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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.77.769" ID-GBIF-Dataset="041b36c9-43f0-42e0-a8fd-0e7825af491c" ID-PMC="PMC3088056" ID-Pensoft-Pub="1313-2970-77-17" ID-PubMed="21594149" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-77-17" ModsDocOrigin="ZooKeys 77" ModsDocTitle="On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera, Ensifera)" checkinTime="1451250827295" checkinUser="pensoft" docAuthor="Heads, Sam W. &amp; Leuzinger, Lea" docDate="2011" docId="325EB51A702275FDE7BA91869E4425A8" docLanguage="en" docName="ZooKeys 77: 17-30" docOrigin="ZooKeys 77" docSource="http://dx.doi.org/10.3897/zookeys.77.769" docTitle="Schizodactylus Brulle 1835" docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="FFB9FFCFF633200CFFE2E534FF9D5F4F" masterDocTitle="On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera, Ensifera)" masterLastPageNumber="30" masterPageNumber="17" pageNumber="18" updateTime="1668165983640" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera, Ensifera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heads, Sam W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Leuzinger, Lea</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>77</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>17</mods:start>
<mods:end>30</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.77.769</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.77.769</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-77-17</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159361132" LSID="urn:lsid:plazi:treatment:325EB51A702275FDE7BA91869E4425A8" httpUri="http://treatment.plazi.org/id/325EB51A702275FDE7BA91869E4425A8" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
<subSubSection pageId="1" pageNumber="18" type="nomenclature">
<paragraph pageId="1" pageNumber="18">
Genus
<taxonomicName authority="Brulle, 1835" authorityName="Brulle" authorityYear="1835" class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">
Schizodactylus
<normalizedToken originalValue="Brullé">Brulle</normalizedToken>
, 1835
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="18" type="reference_group">
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Schizodactylus</taxonomicName>
<bibRefCitation author="Brulle, GA" journalOrPublisher="In: Audouin JV, Brulle GA (Eds) Histoire Naturelle des Insectes" pageId="7" pageNumber="24" pagination="1 - 225" title="Cinquieme Ordre. Orthopteres." volume="9" year="1835">
<normalizedToken originalValue="Brullé">Brulle</normalizedToken>
1835
</bibRefCitation>
: 161. Type species:
<taxonomicName class="Insecta" family="Gryllidae" genus="Gryllus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gryllus monstrosus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="monstrosus">Gryllus monstrosus</taxonomicName>
Drury, 1773.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Insecta" genus="Schizocephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizocephalus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Schizocephalus</taxonomicName>
<bibRefCitation author="Brunner von Wattenwyl, C" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglicen Zoologisch-Botanischen Gesellschaft in Wien" pageId="7" pageNumber="24" pagination="247 - 394" title="Monographie der Stenopelmatiden und Gryllacriden." volume="38" year="1888">Brunner von Wattenwyl 1888</bibRefCitation>
: 313, lapsus calami.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Dactylocomicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylocomicus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Dactylocomicus</taxonomicName>
<bibRefCitation author="Karny, HH" journalOrPublisher="Transactions of the Royal Society of South Africa" pageId="9" pageNumber="26" pagination="99 - 105" title="On the geographical distribution of the Indo-African and Mediterranean Gryllacrids" volume="19" year="1931">Karny 1931</bibRefCitation>
: 102. Type species:
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Comicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Comicus inexpectatus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="inexpectatus">Comicus inexpectatus</taxonomicName>
Werner, 1901.
</paragraph>
<paragraph pageId="1" pageNumber="18">
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Brauckmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brauckmannia" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Brauckmannia</taxonomicName>
<bibRefCitation author="Martins-Neto, RG" journalOrPublisher="Gaea" pageId="9" pageNumber="26" pagination="3 - 8" title="New Orthoptera Stenopelmatoidea and Hagloidea (Ensifera) from the Santana Formation (Lower Cretaceous, northeast Brazil) with description of new taxa." volume="3" year="2007">Martins-Neto 2007</bibRefCitation>
: 4. Type species:
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Brauckmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brauckmannia groeningae" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="groeningae">Brauckmannia groeningae</taxonomicName>
Martins-Neto, 2007, syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="18" type="diagnosis">
<paragraph pageId="1" pageNumber="18">Diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="18">
The genus
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Schizodactylus</taxonomicName>
as presently defined comprises all large and robust schizodactylids with wings developed and, with the exception of
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus inexpectatus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="inexpectatus">Schizodactylus inexpectatus</taxonomicName>
in which the wings are reduced, extending beyond the apex of the abdomen where they terminate in a conspicuous coil.
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Schizodactylus</taxonomicName>
species are further characterised by greatly enlarged mouthparts, a broad diamond-shaped labrum, and strong laterally compressed pro- and mesothoracic legs. The genus is readily separated from
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Comicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Comicus" order="Orthoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">Comicus</taxonomicName>
, the only other genus in the family, which is characterised by a markedly smaller and more gracile body (usually less than 25 mm in length), long, slender legs, and complete reduction of the wings.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="19" type="comments">
<paragraph pageId="2" pageNumber="19">
<pageBreakToken pageId="2" pageNumber="19" start="start">Comments</pageBreakToken>
.
</paragraph>
<paragraph pageId="2" pageNumber="19">
Given the presence of distinctively coiled wings and well-developed lateral processes on the tarsi, there can be no doubt as to the placement of
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Brauckmannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brauckmannia groeningae" order="Orthoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="groeningae">Brauckmannia groeningae</taxonomicName>
in
<taxonomicName family="Schizodactylidae" lsidName="" pageId="2" pageNumber="19" rank="family">Schizodactylidae</taxonomicName>
. In addition, there are no characters preserved in either the holotype (so far as can be seen from
<normalizedToken originalValue="Martins-Netos">Martins-Neto's</normalizedToken>
illustrations) or the new material described below to exclude the species from the genus
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus" order="Orthoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">Schizodactylus</taxonomicName>
. Both fossil specimens match closely the general habitus, tarsal morphology and metatibial armature of extant
<taxonomicName class="Insecta" family="Schizodactylidae" genus="Schizodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schizodactylus" order="Orthoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">Schizodactylus</taxonomicName>
species, and there are no convincing apomorphies to support separate generic placement.
</paragraph>
</subSubSection>
</treatment>
</document>