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<document id="3A94DE26B5488A349630049C5B898FB0" ID-DOI="10.5281/zenodo.173372" ID-GBIF-Dataset="f6db05a5-a22e-4b7a-8777-969c7c6e3dc1" ID-ISSN="1175­5326" ID-Zenodo-Dep="173372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459774100886" checkinUser="plazi" docAuthor="Fierros-López, Hugo Eduardo" docDate="2006" docId="291587E8FFE3A348326FFE951C40922B" docLanguage="en" docName="zt01279p068.pdf" docOrigin="Zootaxa 1279" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Scaphisoma" docType="key" docVersion="8" lastPageNumber="67" masterDocId="D52CFF90FFEDA3463367FF8E196A966D" masterDocTitle="Four new species of Scaphisoma Leach with maculate elytra (Coleoptera: Staphylinidae: Scaphidiinae) from Mexico, with new records and comments on S. balteatum Matthews" masterLastPageNumber="68" masterPageNumber="53" pageNumber="67" updateTime="1698217572311" updateUser="plazi">
<mods:mods id="6516ABBB8D264B2F5A856FA5FD1EAEC3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7ECCD5251046E59988140BFD0E6413D8">
<mods:title id="4F498E319CE7866A07D9974A695B2616">Four new species of Scaphisoma Leach with maculate elytra (Coleoptera: Staphylinidae: Scaphidiinae) from Mexico, with new records and comments on S. balteatum Matthews</mods:title>
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<mods:name id="77E13F81EBADC96602AD2C295A2F34F2" type="personal">
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<mods:namePart id="45E89C8DE3C36A05C586E0272F1AF34D">Fierros-López, Hugo Eduardo</mods:namePart>
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<mods:date id="8F8C0B94C789AFB6A52578B26489ACA7">2006</mods:date>
<mods:detail id="CDAB72DF81025F8C806ABBAA6DB2EACF" type="volume">
<mods:number id="4F2B68CE15F47EDECE0A709DFE6D630A">1279</mods:number>
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<mods:classification id="F1858C2976A9B7712474CA1DDD2B8731">journal article</mods:classification>
<mods:identifier id="CEA91DA4CA9219BE736BF770DCBB03F7" type="DOI">10.5281/zenodo.173372</mods:identifier>
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<mods:identifier id="C5BBC27D403D681E1767085D9B83F1A2" type="ISSN">1175­5326</mods:identifier>
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<treatment id="291587E8FFE3A348326FFE951C40922B" ID-DOI="http://doi.org/10.5281/zenodo.6253056" ID-GBIF-Taxon="119347646" ID-Zenodo-Dep="6253056" LSID="urn:lsid:plazi:treatment:291587E8FFE3A348326FFE951C40922B" httpUri="http://treatment.plazi.org/id/291587E8FFE3A348326FFE951C40922B" lastPageNumber="67" pageId="14" pageNumber="67">
<subSubSection id="E9A66575FFE3A348326FFE951D419758" box="[264,1067,283,310]" pageId="14" pageNumber="67" type="nomenclature">
<paragraph id="A10336FEFFE3A348326FFE951D419758" blockId="14.[264,1067,283,310]" box="[264,1067,283,310]" pageId="14" pageNumber="67">
<heading id="FA4B8192FFE3A348326FFE951D419758" bold="true" box="[264,1067,283,310]" fontSize="11" level="1" pageId="14" pageNumber="67" reason="1">
<emphasis id="93C8EAECFFE3A348326FFE951D419758" bold="true" box="[264,1067,283,310]" pageId="14" pageNumber="67">
Key to the Mexican species of
<taxonomicName id="66BC4D7DFFE3A34831E7FE921A79975B" ID-CoL="8VS35" box="[640,787,284,310]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="93C8EAECFFE3A34831E7FE921A79975B" bold="true" box="[640,787,284,310]" italics="true" pageId="14" pageNumber="67">Scaphisoma</emphasis>
</taxonomicName>
with maculated elytra
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<subSubSection id="E9A66575FFE3A348326FFED21C40922B" pageId="14" pageNumber="67" type="key">
<keyStep id="1A482E5BFFE3A348326FFED21C4197F3" pageId="14" pageNumber="67">
<paragraph id="A10336FEFFE3A348326FFED21C41971B" blockId="14.[264,1325,348,1094]" box="[264,1323,348,374]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFED21C41971B" box="[264,1323,348,374]" pageId="14" pageNumber="67">
1 Mesocoxal lines parallel (
<figureCitation id="39872A7BFFE3A3483101FED21BDC971B" box="[614,694,348,374]" captionStart="FIGURE 3" captionStartId="5.[264,367,1571,1595]" captionTargetBox="[444,1151,326,1557]" captionTargetId="figure@5.[444,1151,326,1557]" captionTargetPageId="5" captionText="FIGURE 3. Thorax of Scaphisoma, ventral view showing meso and metacoxal lines (other punctures are omitted). a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. Scale bar 1 mm." httpUri="https://zenodo.org/record/173375/files/figure.png" pageId="14" pageNumber="67">Figs. 3</figureCitation>
b, c); elytra with yellow apex ................................... 2
</keyLead>
</paragraph>
<paragraph id="A10336FEFFE3A348326FFE0A1C4197F3" blockId="14.[264,1325,348,1094]" box="[264,1323,388,414]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFE0A1C4197F3" box="[264,1323,388,414]" pageId="14" pageNumber="67">
­ Mesocoxal lines parabolic (
<figureCitation id="39872A7BFFE3A348311CFE0A1BA697F3" box="[635,716,388,414]" captionStart="FIGURE 3" captionStartId="5.[264,367,1571,1595]" captionTargetBox="[444,1151,326,1557]" captionTargetId="figure@5.[444,1151,326,1557]" captionTargetPageId="5" captionText="FIGURE 3. Thorax of Scaphisoma, ventral view showing meso and metacoxal lines (other punctures are omitted). a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. Scale bar 1 mm." httpUri="https://zenodo.org/record/173375/files/figure.png" pageId="14" pageNumber="67">Figs. 3</figureCitation>
a, d); elytra with red or reddish brown apex .......... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A482E5BFFE3A348326FFE221C4094E0" pageId="14" pageNumber="67">
<paragraph id="A10336FEFFE3A348326FFE221C409478" blockId="14.[264,1325,348,1094]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFE221C409478" pageId="14" pageNumber="67">
2 Metacoxal lines sinuate (
<figureCitation id="39872A7BFFE3A348313AFE221BC997AB" box="[605,675,428,454]" captionStart="FIGURE 3" captionStartId="5.[264,367,1571,1595]" captionTargetBox="[444,1151,326,1557]" captionTargetId="figure@5.[444,1151,326,1557]" captionTargetPageId="5" captionText="FIGURE 3. Thorax of Scaphisoma, ventral view showing meso and metacoxal lines (other punctures are omitted). a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. Scale bar 1 mm." httpUri="https://zenodo.org/record/173375/files/figure.png" pageId="14" pageNumber="67">Fig. 3</figureCitation>
d); pronotum reddish orange; parameres with preapical cavity; internal sac with flagellum loosely coiled in the median lobe and bearing small asperities (
<figureCitation id="39872A7BFFE3A34832D1FE721B6D947B" box="[438,519,508,534]" captionStart="FIGURE 4" captionStartId="9.[264,368,1574,1598]" captionTargetBox="[264,1310,436,1562]" captionTargetId="figure@9.[264,1323,436,1562]" captionTargetPageId="9" captionText="FIGURE 4. Aedeagi of Scaphisoma, dorsal, ventral and lateral views respectively. a – c) S. jaliscanum sp. nov.; d – f) S. cortesaguilari sp. nov.; g – i) S. opochtli sp. nov.; j – l) S. versicolor sp. nov.; m – o) S. balteatum Matthews. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/173376/files/figure.png" pageId="14" pageNumber="67">Figs. 4</figureCitation>
j–l)
<emphasis id="93C8EAECFFE3A348315CFE721DAC9478" box="[571,1222,508,533]" italics="true" pageId="14" pageNumber="67">
....................................................
<taxonomicName id="66BC4D7DFFE3A34830DCFE721DAC9478" ID-CoL="79SV4" box="[955,1222,508,533]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="versicolor" status="sp. nov.">Scaphisoma versicolor</taxonomicName>
</emphasis>
<taxonomicNameLabel id="88FB5797FFE3A34837AAFE751C409478" box="[1229,1322,507,533]" pageId="14" pageNumber="67" rank="species">
<emphasis id="93C8EAECFFE3A34837AAFE751C409478" bold="true" box="[1229,1322,507,533]" pageId="14" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="A10336FEFFE3A348326FFDAA1C4094E0" blockId="14.[264,1325,348,1094]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFDAA1C4094E0" pageId="14" pageNumber="67">
<emphasis id="93C8EAECFFE3A348326FFDAA18789450" box="[264,274,548,573]" italics="true" pageId="14" pageNumber="67">­</emphasis>
Metacoxal lines parallel (
<figureCitation id="39872A7BFFE3A3483102FDAA1BC79453" box="[613,685,548,574]" captionStart="FIGURE 3" captionStartId="5.[264,367,1571,1595]" captionTargetBox="[444,1151,326,1557]" captionTargetId="figure@5.[444,1151,326,1557]" captionTargetPageId="5" captionText="FIGURE 3. Thorax of Scaphisoma, ventral view showing meso and metacoxal lines (other punctures are omitted). a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. Scale bar 1 mm." httpUri="https://zenodo.org/record/173375/files/figure.png" pageId="14" pageNumber="67">Fig. 3</figureCitation>
a); pronotum dark brown or black; parameres without preapical modification; internal sac with flagellum closely coiled and striate (
<figureCitation id="39872A7BFFE3A3483797FDC2182D94E3" captionStart="FIGURE 4" captionStartId="9.[264,368,1574,1598]" captionTargetBox="[264,1310,436,1562]" captionTargetId="figure@9.[264,1323,436,1562]" captionTargetPageId="9" captionText="FIGURE 4. Aedeagi of Scaphisoma, dorsal, ventral and lateral views respectively. a – c) S. jaliscanum sp. nov.; d – f) S. cortesaguilari sp. nov.; g – i) S. opochtli sp. nov.; j – l) S. versicolor sp. nov.; m – o) S. balteatum Matthews. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/173376/files/figure.png" pageId="14" pageNumber="67">Figs. 4</figureCitation>
a–c)
<emphasis id="93C8EAECFFE3A348321BFDFA1DAF94E0" box="[380,1221,628,653]" italics="true" pageId="14" pageNumber="67">
............................................................................
<taxonomicName id="66BC4D7DFFE3A34830D6FDFA1DAF94E0" ID-CoL="6XTBP" box="[945,1221,628,653]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="jaliscanum" status="sp. nov.">Scaphisoma jaliscanum</taxonomicName>
</emphasis>
<taxonomicNameLabel id="88FB5797FFE3A34837AAFDFD1C4094E0" box="[1229,1322,627,653]" pageId="14" pageNumber="67" rank="species">
<emphasis id="93C8EAECFFE3A34837AAFDFD1C4094E0" bold="true" box="[1229,1322,627,653]" pageId="14" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A482E5BFFE3A348326FFD121C41953B" pageId="14" pageNumber="67">
<paragraph id="A10336FEFFE3A348326FFD121C40956B" blockId="14.[264,1325,348,1094]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFD121C40956B" pageId="14" pageNumber="67">
3 Metacoxal line convex (
<figureCitation id="39872A7BFFE3A348313AFD121BCC94DB" box="[605,678,668,694]" captionStart="FIGURE 3" captionStartId="5.[264,367,1571,1595]" captionTargetBox="[444,1151,326,1557]" captionTargetId="figure@5.[444,1151,326,1557]" captionTargetPageId="5" captionText="FIGURE 3. Thorax of Scaphisoma, ventral view showing meso and metacoxal lines (other punctures are omitted). a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. Scale bar 1 mm." httpUri="https://zenodo.org/record/173375/files/figure.png" pageId="14" pageNumber="67">Fig. 3</figureCitation>
b); central area of pronotum with with black macula; aedeagus with symmetrical median lobe, which has lateral subapical projections (
<figureCitation id="39872A7BFFE3A3483797FD4A182C956B" captionStart="FIGURE 4" captionStartId="9.[264,368,1574,1598]" captionTargetBox="[264,1310,436,1562]" captionTargetId="figure@9.[264,1323,436,1562]" captionTargetPageId="9" captionText="FIGURE 4. Aedeagi of Scaphisoma, dorsal, ventral and lateral views respectively. a – c) S. jaliscanum sp. nov.; d – f) S. cortesaguilari sp. nov.; g – i) S. opochtli sp. nov.; j – l) S. versicolor sp. nov.; m – o) S. balteatum Matthews. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/173376/files/figure.png" pageId="14" pageNumber="67">Figs. 4</figureCitation>
d–f)
<emphasis id="93C8EAECFFE3A34832E3FD621DAD9568" box="[388,1223,748,773]" italics="true" pageId="14" pageNumber="67">
......................................................................
<taxonomicName id="66BC4D7DFFE3A34830EAFD621DAD9568" ID-CoL="6XTN5" box="[909,1223,748,773]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="cortesaguilari" status="sp. nov.">Scaphisoma cortesaguilari</taxonomicName>
</emphasis>
<taxonomicNameLabel id="88FB5797FFE3A34837AAFD621C40956B" box="[1229,1322,748,774]" pageId="14" pageNumber="67" rank="species">
<emphasis id="93C8EAECFFE3A34837AAFD621C40956B" bold="true" box="[1229,1322,748,774]" pageId="14" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="A10336FEFFE3A348326FFC9A1C41953B" blockId="14.[264,1325,348,1094]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFC9A1C41953B" pageId="14" pageNumber="67">
­ Metacoxal lines sinuate (
<figureCitation id="39872A7BFFE3A3483102FC9A1BC79543" box="[613,685,788,814]" captionStart="FIGURE 3" captionStartId="5.[264,367,1571,1595]" captionTargetBox="[444,1151,326,1557]" captionTargetId="figure@5.[444,1151,326,1557]" captionTargetPageId="5" captionText="FIGURE 3. Thorax of Scaphisoma, ventral view showing meso and metacoxal lines (other punctures are omitted). a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. Scale bar 1 mm." httpUri="https://zenodo.org/record/173375/files/figure.png" pageId="14" pageNumber="67">Fig. 3</figureCitation>
c); pronotum uniformly colored; aedeagus with other characters ...................................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1A482E5BFFE3A348326FFCEA1C40922B" pageId="14" pageNumber="67">
<paragraph id="A10336FEFFE3A348326FFCEA1C4095A3" blockId="14.[264,1325,348,1094]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFCEA1C4095A3" pageId="14" pageNumber="67">
4 Pronotum black; ventrites dark brown; humeral area of elytra with black or brown macula (
<figureCitation id="39872A7BFFE3A34832F9FC02188895CB" box="[414,482,908,934]" captionStart="FIGURE 1" captionStartId="3.[264,368,1676,1700]" captionTargetBox="[275,1305,326,1657]" captionTargetId="figure@3.[264,1323,307,1672]" captionTargetPageId="3" captionText="FIGURE 1. Habitus drawings of Scaphisoma. a) S. jaliscanum sp. nov.; b) S. cortesaguilari sp. nov.; c) S. opochtli sp. nov.; d) S. versicolor sp. nov. (all holotypes). Scale bar 1 mm." httpUri="https://zenodo.org/record/173373/files/figure.png" pageId="14" pageNumber="67">Fig. 1</figureCitation>
c); Aedeagus with asymmetrical median lobe, in dorsal view curved to the left (
<figureCitation id="39872A7BFFE3A34832FDFC3A188B95A3" box="[410,481,948,974]" captionStart="FIGURE 4" captionStartId="9.[264,368,1574,1598]" captionTargetBox="[264,1310,436,1562]" captionTargetId="figure@9.[264,1323,436,1562]" captionTargetPageId="9" captionText="FIGURE 4. Aedeagi of Scaphisoma, dorsal, ventral and lateral views respectively. a – c) S. jaliscanum sp. nov.; d – f) S. cortesaguilari sp. nov.; g – i) S. opochtli sp. nov.; j – l) S. versicolor sp. nov.; m – o) S. balteatum Matthews. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/173376/files/figure.png" pageId="14" pageNumber="67">
Fig.
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g)
<emphasis id="93C8EAECFFE3A3483167FC3A1DAF95A0" box="[512,1221,948,973]" italics="true" pageId="14" pageNumber="67">
...............................................................
<taxonomicName id="66BC4D7DFFE3A34830B6FC3A1DAF95A0" ID-CoL="6XTDR" box="[977,1221,948,973]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="opochtli" status="sp. nov.">Scaphisoma opochtli</taxonomicName>
</emphasis>
<taxonomicNameLabel id="88FB5797FFE3A34837AAFC3A1C4095A3" box="[1229,1322,948,974]" pageId="14" pageNumber="67" rank="species">
<emphasis id="93C8EAECFFE3A34837AAFC3A1C4095A3" bold="true" box="[1229,1322,948,974]" pageId="14" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="A10336FEFFE3A348326FFC521C40922B" blockId="14.[264,1325,348,1094]" pageId="14" pageNumber="67">
<keyLead id="1A4D95CBFFE3A348326FFC521C40922B" pageId="14" pageNumber="67">
­ Pronotum orange­red; ventrites reddish; humeral area of elytra red; aedeagus with symmetrical, straight and apically pointed median lobe (
<figureCitation id="39872A7BFFE3A34830AAFB8A1D789273" box="[973,1042,1028,1054]" captionStart="FIGURE 4" captionStartId="9.[264,368,1574,1598]" captionTargetBox="[264,1310,436,1562]" captionTargetId="figure@9.[264,1323,436,1562]" captionTargetPageId="9" captionText="FIGURE 4. Aedeagi of Scaphisoma, dorsal, ventral and lateral views respectively. a – c) S. jaliscanum sp. nov.; d – f) S. cortesaguilari sp. nov.; g – i) S. opochtli sp. nov.; j – l) S. versicolor sp. nov.; m – o) S. balteatum Matthews. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/173376/files/figure.png" pageId="14" pageNumber="67">
Fig.
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m) ................................
<emphasis id="93C8EAECFFE3A348325CFBA21DDA9228" box="[315,1200,1068,1093]" italics="true" pageId="14" pageNumber="67">
....................................................................................
<taxonomicName id="66BC4D7DFFE3A34830C1FBA21DDA9228" ID-CoL="79TNP" authority="Matthews" authorityName="Matthews" box="[934,1200,1068,1093]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="67" phylum="Arthropoda" rank="species" species="balteatum">Scaphisoma balteatum</taxonomicName>
</emphasis>
Matthews
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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