref4677" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44 (3), 185 - 234." type="journal article" year="2001">Paśnik 2001: 189</bibRefCitation>
</treatmentCitation>
(record from
<collectingCountry id="C2CD19E9FFD7FFA27E47FAEA84F50AC6" box="[334,466,1400,1425]" name="North Korea" pageId="1" pageNumber="374">North Korea</collectingCountry>
);
<treatmentCitation id="3B7B7F68FFD7FFA27EEAFAEB87800AC7" author="Smetana, A." box="[483,679,1400,1425]" page="465" pageId="1" pageNumber="374" year="2004">
<bibRefCitation id="DE4B2488FFD7FFA27EEAFAEB87800AC7" author="Smetana, A." box="[483,679,1400,1425]" pageId="1" pageNumber="374" refId="ref4957" refString="Smetana, A. (2004) Subfamily Aleocharinae Fleming, 1821. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Histeroidea, Staphylinoidea. Apollo Books, Stenstrup, pp. 353 - 494." type="book" year="2004">Smetana 2004: 465</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="3B7B7F68FFD7FFA27C24FAEA81450AC7" author="Schulke, M. &amp; Smetana, A." box="[813,1122,1400,1425]" page="678" pageId="1" pageNumber="374" year="2015">
<bibRefCitation id="DE4B2488FFD7FFA27C24FAEA81450AC7" author="Schulke, M. &amp; Smetana, A." box="[813,1122,1400,1425]" pageId="1" pageNumber="374" pagination="304 - 1134" refId="ref4711" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 - 2. Revised and updated edition. Hydrophiloidea Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana 2015: 678</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F2C00AF2FFD7FFA27F9EFA2D864D09E6" pageId="1" pageNumber="374" type="materials_examined">
<paragraph id="BA655979FFD7FFA27F9EFA2D87560912" blockId="1.[151,1437,1470,2037]" pageId="1" pageNumber="374">
<emphasis id="88AE856BFFD7FFA27F9EFA2D84630A8F" bold="true" box="[151,324,1470,1497]" pageId="1" pageNumber="374">Type material.</emphasis>
<materialsCitation id="0AB25324FFD7FFA27E42FA2D87560912" ID-GBIF-Occurrence="3436476307" collectingDate="1880-05" collectionCode="MM, BMNH" collectorName="G. Lewis." country="Japan" county="Maruyama" location="Lewis" municipality="Sharp" pageId="1" pageNumber="374" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="6561E7DBFFD7FFA27E42FA2D84990A8E" box="[331,446,1471,1497]" pageId="1" pageNumber="374" type="lectotype">Lectotype</typeStatus>
, here designated,
<specimenCount id="ACDC92F0FFD7FFA27D8DFA2D87BC0A8E" box="[644,667,1471,1497]" pageId="1" pageNumber="374" type="male"></specimenCount>
, “Type [RRL]//
<taxonomicName id="7DDA22FAFFD7FFA27C4EFA2C813F0A8E" authorityName="&quot; Sharp" authorityYear="1888" box="[839,1048,1470,1497]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="japonica">Myllaena japonica</taxonomicName>
Type/D.S./Miyanoshita/
<date id="CE647FB9FFD7FFA27A23FA2D80BB0A8E" box="[1322,1436,1471,1497]" pageId="1" pageNumber="374" value="1880-05">
<collectingDate id="DE208651FFD7FFA27A23FA2D80BB0A8E" box="[1322,1436,1471,1497]" pageId="1" pageNumber="374" value="1880-05">May 1880</collectingDate>
</date>
<location id="BF050FA2FFD7FFA27F9EFA7185FC0AAA" LSID="urn:lsid:plazi:treatment:3273E86FFFD7FFA17F9EFB3C86080DA6:BF050FA2FFD7FFA27F9EFA7185FC0AAA" box="[151,219,1507,1533]" country="Japan" county="Maruyama" municipality="Sharp" name="Lewis" pageId="1" pageNumber="374">Lewis</location>
//
<location id="BF050FA2FFD7FFA27FE4FA71845B0AAA" LSID="urn:lsid:plazi:treatment:3273E86FFFD7FFA17F9EFB3C86080DA6:BF050FA2FFD7FFA27FE4FA71845B0AAA" box="[237,380,1507,1533]" country="Japan" county="Maruyama" municipality="Sharp" name="Miyanoshita" pageId="1" pageNumber="374">Miyanoshita</location>
//
<collectingCountry id="C2CD19E9FFD7FFA27E84FA7184E90AAA" box="[397,462,1507,1533]" name="Japan" pageId="1" pageNumber="374">Japan</collectingCountry>
./
<collectorName id="172F3CAFFFD7FFA27ED0FA7187760AAA" box="[473,593,1507,1533]" pageId="1" pageNumber="374">G. Lewis.</collectorName>
//Sharp Coll./1905
<emphasis id="88AE856BFFD7FFA27C3BFA7186670AAA" bold="true" box="[818,832,1507,1533]" pageId="1" pageNumber="374"></emphasis>
313.//
<typeStatus id="6561E7DBFFD7FFA27C74FA7186C80AAA" box="[893,1007,1507,1533]" pageId="1" pageNumber="374" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="7DDA22FAFFD7FFA27CFEFA7181470AAA" authorityName="Erichson" authorityYear="1837" box="[1015,1120,1507,1533]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="genus">Myllaena</taxonomicName>
/japonica/
<collectingMunicipality id="5A01C303FFD7FFA27BDDFA7180320AAA" box="[1236,1301,1507,1533]" pageId="1" pageNumber="374">Sharp</collectingMunicipality>
, 1874/des.
<collectingCounty id="530421F5FFD7FFA27F9EF99584340976" box="[151,275,1543,1569]" pageId="1" pageNumber="374">Maruyama</collectingCounty>
, 2011[RPC]” (considerably broken, probably by DS; abdominal segments VIII
<emphasis id="88AE856BFFD7FFA27B93F995818F0976" bold="true" box="[1178,1192,1543,1569]" pageId="1" pageNumber="374"></emphasis>
X and aedeagus were dissected and mounted by
<collectionCode id="DCCBC1BCFFD7FFA27EC9F9B984D30912" box="[448,500,1579,1605]" country="France" name="University of Montpellier" pageId="1" pageNumber="374">MM</collectionCode>
) (
<collectionCode id="DCCBC1BCFFD7FFA27D03F9B987430912" box="[522,612,1579,1605]" pageId="1" pageNumber="374">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="BA655979FFD7FFA27FCEF9DD864D09E6" blockId="1.[151,1437,1470,2037]" pageId="1" pageNumber="374">
<emphasis id="88AE856BFFD7FFA27FCEF9DD8712093F" bold="true" box="[199,565,1614,1641]" pageId="1" pageNumber="374">Non-type specimens examined.</emphasis>
[
<collectingCountry id="C2CD19E9FFD7FFA27D4BF9DC87AC093E" box="[578,651,1614,1641]" name="Japan" pageId="1" pageNumber="374">Japan</collectingCountry>
]:
<emphasis id="88AE856BFFD7FFA27D92F9DC8625093E" bold="true" box="[667,770,1614,1641]" pageId="1" pageNumber="374">Honshû:</emphasis>
<materialsCitation id="0AB25324FFD7FFA27C01F9DD877D09DA" ID-GBIF-Occurrence="3436476309" collectingDate="1996-03-16" collectionCode="KUM" collectorName=" M. Maruyama" pageId="1" pageNumber="374" specimenCount="10" specimenCount-female="4" specimenCount-male="6">
<specimenCount id="ACDC92F0FFD7FFA27C01F9DD8614093E" box="[776,819,1615,1641]" pageId="1" pageNumber="374" type="male">6 ♂</specimenCount>
,
<specimenCount id="ACDC92F0FFD7FFA27C37F9DD8640093E" box="[830,871,1615,1641]" pageId="1" pageNumber="374" type="female">4 ♀</specimenCount>
, 61 unsexed,
<collectorName id="172F3CAFFFD7FFA27CF4F9DD81A1093E" box="[1021,1158,1615,1641]" pageId="1" pageNumber="374">Amagi-tôge</collectorName>
,
<collectorName id="172F3CAFFFD7FFA27B99F9DD8007093E" box="[1168,1312,1615,1641]" pageId="1" pageNumber="374">Kawazu-chô</collectorName>
,
<collectorName id="172F3CAFFFD7FFA27A22F9DD85E509DA" pageId="1" pageNumber="374">Shizuokaken</collectorName>
,
<date id="CE647FB9FFD7FFA27FC7F9E1846809DA" box="[206,335,1651,1677]" pageId="1" pageNumber="374" value="1996-03-16">
<collectingDate id="DE208651FFD7FFA27FC7F9E1846809DA" box="[206,335,1651,1677]" pageId="1" pageNumber="374" value="1996-03-16">16 III 1996</collectingDate>
</date>
,
<collectorName id="172F3CAFFFD7FFA27E52F9E184DC09DA" box="[347,507,1651,1677]" pageId="1" pageNumber="374">M. Maruyama</collectorName>
(
<collectionCode id="DCCBC1BCFFD7FFA27D03F9E1877609DA" box="[522,593,1651,1677]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="374" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AB25324FFD7FFA27D6CF9E180A409DA" ID-GBIF-Occurrence="3436476311" box="[613,1411,1651,1677]" collectingDate="1996-03-14" collectionCode="KUM" collectorName="M. Maruyama" pageId="1" pageNumber="374" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="ACDC92F0FFD7FFA27D6CF9E187B509DA" box="[613,658,1651,1677]" pageId="1" pageNumber="374" type="male">1 ♂</specimenCount>
,
<specimenCount id="ACDC92F0FFD7FFA27D97F9E187EF09DA" box="[670,712,1651,1677]" pageId="1" pageNumber="374" type="female">1 ♀</specimenCount>
, 8 unsexed, same locality,
<date id="CE647FB9FFD7FFA27CFEF9E1815F09DA" box="[1015,1144,1651,1677]" pageId="1" pageNumber="374" value="1996-03-14">
<collectingDate id="DE208651FFD7FFA27CFEF9E1815F09DA" box="[1015,1144,1651,1677]" pageId="1" pageNumber="374" value="1996-03-14">14 III 1996</collectingDate>
</date>
,
<collectorName id="172F3CAFFFD7FFA27B8AF9E1800309DA" box="[1155,1316,1651,1677]" pageId="1" pageNumber="374">M. Maruyama</collectorName>
(
<collectionCode id="DCCBC1BCFFD7FFA27A3AF9E1805D09DA" box="[1331,1402,1651,1677]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="374" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0AB25324FFD7FFA27A87F9E1864109E6" ID-GBIF-Occurrence="3436476304" collectingDate="1996-03-14" collectingDateMax="1996-03-16" collectingDateMin="1996-03-14" collectionCode="KUM" collectorName="S. Naomi" pageId="1" pageNumber="374" specimenCount="2" specimenCount-male="2">
<specimenCount id="ACDC92F0FFD7FFA27A87F9E1858909E6" pageId="1" pageNumber="374" type="male">2 ♂</specimenCount>
, 17 unsexed, same locality,
<date id="CE647FB9FFD7FFA27EE1F90587B409E6" box="[488,659,1687,1713]" pageId="1" pageNumber="374" value="1996-03-14" valueMax="1996-03-16" valueMin="1996-03-14">
<collectingDate id="DE208651FFD7FFA27EE1F90587B409E6" box="[488,659,1687,1713]" pageId="1" pageNumber="374" value="1996-03-14" valueMax="1996-03-16" valueMin="1996-03-14">1416 III 1996</collectingDate>
</date>
,
<collectorName id="172F3CAFFFD7FFA27D97F905862009E6" box="[670,775,1687,1713]" pageId="1" pageNumber="374">S. Naomi</collectorName>
(
<collectionCode id="DCCBC1BCFFD7FFA27C1FF905867A09E6" box="[790,861,1687,1713]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="1" pageNumber="374" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2C00AF2FFD7FFA27FCEF928870309AE" pageId="1" pageNumber="374" type="distribution">
<paragraph id="BA655979FFD7FFA27FCEF928870309AE" blockId="1.[151,1437,1470,2037]" pageId="1" pageNumber="374">
<emphasis id="88AE856BFFD7FFA27FCEF92884460983" bold="true" box="[199,353,1722,1748]" pageId="1" pageNumber="374">Distribution.</emphasis>
<collectingCountry id="C2CD19E9FFD7FFA27E62F92984890982" box="[363,430,1723,1749]" name="China" pageId="1" pageNumber="374">China</collectingCountry>
(
<collectingRegion id="781E979BFFD7FFA27EC9F92987700982" box="[448,599,1723,1749]" country="China" name="Heilongjiang" pageId="1" pageNumber="374">Heilongjiang</collectingRegion>
,
<collectingRegion id="781E979BFFD7FFA27D6FF92987BB0981" box="[614,668,1723,1750]" country="China" name="Jilin" pageId="1" pageNumber="374">Jilin</collectingRegion>
)?;
<collectingCountry id="C2CD19E9FFD7FFA27DB7F92986730982" box="[702,852,1723,1749]" name="North Korea" pageId="1" pageNumber="374">North Korea</collectingCountry>
?;
<collectingCountry id="C2CD19E9FFD7FFA27C67F92986890982" box="[878,942,1723,1749]" name="Japan" pageId="1" pageNumber="374">Japan</collectingCountry>
(
<emphasis id="88AE856BFFD7FFA27CCBF928810F0982" bold="true" box="[962,1064,1722,1749]" pageId="1" pageNumber="374">Honshû:</emphasis>
<collectingRegion id="781E979BFFD7FFA27B3AF929818B0982" box="[1075,1196,1723,1749]" country="Japan" name="Kanagawa" pageId="1" pageNumber="374">Kanagawa</collectingRegion>
,
<collectingRegion id="781E979BFFD7FFA27BB2F92980000982" box="[1211,1319,1723,1749]" country="Japan" name="Shizuoka" pageId="1" pageNumber="374">Shizuoka</collectingRegion>
;
<emphasis id="88AE856BFFD7FFA27A3FF92880BB0982" bold="true" box="[1334,1436,1722,1749]" pageId="1" pageNumber="374">Kyûshû:</emphasis>
<collectingRegion id="781E979BFFD7FFA27F9EF94D842109AD" box="[151,262,1759,1786]" country="Japan" name="Nagasaki" pageId="1" pageNumber="374">Nagasaki</collectingRegion>
?). See, Comments below.
</paragraph>
</subSubSection>
<subSubSection id="F2C00AF2FFD7FFA17FCEF890878C0E86" lastPageId="2" lastPageNumber="375" pageId="1" pageNumber="374" type="description">
<paragraph id="BA655979FFD7FFA27FCEF89080A30816" blockId="1.[151,1437,1470,2037]" pageId="1" pageNumber="374">
<emphasis id="88AE856BFFD7FFA27FCEF8908452084B" bold="true" box="[199,373,1794,1820]" pageId="1" pageNumber="374">Redescription.</emphasis>
Body (Figs. 1, 5) large, robust, about 2.23.0 mm, widest at elytra. Color darker brown, but mouthparts, basal segments of antennae, and legs reddish brown, similarly colored, a little paler than body color.
</paragraph>
<paragraph id="BA655979FFD7FFA27FCEF8D986160886" blockId="1.[151,1437,1470,2037]" pageId="1" pageNumber="374">Head subtriangular, more or less produced anteriorly, wider than length, shorter than pronotum and elytra; eyes large, almost as long as temporal length; antenna reddish brown, longer than total length from head to pronotum; antennal segments all longer than width; segment I shortest, about half shorter than II, almost 0.6 times shorter than III; XI almost as long as II, and about 2 times longer than I.</paragraph>
<paragraph id="BA655979FFD7FFA17FCEF84985F80F82" blockId="1.[151,1437,1470,2037]" lastBlockId="2.[151,1437,151,753]" lastPageId="2" lastPageNumber="375" pageId="1" pageNumber="374">Pronotum shorter than elytra and wider than head, semicircular, trapezoid, and wider than length, and widest at base; posterior-lateral angle acute, slightly protrude posteriorly; basal width about 1.8 times wider than narrowest width.</paragraph>
<paragraph id="BA655979FFD4FFA17FCEFF4D86380FAE" blockId="2.[151,1437,151,753]" box="[199,799,223,249]" pageId="2" pageNumber="375">Elytra wider than length; hind wings fully developed.</paragraph>
<paragraph id="BA655979FFD4FFA17FCEFE9186C30E16" blockId="2.[151,1437,151,753]" pageId="2" pageNumber="375">
Median lobe of aedeagus (Figs. 6
<emphasis id="88AE856BFFD4FFA17D30FE9187600E4A" bold="true" box="[569,583,259,285]" pageId="2" pageNumber="375"></emphasis>
7) rather simplified; flagellum long; basal sclerites separated; apical sclerites separated, semicircular. Spermatheca (Fig. 8) about 17 times coiled at base.
</paragraph>
<paragraph id="BA655979FFD4FFA17FCEFED986D70E32" blockId="2.[151,1437,151,753]" box="[199,1008,330,357]" pageId="2" pageNumber="375">
<emphasis id="88AE856BFFD4FFA17FCEFED987290E32" bold="true" box="[199,526,330,357]" pageId="2" pageNumber="375">
Measurements of
<typeStatus id="6561E7DBFFD4FFA17E95FED9872E0E32" box="[412,521,331,357]" pageId="2" pageNumber="375" type="lectotype">lectotype</typeStatus>
.
</emphasis>
BL, ≈ 3.0; PL, 0.59; PW, 0.83; HTL, 0.48.
</paragraph>
<paragraph id="BA655979FFD4FFA17FCEFEFD878C0E86" blockId="2.[151,1437,151,753]" pageId="2" pageNumber="375">
<emphasis id="88AE856BFFD4FFA17FCEFEFD87F70EDF" bold="true" box="[199,720,366,393]" pageId="2" pageNumber="375">
Measurements of specimens from
<collectingRegion id="781E979BFFD4FFA17D52FEFC87EB0EDF" box="[603,716,366,392]" country="Japan" name="Shizuoka" pageId="2" pageNumber="375">Shizuoka</collectingRegion>
.
</emphasis>
BL, 2.23.0; FBL, 1.41.5; HL, 0.300.45; HW, 0.480.55; PL, 0.550.61; PW, 0.850.90; EL, 0.680.80; EW, 0.930.95; HTL, 0.480.55; AL, 1.151.25; AS-I, 0.070.09; AS-II, 0.140.15; AS-III, 0.120.13; AS-XI, 0.170.2.
</paragraph>
</subSubSection>
<subSubSection id="F2C00AF2FFD4FFA17FCEFE4886080DA6" pageId="2" pageNumber="375" type="discussion">
<paragraph id="BA655979FFD4FFA17FCEFE4881E70D6A" blockId="2.[151,1437,151,753]" pageId="2" pageNumber="375">
<emphasis id="88AE856BFFD4FFA17FCEFE4884680EA3" bold="true" box="[199,335,474,500]" pageId="2" pageNumber="375">Comments.</emphasis>
This species was recorded from
<collectingCountry id="C2CD19E9FFD4FFA17DCFFE49862E0EA2" box="[710,777,475,501]" name="China" pageId="2" pageNumber="375">China</collectingCountry>
and
<collectingCountry id="C2CD19E9FFD4FFA17C4BFE4986F70EA2" box="[834,976,475,501]" name="North Korea" pageId="2" pageNumber="375">North Korea</collectingCountry>
(
<bibRefCitation id="DE4B2488FFD4FFA17CE9FE4981190EA2" author="Li, J." box="[992,1086,475,501]" pageId="2" pageNumber="375" refId="ref4394" refString="Li, J. (1992) The Coleoptera fauna of Northeast China. Jilin Education Publishing House, Jinlin, 208 pp. (In Chinese, with English title.)" type="book" year="1992">Li 1992</bibRefCitation>
;
<bibRefCitation id="DE4B2488FFD4FFA17B42FE4981FE0EA2" author="Pasnik, G." box="[1099,1241,475,501]" pageId="2" pageNumber="375" pagination="185 - 234" refId="ref4677" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoologica Cracoviensia, 44 (3), 185 - 234." type="journal article" year="2001">Paśnik 2001</bibRefCitation>
). However, these records are doubtful because they are not based on observation of the genitalia of
<taxonomicName id="7DDA22FAFFD4FFA17B3BFE6D819D0D4F" authorityName="&quot; Sharp" authorityYear="1888" box="[1074,1210,511,536]" class="Insecta" family="Staphylinidae" genus="Myllaena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="88AE856BFFD4FFA17B3BFE6D819D0D4F" box="[1074,1210,511,536]" italics="true" pageId="2" pageNumber="375">M. japonica</emphasis>
</taxonomicName>
, that is essential for species-level identification for this group. Re-examination of the voucher specimens is needed.
</paragraph>
<paragraph id="BA655979FFD4FFA17FCEFDD586080DA6" blockId="2.[151,1437,151,753]" pageId="2" pageNumber="375">
<materialsCitation id="0AB25324FFD4FFA17FCEFDD5860C0DA6" ID-GBIF-Occurrence="3436476302" country="Japan" county="Amagi-toge" location="Miyanoshita" municipality="In" pageId="2" pageNumber="375" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nagasaki" typeStatus="syntype">
This was originally described from
<collectingRegion id="781E979BFFD4FFA17D4FFDD587890D36" box="[582,686,583,609]" country="Japan" name="Nagasaki" pageId="2" pageNumber="375">Nagasaki</collectingRegion>
and Miyanoshita(possibly in
<collectingRegion id="781E979BFFD4FFA17CF8FDD5814B0D36" box="[1009,1132,583,609]" country="Japan" name="Kanagawa" pageId="2" pageNumber="375">Kanagawa</collectingRegion>
).The
<typeStatus id="6561E7DBFFD4FFA17BA3FDD580260D36" box="[1194,1281,583,609]" pageId="2" pageNumber="375" type="syntype">syntype</typeStatus>
are composed of
<specimenCount id="ACDC92F0FFD4FFA17FBCFDF9841A0DD2" box="[181,317,619,645]" pageId="2" pageNumber="375" type="female">two females</specimenCount>
from
<collectingRegion id="781E979BFFD4FFA17E89FDF984CF0DD2" box="[384,488,619,645]" country="Japan" name="Nagasaki" pageId="2" pageNumber="375">Nagasaki</collectingRegion>
and
<specimenCount id="ACDC92F0FFD4FFA17D17FDF987A40DD2" box="[542,643,619,645]" pageId="2" pageNumber="375" type="male">one male</specimenCount>
from
<location id="BF050FA2FFD4FFA17DCEFDF986710DD2" LSID="urn:lsid:plazi:treatment:3273E86FFFD7FFA17F9EFB3C86080DA6:BF050FA2FFD4FFA17DCEFDF986710DD2" box="[711,854,619,645]" country="Japan" county="Amagi-toge" municipality="In" name="Miyanoshita" pageId="2" pageNumber="375" stateProvince="Nagasaki">Miyanoshita</location>
, and the latter is designated as
<typeStatus id="6561E7DBFFD4FFA17BA7FDF980320DD2" box="[1198,1301,619,645]" pageId="2" pageNumber="375" type="lectotype">lectotype</typeStatus>
because the specimens from
<collectingRegion id="781E979BFFD4FFA17E58FD1D849E0DFE" box="[337,441,655,681]" country="Japan" name="Nagasaki" pageId="2" pageNumber="375">Nagasaki</collectingRegion>
belong to a different species (see,
<location id="BF050FA2FFD4FFA17C4BFD1D869C0DFE" LSID="urn:lsid:plazi:treatment:3273E86FFFD7FFA17F9EFB3C86080DA6:BF050FA2FFD4FFA17C4BFD1D869C0DFE" box="[834,955,655,681]" country="Japan" county="Amagi-toge" municipality="In" name="Comments" pageId="2" pageNumber="375" stateProvince="Nagasaki">Comments</location>
of the next species). A distribution map of the examined specimens of this species in
<collectingCountry id="C2CD19E9FFD4FFA17D7CFD2187920D9A" box="[629,693,691,717]" name="Japan" pageId="2" pageNumber="375">Japan</collectingCountry>
is illustrated in
<figureCitation id="22E145FCFFD4FFA17C62FD2186990D9A" box="[875,958,691,717]" captionStart="FIGURES 18" captionStartId="6.[151,264,1655,1679]" captionTargetBox="[151,1435,453,1631]" captionTargetId="figure-121@6.[151,1436,453,1632]" captionTargetPageId="6" captionText="FIGURES 18 Locality map of examined specimens of Myllaena spp. from Japan: white triangle, M. japonica Sharp, 1888; black triangle, M. lewisi Cameron, 1933; white inverted triangle, M. rufotestacea Cameron, 1933; black inverted triangle, M. torrentum Cameron, 1933." figureDoi="http://doi.org/10.5281/zenodo.5843835" httpUri="https://zenodo.org/record/5843835/files/figure.png" pageId="2" pageNumber="375">Fig. 18</figureCitation>
.
<collectingMunicipality id="5A01C303FFD4FFA17CC0FD2186C70D9A" box="[969,992,691,717]" pageId="2" pageNumber="375">In</collectingMunicipality>
<collectingRegion id="781E979BFFD4FFA17CE1FD2181740D9A" box="[1000,1107,691,717]" country="Japan" name="Shizuoka" pageId="2" pageNumber="375">Shizuoka</collectingRegion>
, this species was collected in leaf litter near a wet cliff along the old road of
<collectingCounty id="530421F5FFD4FFA17DABFD45860C0DA6" box="[674,811,727,753]" pageId="2" pageNumber="375">Amagi-tôge</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>