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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.594.8768" ID-GBIF-Dataset="56393d29-ddfc-497f-be95-070616865240" ID-PMC="PMC4926701" ID-Pensoft-Pub="1313-2970-594-99" ID-PubMed="27408560" ID-ZBK="4D2882008FF2422697EC3984E8EF56DD" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-594-99" ModsDocOrigin="ZooKeys 594" ModsDocTitle="A review of the sarawakensis species group of the ground beetle genus Orthogonius MacLeay, 1825 (Coleoptera, Carabidae, Orthogoniini)" checkinTime="1464725703941" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2016" docId="1082268BB71849AD0EB1FE919A158ABB" docLanguage="en" docName="ZooKeys 594: 99-110" docOrigin="ZooKeys 594" docSource="http://dx.doi.org/10.3897/zookeys.594.8768" docTitle="Orthogonius sabahicus Tian &amp; Deuve, 2016, sp. n." docType="treatment" docUuid="C8EA2EDC-08D7-42E2-962E-347D379DCF29" docUuidSource="ZooBank" docVersion="5" lastPageNumber="106" masterDocId="37113C2EFFB2286AFFB3FFF6FFD9DD1C" masterDocTitle="A review of the sarawakensis species group of the ground beetle genus Orthogonius MacLeay, 1825 (Coleoptera, Carabidae, Orthogoniini)" masterLastPageNumber="110" masterPageNumber="99" pageNumber="104" updateTime="1668163168040" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the sarawakensis species group of the ground beetle genus Orthogonius MacLeay, 1825 (Coleoptera, Carabidae, Orthogoniini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deuve, Thierry</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>594</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>99</mods:start>
<mods:end>110</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.594.8768</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.594.8768</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-594-99</mods:identifier>
<mods:identifier type="ZBK">4D2882008FF2422697EC3984E8EF56DD</mods:identifier>
<mods:identifier type="ZooBank">4D2882008FF2422697EC3984E8EF56DD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127901854" LSID="urn:lsid:zoobank.org:act:C8EA2EDC-08D7-42E2-962E-347D379DCF29" httpUri="http://treatment.plazi.org/id/1082268BB71849AD0EB1FE919A158ABB" lastPageId="7" lastPageNumber="106" pageId="5" pageNumber="104">
<subSubSection pageId="5" pageNumber="104" type="multiple">
<paragraph pageId="5" pageNumber="104">
<pageBreakToken pageId="5" pageNumber="104" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Carabidae
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="104" type="nomenclature">
<paragraph pageId="5" pageNumber="104">
<taxonomicName LSID="http://zoobank.org/C8EA2EDC-08D7-42E2-962E-347D379DCF29" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius sabahicus" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="sabahicus">Orthogonius sabahicus</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="104">sp. n.</taxonomicNameLabel>
Fig. 5
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="105" type="materials_examined">
<paragraph pageId="6" pageNumber="105">
<pageBreakToken pageId="6" pageNumber="105" start="start">Holotype</pageBreakToken>
.
</paragraph>
<paragraph pageId="6" pageNumber="105">male, &quot;Malaisie, Sabah, Crocker Range, Avril 94, Chew&quot;, in MNHN; paratypes: 1 male, &quot;Malaysia, N. Borneo, Sabah, Keningau distr. Trus Madi Mt. 1150 m, 7.VII.2011, A. Klimenko legit&quot;, in CIB; 1 female, IBID, in SCAU.</paragraph>
<paragraph pageId="6" pageNumber="105">Length: 16.0-17.0 mm; width: 6.5-6.6 mm. Habitus as in Fig. 5A.</paragraph>
<caption pageId="6" pageNumber="105">
<paragraph pageId="6" pageNumber="105">
Figure 5.
<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius sabahicus" order="Coleoptera" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="sabahicus">Orthogonius sabahicus</taxonomicName>
sp. n. A habitus, male, holotype B median lobe, right lateral view C apical lamella, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="106" pageId="6" pageNumber="105" type="description">
<paragraph pageId="6" pageNumber="105">Description.</paragraph>
<paragraph pageId="6" pageNumber="105">Head dark brown, other parts reddish brown; smooth and glabrous, impunctate, elytral intervals covered with fine and sparse punctures; head and pronotum faintly wrinkled; microsculptural engraved meshes isodiametric on head, pronotum and elytra.</paragraph>
<paragraph pageId="6" pageNumber="105">
Head stout, slightly longer than wide, HL/HW = 1.05-1.10; vertex convex; labrum quite long, nearly as long as wide, widely and feebly emarginated at frontal margin, 6-setose; clypeus bisetose, base raised and sides longitudinal furrowed, joining to the frontal impressions; frontal impressions pit-like; eyes moderately developed and prominent; palps slender, subcylindrical, the 3rd maxillary palpomere as long as the 4th, both glabrous, with two short setae at apex of the 3rd; the 2nd labial palpomere slightly longer than the 3rd, bisetose on inner margin, the 3rd faintly and sparsely setose; ligula narrow, bisetose at apex; mentum and palpigers unsetose, submentum bisetose; antennae extending beyond basal 1/5 of elytra, pubescent from the 4th antennomere, starting from basal 1/4; the 1
<normalizedToken originalValue="st">st-</normalizedToken>
3rd antennomeres and basal 1/4 of the 4th antennomere glabrous; the 1st antennomere stouter than other, with a long seta at subapex, several shorter setae present at apexes of each joint from the 2nd antennomere; the 1st antennomere less than twice as long as the 2nd, and slightly longer than the 3rd.
</paragraph>
<paragraph pageId="6" pageNumber="105">Pronotum transverse, notably wider than long, PW/PL = 1.66-1.68, widest at about middle; lateral expanded margins flat, even, feebly reflexed throughout; front and base well bordered; fore transversal impression faint, basal one evident; basal foveae small; disc markedly convex; front margin nearly straight at middle, basal margin feebly bisinuate; hind and front angles completely rounded off; base slightly shorter than front.</paragraph>
<paragraph pageId="6" pageNumber="105">
Elytra elongate-ovate, distinctly longer than wide, EL/EW = 1.61-1.63; nearly parallel-sided, widest at middle; basal border interrupted against the 1
<normalizedToken originalValue="st">st-</normalizedToken>
3rd intervals; humeri rather square; apex roundly truncate, without outer angles, inner angle broadly blunt; disc convex; striae finely and deeply striate; intervals convex, subequal in width; the 3rd interval with median and posterior setiferous pores (anterior one wanted), both are located on middle of the interval.
</paragraph>
<paragraph pageId="6" pageNumber="105">Femora moderately expanded, hind femur bisetose posteriorly; for tibia dilated at apex, apical margin obtusely truncate or sinuate towards outer angle, outer margin notably serrate; middle tibia slightly curved; hind legs slender, apical spurs long and sharp, the 1st and 3rd tarsomeres much longer than the 2nd and 4th respectively, the 4th tarsomere deeply emarginated at apex, with lobes about half as long as the joint; fore and middle tarsal claws weakly pectinate, hind claws simple.</paragraph>
<paragraph pageId="7" pageNumber="106">
<pageBreakToken pageId="7" pageNumber="106" start="start">Prosternal</pageBreakToken>
process bordered at apex; abdominal ventrite VII finely and shallowly emarginate at apical margin in male.
</paragraph>
<paragraph pageId="7" pageNumber="106">Male genitalia (Fig. 5B, C): Stout and robust, apex suddenly narrowed and pointed in lateral view; the apical lamella short, wider than long.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="106" type="remarks">
<paragraph pageId="7" pageNumber="106">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="106">It is a very peculiar species, differing from other congeners by its mentum asetose, elytral base incompletely bordered, anterior dorsal pores missed on the 3rd interval, and hind tarsal claws simple.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="106" type="etymology">
<paragraph pageId="7" pageNumber="106">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="106">To refer to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="106" type="distribution">
<paragraph pageId="7" pageNumber="106">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="106">Malaysia (Sabah).</paragraph>
</subSubSection>
</treatment>
</document>