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<document id="1318B4B9BB92F671FA49B25298887E04" ID-DOI="10.11646/zootaxa.5424.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10808123" ID-ZooBank="EFF99EA9-03C3-4C9F-B4E0-22684DE34143" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710236759529" checkinUser="plazi" docAuthor="Yamada, Kazutaka" docDate="2024" docId="1E3F87F75209FFA3FF09FE2CFB9B9CFC" docLanguage="en" docName="zootaxa.5424.1.7.pdf" docOrigin="Zootaxa 5424 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiastethus longiceps Poppius 1915" docType="treatment" docVersion="2" lastPageNumber="130" masterDocId="E206FF8F520BFFA0FF9EFFF6FF879D09" masterDocTitle="Taxonomic corrections on Oriental Anthocoridae (Hemiptera: Heteroptera) described by Bertil Poppius" masterLastPageNumber="135" masterPageNumber="127" pageNumber="129" updateTime="1710237684799" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3F25392E0BFC0DFD581F088BC26D38FB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E958ACC105EDDF120619BAA70E06C088">Taxonomic corrections on Oriental Anthocoridae (Hemiptera: Heteroptera) described by Bertil Poppius</mods:title>
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<mods:roleTerm id="21F306B2A82BC60A11A3E688BE94995D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70CE635E72C970CB73C6FA1FFC41CA29">Yamada, Kazutaka</mods:namePart>
<mods:affiliation id="C728C91F7C60781ED2C6B49E227D9863">Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6 - chome, Sanda-shi, Hyogo 669 - 1546, Japan. &amp; Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6 - chome, Sanda-shi, Hyogo 669 - 1546, Japan.</mods:affiliation>
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<mods:title id="743EBC8C13064DEA1CBBF418F73CD6C2">Zootaxa</mods:title>
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<mods:date id="F63540D56A14135ECC8495CB3F33382F">2024</mods:date>
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<mods:number id="3FF2311C6ABCDFB8370B8D4653204F99">2024-03-12</mods:number>
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<mods:number id="A086FF26DD3679B3E8ED7F7F3069EB9F">5424</mods:number>
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<mods:url id="93CB54E0B67833F4D44DE6A4B0E21AEA">http://dx.doi.org/10.11646/zootaxa.5424.1.7</mods:url>
</mods:location>
<mods:classification id="7AB2F006C060A305AB1D6F4296665F04">journal article</mods:classification>
<mods:identifier id="49AA0C96D0137021ACAFACF4ADA794F7" type="DOI">10.11646/zootaxa.5424.1.7</mods:identifier>
<mods:identifier id="F7AB339B6BA0FA03DA152FDB2742CC23" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F337AF211BF573355BD848D0770A11A1" type="Zenodo-Dep">10808123</mods:identifier>
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<treatment id="1E3F87F75209FFA3FF09FE2CFB9B9CFC" ID-DOI="http://doi.org/10.5281/zenodo.10808131" ID-Zenodo-Dep="10808131" LSID="urn:lsid:plazi:treatment:1E3F87F75209FFA3FF09FE2CFB9B9CFC" httpUri="http://treatment.plazi.org/id/1E3F87F75209FFA3FF09FE2CFB9B9CFC" lastPageId="3" lastPageNumber="130" pageId="2" pageNumber="129">
<subSubSection id="DE8C656A5209FFA2FF09FE2CFDE89CFD" box="[151,623,474,501]" pageId="2" pageNumber="129" type="nomenclature">
<paragraph id="962936E15209FFA2FF09FE2CFDE89CFD" blockId="2.[151,623,474,537]" box="[151,623,474,501]" pageId="2" pageNumber="129">
<heading id="CD61818D5209FFA2FF09FE2CFDE89CFD" bold="true" box="[151,623,474,501]" fontSize="11" level="1" pageId="2" pageNumber="129" reason="1">
<taxonomicName id="51964D625209FFA2FF09FE2CFDE89CFD" authority="Poppius, 1915" authorityName="Poppius" authorityYear="1915" box="[151,623,474,501]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="A4E2EAF35209FFA2FF09FE2CFDE89CFD" bold="true" box="[151,623,474,501]" pageId="2" pageNumber="129">
<emphasis id="A4E2EAF35209FFA2FF09FE2CFE3F9CFC" bold="true" box="[151,440,474,501]" italics="true" pageId="2" pageNumber="129">Cardiastethus longiceps</emphasis>
<bibRefCitation id="F2074B105209FFA2FE5EFE2CFDE89CFD" author="Poppius, B." box="[448,623,474,500]" pageId="2" pageNumber="129" pagination="1 - 80" refId="ref5294" refString="Poppius, B. (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8), 1 - 80. [1914, March 1915]" type="journal article" year="1915">Poppius, 1915</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8C656A5209FFA2FF09FE09FE959F10" box="[151,274,511,537]" pageId="2" pageNumber="129" type="description">
<paragraph id="962936E15209FFA2FF09FE09FE959F10" blockId="2.[151,623,474,537]" box="[151,274,511,537]" pageId="2" pageNumber="129">
(
<figureCitation id="0EAD2A645209FFA2FF01FE09FE8D9F10" box="[159,266,511,537]" captionStart="FIGURES 4–6" captionStartId="2.[151,264,1746,1770]" captionTargetBox="[151,1387,1022,1723]" captionTargetId="figure-286@2.[151,1436,1022,1723]" captionTargetPageId="2" captionText="FIGURES 4–6. Holotype of Cardiastethus longiceps Poppius, 1915 and its labels. 4–5, dorsal and lateral views; 6, labels. Scales: 0.5 mm. (©SDEI, courtesy of J. F. Tsai and NMNS, published with permission.)" figureDoi="http://doi.org/10.5281/zenodo.10808129" httpUri="https://zenodo.org/record/10808129/files/figure.png" pageId="2" pageNumber="129">Figs. 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DE8C656A5209FFA2FF09FDB5FADE9E3B" pageId="2" pageNumber="129" type="reference_group">
<paragraph id="962936E15209FFA2FF09FDB5FC509F52" blockId="2.[151,1436,579,818]" box="[151,983,579,603]" pageId="2" pageNumber="129">
<treatmentCitationGroup id="B68611CF5209FFA2FF09FDB5FC509F52" box="[151,983,579,603]" pageId="2" pageNumber="129">
<taxonomicName id="51964D625209FFA2FF09FDB5FE059F52" authorityName="Poppius" authorityYear="1915" box="[151,386,579,603]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="longicep">
<emphasis id="A4E2EAF35209FFA2FF09FDB5FE059F52" box="[151,386,579,603]" italics="true" pageId="2" pageNumber="129">Cardiastethus longicep</emphasis>
</taxonomicName>
<materialsCitation id="26FE3CBC5209FFA2FE1CFDB2FC509F52" box="[386,983,579,603]" collectionCode="SDEI" country="Taiwan" location="Taihorin" pageId="2" pageNumber="129" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
s
<treatmentCitation id="173710F05209FFA2FE0CFDB5FDC79F52" author="Poppius, B." box="[402,576,579,603]" page="8" pageId="2" pageNumber="129" year="1915">
<bibRefCitation id="F2074B105209FFA2FE0CFDB5FDC79F52" author="Poppius, B." box="[402,576,579,603]" pageId="2" pageNumber="129" pagination="1 - 80" refId="ref5294" refString="Poppius, B. (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8), 1 - 80. [1914, March 1915]" type="journal article" year="1915">Poppius, 1915: 8</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="492D88435209FFA2FDD4FDB5FD2B9F52" box="[586,684,579,603]" pageId="2" pageNumber="129" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8090FD685209FFA2FD28FDB2FD4E9F52" box="[694,713,580,603]" count="1" pageId="2" pageNumber="129" type="female">♀</specimenCount>
,
<collectingCountry id="EE8176715209FFA2FD4BFDB5FCA59F52" box="[725,802,579,603]" name="Taiwan" pageId="2" pageNumber="129">Taiwan</collectingCountry>
,
<location id="9349603A5209FFA2FCB5FDB5FC079F52" LSID="urn:lsid:plazi:treatment:1E3F87F75209FFA3FF09FE2CFB9B9CFC:9349603A5209FFA2FCB5FDB5FC079F52" box="[811,896,579,603]" country="Taiwan" name="Taihorin" pageId="2" pageNumber="129">Taihorin</location>
(
<collectionCode id="F087AE245209FFA2FC10FDB5FC4B9F52" box="[910,972,579,603]" pageId="2" pageNumber="129">SDEI</collectionCode>
).
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="962936E15209FFA2FF09FD94FE269F90" blockId="2.[151,1436,579,818]" pageId="2" pageNumber="129">
<taxonomicName id="51964D625209FFA2FF09FD94FD5C9F73" authority="Yamada &amp; Hirowatari, 2004: 57" authorityName="Yamada &amp; Hirowatari" authorityPageNumber="57" authorityYear="2004" box="[151,731,610,634]" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="A4E2EAF35209FFA2FF09FD94FE0A9F73" box="[151,397,610,634]" italics="true" pageId="2" pageNumber="129">Lippomanus brevicornis</emphasis>
<materialsCitation id="26FE3CBC5209FFA2FE0DFD94FD5C9F73" box="[403,731,610,634]" collectionCode="OMU" country="Japan" county="New" location="Ryukyus" municipality="Uehara" pageId="2" pageNumber="129" specimenCount="1" specimenCount-male="1" typeStatus="holotype">Yamada &amp; Hirowatari, 2004: 57</materialsCitation>
</taxonomicName>
.
<typeStatus id="492D88435209FFA2FD7BFD94FCC09F73" box="[741,839,610,634]" pageId="2" pageNumber="129" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8090FD685209FFA2FCCFFD94FCE19F70" box="[849,870,610,633]" count="1" pageId="2" pageNumber="129" type="male">♂</specimenCount>
,
<collectingCountry id="EE8176715209FFA2FCEEFD94FC2B9F70" box="[880,940,610,633]" name="Japan" pageId="2" pageNumber="129">Japan</collectingCountry>
,
<location id="9349603A5209FFA2FC29FD94FB949F73" LSID="urn:lsid:plazi:treatment:1E3F87F75209FFA3FF09FE2CFB9B9CFC:9349603A5209FFA2FC29FD94FB949F73" box="[951,1043,610,634]" country="Japan" county="New" municipality="Uehara" name="Ryukyus" pageId="2" pageNumber="129">Ryukyus</location>
,
<location id="9349603A5209FFA2FB82FD94FB3D9F73" LSID="urn:lsid:plazi:treatment:1E3F87F75209FFA3FF09FE2CFB9B9CFC:9349603A5209FFA2FB82FD94FB3D9F73" box="[1052,1210,610,634]" country="Japan" county="New" municipality="Uehara" name="Iriomote Island" pageId="2" pageNumber="129">Iriomote Island</location>
,
<collectingMunicipality id="764DAC9B5209FFA2FB5AFD94FA8A9F73" box="[1220,1293,610,634]" pageId="2" pageNumber="129">Uehara</collectingMunicipality>
(OMU).
<emphasis id="A4E2EAF35209FFA2FAF2FD94FE269F90" bold="true" pageId="2" pageNumber="129">
<collectingCounty id="7F484E6D5209FFA2FAF2FD94FA1B9F73" box="[1388,1436,610,634]" pageId="2" pageNumber="129">New</collectingCounty>
subjective synonym.
</emphasis>
</paragraph>
<paragraph id="962936E15209FFA2FF09FD69FD8B9FFD" blockId="2.[151,1436,579,818]" pageId="2" pageNumber="129">
<treatmentCitationGroup id="B68611CF5209FFA2FF09FD69FD8B9FFD" pageId="2" pageNumber="129">
<taxonomicName id="51964D625209FFA2FF09FD69FE0B9FBE" authorityName="Poppius" authorityYear="1915" box="[151,396,671,695]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="A4E2EAF35209FFA2FF09FD69FE0B9FBE" box="[151,396,671,695]" italics="true" pageId="2" pageNumber="129">Cardiastethus longiceps</emphasis>
</taxonomicName>
:
<treatmentCitation id="173710F05209FFA2FE04FD69FDD59FBE" author="Esaki, T." box="[410,594,671,695]" page="171" pageId="2" pageNumber="129" year="1926">
<bibRefCitation id="F2074B105209FFA2FE04FD69FDD59FBE" author="Esaki, T." box="[410,594,671,695]" pageId="2" pageNumber="129" pagination="136 - 189" refId="ref4589" refString="Esaki, T. (1926) Verzeichnis der Hemiptera-Heteroptera der Insel Formosa. Annales Historico-Naturales Musei Nationalis Hungarici, 24, 136 - 189." type="journal article" year="1926">Esaki (1926: 171)</bibRefCitation>
</treatmentCitation>
(listed, distribution),
<treatmentCitation id="173710F05209FFA2FCB1FD69FC5C9FBE" author="Hiura, I." box="[815,987,671,695]" page="43" pageId="2" pageNumber="129" year="1960">
<bibRefCitation id="F2074B105209FFA2FCB1FD69FC5C9FBE" author="Hiura, I." box="[815,987,671,695]" pageId="2" pageNumber="129" pagination="43 - 55" refId="ref4818" refString="Hiura, I. (1960) Contribution to the knowledge of Anthocoridae from Japan and its adjacent territories (Hemiptera-Heteroptera) 2. Bulletin of the Osaka Museum of Natural History, 12, 43 - 55." type="journal article" year="1960">Hiura (1960: 43)</bibRefCitation>
</treatmentCitation>
(listed),
<treatmentCitation id="173710F05209FFA2FBA9FD69FB549FBE" author="Ford, L. J." box="[1079,1235,671,695]" page="52" pageId="2" pageNumber="129" year="1979">
<bibRefCitation id="F2074B105209FFA2FBA9FD69FB549FBE" author="Ford, L. J." box="[1079,1235,671,695]" pageId="2" pageNumber="129" refId="ref4615" refString="Ford, L. J. (1979) The Phylogeny and Biogeography of the Cimicoidea (Insecta: Hemiptera). MSc. Thesis. University of Connecticut, Storrs, 138 pp." type="book" year="1979">Ford (1979: 52</bibRefCitation>
</treatmentCitation>
, 88) (listed),
<treatmentCitation id="173710F05209FFA2FAC5FD69FEF79FDF" author="Zheng, L. - Y. &amp; Bu, W. - J." page="27" pageId="2" pageNumber="129" year="1990">
<bibRefCitation id="F2074B105209FFA2FAC5FD69FEF79FDF" author="Zheng, L. - Y. &amp; Bu, W. - J." pageId="2" pageNumber="129" pagination="23 - 27" refId="ref5594" refString="Zheng, L. - Y. &amp; Bu, W. - J. (1990) A list of Anthocoridae from China. Contributions from the Tianjin Natural History Museum, 7, 23 - 27. [in Chinese]" type="journal article" year="1990">Zheng &amp; Bu (1990: 27)</bibRefCitation>
</treatmentCitation>
(listed, distribution),
<treatmentCitation id="173710F05209FFA2FDD3FD48FC9D9FDF" author="Pericart, J." box="[589,794,702,726]" page="130" pageId="2" pageNumber="129" year="1996">
<bibRefCitation id="F2074B105209FFA2FDD3FD48FC9D9FDF" author="Pericart, J." box="[589,794,702,726]" pageId="2" pageNumber="129" refId="ref5038" refString="Pericart, J. (1996) Family Anthocoridae Fieber, 1836 - flower bugs, minute pirate bugs. In: Aukema, B. &amp; Rieger, Ch. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 2. Cimicomorpha I. The Netherlands Entomological Society, Amsterdam, pp. 108 - 140." type="book" year="1996">Péricart (1996: 130)</bibRefCitation>
</treatmentCitation>
(catalog, distribution),
<treatmentCitation id="173710F05209FFA2FB97FD48FB749FDF" author="Bu, W. - J. &amp; Zheng, L. - Y." box="[1033,1267,702,726]" page="53" pageId="2" pageNumber="129" year="2001">
<bibRefCitation id="F2074B105209FFA2FB97FD48FB749FDF" author="Bu, W. - J. &amp; Zheng, L. - Y." box="[1033,1267,702,726]" pageId="2" pageNumber="129" refId="ref4305" refString="Bu, W. - J. &amp; Zheng, L. - Y. (2001) Hemiptera: Lasiochilidae, Lyctocoridae, Anthocoridae. Fauna Sinica. Insecta. Vol. 24. Science Press, Beijing, 267 pp. [in Chinese, English summary]" type="book" year="2001">Bu &amp; Zheng (2001: 53</bibRefCitation>
</treatmentCitation>
, 93, 95, 230) (in key, redescription, distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="962936E15209FFA2FF09FD0AFADE9E3B" blockId="2.[151,1436,579,818]" pageId="2" pageNumber="129">
<treatmentCitationGroup id="B68611CF5209FFA2FF09FD0AFADE9E3B" pageId="2" pageNumber="129">
<taxonomicName id="51964D625209FFA2FF09FD0AFE169E1A" authorityName="Yamada &amp; Hirowatari" authorityYear="2004" box="[151,401,763,787]" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="A4E2EAF35209FFA2FF09FD0AFE169E1A" box="[151,401,763,787]" italics="true" pageId="2" pageNumber="129">Lippomanus brevicornis</emphasis>
</taxonomicName>
:
<treatmentCitation id="173710F05209FFA2FE3CFD0AFD709E1A" author="Yasunaga, T. &amp; Yamada, K." box="[418,759,763,788]" page="35" pageId="2" pageNumber="129" year="2014">
<bibRefCitation id="F2074B105209FFA2FE3CFD0AFD709E1A" author="Yasunaga, T. &amp; Yamada, K." box="[418,759,763,788]" pageId="2" pageNumber="129" pagination="25 - 38" refId="ref6000" refString="Yasunaga, T. &amp; Yamada, K. (2014) Japanese or scientific names proposed for mirid and anthocorid species, recently described or recorded from Japan (Heteroptera, Miridae and Anthocoridae). Rostria, 57, 25 - 38. [in Japanese, English summary]" type="journal article" year="2014">Yasunaga &amp; Yamada (2014: 35)</bibRefCitation>
</treatmentCitation>
(diagnosis, bionomics, distribution),
<treatmentCitation id="173710F05209FFA2FB1DFD0DFA1B9E1A" author="Yamada, K. &amp; Yasunaga, T. &amp; Ishikawa, T." box="[1155,1436,763,787]" page="425" pageId="2" pageNumber="129" year="2016">
<bibRefCitation id="F2074B105209FFA2FB1DFD0DFA1B9E1A" author="Yamada, K. &amp; Yasunaga, T. &amp; Ishikawa, T." box="[1155,1436,763,787]" pageId="2" pageNumber="129" refId="ref5846" refString="Yamada, K., Yasunaga, T. &amp; Ishikawa, T. (2016) Family Anthocoridae. In: Hayashi, M., Tomokuni, M., Yoshizawa, K. &amp; Ishikawa, T. (Eds.), Catalogue of the insects of Japan. Vol. 4. Paraneoptera. The Entomological Society of Japan and Touka-shobo, Fukuoka, pp. 422 - 428. [in Japanese]" type="book" year="2016">
Yamada
<emphasis id="A4E2EAF35209FFA2FB41FD0AFA909E1A" box="[1247,1303,763,787]" italics="true" pageId="2" pageNumber="129">et al.</emphasis>
(2016: 425)
</bibRefCitation>
</treatmentCitation>
(catalog, distribution),
<treatmentCitation id="173710F05209FFA2FE31FCECFDF19E3B" author="Devi, N. &amp; Gupta, P. R. &amp; Ram, B." box="[431,630,794,818]" page="2" pageId="2" pageNumber="129" year="2017">
<bibRefCitation id="F2074B105209FFA2FE31FCECFDF19E3B" author="Devi, N. &amp; Gupta, P. R. &amp; Ram, B." box="[431,630,794,818]" pageId="2" pageNumber="129" pagination="1 - 8" refId="ref4468" refString="Devi, N., Gupta, P. R. &amp; Ram, B. (2017) Distribution and seasonal activity of anthocorid bugs (Hemiptera: Anthocoridae) in subtemperate zone of Himachal Pradesh (India). Research Journal of Chemical and Environmental Sciences, 5 (4), 1 - 8." type="journal article" year="2017">
Devi
<emphasis id="A4E2EAF35209FFA2FE79FCEDFD9D9E3B" box="[487,538,794,818]" italics="true" pageId="2" pageNumber="129">et al.</emphasis>
(2017: 2
</bibRefCitation>
</treatmentCitation>
, 4) (records, bionomics),
<treatmentCitation id="173710F05209FFA2FCE5FCECFBE99E3B" ID-DOI="http://doi.org/10.5281/zenodo.3704466" ID-Zenodo-Dep="3704466" author="Ballal, C. R. &amp; Akbar, S. A. &amp; Yamada, K. &amp; Wachkoo, A. A. &amp; Varshney, R." box="[891,1134,794,818]" httpUri="http://treatment.plazi.org/id/F25E8920856C9609FCA87CD2FD4AFEA3" page="208" pageId="2" pageNumber="129" year="2018">
<bibRefCitation id="F2074B105209FFA2FCE5FCECFBE99E3B" author="Ballal, C. R. &amp; Akbar, S. A. &amp; Yamada, K. &amp; Wachkoo, A. A. &amp; Varshney, R." box="[891,1134,794,818]" pageId="2" pageNumber="129" pagination="207 - 226" refId="ref4231" refString="Ballal, C. R., Akbar, S. A., Yamada, K., Wachkoo, A. A. &amp; Varshney, R. (2018) Annotated catalogue of the flower bugs from India (Heteroptera: Anthocoridae, Lasiochilidae). Acta Entomologica Musei Nationalis Pragae, 58, 207 - 226. https: // doi. org / 10.2478 / aemnp- 2018 - 0018" type="journal article" year="2018">
Ballal
<emphasis id="A4E2EAF35209FFA2FC20FCEDFC759E3B" box="[958,1010,794,818]" italics="true" pageId="2" pageNumber="129">et al.</emphasis>
(2018: 208)
</bibRefCitation>
</treatmentCitation>
(records, distribution).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DE8C656A5209FFA2FF09FC96FEE89AEE" pageId="2" pageNumber="129" type="materials_examined">
<paragraph id="962936E15209FFA2FF09FC96FEFC9EEF" blockId="2.[151,1436,863,998]" pageId="2" pageNumber="129">
<emphasis id="A4E2EAF35209FFA2FF09FC96FD5A9E73" bold="true" box="[151,733,863,890]" pageId="2" pageNumber="129">
Type material examined.
<taxonomicName id="51964D625209FFA2FE5CFCA9FD4C9E73" authorityName="Poppius" authorityYear="1915" box="[450,715,863,890]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="longicep">
<emphasis id="A4E2EAF35209FFA2FE5CFCA9FD4C9E73" bold="true" box="[450,715,863,890]" italics="true" pageId="2" pageNumber="129">Cardiastethus longicep</emphasis>
</taxonomicName>
s.
</emphasis>
<materialsCitation id="26FE3CBC5209FFA2FD7CFC96FEFC9EEF" collectingDate="1911-05" collectorName="H. Sauter &amp; Holotypus'" country="Taiwan" determinerName="Poppius" location="Taihorin" pageId="2" pageNumber="129" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<emphasis id="A4E2EAF35209FFA2FD7CFC96FCDE9E73" bold="true" box="[738,857,864,890]" pageId="2" pageNumber="129">
<typeStatus id="492D88435209FFA2FD7CFC96FCD49E73" box="[738,851,864,890]" pageId="2" pageNumber="129" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="8090FD685209FFA2FCC0FC97FCF49E72" box="[862,883,865,891]" count="1" pageId="2" pageNumber="129" type="female">♀</specimenCount>
, ‘
<location id="9349603A5209FFA2FC1AFC96FC6F9E73" LSID="urn:lsid:plazi:treatment:1E3F87F75209FFA3FF09FE2CFB9B9CFC:9349603A5209FFA2FC1AFC96FC6F9E73" box="[900,1000,864,890]" country="Taiwan" name="Taihorin" pageId="2" pageNumber="129">Taihorin</location>
\
<collectingCountry id="EE8176715209FFA2FC62FC97FBD99E72" box="[1020,1118,865,891]" name="Taiwan" pageId="2" pageNumber="129">Formosa</collectingCountry>
\
<collectorName id="3B6353375209FFA2FBEFFC97FB5D9E73" box="[1137,1242,864,891]" pageId="2" pageNumber="129">H. Sauter</collectorName>
<date id="E22810215209FFA2FB41FC97FAB49E73" box="[1247,1331,864,891]" pageId="2" pageNumber="129" value="1911-05">
<collectingDate id="F26CE9C95209FFA2FB41FC97FAB49E73" box="[1247,1331,864,891]" pageId="2" pageNumber="129" value="1911-05">V 1911</collectingDate>
</date>
’ [printed, plus ‘VII’ by handwritten at right anterior part], ‘
<taxonomicName id="51964D625209FFA2FD55FC72FCEE9E97" authorityName="Fieber" authorityYear="1860" box="[715,873,900,926]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="134" phylum="Arthropoda" rank="genus">Cardiastethus</taxonomicName>
\ longiceps \ n. sp.’ [handwritten], ‘
<specimenCount id="8090FD685209FFA2FA96FC72FAA79E97" box="[1288,1312,900,926]" count="1" pageId="2" pageNumber="129" type="female">♀</specimenCount>
’, ‘
<determinerName id="63F5DA255209FFA2FAA3FC72FA1B9E97" box="[1341,1436,900,926]" pageId="2" pageNumber="129">Poppius</determinerName>
det.’, ‘
<collectorName id="3B6353375209FFA2FF43FC5EFED99ECB" box="[221,350,936,962]" pageId="2" pageNumber="129">Holotypus’</collectorName>
[red square], ‘TYPUS’ [light red square]; mounted on card, segments III–IV lacking; deposited in SDEI (
<figureCitation id="0EAD2A645209FFA2FE9FFC3AFEE89EEF" box="[257,367,972,998]" captionStart="FIGURES 4–6" captionStartId="2.[151,264,1746,1770]" captionTargetBox="[151,1387,1022,1723]" captionTargetId="figure-286@2.[151,1436,1022,1723]" captionTargetPageId="2" captionText="FIGURES 4–6. Holotype of Cardiastethus longiceps Poppius, 1915 and its labels. 4–5, dorsal and lateral views; 6, labels. Scales: 0.5 mm. (©SDEI, courtesy of J. F. Tsai and NMNS, published with permission.)" figureDoi="http://doi.org/10.5281/zenodo.10808129" httpUri="https://zenodo.org/record/10808129/files/figure.png" pageId="2" pageNumber="129">Figs. 4–6</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="C2E966695209FFA2FF09F924FB959A06" ID-DOI="http://doi.org/10.5281/zenodo.10808129" ID-Zenodo-Dep="10808129" httpUri="https://zenodo.org/record/10808129/files/figure.png" pageId="2" pageNumber="129" startId="2.[151,264,1746,1770]" targetBox="[151,1387,1022,1723]" targetPageId="2" targetType="figure">
<paragraph id="962936E15209FFA2FF09F924FB959A06" blockId="2.[151,1437,1746,1809]" pageId="2" pageNumber="129">
<emphasis id="A4E2EAF35209FFA2FF09F924FEBA9BE3" bold="true" box="[151,317,1746,1770]" pageId="2" pageNumber="129">FIGURES 4–6.</emphasis>
Holotype of
<taxonomicName id="51964D625209FFA2FE52F925FCE79BE3" authority="Poppius, 1915" authorityName="Poppius" authorityYear="1915" box="[460,864,1746,1771]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="A4E2EAF35209FFA2FE52F925FD449BE2" box="[460,707,1747,1771]" italics="true" pageId="2" pageNumber="129">Cardiastethus longiceps</emphasis>
<bibRefCitation id="F2074B105209FFA2FD52F925FCE79BE3" author="Poppius, B." box="[716,864,1746,1771]" pageId="2" pageNumber="129" pagination="1 - 80" refId="ref5294" refString="Poppius, B. (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8), 1 - 80. [1914, March 1915]" type="journal article" year="1915">Poppius, 1915</bibRefCitation>
</taxonomicName>
and its labels. 4–5, dorsal and lateral views; 6, labels. Scales: 0.5 mm. (©SDEI, courtesy of J. F. Tsai and NMNS, published with permission.)
</paragraph>
</caption>
<paragraph id="962936E15209FFA2FF59F8CBFEE89AEE" blockId="2.[151,1436,1852,2023]" pageId="2" pageNumber="129">
<emphasis id="A4E2EAF35209FFA2FF59F8CBFE619A5E" bold="true" box="[199,486,1852,1879]" pageId="2" pageNumber="129">
<taxonomicName id="51964D625209FFA2FF59F8CBFE589A5E" authorityName="Yamada &amp; Hirowatari" authorityYear="2004" box="[199,479,1852,1879]" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="A4E2EAF35209FFA2FF59F8CBFE589A5E" bold="true" box="[199,479,1852,1879]" italics="true" pageId="2" pageNumber="129">Lippomanus brevicornis</emphasis>
</taxonomicName>
.
</emphasis>
<materialsCitation id="26FE3CBC5209FFA2FE72F8CBFEC09A72" collectingDate="2002-03-26" collectorName="Iriomote Is. &amp; K. Yamada" location="Uehara" pageId="2" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="A4E2EAF35209FFA2FE72F8CBFDE49A5E" bold="true" box="[492,611,1853,1879]" pageId="2" pageNumber="129">
<typeStatus id="492D88435209FFA2FE72F8CBFDDA9A5E" box="[492,605,1853,1879]" pageId="2" pageNumber="129" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="8090FD685209FFA2FDF7F8CBFD079A5E" box="[617,640,1853,1879]" count="1" pageId="2" pageNumber="129" type="male">♂</specimenCount>
, ‘[Ryukyus] \
<location id="9349603A5209FFA2FC80F8CBFCE89A5E" LSID="urn:lsid:plazi:treatment:1E3F87F75209FFA3FF09FE2CFB9B9CFC:9349603A5209FFA2FC80F8CBFCE89A5E" box="[798,879,1853,1879]" name="Uehara" pageId="2" pageNumber="129" stateProvince="Okinawa">Uehara</location>
\
<collectorName id="3B6353375209FFA2FC1DF8CBFB829A5E" box="[899,1029,1853,1879]" pageId="2" pageNumber="129">Iriomote Is.</collectorName>
\
<collectingRegion id="5452F8035209FFA2FB87F8CBFB3C9A5E" box="[1049,1211,1853,1879]" country="Japan" name="Okinawa" pageId="2" pageNumber="129">Okinawa Pref.</collectingRegion>
\
<date id="E22810215209FFA2FB50F8CBFAD39A5E" box="[1230,1364,1853,1879]" pageId="2" pageNumber="129" value="2002-03-26">
<collectingDate id="F26CE9C95209FFA2FB50F8CBFAD39A5E" box="[1230,1364,1853,1879]" pageId="2" pageNumber="129" value="2002-03-26">26. iii. 2002</collectingDate>
</date>
L. T. \
<collectorName id="3B6353375209FFA2FF09F897FE959A72" box="[151,274,1889,1915]" pageId="2" pageNumber="129">K. Yamada</collectorName>
leg.
</materialsCitation>
’, ‘
<materialsCitation id="26FE3CBC5209FFA2FEC2F897FEEC9AEE" collectingDate="2009-03" collectionCode="OMU" pageId="2" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="492D88435209FFA2FEC2F897FE4E9A72" box="[348,457,1889,1915]" pageId="2" pageNumber="129" type="holotype">Holotype</typeStatus>
<specimenCount id="8090FD685209FFA2FE4EF897FE629A72" box="[464,485,1889,1915]" count="1" pageId="2" pageNumber="129" type="male">♂</specimenCount>
\
<taxonomicName id="51964D625209FFA2FE67F897FC8E9A72" authorityName="Yamada &amp; Hirowatari" authorityYear="2004" box="[505,777,1889,1915]" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="brevicornis">Lippomanus brevicornis</taxonomicName>
\ Yamada et Hirowatari, 2004’ [pink squre], ‘OPU-IN-HE \
<date id="E22810215209FFA2FF09F873FF6A9A96" box="[151,237,1925,1951]" pageId="2" pageNumber="129" value="2009-03">
<collectingDate id="F26CE9C95209FFA2FF09F873FF6A9A96" box="[151,237,1925,1951]" pageId="2" pageNumber="129" value="2009-03">2009III</collectingDate>
</date>
0003’ [with black frame]; mounted on triangle card, clavus of left hemelytron removed (glued on same card as a specimen), and abdominal segments VI–IX removed and preserved on slide (glass slide No. OPU-KY077); deposited in
<collectionCode id="F087AE245209FFA2FEB8F83BFEEC9AEE" box="[294,363,1997,2023]" pageId="2" pageNumber="129">OMU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DE8C656A5208FFA3FF59FF61FB9B9CFC" pageId="3" pageNumber="130" type="discussion">
<paragraph id="962936E15208FFA3FF59FF61FD1A9CD8" blockId="3.[151,1437,151,501]" pageId="3" pageNumber="130">
<emphasis id="A4E2EAF35208FFA3FF59FF61FEBC9DB8" bold="true" box="[199,315,151,177]" pageId="3" pageNumber="130">Remarks.</emphasis>
As for the previous species, no new information on the taxonomy and morphology of this species has been published since the original description. Based on the examination of superior digital images of the
<typeStatus id="492D88435208FFA3FAA4FF4DFA1B9DDC" box="[1338,1436,187,213]" pageId="3" pageNumber="130" type="holotype">holotype</typeStatus>
deposited in SDEI brought me to two conclusions. Firstly,
<taxonomicName id="51964D625208FFA3FC8FFF29FC1D9DF0" authorityName="Poppius" authorityYear="1915" box="[785,922,223,249]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="A4E2EAF35208FFA3FC8FFF29FC1D9DF0" box="[785,922,223,249]" italics="true" pageId="3" pageNumber="130">C. longiceps</emphasis>
</taxonomicName>
should be transferred to the genus
<taxonomicName id="51964D625208FFA3FA8CFF29FEA89C14" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="A4E2EAF35208FFA3FA8CFF29FA1B9DF1" box="[1298,1436,223,248]" italics="true" pageId="3" pageNumber="130">Lippomanus</emphasis>
Distant, 1904
</taxonomicName>
based on the following generic level diagnostic characters: antennal segment I not reaching apex of head; distinct setigerous punctures present on posterior half of pronotum, scutellum, and clavus; and lateral margins of hemelytra concavely sinuate on basal halves. Secondly,
<taxonomicName id="51964D625208FFA3FCB2FEBDFA059C6C" authority="Yamada &amp; Hirowatari, 2004" authorityName="Yamada &amp; Hirowatari" authorityYear="2004" box="[812,1410,331,357]" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="A4E2EAF35208FFA3FCB2FEBDFBBA9C6C" box="[812,1085,331,357]" italics="true" pageId="3" pageNumber="130">Lippomanus brevicornis</emphasis>
Yamada &amp; Hirowatari, 2004
</taxonomicName>
is confirmed to be a new subjective synonym of
<taxonomicName id="51964D625208FFA3FD02FE99FCA09C80" authorityName="Poppius" authorityYear="1915" box="[668,807,367,393]" class="Insecta" family="Anthocoridae" genus="Cardiastethus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="A4E2EAF35208FFA3FD02FE99FCA09C80" box="[668,807,367,393]" italics="true" pageId="3" pageNumber="130">C. longiceps</emphasis>
</taxonomicName>
based on the general coloration (body overall yellowish brown, with widely darkened pronotal posterior angles and widely darkened cuneus) and length of antennal segment II (much shorter than head width across eyes).
</paragraph>
<paragraph id="962936E15208FFA3FF59FE2CFB9B9CFC" blockId="3.[151,1437,151,501]" box="[199,1052,474,501]" pageId="3" pageNumber="130">
<emphasis id="A4E2EAF35208FFA3FF59FE2CFEFA9CFC" bold="true" box="[199,381,474,501]" pageId="3" pageNumber="130">Current status.</emphasis>
<taxonomicName id="51964D625208FFA3FE1AFE2DFCBB9CFC" authority="(Poppius, 1915)" authorityName="Yamada" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[388,828,474,501]" class="Insecta" family="Anthocoridae" genus="Lippomanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="longiceps" status="comb. nov.">
<emphasis id="A4E2EAF35208FFA3FE1AFE2DFD079CFC" box="[388,640,475,501]" italics="true" pageId="3" pageNumber="130">Lippomanus longiceps</emphasis>
(
<bibRefCitation id="F2074B105208FFA3FD11FE2DFCB39CFC" author="Poppius, B." box="[655,820,474,501]" pageId="3" pageNumber="130" pagination="1 - 80" refId="ref5294" refString="Poppius, B. (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8), 1 - 80. [1914, March 1915]" type="journal article" year="1915">Poppius, 1915</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="A4E2EAF35208FFA3FCD6FE2DFB929CFD" bold="true" box="[840,1045,474,501]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="BFD157885208FFA3FCD6FE2DFB929CFD" box="[840,1045,474,501]" pageId="3" pageNumber="130" rank="species">new combination</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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