449 lines
88 KiB
XML
449 lines
88 KiB
XML
<document id="9C00F1232AF40391BA0454706895CC8B" ID-DOI="10.1649/0010-065X-71.2.289" ID-ISSN="1938-4394" ID-Zenodo-Dep="10109515" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630240527336" checkinUser="diego" docAuthor="Sokolov, Igor M., Carlton, Christopher E., Watrous, Larry E. & Robison, Henry W." docDate="2017" docId="10340D1FFFC2933AFD378C46FE6F286D" docLanguage="en" docName="0010-065X-71.2.289.pdf" docOrigin="The Coleopterists Bulletin 71 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-71.2.289" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Anillinus alleni SOKOLOV AND CARLTON 2017, new species" docType="treatment" docVersion="3" lastPageNumber="295" masterDocId="EC0D7567FFC0933DFFD68859FFD22D20" masterDocTitle="Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a New Species from the Ozark Interior Highlands of Arkansas, USA" masterLastPageNumber="297" masterPageNumber="289" pageNumber="290" updateTime="1699664608065" updateUser="ExternalLinkService">
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<mods:titleInfo id="B7E594AF77B6926B776E29B90DB7464F">
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<mods:title id="DF0A1299CAD89471837010D548C0FA60">Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a New Species from the Ozark Interior Highlands of Arkansas, USA</mods:title>
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<mods:name id="6527E26F96127BDC2CCF18F585B79BF7" type="personal">
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<mods:namePart id="BE10F1D2826A2A20A37E2FF2BA23B139">Sokolov, Igor M.</mods:namePart>
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<mods:roleTerm id="50E00146EAAD3D0B72E6A0C7F6105174">Author</mods:roleTerm>
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<mods:namePart id="3FFDB91098C80D1C028738DBD7E1BC31">Carlton, Christopher E.</mods:namePart>
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</mods:name>
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<mods:name id="E8ECC1A6CCD3132D65BC987576B90B76" type="personal">
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<mods:roleTerm id="D0E5DDE916A4A91291036C39EB6331E4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="000FEC469A55DD9E3DC87E481080B07A">Watrous, Larry E.</mods:namePart>
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</mods:name>
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<mods:name id="F32581E67EA0F472D2C565FF49F6EB68" type="personal">
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<mods:roleTerm id="30536791F3BCCEF70502FA175D729FF6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2B4E286D475F2C8BEC081F2E2B9FA947">Robison, Henry W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B929293F6ABD4FC26E2EC8D4CBDDE171">text</mods:typeOfResource>
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<mods:title id="C990055E99F1A1D834BDEC349AE42AD5">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="3AE692B1ABBBC56F5315DF68434A4780">
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<mods:date id="651F4FF7E79E1B46B066664825952D5E">2017</mods:date>
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<mods:detail id="0FD11BEEF2D8460D57DA8AAF86EF8185" type="pubDate">
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<mods:number id="AF77ECA61CA49FF8A02F922E2BF15274">2017-06-27</mods:number>
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<mods:number id="393AFCF66416A1D454620C5336C92150">71</mods:number>
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<mods:detail id="717B6F554FD8F28B674852AB0722FC2F" type="issue">
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<mods:number id="9977656A1275E446A096899A9B58D393">2</mods:number>
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<mods:url id="E70487E7DBDFD6DC6E7D984556E461A5">http://dx.doi.org/10.1649/0010-065x-71.2.289</mods:url>
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<mods:classification id="94272F9CBDEE93180E894E21985DAC1E">journal article</mods:classification>
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<mods:identifier id="9ED8B7C1A56198392AD384E6294ACDC0" type="DOI">10.1649/0010-065X-71.2.289</mods:identifier>
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<mods:identifier id="B3AFB5E7A32AA8CA00B648DE4ABB27DE" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="60AED9A52E69AA73DAC7C3F792B9478D" type="ZooBank">7E7DD012-1930-4140-9948-FDF3FC58308A</mods:identifier>
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</mods:mods>
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<treatment id="10340D1FFFC2933AFD378C46FE6F286D" LSID="urn:lsid:plazi:treatment:10340D1FFFC2933AFD378C46FE6F286D" httpUri="http://treatment.plazi.org/id/10340D1FFFC2933AFD378C46FE6F286D" lastPageId="7" lastPageNumber="295" pageId="2" pageNumber="290">
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<subSubSection id="D087EF82FFC2933FFD378C46FC09296F" pageId="2" pageNumber="290" type="nomenclature">
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<paragraph id="9822BC09FFC2933FFD378C46FC09296F" blockId="2.[737,1120,1055,1103]" pageId="2" pageNumber="290">
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<heading id="C36A0B65FFC2933FFD378C46FC09296F" centered="true" fontSize="9" level="2" pageId="2" pageNumber="290" reason="2">
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<emphasis id="AAE9601BFFC2933FFD378C46FC09296F" bold="true" pageId="2" pageNumber="290">
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<taxonomicName id="5F9DC78AFFC2933FFD378C46FB892914" ID-CoL="8MG29" authority="Sokolov and Carlton" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[737,1115,1055,1077]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="290" phylum="Arthropoda" rank="species" species="alleni" status="sp. nov.">
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<emphasis id="AAE9601BFFC2933FFD378C46FCAF2915" bold="true" box="[737,893,1055,1077]" italics="true" pageId="2" pageNumber="290">Anillinus alleni</emphasis>
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Sokolov and Carlton
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</taxonomicName>
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,
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<taxonomicNameLabel id="B1DADD60FFC2933FFCB08C63FC09296F" box="[870,987,1082,1103]" pageId="2" pageNumber="290" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D087EF82FFC2933FFD138C3DFCC12898" pageId="2" pageNumber="290" type="materials_examined">
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<paragraph id="9822BC09FFC2933FFD138C3DFC5729DE" blockId="2.[685,1172,1124,1730]" pageId="2" pageNumber="290">
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<emphasis id="AAE9601BFFC2933FFD138C3DFC882959" bold="true" box="[709,858,1124,1145]" pageId="2" pageNumber="290">Type Material.</emphasis>
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<materialsCitation id="28F5B654FFC2933FFCB68C3DFC5729DE" collectingDate="2014-03-18" collectingDateMax="2014-03-19" collectingDateMin="2014-03-18" collectorName="L. Watrous & H. Robison" country="United States of America" county="Stone Co." latitude="35.956" location="Smithsonian Institution" longLatPrecision="6" longitude="-92.1778" municipality="National Collection" pageId="2" pageNumber="290" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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<typeStatus id="472602ABFFC2933FFCB68C3DFC322959" box="[864,992,1124,1145]" pageId="2" pageNumber="290" type="holotype">HOLOTYPE</typeStatus>
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, male, deposited in
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<collectingCountry id="E08AFC99FFC2933FFD7B8C26FD0A29B4" box="[685,728,1151,1172]" name="United States of America" pageId="2" pageNumber="290">U.S.</collectingCountry>
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<collectingMunicipality id="78462673FFC2933FFD378C26FC7229B4" box="[737,928,1151,1172]" pageId="2" pageNumber="290">National Collection</collectingMunicipality>
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,
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<location id="9D42EAD2FFC2933FFC7B8C26FB4329B4" LSID="urn:lsid:plazi:treatment:10340D1FFFC2933AFD378C46FE6F286D:9D42EAD2FFC2933FFC7B8C26FB4329B4" box="[941,1169,1151,1172]" country="United States of America" county="Stone Co." latitude="35.956" longLatPrecision="6" longitude="-92.1778" municipality="National Collection" name="Smithsonian Institution" pageId="2" pageNumber="290" stateProvince="Washington">Smithsonian Institution</location>
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,
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<collectingRegion id="5A5972EBFFC2933FFD7B8CC3FCF3298F" box="[685,801,1178,1199]" country="United States of America" name="Washington" pageId="2" pageNumber="290">Washington</collectingRegion>
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, DC, glued on board, dissected, labeled: \
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<collectingCountry id="E08AFC99FFC2933FFD2F8CEDFCF929E9" box="[761,811,1204,1225]" name="United States of America" pageId="2" pageNumber="290">USA</collectingCountry>
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: AR:
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<collectingCounty id="7143C485FFC2933FFCBF8CEDFC1F29E9" box="[873,973,1204,1225]" pageId="2" pageNumber="290">Stone Co.</collectingCounty>
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,
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<location id="9D42EAD2FFC2933FFC0C8CEDFB4229E9" LSID="urn:lsid:plazi:treatment:10340D1FFFC2933AFD378C46FE6F286D:9D42EAD2FFC2933FFC0C8CEDFB4229E9" box="[986,1168,1204,1225]" country="United States of America" county="Stone Co." latitude="35.956" longLatPrecision="6" longitude="-92.1778" municipality="National Collection" name="Blanchard Springs" pageId="2" pageNumber="290" stateProvince="Washington">Blanchard Springs</location>
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,
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<geoCoordinate id="FDA9DACEFFC2933FFD7B8C96FD2929C4" box="[685,763,1231,1252]" degrees="35.9560" direction="north" orientation="latitude" pageId="2" pageNumber="290" precision="5" value="35.956">35.9560</geoCoordinate>
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<geoCoordinate id="FDA9DACEFFC2933FFCDC8C96FCB029C4" box="[778,866,1231,1252]" degrees="92.1778" direction="west" orientation="longitude" pageId="2" pageNumber="290" precision="5" value="-92.1778">-92.1778</geoCoordinate>
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,
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<date id="EC239AC9FFC2933FFCA38C96FBC129C4" box="[885,1043,1231,1252]" pageId="2" pageNumber="290" value="2014-03-18" valueMax="2014-03-19" valueMin="2014-03-18">
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<collectingDate id="FC676321FFC2933FFCA38C96FBC129C4" box="[885,1043,1231,1252]" pageId="2" pageNumber="290" value="2014-03-18" valueMax="2014-03-19" valueMin="2014-03-18">18-19 III 2014</collectingDate>
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</date>
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ex. soil
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<collectorName id="3568D9DFFFC2933FFBA98C96FD2C29DE" pageId="2" pageNumber="290">L. Watrous</collectorName>
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,
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<collectorName id="3568D9DFFFC2933FFCDC8CB0FCA829DE" box="[778,890,1257,1278]" pageId="2" pageNumber="290">H. Robison</collectorName>
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\.
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</materialsCitation>
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</paragraph>
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<paragraph id="9822BC09FFC2933FFD138D5DFCDA2849" blockId="2.[685,1172,1124,1730]" pageId="2" pageNumber="290">
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<materialsCitation id="28F5B654FFC2933FFD138D5DFCD12849" collectorName="Baton Rouge, LA" location="Arthropod Museum" pageId="2" pageNumber="290" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Louisiana" typeStatus="paratype">
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<typeStatus id="472602ABFFC2933FFD138D5DFC9E2839" box="[709,844,1284,1305]" pageId="2" pageNumber="290" type="paratype">PARATYPES</typeStatus>
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:
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||
<specimenCount id="8E9B7780FFC2933FFC8E8D5DFC492839" box="[856,923,1284,1305]" pageId="2" pageNumber="290" type="male">1 male</specimenCount>
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and
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<specimenCount id="8E9B7780FFC2933FFC068D5DFBFC2839" box="[976,1070,1284,1305]" pageId="2" pageNumber="290" type="female">2 females</specimenCount>
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||
(
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<specimenCount id="8E9B7780FFC2933FFB968D5DFB462839" box="[1088,1172,1284,1305]" pageId="2" pageNumber="290" type="female">1 female</specimenCount>
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dissected), deposited in
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<collectingRegion id="5A5972EBFFC2933FFC438D47FBFB2813" box="[917,1065,1310,1331]" country="United States of America" name="Louisiana" pageId="2" pageNumber="290">Louisiana State</collectingRegion>
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<location id="9D42EAD2FFC2933FFBE78D47FCD1286E" LSID="urn:lsid:plazi:treatment:10340D1FFFC2933AFD378C46FE6F286D:9D42EAD2FFC2933FFBE78D47FCD1286E" name="Arthropod Museum" pageId="2" pageNumber="290" stateProvince="Louisiana">Arthropod Museum</location>
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,
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<collectorName id="3568D9DFFFC2933FFCC18D60FC04286E" box="[791,982,1337,1358]" pageId="2" pageNumber="290">Baton Rouge, LA</collectorName>
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, labeled same as
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<typeStatus id="472602ABFFC2933FFD7B8D0DFCD12849" box="[685,771,1364,1385]" pageId="2" pageNumber="290" type="holotype">holotype</typeStatus>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="9822BC09FFC2933FFD138D37FCC12898" blockId="2.[685,1172,1124,1730]" pageId="2" pageNumber="290">
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<emphasis id="AAE9601BFFC2933FFD138D37FC4A28A3" bold="true" box="[709,920,1390,1411]" pageId="2" pageNumber="290">
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Type Locality.
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<collectingCountry id="E08AFC99FFC2933FFCB48D37FC4728A3" box="[866,917,1390,1411]" name="United States of America" pageId="2" pageNumber="290">USA</collectingCountry>
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:
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</emphasis>
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<collectingRegion id="5A5972EBFFC2933FFC748D37FC2F28A3" box="[930,1021,1390,1411]" country="United States of America" name="Arkansas" pageId="2" pageNumber="290">Arkansas</collectingRegion>
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, Stone County, Blanchard Springs, latitude 35.9560°, longitude -92.1778°.
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</paragraph>
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</subSubSection>
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<subSubSection id="D087EF82FFC2933EFD138DE7FD5528CB" lastPageId="3" lastPageNumber="291" pageId="2" pageNumber="290" type="etymology">
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<paragraph id="9822BC09FFC2933FFD138DE7FBF02B52" blockId="2.[685,1172,1124,1730]" pageId="2" pageNumber="290">
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<emphasis id="AAE9601BFFC2933FFD138DE7FCEA28F3" bold="true" box="[709,824,1470,1491]" pageId="2" pageNumber="290">Etymology.</emphasis>
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The specific epithet is a Latinized eponym in the genitive case. It is based on the surname of Robert T. (Tommy) Allen (1939–2016), the second author’ s graduate advisor, in honor of his contributions to knowledge of the biota of the Interior Highlands and for setting the authors on the path to rediscover this elusive species.
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</paragraph>
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<paragraph id="9822BC09FFC2933EFD138E21FD5528CB" blockId="2.[685,1172,1124,1730]" lastBlockId="3.[173,660,1413,1730]" lastPageId="3" lastPageNumber="291" pageId="2" pageNumber="290">
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<emphasis id="AAE9601BFFC2933FFD138E21FC962BAD" bold="true" box="[709,836,1656,1677]" pageId="2" pageNumber="290">Recognition.</emphasis>
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Adults of this new species are distinguished easily from those of other species of the genus by the structure of the hind legs, specifically the modified metafemora in both sexes (
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<figureCitation id="00A6A08CFFC3933EFF628DF9FEF92895" box="[180,299,1440,1461]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="3" pageNumber="291">Fig. 1G, H</figureCitation>
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), especially the extremely elongate spinose processes of the metafemora of the males, and the male metatibiae covered with long setae.
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</paragraph>
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</subSubSection>
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||
<caption id="CCE2EC81FFC3933EFF138C17FDFF2870" ID-DOI="http://doi.org/10.5281/zenodo.10109517" ID-Zenodo-Dep="10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="3" pageNumber="291" startId="3.[197,228,1102,1121]" targetBox="[173,1194,195,1086]" targetPageId="3">
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<paragraph id="9822BC09FFC3933EFF138C17FDFF2870" blockId="3.[173,1194,1102,1360]" pageId="3" pageNumber="291">
|
||
<emphasis id="AAE9601BFFC3933EFF138C17FF2C2941" bold="true" box="[197,254,1102,1121]" pageId="3" pageNumber="291">Fig. 1.</emphasis>
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SEM images of body parts of
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<taxonomicName id="5F9DC78AFFC3933EFDCF8C17FD4E2941" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[537,668,1102,1121]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="species" species="alleni">
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<emphasis id="AAE9601BFFC3933EFDCF8C17FD4E2941" box="[537,668,1102,1121]" italics="true" pageId="3" pageNumber="291">Anillinus alleni</emphasis>
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||
</taxonomicName>
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. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1
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<superScript id="6FE81141FFC3933EFD0F8CC8FD3129BD" attach="none" box="[729,739,1169,1181]" fontSize="5" pageId="3" pageNumber="291">st</superScript>
|
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, 2
|
||
<superScript id="6FE81141FFC3933EFD2A8CC8FCDE29BD" attach="none" box="[764,780,1169,1181]" fontSize="5" pageId="3" pageNumber="291">nd</superScript>
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||
, and 3
|
||
<superScript id="6FE81141FFC3933EFC9A8CC8FC8B29BD" attach="left" box="[844,857,1169,1181]" fontSize="5" pageId="3" pageNumber="291">rd</superScript>
|
||
fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D087EF82FFC3933AFF138DA8FE6F286D" lastPageId="7" lastPageNumber="295" pageId="3" pageNumber="291" type="description">
|
||
<paragraph id="9822BC09FFC3933BFF138DA8FC4528A3" blockId="3.[173,660,1413,1730]" lastBlockId="6.[685,1172,1257,1730]" lastPageId="6" lastPageNumber="294" pageId="3" pageNumber="291">
|
||
<emphasis id="AAE9601BFFC3933EFF138DA8FE9A2B26" bold="true" box="[197,328,1521,1542]" pageId="3" pageNumber="291">Description.</emphasis>
|
||
Large-sized for genus (SBL
|
||
<quantity id="5F6511ECFFC3933EFF7B8E55FEED2B01" box="[173,319,1547,1569]" metricMagnitude="-3" metricUnit="m" metricValue="2.505" metricValueMax="2.51" metricValueMin="2.5" pageId="3" pageNumber="291" unit="mm" value="2.505" valueMax="2.51" valueMin="2.5">2.50–2.51 mm</quantity>
|
||
, mean 2.51±
|
||
<quantity id="5F6511ECFFC3933EFE1D8E55FDE12B01" box="[459,563,1548,1569]" metricMagnitude="-5" metricUnit="m" metricValue="1.0999999999999999" pageId="3" pageNumber="291" unit="mm" value="0.011">0.011 mm</quantity>
|
||
,
|
||
<emphasis id="AAE9601BFFC3933EFD948E55FD9C2B01" box="[578,590,1548,1569]" italics="true" pageId="3" pageNumber="291">n</emphasis>
|
||
= 2).
|
||
<emphasis id="AAE9601BFFC3933EFF7B8E7FFED62B1B" bold="true" box="[173,260,1574,1595]" pageId="3" pageNumber="291">Habitus:</emphasis>
|
||
Body form (
|
||
<figureCitation id="00A6A08CFFC3933EFE778E7FFE242B1B" box="[417,502,1574,1595]" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="3" pageNumber="291">Fig. 3A</figureCitation>
|
||
) subdepressed, slightly ovate, elongate (WE/SBL 0.37±0.005), head normally proportioned for genus (WH/WPm 0.74±0.054), pronotum moderately narrow in comparison to elytra (WPm/WE 0.77±0.039).
|
||
<emphasis id="AAE9601BFFC3933EFF7B8EF4FEF62BE2" bold="true" box="[173,292,1709,1730]" pageId="3" pageNumber="291">Integument:</emphasis>
|
||
Body color brunneo-rufous, appendages testaceous. Microsculpture absent from clypeus and vertex, obliterated on pronotal disc, distinct over frontal area of head, sides and base of pronotum, and elytra. Proepisternum (pes) with distinct microsculpture, prosternum (ps) smooth (
|
||
<figureCitation id="00A6A08CFFC3933EFBC18DA8FBB32B26" box="[1047,1121,1521,1542]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="3" pageNumber="291">Fig. 1B</figureCitation>
|
||
). Body surface shiny, surface sparsely and finely punctate, covered with sparse, yellowish, short setae. Vestiture of elytra moderately long (0.3–0.4 length of discal setae).
|
||
<emphasis id="AAE9601BFFC3933EFC828E05FC072B51" bold="true" box="[852,981,1628,1649]" pageId="3" pageNumber="291">Fixed setae:</emphasis>
|
||
Primary head setae typical of
|
||
<taxonomicName id="5F9DC78AFFC3933EFCFA8E2EFCA92BAC" authorityName="Jeannel" authorityYear="1937" box="[812,891,1655,1676]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="291" phylum="Arthropoda" rank="subTribe" subTribe="Anillina">Anillina</taxonomicName>
|
||
and include a pair of clypeal (cs), a pair of frontal (fs), and 2 pairs of supraorbital (ssa and ssp) setae (
|
||
<figureCitation id="00A6A08CFFC3933EFC578EF4FC1F2BE2" box="[897,973,1709,1730]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="3" pageNumber="291">Fig. 1A</figureCitation>
|
||
). Mentum with 2 pairs of long primary (paramedial and lateral) setae and 2 shorter paramedial setulae (
|
||
<figureCitation id="00A6A08CFFC49339FE718B9CFE272EFA" box="[423,501,965,986]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1D</figureCitation>
|
||
). Submentum with 2 pairs of long primary setae in 2 rows (paramedial and lateral) and few additional shorter setulae (
|
||
<figureCitation id="00A6A08CFFC49339FF3C8C4CFEEB290A" box="[234,313,1045,1066]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1D</figureCitation>
|
||
). Pronotum with 2 long primary lateral setae (midlateral, ls, and basilateral, bs) on each side (
|
||
<figureCitation id="00A6A08CFFC49339FF2B8C13FE95297F" box="[253,327,1098,1119]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1B</figureCitation>
|
||
). Elytra with 3 discal setae (
|
||
<figureCitation id="00A6A08CFFC49339FD808C13FF672959" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1C</figureCitation>
|
||
, ed3-4, 6), with scutellar (ed2) and apical (ed8) setae. Last 2 (8
|
||
<superScript id="6FE81141FFC49339FEE68C23FEEC29A8" attach="left" box="[304,318,1146,1160]" fontSize="6" pageId="4" pageNumber="292">th</superScript>
|
||
and 9
|
||
<superScript id="6FE81141FFC49339FEA88C23FE5E29A8" attach="right" box="[382,396,1146,1160]" fontSize="6" pageId="4" pageNumber="292">th</superScript>
|
||
) pores (eo8 and eo9) of umbilicate series much closer to each other than 7
|
||
<superScript id="6FE81141FFC49339FDB98CCDFDAF2982" attach="left" box="[623,637,1172,1186]" fontSize="6" pageId="4" pageNumber="292">th</superScript>
|
||
(eo7) to 8
|
||
<superScript id="6FE81141FFC49339FF238CF6FED1299D" attach="left" box="[245,259,1199,1213]" fontSize="6" pageId="4" pageNumber="292">th</superScript>
|
||
(
|
||
<figureCitation id="00A6A08CFFC49339FEC68CEDFE8929E9" box="[272,347,1204,1225]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1C</figureCitation>
|
||
). Fifth visible sternite of male with 2 setae and of female with 4 setae along posterior margin.
|
||
<emphasis id="AAE9601BFFC49339FEEB8CB0FEA829DE" bold="true" box="[317,378,1257,1278]" pageId="4" pageNumber="292">Head:</emphasis>
|
||
Anterior margin of clypeus (cl) straight (
|
||
<figureCitation id="00A6A08CFFC49339FEC38D5DFEB02839" box="[277,354,1284,1305]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1A</figureCitation>
|
||
). Frontal area with subtle tubercle (ft) medially near frontoclypeal suture. Fronto-lateral carinae distinct and long. Labrum (l) transverse with concave anterior margin with 6 setae apically, increasing in size from central pair outwards. Labium (
|
||
<figureCitation id="00A6A08CFFC49339FEF98DD0FEA428BE" box="[303,374,1417,1438]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1D</figureCitation>
|
||
) with mental tooth; mentum (m) and submentum (sm) split, with distinct mentalsubmental suture (ms). Glossal sclerite (gsc) with distinct paraglossae (pg) laterally and 2 setae medially.
|
||
<emphasis id="AAE9601BFFC49339FF0B8DAAFE982B28" bold="true" box="[221,330,1523,1544]" pageId="4" pageNumber="292">Prothorax:</emphasis>
|
||
Pronotum (
|
||
<figureCitation id="00A6A08CFFC49339FE108DAAFDC02B28" box="[454,530,1523,1544]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1B</figureCitation>
|
||
) relatively small (LP/LE 0.36±0.019) and slightly transverse for genus (WPm/LP 1.30±0.084), with lateral margins shallowly sinuate and moderately constricted posteriorly (WPm/WPp 1.29±0.020). Anterior angles indistinct, posterior angles nearly rectangular (90–100°). Width between posterior angles much greater than between anterior angles (WPa/WPp 0.91±0.017). Basal margin straight. Prosternum (
|
||
<figureCitation id="00A6A08CFFC49339FCF88B9CFCAF2EFA" box="[814,893,965,986]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1E</figureCitation>
|
||
, ps) slightly protruding on anterior margin medially, there with a group of longer setae relative to other prosternal vestiture, without long ambulatory setae at middle of sclerite. Prosternal intercoxal process (psp) unmargined, slightly dilated apically, and semicircularly rounded at apex, with a row of sparse setulae along midline.
|
||
<emphasis id="AAE9601BFFC49339FD7B8C26FCC829B4" bold="true" box="[685,794,1151,1172]" pageId="4" pageNumber="292">Scutellum:</emphasis>
|
||
Externally visible, widely triangular, with pointed apex (
|
||
<figureCitation id="00A6A08CFFC49339FCA68CC3FC6F298F" box="[880,957,1178,1199]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1C</figureCitation>
|
||
, sct).
|
||
<emphasis id="AAE9601BFFC49339FC2B8CC3FB97298F" bold="true" box="[1021,1093,1178,1199]" pageId="4" pageNumber="292">Elytra:</emphasis>
|
||
Widely depressed along suture, comparatively long (LE/ SBL 0.61±0.018) and narrow for genus (WE/LE 0.61±0.023), with traces of 4–5 striae (
|
||
<figureCitation id="00A6A08CFFC49339FBEC8CB0FB5529DE" box="[1082,1159,1257,1278]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1C</figureCitation>
|
||
). Humeri indistinct, markedly rounded, in outline forming obtuse angle with longitudinal axis of body. Lateral margins subparallel, slightly divergent at basal fifth, evenly rounded to apex in apical third, without subapical sinuation. Basal margination distinct (
|
||
<figureCitation id="00A6A08CFFC49339FD2B8DD0FCB328BE" box="[765,865,1416,1438]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1B–C</figureCitation>
|
||
, bm).
|
||
<emphasis id="AAE9601BFFC49339FC4C8DD0FBCB28BE" bold="true" box="[922,1049,1417,1438]" pageId="4" pageNumber="292">Pterothorax:</emphasis>
|
||
Metaventrite (mtv) short (
|
||
<figureCitation id="00A6A08CFFC49339FCFE8DFAFCA22898" box="[808,880,1443,1464]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="4" pageNumber="292">Fig. 1F</figureCitation>
|
||
), distance between meso- and metacoxae (mtvx) about diameter of mesocoxa (mscx). Metendoventrite linear without lateral arms.
|
||
<emphasis id="AAE9601BFFC49339FD7B8DAAFD312B28" bold="true" box="[685,739,1523,1544]" pageId="4" pageNumber="292">Legs:</emphasis>
|
||
Legs of moderate length, not elongate. Prothoracic legs of males with first tarsomere (ta1) markedly dilated apico-laterally with 2 rows of oval articulo-setae (as) (
|
||
<bibRefCitation id="FC0CC1F8FFC49339FCB08E1AFC062B78" author="Stork, N. E." box="[870,980,1603,1624]" pageId="4" pageNumber="292" pagination="175 - 306" refId="ref6057" refString="Stork, N. E. 1980. A scanning electron microscope study of tarsal adhesive setae in the Coleoptera. Zoological Journal of the Linnean Society 68: 175 - 306." type="journal article" year="1980">Stork 1980</bibRefCitation>
|
||
) on ventral surface (
|
||
<figureCitation id="00A6A08CFFC49339FD628E04FCD52B52" box="[692,775,1629,1650]" captionStart="Fig" captionStartId="4.[175,206,746,765]" captionTargetBox="[151,1172,194,730]" captionTargetId="figure-467@4.[151,1172,194,730]" captionTargetPageId="4" captionText="Fig. 2. SEM illustrations of structural features of the legs of Anillinus alleni. A) Left foreleg, male, dorsal view, B) Right mesotibia, female, ventral view, C) Left metatibia, male, ventro-lateral view, D) Left metatibia, female, medial view. as = adhesive seta; msb = mesotibial brush; msms = mesotibial modified seta; mss = mesotibial spur; mtb = metatibial brush; mts = metatibial spur; mtts = metatibial modified setae; ta1-ta3 = protarsomeres 1–3; tbn = tibial notch. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10109519" httpUri="https://zenodo.org/record/10109519/files/figure.png" pageId="4" pageNumber="292">Fig. 2A</figureCitation>
|
||
). Mesotibiae (
|
||
<figureCitation id="00A6A08CFFC49339FC4C8E04FC382B52" box="[922,1002,1629,1650]" captionStart="Fig" captionStartId="4.[175,206,746,765]" captionTargetBox="[151,1172,194,730]" captionTargetId="figure-467@4.[151,1172,194,730]" captionTargetPageId="4" captionText="Fig. 2. SEM illustrations of structural features of the legs of Anillinus alleni. A) Left foreleg, male, dorsal view, B) Right mesotibia, female, ventral view, C) Left metatibia, male, ventro-lateral view, D) Left metatibia, female, medial view. as = adhesive seta; msb = mesotibial brush; msms = mesotibial modified seta; mss = mesotibial spur; mtb = metatibial brush; mts = metatibial spur; mtts = metatibial modified setae; ta1-ta3 = protarsomeres 1–3; tbn = tibial notch. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10109519" httpUri="https://zenodo.org/record/10109519/files/figure.png" pageId="4" pageNumber="292">Fig. 2B</figureCitation>
|
||
) with 1 row of modified posterodorsal setae (msms) at apical twothirds, with 2 terminal spurs (mss) and tibial brush (msb). Metafemora (mtf) modified in both genders: strongly curved with long, stout spine (sp) at apical third in males (
|
||
<figureCitation id="00A6A08CFFC59338FE978E52FE592B00" box="[321,395,1547,1568]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="5" pageNumber="293">Fig. 1G</figureCitation>
|
||
) and of normal shape with triangular enlargement at apical half in females (
|
||
<figureCitation id="00A6A08CFFC59338FDBB8E7FFF1E2B76" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="5" pageNumber="293">Fig. 1H</figureCitation>
|
||
). Metatibiae with modified hairy posterodorsal setae (mtms) in apical two-thirds in males (
|
||
<figureCitation id="00A6A08CFFC59338FD928E05FD5E2B51" box="[580,652,1628,1649]" captionStart="Fig" captionStartId="4.[175,206,746,765]" captionTargetBox="[151,1172,194,730]" captionTargetId="figure-467@4.[151,1172,194,730]" captionTargetPageId="4" captionText="Fig. 2. SEM illustrations of structural features of the legs of Anillinus alleni. A) Left foreleg, male, dorsal view, B) Right mesotibia, female, ventral view, C) Left metatibia, male, ventro-lateral view, D) Left metatibia, female, medial view. as = adhesive seta; msb = mesotibial brush; msms = mesotibial modified seta; mss = mesotibial spur; mtb = metatibial brush; mts = metatibial spur; mtts = metatibial modified setae; ta1-ta3 = protarsomeres 1–3; tbn = tibial notch. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10109519" httpUri="https://zenodo.org/record/10109519/files/figure.png" pageId="5" pageNumber="293">Fig. 2C</figureCitation>
|
||
) and without modified setae in females (
|
||
<figureCitation id="00A6A08CFFC59338FDEE8E2EFD552BAC" box="[568,647,1655,1676]" captionStart="Fig" captionStartId="4.[175,206,746,765]" captionTargetBox="[151,1172,194,730]" captionTargetId="figure-467@4.[151,1172,194,730]" captionTargetPageId="4" captionText="Fig. 2. SEM illustrations of structural features of the legs of Anillinus alleni. A) Left foreleg, male, dorsal view, B) Right mesotibia, female, ventral view, C) Left metatibia, male, ventro-lateral view, D) Left metatibia, female, medial view. as = adhesive seta; msb = mesotibial brush; msms = mesotibial modified seta; mss = mesotibial spur; mtb = metatibial brush; mts = metatibial spur; mtts = metatibial modified setae; ta1-ta3 = protarsomeres 1–3; tbn = tibial notch. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10109519" httpUri="https://zenodo.org/record/10109519/files/figure.png" pageId="5" pageNumber="293">Fig. 2D</figureCitation>
|
||
), with 2 terminal spurs (mts) and tibial brush (mtb). Tarsi pentamerous, last and 1
|
||
<superScript id="6FE81141FFC59338FE338EF1FE202B96" attach="left" box="[485,498,1704,1718]" fontSize="6" pageId="5" pageNumber="293">st</superScript>
|
||
tarsomeres the longest, tarsomeres 2–4 of equal length on tarsi of all legs, 1
|
||
<superScript id="6FE81141FFC59338FCC88E5FFCF82B34" attach="left" box="[798,810,1542,1556]" fontSize="6" pageId="5" pageNumber="293">st</superScript>
|
||
tarsomere shorter than combined length of tarsomeres 2–4. Tarsal claws simple, edentate.
|
||
<emphasis id="AAE9601BFFC59338FD158E18FC772B76" bold="true" box="[707,933,1601,1622]" pageId="5" pageNumber="293">Abdominal ventrites:</emphasis>
|
||
Five visible abdominal ventrites: 2
|
||
<superScript id="6FE81141FFC59338FCFA8E0EFCEC2B45" attach="left" box="[812,830,1623,1637]" fontSize="6" pageId="5" pageNumber="293">nd</superScript>
|
||
ventrite longest, 3.9 times longer than 3
|
||
<superScript id="6FE81141FFC59338FD198E2BFD0C2BA0" attach="left" box="[719,734,1650,1664]" fontSize="6" pageId="5" pageNumber="293">rd</superScript>
|
||
or 4
|
||
<superScript id="6FE81141FFC59338FCDE8E2BFCC42BA0" attach="none" box="[776,790,1650,1664]" fontSize="6" pageId="5" pageNumber="293">th</superScript>
|
||
; 3
|
||
<superScript id="6FE81141FFC59338FCFB8E2BFCEE2BA0" attach="left" box="[813,828,1650,1664]" fontSize="6" pageId="5" pageNumber="293">rd</superScript>
|
||
and 4
|
||
<superScript id="6FE81141FFC59338FCA38E2BFC512BA0" attach="left" box="[885,899,1650,1664]" fontSize="6" pageId="5" pageNumber="293">th</superScript>
|
||
equal in length; 5
|
||
<superScript id="6FE81141FFC59338FBFD8E2BFBEB2BA0" attach="left" box="[1067,1081,1650,1664]" fontSize="6" pageId="5" pageNumber="293">th</superScript>
|
||
ventrite 2.3 times longer than 4
|
||
<superScript id="6FE81141FFC59338FCA18ED4FC572BBB" attach="none" box="[887,901,1677,1691]" fontSize="6" pageId="5" pageNumber="293">th</superScript>
|
||
. Intercoxal process (ipa) of 2
|
||
<superScript id="6FE81141FFC59338FB418ED4FB7B2BBB" attach="none" box="[1175,1193,1677,1691]" fontSize="6" pageId="5" pageNumber="293">nd</superScript>
|
||
ventrite broad, tapering and truncated anteriorly (
|
||
<figureCitation id="00A6A08CFFC6933BFF488CB0FF3B29DE" box="[158,233,1257,1278]" captionStart="Fig" captionStartId="3.[197,228,1102,1121]" captionTargetBox="[173,1194,195,1086]" captionTargetId="figure-182@3.[173,1194,194,1086]" captionTargetPageId="3" captionText="Fig. 1. SEM images of body parts of Anillinus alleni. A) Head, dorsal view, B) Pronotum, dorsal view, C) Elytra, dorsal view, D) Head, ventral view, E) Prosternum, ventral view, F) Metasternum, ventral view, G) Left and right metafemora, male, H) Right metafemur, female. bm = basal emargination; bs = basolateral pronotal seta; cl = clypeus; cs = clypeal seta; ed2 = scutellar seta; ed3-4, 6 = 1st, 2nd, and 3rd fixed discal setae; ed8 = apical seta; eo1-9 = setae 1–9 from the umbilicate series; fs = frontal seta; ft = frontal tubercle; gsc = glossal sclerite; ipa = intercoxal process of abdominal ventrite 2; lb = labrum; ls = midlaterall pronotal seta; m = mentum; mp3 = maxillary palpomere 3; mp4 = maxillary palpomere 4; ms = mental-submental suture; mscx = mesocoxa; mtcx = metacoxa; mtf = metafemur; mtt = metatibia; mttr = metatrochanter; mtv = metaventrite; pep = proepipleuron; pes = proepisternum; pg = paraglossa; prcx = procoxa; ps = prosternum; psp = prosternal process; sct = scutellum; sm = submentum; sp = metafemoral spine; ssa = anterior supraorbital seta; ssp = posterior supraorbital seta. Scale bars: A–B, E–H = 0.2 mm; C = 0.5 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109517" httpUri="https://zenodo.org/record/10109517/files/figure.png" pageId="6" pageNumber="294">Fig. 1F</figureCitation>
|
||
). Width of process more than half (0.64) length of metacoxa.
|
||
<emphasis id="AAE9601BFFC6933BFE838D5DFE392839" bold="true" box="[341,491,1284,1305]" pageId="6" pageNumber="294">Male genitalia:</emphasis>
|
||
Median lobe of aedeagus (
|
||
<figureCitation id="00A6A08CFFC6933BFF288D47FE982813" box="[254,330,1310,1331]" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="6" pageNumber="294">Fig. 3B</figureCitation>
|
||
) anopic, with short basal lobe, long arcuate shaft with markedly dilated apical half, and apex truncate at tip (may be broken). Dorsal margin strongly sclerotized along almost entire length and in form of pronounced ridge (dr). Ventral margin enlarged and convex in apical half. Dorsal sclerite (ds) in form of a semicircular blade-like structure with short basal prolongations. Internal sac also with 2 spine-like structures and plates of distinctive shape (esp) in apicoventral area. Left paramere slightly enlarged apically and greatly so basally, (
|
||
<figureCitation id="00A6A08CFFC6933BFF3A8E1AFEE42B78" box="[236,310,1603,1624]" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="6" pageNumber="294">Fig. 3C</figureCitation>
|
||
), with 1 long seta. Right paramere long and moderately narrow, with 4–5 long setae (
|
||
<figureCitation id="00A6A08CFFC6933BFF088E21FEE62BAD" box="[222,308,1656,1677]" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="6" pageNumber="294">Fig. 3D</figureCitation>
|
||
).
|
||
<emphasis id="AAE9601BFFC6933BFE868E21FDDA2BAD" bold="true" box="[336,520,1656,1677]" pageId="6" pageNumber="294">Female genitalia:</emphasis>
|
||
Ovipositor sclerites (
|
||
<figureCitation id="00A6A08CFFC6933BFF248ECBFEEB2B87" box="[242,313,1682,1703]" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="6" pageNumber="294">Fig. 3F</figureCitation>
|
||
) with gonocoxite 1 asetose (gc1). Gonocoxite 2 falciform (gc2), 1.7–1.8 times longer than basal width, markedly curved, with tiny ensiform (es) and normal nematiform (ns) setae. Laterotergite (lt) with 10–12 setae. Spermatheca (
|
||
<figureCitation id="00A6A08CFFC6933BFBBA8D47FD19286E" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="6" pageNumber="294">Fig. 3E</figureCitation>
|
||
, sp) with distal part of cornu (dpc) markedly dilated. Nodulus (n) short, ramus practically undifferentiated (
|
||
<figureCitation id="00A6A08CFFC6933BFCE88D37FC5928A3" box="[830,907,1390,1411]" captionStart="Fig" captionStartId="5.[197,228,1304,1323]" captionTargetBox="[185,1177,196,1286]" captionTargetId="figure-147@5.[183,1182,194,1288]" captionTargetPageId="5" captionText="Fig. 3. Habitus and internal genitalia of Anillinus alleni. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="6" pageNumber="294">Fig. 3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="CCE2EC81FFC49339FF798AB3FE692E7C" ID-DOI="http://doi.org/10.5281/zenodo.10109519" ID-Zenodo-Dep="10109519" httpUri="https://zenodo.org/record/10109519/files/figure.png" pageId="4" pageNumber="292" startId="4.[175,206,746,765]" targetBox="[151,1172,194,730]" targetPageId="4">
|
||
<paragraph id="9822BC09FFC49339FF798AB3FE692E7C" blockId="4.[151,1173,746,860]" pageId="4" pageNumber="292">
|
||
<emphasis id="AAE9601BFFC49339FF798AB3FF3B2FDD" bold="true" box="[175,233,746,765]" pageId="4" pageNumber="292">Fig. 2.</emphasis>
|
||
SEM illustrations of structural features of the legs of
|
||
<taxonomicName id="5F9DC78AFFC49339FD028AB3FC8A2FDD" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[724,856,746,765]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="292" phylum="Arthropoda" rank="species" species="alleni">
|
||
<emphasis id="AAE9601BFFC49339FD028AB3FC8A2FDD" box="[724,856,746,765]" italics="true" pageId="4" pageNumber="292">Anillinus alleni</emphasis>
|
||
</taxonomicName>
|
||
. A) Left foreleg, male, dorsal view, B) Right mesotibia, female, ventral view, C) Left metatibia, male, ventro-lateral view, D) Left metatibia, female, medial view. as = adhesive seta; msb = mesotibial brush; msms = mesotibial modified seta; mss = mesotibial spur; mtb = metatibial brush; mts = metatibial spur; mtts = metatibial modified setae; ta1-ta3 = protarsomeres 1–3; tbn = tibial notch. Scale bar = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CCE2EC81FFC59338FF138D41FE54289B" ID-DOI="http://doi.org/10.5281/zenodo.10109521" ID-Zenodo-Dep="10109521" httpUri="https://zenodo.org/record/10109521/files/figure.png" pageId="5" pageNumber="293" startId="5.[197,228,1304,1323]" targetBox="[185,1177,196,1286]" targetPageId="5">
|
||
<paragraph id="9822BC09FFC59338FF138D41FE54289B" blockId="5.[173,1194,1304,1467]" pageId="5" pageNumber="293">
|
||
<emphasis id="AAE9601BFFC59338FF138D41FF2C280B" bold="true" box="[197,254,1304,1323]" pageId="5" pageNumber="293">Fig. 3.</emphasis>
|
||
Habitus and internal genitalia of
|
||
<taxonomicName id="5F9DC78AFFC59338FDE68D41FD61280B" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[560,691,1304,1323]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="293" phylum="Arthropoda" rank="species" species="alleni">
|
||
<emphasis id="AAE9601BFFC59338FDE68D41FD61280B" box="[560,691,1304,1323]" italics="true" pageId="5" pageNumber="293">Anillinus alleni</emphasis>
|
||
</taxonomicName>
|
||
. A) Habitus, B) Aedeagal median lobe, right lateral view, C) Left paramere, left lateral view, D) Right paramere, right lateral view, E) Spermatheca and apical part of spermathecal gland, F) Female right ovipositor sclerites, dorsal view. bl = blade of gonocoxite 2; dpc = distal part of cornu; dr = dorsal ridge of median lobe; ds = dorsal sclerites of median lobe; es = ensiform setae; esp = endofallic spines of median lobe; gc1 = gonoxocite 1; gc2 = gonocoxite 2; lt = laterotergite; n = nodulus; ns = nematiform seta; ppc = proximal part of cornu; sg = spermathecal gland; sp = spermatheca. Scale bars: A = 0.5 mm; B = 0.2 mm; E–F = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CCE2EC81FFC6933BFF798C08FB87298C" ID-DOI="http://doi.org/10.5281/zenodo.10109523" ID-Zenodo-Dep="10109523" httpUri="https://zenodo.org/record/10109523/files/figure.png" pageId="6" pageNumber="294" startId="6.[175,206,1105,1124]" targetBox="[151,1173,194,1089]" targetPageId="6">
|
||
<paragraph id="9822BC09FFC6933BFF798C08FB87298C" blockId="6.[151,1173,1105,1196]" pageId="6" pageNumber="294">
|
||
<emphasis id="AAE9601BFFC6933BFF798C08FF3E2944" bold="true" box="[175,236,1105,1124]" pageId="6" pageNumber="294">Fig. 4.</emphasis>
|
||
Locality records for
|
||
<taxonomicName id="5F9DC78AFFC6933BFE688C08FD802944" box="[446,594,1105,1124]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="AAE9601BFFC6933BFE688C08FDD82944" box="[446,522,1105,1124]" italics="true" pageId="6" pageNumber="294">Anillinus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from the Ozark and Ouachita Mountains:
|
||
<taxonomicName id="5F9DC78AFFC6933BFC0B8C08FBF82944" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[989,1066,1105,1124]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="alleni">
|
||
<emphasis id="AAE9601BFFC6933BFC0B8C08FBF82944" box="[989,1066,1105,1124]" italics="true" pageId="6" pageNumber="294">A. alleni</emphasis>
|
||
</taxonomicName>
|
||
– red star;
|
||
<taxonomicName id="5F9DC78AFFC6933BFF418C30FF39295C" box="[151,235,1129,1148]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="aleyae">
|
||
<emphasis id="AAE9601BFFC6933BFF418C30FF39295C" box="[151,235,1129,1148]" italics="true" pageId="6" pageNumber="294">A. aleyae</emphasis>
|
||
</taxonomicName>
|
||
– white diamonds;
|
||
<taxonomicName id="5F9DC78AFFC6933BFE738C30FD24295C" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[421,758,1129,1148]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="magazinensis">
|
||
<emphasis id="AAE9601BFFC6933BFE738C30FDE1295C" box="[421,563,1129,1148]" italics="true" pageId="6" pageNumber="294">A. magazinensis</emphasis>
|
||
Sokolov and Carlton
|
||
</taxonomicName>
|
||
– white circles;
|
||
<taxonomicName id="5F9DC78AFFC6933BFC438C30FC24295C" box="[917,1014,1129,1148]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="AAE9601BFFC6933BFC438C30FC24295C" box="[917,1014,1129,1148]" italics="true" pageId="6" pageNumber="294">A. lescheni</emphasis>
|
||
</taxonomicName>
|
||
– white triangle;
|
||
<taxonomicName id="5F9DC78AFFC6933BFF418CD8FE6829B4" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[151,442,1153,1172]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="robisoni">
|
||
<emphasis id="AAE9601BFFC6933BFF418CD8FF2B29B4" box="[151,249,1153,1172]" italics="true" pageId="6" pageNumber="294">A. robisoni</emphasis>
|
||
Sokolov and Carlton
|
||
</taxonomicName>
|
||
– white flowers;
|
||
<taxonomicName id="5F9DC78AFFC6933BFD888CD8FC5029B4" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[606,898,1153,1172]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="stephani">
|
||
<emphasis id="AAE9601BFFC6933BFD888CD8FD1329B4" box="[606,705,1153,1172]" italics="true" pageId="6" pageNumber="294">A. stephani</emphasis>
|
||
Sokolov and Carlton
|
||
</taxonomicName>
|
||
– black square;
|
||
<taxonomicName id="5F9DC78AFFC6933BFBCB8CD8FE9B298C" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="294" phylum="Arthropoda" rank="species" species="tishechkini">
|
||
<emphasis id="AAE9601BFFC6933BFBCB8CD8FB4629B4" box="[1053,1172,1153,1172]" italics="true" pageId="6" pageNumber="294">A. tishechkini</emphasis>
|
||
Sokolov and Carlton
|
||
</taxonomicName>
|
||
– white squares (
|
||
<bibRefCitation id="FC0CC1F8FFC6933BFE348CC0FD5D298C" author="Sokolov, I. M." box="[482,655,1177,1196]" pageId="6" pageNumber="294" pagination="185 - 233" refId="ref5826" refString="Sokolov, I. M., C. E. Carlton, and J. F. Cornell. 2004. Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini). The Coleopterists Bulletin 58: 185 - 233." type="journal article" year="2004">
|
||
Sokolov
|
||
<emphasis id="AAE9601BFFC6933BFDE68CC0FD8F298C" box="[560,605,1177,1196]" italics="true" pageId="6" pageNumber="294">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
; Sokolov and Watrous 2008; personal observations).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9822BC09FFC6933BFD138DD0FC6628F3" blockId="6.[685,1172,1257,1730]" pageId="6" pageNumber="294">
|
||
<emphasis id="AAE9601BFFC6933BFD138DD0FC1028BE" bold="true" box="[709,962,1417,1438]" pageId="6" pageNumber="294">Geographic Distribution.</emphasis>
|
||
This species is known from the
|
||
<typeStatus id="472602ABFFC6933BFCD48DFAFCF92898" box="[770,811,1443,1464]" pageId="6" pageNumber="294">type</typeStatus>
|
||
locality in the Ozark Mountains of the Interior Highlands (
|
||
<figureCitation id="00A6A08CFFC6933BFCBA8DE7FC7A28F3" box="[876,936,1470,1491]" captionStart="Fig" captionStartId="6.[175,206,1105,1124]" captionTargetBox="[151,1173,194,1089]" captionTargetId="figure-287@6.[151,1173,194,1089]" captionTargetPageId="6" captionText="Fig. 4. Locality records for Anillinus species from the Ozark and Ouachita Mountains: A. alleni – red star; A. aleyae – white diamonds; A. magazinensis Sokolov and Carlton – white circles; A. lescheni – white triangle; A. robisoni Sokolov and Carlton – white flowers; A. stephani Sokolov and Carlton – black square; A. tishechkini Sokolov and Carlton – white squares (Sokolov et al. 2004; Sokolov and Watrous 2008; personal observations)." figureDoi="http://doi.org/10.5281/zenodo.10109523" httpUri="https://zenodo.org/record/10109523/files/figure.png" pageId="6" pageNumber="294">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9822BC09FFC6933AFD138D81FF392C06" blockId="6.[685,1172,1257,1730]" lastBlockId="7.[173,660,194,1357]" lastPageId="7" lastPageNumber="295" pageId="6" pageNumber="294">
|
||
<emphasis id="AAE9601BFFC6933BFD138D81FCA028CD" bold="true" box="[709,882,1496,1517]" pageId="6" pageNumber="294">Natural History.</emphasis>
|
||
Prior to his death, Thomas C. Barr, Jr. corresponded with HWR and shared his recollections about collecting the original, apparently lost, specimen of this species
|
||
<emphasis id="AAE9601BFFC6933BFBF48E71FB802B1D" box="[1058,1106,1576,1597]" italics="true" pageId="6" pageNumber="294">circa</emphasis>
|
||
1971. Relevant excerpts include: “The Blanchard Springs anilline will be under flat rocks in talus along the path leading to Blanchard Springs, itself, about 75–100 yd before you get to the springs…..to the right of the path as you walk to the spring outlet but not very far up the hill, because you are looking for as moist a well-drained microenvironment as possible.” (T. C. Barr email correspondence with HWR, 2005).
|
||
</paragraph>
|
||
<paragraph id="9822BC09FFC7933AFF138975FEFC2CDB" blockId="7.[173,660,194,1357]" pageId="7" pageNumber="295">
|
||
Specimens in the
|
||
<typeStatus id="472602ABFFC7933AFEBA8975FE472C61" box="[364,405,300,321]" pageId="7" pageNumber="295">type</typeStatus>
|
||
series were extracted from soil using a soil-washing technique, thus implying an endogean habitat association. Their absence from numerous surface litter samples taken during previous years suggests a true specialization to a subterranean lifestyle that may include parts of the adjacent cavern system and surrounding karst topographies.
|
||
</paragraph>
|
||
<paragraph id="9822BC09FFC7933AFF138A59FE6F286D" blockId="7.[173,660,194,1357]" pageId="7" pageNumber="295">
|
||
<emphasis id="AAE9601BFFC7933AFF138A59FD892F35" bold="true" box="[197,603,512,533]" pageId="7" pageNumber="295">Comparison among Other Species.</emphasis>
|
||
The structure of the internal genitalia unequivocally indicates relatedness between the new species and a complex of endogean species of
|
||
<taxonomicName id="5F9DC78AFFC7933AFE3A8A09FD932F45" authorityName="Casey" authorityYear="1918" box="[492,577,592,613]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AAE9601BFFC7933AFE3A8A09FD932F45" box="[492,577,592,613]" italics="true" pageId="7" pageNumber="295">Anillinus</emphasis>
|
||
</taxonomicName>
|
||
that occurs west of the
|
||
<collectingRegion id="5A5972EBFFC7933AFE908A32FE602FA0" box="[326,434,619,640]" country="United States of America" name="Mississippi" pageId="7" pageNumber="295">Mississippi</collectingRegion>
|
||
River. The shape of the apical part of the median lobe, specifically the enlarged and convex ventral margin, closely resembles that of
|
||
<taxonomicName id="5F9DC78AFFC7933AFEE88AE3FDB72FEF" authority="Sokolov and Watrous" authorityName="Sokolov and Watrous" box="[318,613,698,719]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="aleyae">
|
||
<emphasis id="AAE9601BFFC7933AFEE88AE3FE472FEF" box="[318,405,698,719]" italics="true" pageId="7" pageNumber="295">A. aleyae</emphasis>
|
||
Sokolov and Watrous
|
||
</taxonomicName>
|
||
, also from the Ozark Mountains, and
|
||
<taxonomicName id="5F9DC78AFFC7933AFE0E8A8CFECC2E25" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="AAE9601BFFC7933AFE0E8A8CFDEC2FCA" box="[472,574,725,746]" italics="true" pageId="7" pageNumber="295">A. lescheni</emphasis>
|
||
Sokolov and Carlton
|
||
</taxonomicName>
|
||
, from the adjacent Ouachita Mountains to the south (
|
||
<bibRefCitation id="FC0CC1F8FFC7933AFEEE8B53FDD52E3F" author="Sokolov, I. M." box="[312,519,778,799]" pageId="7" pageNumber="295" pagination="185 - 233" refId="ref5826" refString="Sokolov, I. M., C. E. Carlton, and J. F. Cornell. 2004. Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini). The Coleopterists Bulletin 58: 185 - 233." type="journal article" year="2004">
|
||
Sokolov
|
||
<emphasis id="AAE9601BFFC7933AFE428B53FE182E3F" box="[404,458,778,799]" italics="true" pageId="7" pageNumber="295">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
; Sokolov and Watrous 2008), suggesting monophyly of the three species. The median lobe structure is less similar to members of the
|
||
<collectingRegion id="5A5972EBFFC7933AFE958B03FEA82E4F" box="[323,378,858,879]" country="United States of America" name="Texas" pageId="7" pageNumber="295">Texas</collectingRegion>
|
||
assemblage of troglobitic and endogean species (Sokolov
|
||
<emphasis id="AAE9601BFFC7933AFE128B2DFE2B2EA9" box="[452,505,884,905]" italics="true" pageId="7" pageNumber="295">et al.</emphasis>
|
||
2014). Among endogean species from the eastern part of the genus’ range, the Appalachian
|
||
<emphasis id="AAE9601BFFC7933AFE588BF3FE232E9F" box="[398,497,938,959]" italics="true" pageId="7" pageNumber="295">moseleyae</emphasis>
|
||
group of species (
|
||
<bibRefCitation id="FC0CC1F8FFC7933AFF658B9DFE932EF9" author="Sokolov, I. M." box="[179,321,964,985]" pageId="7" pageNumber="295" pagination="1 - 14" refId="ref5686" refString="Sokolov, I. M. 2011. Five new species of Anillinus Casey from the southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini). Insecta Mundi 164: 1 - 14." type="journal article" year="2011">Sokolov 2011</bibRefCitation>
|
||
) is most similar to
|
||
<taxonomicName id="5F9DC78AFFC7933AFDCA8B9DFDA12EF9" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[540,627,964,985]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="alleni">
|
||
<emphasis id="AAE9601BFFC7933AFDCA8B9DFDA12EF9" box="[540,627,964,985]" italics="true" pageId="7" pageNumber="295">A. alleni</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the median lobe. The shape of the spermatheca, namely the markedly dilated distal part of the cornu of the new species, is similar to that of the
|
||
<collectingRegion id="5A5972EBFFC7933AFF3A8C77FEF12963" box="[236,291,1070,1091]" country="United States of America" name="Texas" pageId="7" pageNumber="295">Texas</collectingRegion>
|
||
species
|
||
<taxonomicName id="5F9DC78AFFC7933AFEA08C77FF25297E" authority="Sokolov and Reddell" authorityName="Sokolov and Reddell" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="forthoodensis">
|
||
<emphasis id="AAE9601BFFC7933AFEA08C77FDC02963" box="[374,530,1070,1091]" italics="true" pageId="7" pageNumber="295">A. forthoodensis</emphasis>
|
||
Sokolov and Reddell
|
||
</taxonomicName>
|
||
(Sokolov
|
||
<emphasis id="AAE9601BFFC7933AFE8D8C10FE5E297E" box="[347,396,1097,1118]" italics="true" pageId="7" pageNumber="295">et al.</emphasis>
|
||
2014). The combination of external characteristics of
|
||
<taxonomicName id="5F9DC78AFFC7933AFE748C3AFE202958" authorityName="SOKOLOV AND CARLTON" authorityYear="2017" box="[418,498,1123,1144]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="alleni">
|
||
<emphasis id="AAE9601BFFC7933AFE748C3AFE202958" box="[418,498,1123,1144]" italics="true" pageId="7" pageNumber="295">A. alleni</emphasis>
|
||
</taxonomicName>
|
||
is unique among congeners from the western part of the genus’ range (
|
||
<emphasis id="AAE9601BFFC7933AFF628CC0FF18298E" box="[180,202,1177,1198]" italics="true" pageId="7" pageNumber="295">i.e</emphasis>
|
||
., west of the
|
||
<collectingRegion id="5A5972EBFFC7933AFE9C8CC0FE61298E" box="[330,435,1177,1198]" country="United States of America" name="Mississippi" pageId="7" pageNumber="295">Mississippi</collectingRegion>
|
||
River), whereas among the species from the east the new species resembles the
|
||
<collectingRegion id="5A5972EBFFC7933AFED58C97FE8529C3" box="[259,343,1230,1251]" country="United States of America" name="Alabama" pageId="7" pageNumber="295">Alabama</collectingRegion>
|
||
troglobitic species
|
||
<taxonomicName id="5F9DC78AFFC7933AFDCF8C97FED329DD" authority="(Jeannel)" baseAuthorityName="Jeannel" class="Insecta" family="Carabidae" genus="Troglanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="valentinei">
|
||
<emphasis id="AAE9601BFFC7933AFDCF8C97FD4129C3" box="[537,659,1230,1251]" italics="true" pageId="7" pageNumber="295">A. valentinei</emphasis>
|
||
(Jeannel)
|
||
</taxonomicName>
|
||
and, especially,
|
||
<taxonomicName id="5F9DC78AFFC7933AFE6C8CB1FD4229DD" authority="Sokolov" authorityName="Sokolov" authorityYear="2012" box="[442,656,1256,1277]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="295" phylum="Arthropoda" rank="species" species="tombarri">
|
||
<emphasis id="AAE9601BFFC7933AFE6C8CB1FDE329DD" box="[442,561,1256,1277]" italics="true" pageId="7" pageNumber="295">A. tombarri</emphasis>
|
||
Sokolov
|
||
</taxonomicName>
|
||
, sharing with the latter unusually large size, wide basal margin of the pronotum, and spinose male metafemora (
|
||
<bibRefCitation id="FC0CC1F8FFC7933AFEFF8D61FE60286D" author="Sokolov, I. M." box="[297,434,1336,1357]" pageId="7" pageNumber="295" pagination="61 - 71" refId="ref5735" refString="Sokolov, I. M. 2012. Five new species of Anillinus Casey from Alabama with a key to the Alabama species (Carabidae: Trechinae: Bembidiini). Annals of Carnegie Museum 81: 61 - 71." type="journal article" year="2012">Sokolov 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |