300 lines
42 KiB
XML
300 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1187.113473" ID-Pensoft-Pub="1313-2970-1187-341" ID-Pensoft-UUID="787AD6721D9852DFAD6AEB21E125E181" ID-ZooBank="2F7962833187447E88C64B61C50B640C" ModsDocID="1313-2970-1187-341" checkinTime="1703815707905" checkinUser="pensoft" docAuthor="Akkari, Nesrine, Macek, Oliver & Stoev, Pavel" docDate="2023" docId="1033BE23414C51779C61A48CC0360086" docLanguage="en" docName="ZooKeys 1187: 341-399" docOrigin="ZooKeys 1187" docPubDate="2023-12-28" docSource="http://dx.doi.org/10.3897/zookeys.1187.113473" docTitle="Paracortina voluta Wang & Zhang 1993" docType="treatment" docVersion="1" id="787AD6721D9852DFAD6AEB21E125E181" lastPageNumber="341" masterDocId="787AD6721D9852DFAD6AEB21E125E181" masterDocTitle="A revision of the millipede family Paracortinidae (Diplopoda, Callipodida)" masterLastPageNumber="399" masterPageNumber="341" pageNumber="341" updateTime="1703815707905" updateUser="pensoft">
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<mods:titleInfo id="439540D697B4506AA7162711A4BE89C2">
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<mods:title id="54B447DE5E5CC1A3BD19B4CAF0B2CD46">A revision of the millipede family Paracortinidae (Diplopoda, Callipodida)</mods:title>
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<mods:name id="B6E2261E95124009F5A4C824BEB4AD29" type="personal">
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<mods:namePart id="E4451090FFA6F4E6E4C9A341C00DD19D">Akkari, Nesrine</mods:namePart>
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<mods:nameIdentifier id="0D2827D0E7D51AC40D96DBFC68831250" type="ORCID">https://orcid.org/0000-0001-5019-4833</mods:nameIdentifier>
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<mods:affiliation id="A95B4BC479B696666D8B55A4C284FC5F">Naturhistorisches Museum Wien, Burgring 7, 1010 Wien, Austria</mods:affiliation>
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<mods:nameIdentifier id="58D659AADE389D9ABADD3875C8F118F4" type="email">nes.akkari@gmail.com</mods:nameIdentifier>
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<mods:namePart id="903859B0EFC368BE26768C606E016FE8">Macek, Oliver</mods:namePart>
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<mods:nameIdentifier id="575929893CF32B4C2AE242E41744943C" type="ORCID">https://orcid.org/0000-0002-8146-5373</mods:nameIdentifier>
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<mods:affiliation id="6056C2ADFAF1108C98B3E1E8C14C65C6">Naturhistorisches Museum Wien, Burgring 7, 1010 Wien, Austria</mods:affiliation>
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<mods:namePart id="EF32534874A5B421DE0FF4534CD37031">Stoev, Pavel</mods:namePart>
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<mods:nameIdentifier id="2ADD90973A82D56D0F029CBC60D7F77D" type="ORCID">https://orcid.org/0000-0002-5702-5677</mods:nameIdentifier>
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<mods:affiliation id="75137975A0FC9F02D269C82425A2B774">National Museum of Natural History at the Bulgarian Academy of Sciences, Tsar Osvoboditel Blvd. 1, 1000 Sofia, Bulgaria & Pensoft Publishers, Prof. G. Zlatarski Str. 12, Sofia, Bulgaria</mods:affiliation>
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<mods:typeOfResource id="FFB2E426A16D54716F3B05A605C63CBD">text</mods:typeOfResource>
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<mods:title id="E9DB7ED5FA71A049D80F1C5F4718FB3B">ZooKeys</mods:title>
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<mods:part id="D883145F301E6D1851F1F0B2E573EB15">
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<mods:date id="58FC17254D08166E220638FECAA8D8D3">2023</mods:date>
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<mods:number id="AF7E2CB668B7E3FB7C15D8F77B195A5E">2023-12-28</mods:number>
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<mods:number id="EF88F0D420C5FD51F607631CED9B0701">1187</mods:number>
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<mods:start id="A7FB1F2A45FDF2BAB9283079A142D6F9">341</mods:start>
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<mods:classification id="6AEF07B60C011C388DDC42DBD0867723">journal article</mods:classification>
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<treatment id="1033BE23414C51779C61A48CC0360086" LSID="urn:lsid:plazi:treatment:1033BE23414C51779C61A48CC0360086" httpUri="http://treatment.plazi.org/id/1033BE23414C51779C61A48CC0360086" lastPageNumber="341" pageId="0" pageNumber="341">
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<subSubSection id="1FFAF5162CCF352621CD792BE515EB9E" pageId="0" pageNumber="341" type="nomenclature">
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<paragraph id="E882222F9723EECCD7E5BC8869915FC2" pageId="0" pageNumber="341">
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<taxonomicName id="51F43A24E9AE85A6F940E9A6F1EDD5B7" ID-CoL="4CRBM" LSID="1033BE23-414C-5177-9C61-A48CC0360086" authority="Wang & Zhang, 1993" authorityName="Wang & Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina voluta" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="voluta">Paracortina voluta Wang & Zhang, 1993</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0EE7597C5C335C25A04F65E3B06BD9CC" pageId="0" pageNumber="341" type="description">
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<paragraph id="5CCD3AEFF03609434C22C6BD3AAFD2A3" pageId="0" pageNumber="341">
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<figureCitation id="439669C04429C1220A2A1DF84811111B" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">Figs 20</figureCitation>
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<figureCitation id="006625B063E880307414F0338FEA367D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">, 21</figureCitation>
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<figureCitation id="9BC03423AEE46374AFEF7BA47DE54F81" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM), left gonopod A anterior view B posterior view C lateral view D mesal view. Abbreviations: b = falcate mesal process of coxa; Cl = lateral lobe of coxa; dl = distal lamella of telopodite; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure22" httpUri="https://binary.pensoft.net/fig/958797" pageId="0" pageNumber="341">, 22</figureCitation>
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<figureCitation id="67476BFFAA6A4FD6CD723F80AA4A4101" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Modifications on leg-pair 6 and leg-pair 7 in adult males of species of Paracortinidae A, B Paracortina viriosa (Wang & Zhang, 1993), male A leg pair 6 B leg pair 7 C, D Paracortina voluta Wang & Zhang, 1993, male C leg pair 6 D leg pair 7 E Paracortina asciformis Akkari & Stoev, sp. nov., male holotype, leg pair 7 F Paracortina kabaki Akkari & Stoev, sp. nov., male holotype, leg pair 7 G Scotopetalum chinensis (Stoev & Geoffroy, 2004) comb. nov., male paratype, leg pair 7. Abbreviations: c = coxa; pr = prefemur; tr = trochanter." figureDoi="10.3897/zookeys.1187.113473.figure27" httpUri="https://binary.pensoft.net/fig/958802" pageId="0" pageNumber="341">, 27C, D</figureCitation>
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<figureCitation id="C21DF13C4BD959387EA4775FC683C69E" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution map of Paracortinidae in China and Vietnam." figureDoi="10.3897/zookeys.1187.113473.figure28" httpUri="https://binary.pensoft.net/fig/958803" pageId="0" pageNumber="341">, 28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="3D45E4AA675F8577392AA47437A89776" pageId="0" pageNumber="341" type="reference_group">
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<paragraph id="72B430A43F2A8D75EC2B867340BE80FE" pageId="0" pageNumber="341">
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<taxonomicName id="EFE158101D21D8A1A6B91639E6E7C20A" authorityName="Wang & Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina voluta" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="voluta">Paracortina voluta</taxonomicName>
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Wang & Zhang, 1993: 377, figs 6-9;
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<bibRefCitation id="A22A21C8797427B912BBBAEAC5482502" DOI="https://doi.org/10.2476/asjaa.53.93" author="Stoev, P" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="341" pagination="93 - 103" refId="B21" refString="Stoev, P, Geoffroy, JJ, 2004. Review of the millipede family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida). Acta Arachnologica 53 (2): 93 - 103, DOI: https://doi.org/10.2476/asjaa.53.93" title="Review of the millipede family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida)." url="https://doi.org/10.2476/asjaa.53.93" volume="53" year="2004">Stoev and Geoffroy 2004</bibRefCitation>
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: 103, key;
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<bibRefCitation id="EC67637F8B5B7311EDF6324BD31A00D1" DOI="https://doi.org/10.3897/zookeys.517.9949" author="Liu, WX" journalOrPublisher="ZooKeys" pageId="0" pageNumber="341" pagination="123 - 140" refId="B9" refString="Liu, WX, Tian, MY, 2015. Two new cave-dwelling species of the millipede genus Paracortina Wang & Zhang, 1993 from southern China (Diplopoda, Callipodida, Paracortinidae). ZooKeys 517: 123 - 140, DOI: https://doi.org/10.3897/zookeys.517.9949" title="Two new cave-dwelling species of the millipede genus Paracortina Wang & Zhang, 1993 from southern China (Diplopoda, Callipodida, Paracortinidae)." url="https://doi.org/10.3897/zookeys.517.9949" volume="517" year="2015">Liu and Tian 2015</bibRefCitation>
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: 139, key.
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</paragraph>
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</subSubSection>
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<subSubSection id="1991446C066316930BB0C08131F1A36F" pageId="0" pageNumber="341" type="materials_examined">
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<paragraph id="C4C6458BD28DFE6DDD47BB5D5E02DB5F" pageId="0" pageNumber="341">Studied material.</paragraph>
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<paragraph id="722DBBEC871F32AD66ECDB5C887072AF" pageId="0" pageNumber="341">
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<materialsCitation id="8410876F8B9C3178359E21C0BD8A6BCC" collectingDate="2011-07-18" collectorName="NW Pingchuan, I. Belousov, I. Kabak" country="China" determinerName="Akkari" latitude="27.66861" location="Sichuan Province" longLatPrecision="20" longitude="101.73444" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sichuan Province">
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<specimenCount id="3D4114A0D6F8E6B78D1C77B7ADB612EC" type="male">2 males</specimenCount>
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,
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<specimenCount id="94203C5E94F110D6CB0B610E878666DA" type="female">1 female</specimenCount>
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,
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<collectingCountry id="2F2C34899C7CE5532826F87CCA3A38D7" name="China">China</collectingCountry>
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,
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<collectingRegion id="114073A6DD81EE010EF1939521817DB8" country="China" name="Sichuan">Sichuan Province</collectingRegion>
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,
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<collectorName id="EB6D3C7BAD3694CE06A118384CE8BFA6">NW Pingchuan</collectorName>
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,
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<geoCoordinate id="FDAF8FB1028E2ADDA50C8C79CD3129B5" degrees="27" direction="north" minutes="40" orientation="latitude" precision="15" seconds="07" value="27.66861">27°40'07"N</geoCoordinate>
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,
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<geoCoordinate id="98BF5F22AB4268645C523D4674BC6393" degrees="101" direction="east" minutes="44" orientation="longitude" precision="15" seconds="04" value="101.73444">101°44'04"E</geoCoordinate>
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,
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<collectingDate id="63C5D9BD294713C72F6EC460FEA234AE" value="2011-07-18">18.07.2011</collectingDate>
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,
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<collectorName id="2BCF2F03ACA4E4271D4290B714035592">I. Belousov</collectorName>
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&
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<collectorName id="70F92B7EC532D4C081C0F1E75A92EE0E">I. Kabak</collectorName>
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leg. (Rd 5346 ZMUM),
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<determinerName id="13F112EA71FD683CF266037345139E7B">Akkari</determinerName>
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det.
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</materialsCitation>
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2023.
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</paragraph>
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</subSubSection>
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<subSubSection id="0C4845BA9402CB4CCB59E4E648D78305" pageId="0" pageNumber="341" type="diagnosis">
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<paragraph id="F3A6FF0967FE63B4C5BBBBDB65A712B2" pageId="0" pageNumber="341">Diagnosis.</paragraph>
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<paragraph id="A30933DED98A1A5A3AB678223C08C84D" pageId="0" pageNumber="341">
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Most similar to
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<taxonomicName id="1F5C51A385B1FE42A43AAE1DBDF625A5" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. viriosa" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="viriosa">
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<emphasis id="89CDE2207689A237E961419416766D25" italics="true" pageId="0" pageNumber="341">P. viriosa</emphasis>
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</taxonomicName>
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in the shape of the distal part of the telopodite with two main folds and a strong hook-shaped median processes, differing in the earlobe-shaped distal process and the smaller and more distally located anterior lamella.
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</paragraph>
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</subSubSection>
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<subSubSection id="820C4C5719D48F5F4090F64344A9697A" pageId="0" pageNumber="341" type="descriptive notes">
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<paragraph id="AB6F516B7BBE237ED0224406FCA9A771" pageId="0" pageNumber="341">Descriptive notes.</paragraph>
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<paragraph id="7D4D9478DF61E5ADDAD136A146FEB758" pageId="0" pageNumber="341">
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Male with 56PTs+Telson. Length ca 49.5 mm. Live colour unknown. Preserved specimen with a brownish general colour, legs and antennae dark brown (Figs
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<figureCitation id="519AA1F2F9E7FBF154FFF95474DF3BDB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20</figureCitation>
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,
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<figureCitation id="5E1B5849A5135D827B1A78D0288CD2CF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21A</figureCitation>
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), Head with dark pigmentation on the vertex and frons (Fig.
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<figureCitation id="20D07E30042F46BAA0C9597DD13387D6" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20B, C</figureCitation>
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), mandibular stipes and gnathochilarium with yellowish spots. Fields of ommatidia subtriangular, composed of ~ 47 ommatidia in ten rows (Fig.
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<figureCitation id="7124809943C0D22ECAA98E93C7C3E1AC" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20B, C</figureCitation>
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). Organ of
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<normalizedToken id="978F75A7ACA38B55B00EF14748397158" originalValue="Tömösváry">Toemoesvary</normalizedToken>
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very large, ~ 1.2 mm, situated close to and touching anterior side of eye (Fig.
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<figureCitation id="7C81908799CC644CF049E2DEEE999108" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20B</figureCitation>
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). Prozona brownish sputtered with a paler colour; metazona dorsally dark tawny-brown, especially on crests, anterior part pale with brown alveolate spots (Fig.
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<figureCitation id="7512742546E743E3085A5335345C091E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20A, B</figureCitation>
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).
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</paragraph>
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<caption id="8F09A6DB26323B78FC81E9B41E5A7F61" doi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341" start="Figure 20" startId="F20">
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<paragraph id="01CCA1CB7A60CF588D4251E860887321" pageId="0" pageNumber="341">
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<emphasis id="77486F9E0198AE35EFA93E0851398FD5" bold="true" pageId="0" pageNumber="341">Figure 20.</emphasis>
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<taxonomicName id="6DFF14C15FA7C95CE069F3BB2908E11C" authorityName="Wang & Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina voluta" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="voluta">
|
|
<emphasis id="9BEBC090336AD4B1D1C3159E5BC0993B" italics="true" pageId="0" pageNumber="341">Paracortina voluta</emphasis>
|
|
</taxonomicName>
|
|
Wang & Zhang, 1993 (Rd 5346 ZMUM)
|
|
<emphasis id="1197C0542B92ADC0961116BA6F48FF35" bold="true" pageId="0" pageNumber="341">A</emphasis>
|
|
female habitus, lateral view
|
|
<emphasis id="BEA9F92BACF4336A885C28058C4908A5" bold="true" pageId="0" pageNumber="341">B, C</emphasis>
|
|
male head and anterior pleurotergites
|
|
<emphasis id="ECE7C2608B6940B95FF58F613373D08B" bold="true" pageId="0" pageNumber="341">B</emphasis>
|
|
lateral view
|
|
<emphasis id="3DED89C30831B33EA7FCB085F96E1556" bold="true" pageId="0" pageNumber="341">C</emphasis>
|
|
frontal view.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="B3FB329E9D79D1B886E746FD61D2A84E" doi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341" start="Figure 21" startId="F21">
|
|
<paragraph id="DC0F25025D920E47C57B210160EE9DAB" pageId="0" pageNumber="341">
|
|
<emphasis id="5A556B3356ED7FC9D74E4B715F4A7A56" bold="true" pageId="0" pageNumber="341">Figure 21.</emphasis>
|
|
<taxonomicName id="166338A21E3BD1945EE88D02F19B6A28" authorityName="Wang & Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina voluta" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="voluta">
|
|
<emphasis id="A7C78F380C4592CD76E564A41303E690" italics="true" pageId="0" pageNumber="341">Paracortina voluta</emphasis>
|
|
</taxonomicName>
|
|
Wang & Zhang, 1993, male (Rd 5346 ZMUM)
|
|
<emphasis id="BA1823D14A12ECBE18B417101504D1AA" bold="true" pageId="0" pageNumber="341">A</emphasis>
|
|
head, anterior pleurotergites and legs, frontal view
|
|
<emphasis id="AFD0645CFB604ECC4E1C48A747A7CE7C" bold="true" pageId="0" pageNumber="341">B-D</emphasis>
|
|
gonopods
|
|
<emphasis id="9714F4EAAEE22CCAFB1677BEBF2F7748" bold="true" pageId="0" pageNumber="341">B</emphasis>
|
|
anterior view
|
|
<emphasis id="27CBBDE0E4F34EC0E66DF85BE6D1082D" bold="true" pageId="0" pageNumber="341">C</emphasis>
|
|
posterior view
|
|
<emphasis id="1F4C22CE05F77E18E7F001084DFDF2DD" bold="true" pageId="0" pageNumber="341">D</emphasis>
|
|
telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf1 = prefemoroidal process 1; pf2 = prefemoroidal process 2; ps =parasolenomere; s = solenomere; T = telopodite.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="1BD3541FA7379DA2CAF14B86EE5A3F18" pageId="0" pageNumber="341" type="description">
|
|
<paragraph id="8DE7199E4F0CFEBA98C8CC82EAA77D94" pageId="0" pageNumber="341">Male sexual characters.</paragraph>
|
|
<paragraph id="C50D05C51AFB7E7A13EE46B271344DF0" pageId="0" pageNumber="341">
|
|
Head with a protruding triangular projection on vertex (Fig.
|
|
<figureCitation id="B54255019CA69B440842C52F42F1318F" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20B, C</figureCitation>
|
|
,
|
|
<figureCitation id="733337503CBCC010397FF824BE710665" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21A</figureCitation>
|
|
). Leg-pairs 1 and 2 reduced and more setose than the rest, showing prefemoral and tarsal brushes, leg-pair 2 with a large anterior process on coxa and posterior gonopore (Fig.
|
|
<figureCitation id="8D1EE5E5753A783A250F060234380E26" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21A</figureCitation>
|
|
), leg-pair 4 with anterior triangular projection on coxa (Fig.
|
|
<figureCitation id="56CBB1B73EAC1BCC80A6A2DBB842D761" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21A</figureCitation>
|
|
), leg-pair 6 with one short triangular mesal process and a smaller lateral one on coxa, prefemur proximally slightly constricted on the posterior margin (Fig.
|
|
<figureCitation id="F71E2B341E49C9885B61269D9AA5B769" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Modifications on leg-pair 6 and leg-pair 7 in adult males of species of Paracortinidae A, B Paracortina viriosa (Wang & Zhang, 1993), male A leg pair 6 B leg pair 7 C, D Paracortina voluta Wang & Zhang, 1993, male C leg pair 6 D leg pair 7 E Paracortina asciformis Akkari & Stoev, sp. nov., male holotype, leg pair 7 F Paracortina kabaki Akkari & Stoev, sp. nov., male holotype, leg pair 7 G Scotopetalum chinensis (Stoev & Geoffroy, 2004) comb. nov., male paratype, leg pair 7. Abbreviations: c = coxa; pr = prefemur; tr = trochanter." figureDoi="10.3897/zookeys.1187.113473.figure27" httpUri="https://binary.pensoft.net/fig/958802" pageId="0" pageNumber="341">27C</figureCitation>
|
|
), leg-pair 7 with one mesal hyaline pointed process and a rounded projection surmounted by a similar but slightly smaller one, trochanter with a tuft of strong setae (Fig.
|
|
<figureCitation id="861E9E6F5D1131140B59EA442F9EC520" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Modifications on leg-pair 6 and leg-pair 7 in adult males of species of Paracortinidae A, B Paracortina viriosa (Wang & Zhang, 1993), male A leg pair 6 B leg pair 7 C, D Paracortina voluta Wang & Zhang, 1993, male C leg pair 6 D leg pair 7 E Paracortina asciformis Akkari & Stoev, sp. nov., male holotype, leg pair 7 F Paracortina kabaki Akkari & Stoev, sp. nov., male holotype, leg pair 7 G Scotopetalum chinensis (Stoev & Geoffroy, 2004) comb. nov., male paratype, leg pair 7. Abbreviations: c = coxa; pr = prefemur; tr = trochanter." figureDoi="10.3897/zookeys.1187.113473.figure27" httpUri="https://binary.pensoft.net/fig/958802" pageId="0" pageNumber="341">27D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="D27E627F124EC51A9DD96B3F95896E05" pageId="0" pageNumber="341">
|
|
<emphasis id="75FAE5F5D6126A4C54A393A4B58AD018" bold="true" pageId="0" pageNumber="341">
|
|
<emphasis id="B819F59822A19D7ED7090CE5D1D8FC74" italics="true" pageId="0" pageNumber="341">Gonopods</emphasis>
|
|
.
|
|
</emphasis>
|
|
Parallel, slightly converging with the distal solenomeral processes of telopodites crossing (Fig.
|
|
<figureCitation id="5F3275C4A5E1586621B7B230930A27B5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21B-D</figureCitation>
|
|
). Each gonopod with two asymmetrical, short, clavate prefemoroidal processes with (
|
|
<emphasis id="F7ABAA270632B19FEC6D0EA0ABC69684" bold="true" pageId="0" pageNumber="341">pf1</emphasis>
|
|
) broader and more setose than (
|
|
<emphasis id="49E832BCE8505206311552E5D8B1C5E9" bold="true" pageId="0" pageNumber="341">pf2</emphasis>
|
|
) (Figs
|
|
<figureCitation id="AF59182C562C5E92BBEC5520388DE832" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21B, C</figureCitation>
|
|
,
|
|
<figureCitation id="B7949D0700C3633730ABFD03B6D85095" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM), left gonopod A anterior view B posterior view C lateral view D mesal view. Abbreviations: b = falcate mesal process of coxa; Cl = lateral lobe of coxa; dl = distal lamella of telopodite; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure22" httpUri="https://binary.pensoft.net/fig/958797" pageId="0" pageNumber="341">22</figureCitation>
|
|
); coxal anterior lobe lower than the lateral lobes low (Fig.
|
|
<figureCitation id="167D0C7A397D2079BCE2F5748958D0DA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21C</figureCitation>
|
|
), long and falcate coxal process (
|
|
<emphasis id="24D46A6BE989DABAC3FE9C4D8F775DBC" bold="true" pageId="0" pageNumber="341">b</emphasis>
|
|
) reaching the distal part of the telopodite and apically pointed (Figs
|
|
<figureCitation id="54BABC274042EF772DA3EFF7B84A87B9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21C</figureCitation>
|
|
,
|
|
<figureCitation id="DC009C0E4B9B01E6A354A9F072A67794" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM), left gonopod A anterior view B posterior view C lateral view D mesal view. Abbreviations: b = falcate mesal process of coxa; Cl = lateral lobe of coxa; dl = distal lamella of telopodite; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure22" httpUri="https://binary.pensoft.net/fig/958797" pageId="0" pageNumber="341">22A, B, D</figureCitation>
|
|
). Telopodite (
|
|
<emphasis id="7E4EEDE6301305BEF68FCC7F90B026C1" bold="true" pageId="0" pageNumber="341">T</emphasis>
|
|
) with a long stem, distally expanding in an earlobe shape with rounded lateral margin seen in posterior view (Fig.
|
|
<figureCitation id="D17DD4B53FBBCC181CE4A928F6B97D66" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21C, D</figureCitation>
|
|
), in anterior view as an oblique subtrapezoidal plate (Fig.
|
|
<figureCitation id="8EAA7E63C23B7BA7AB9FCCADE357D8CA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21B</figureCitation>
|
|
), with a transparent lamella (
|
|
<emphasis id="82DBF472D9FC8ABE810C4C919D0F3E42" bold="true" pageId="0" pageNumber="341">dl</emphasis>
|
|
) attached on the lower part, marking and S-shape and serrated on the lower margin (Fig.
|
|
<figureCitation id="7A4FE48C94CA66E7557DA984D802280A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21B</figureCitation>
|
|
), mesal process curved upward, twisted and narrowing towards its apex (Figs
|
|
<figureCitation id="11B43EC6E0A6F2F44285D92E34726A88" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21B-D</figureCitation>
|
|
,
|
|
<figureCitation id="B0F19AC9CF9169D7EB1D1128CD0729EE" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM), left gonopod A anterior view B posterior view C lateral view D mesal view. Abbreviations: b = falcate mesal process of coxa; Cl = lateral lobe of coxa; dl = distal lamella of telopodite; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure22" httpUri="https://binary.pensoft.net/fig/958797" pageId="0" pageNumber="341">22A, B</figureCitation>
|
|
), bifurcating into the opening of the solenomere (
|
|
<emphasis id="1348903C4F8901AFD3A19C2CEA2E3945" bold="true" pageId="0" pageNumber="341">s</emphasis>
|
|
) and parasolenomere (
|
|
<emphasis id="A8D992323E05DFC7A4DA59D1960C853C" bold="true" pageId="0" pageNumber="341">ps</emphasis>
|
|
).
|
|
</paragraph>
|
|
<caption id="4C8D14B4DEFE137592171EFD2AF18F91" doi="10.3897/zookeys.1187.113473.figure22" httpUri="https://binary.pensoft.net/fig/958797" pageId="0" pageNumber="341" start="Figure 22" startId="F22">
|
|
<paragraph id="CDEE686E88AA8B858988884AFABD5E63" pageId="0" pageNumber="341">
|
|
<emphasis id="DD34CD8D535A85030E7F500233202EB0" bold="true" pageId="0" pageNumber="341">Figure 22.</emphasis>
|
|
<taxonomicName id="75D74E4B0AAD509C2C2C9161B3D80EB5" authorityName="Wang & Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" genus="Paracortina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracortina voluta" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="voluta">
|
|
<emphasis id="63BFBB292B5F2C861A69BDC517D4FF2D" italics="true" pageId="0" pageNumber="341">Paracortina voluta</emphasis>
|
|
</taxonomicName>
|
|
Wang & Zhang, 1993, male (Rd 5346 ZMUM), left gonopod
|
|
<emphasis id="B9A01A37FC47389952F6B15CBF6D2B97" bold="true" pageId="0" pageNumber="341">A</emphasis>
|
|
anterior view
|
|
<emphasis id="11E6D0F438FCAFE2A301FE25D724AB92" bold="true" pageId="0" pageNumber="341">B</emphasis>
|
|
posterior view
|
|
<emphasis id="FB91F16225246C28EAC900AA0BD918FB" bold="true" pageId="0" pageNumber="341">C</emphasis>
|
|
lateral view
|
|
<emphasis id="D0B77AC7D6B0FD438D21E5F261FF815D" bold="true" pageId="0" pageNumber="341">D</emphasis>
|
|
mesal view. Abbreviations: b = falcate mesal process of coxa; Cl = lateral lobe of coxa; dl =distal lamella of telopodite; pf1 = prefemoroidal process 1; pf2 = prefemoroidal process 2; s = solenomere; T = telopodite.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="783971835FA232C16FC701BA8BB88EBB" pageId="0" pageNumber="341" type="distribution">
|
|
<paragraph id="85E83920E729A30C3E9363D6E72BD241" pageId="0" pageNumber="341">Distribution.</paragraph>
|
|
<paragraph id="1F4781ECF1D3C4FA7CA9256B4EC786E7" pageId="0" pageNumber="341">
|
|
This species was originally described from Ya Jang (= Jajiang) County, Sichuan, China. Here we add a new record from Yanyuan County which is ~ 250 km away in a straight line from the type locality (Fig.
|
|
<figureCitation id="4CCFEA6DE2150B2A76ABEB01F4BD7920" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution map of Paracortinidae in China and Vietnam." figureDoi="10.3897/zookeys.1187.113473.figure28" httpUri="https://binary.pensoft.net/fig/958803" pageId="0" pageNumber="341">28</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C7B6587CFC7225FFD50FEFB3A92589F3" pageId="0" pageNumber="341" type="comments">
|
|
<paragraph id="849918D684F3AE1B821F6D0F989CF511" pageId="0" pageNumber="341">Comments.</paragraph>
|
|
<paragraph id="3393075EDDC8FEA8D33C5A6DC7581ACD" pageId="0" pageNumber="341">
|
|
Although we have no doubt about the identity of the studied specimens, when comparing the gonopod of the new material in mesal view (Fig.
|
|
<figureCitation id="AE925D65E943F3E2696D4DC64AF51003" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Paracortina voluta Wang & Zhang, 1993 (Rd 5346 ZMUM) A female habitus, lateral view B, C male head and anterior pleurotergites B lateral view C frontal view." figureDoi="10.3897/zookeys.1187.113473.figure20" httpUri="https://binary.pensoft.net/fig/958795" pageId="0" pageNumber="341">20D</figureCitation>
|
|
) with the drawing provided in the original description of
|
|
<taxonomicName id="B4350991D2238292BF451092A254B837" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. voluta" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="voluta">
|
|
<emphasis id="D70846C653729794095FBF568B779F50" italics="true" pageId="0" pageNumber="341">P. voluta</emphasis>
|
|
</taxonomicName>
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|
(Wang & Zhang, 1993: fig. 6), we noticed a few differences in the shape of the distal process of the telopodite, with the mesal process more twisted and curved, the anterior lamella more serrated and turned downwards, and the
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<normalizedToken id="DAB9F1132B56F6D090ACDF99392394CB" originalValue="“s-twist”">"s-twist"</normalizedToken>
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more obvious in our specimen (visualised with a rounded notch in mesal view).
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</paragraph>
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<paragraph id="19305C2219BCA9F33617D47710848A34" pageId="0" pageNumber="341">
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Both available identification keys for the family
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<taxonomicName id="8405C76B2B73C7B272BAF511B65AC3A6" authorityName="Wang & Zhang" authorityYear="1993" class="Diplopoda" family="Paracortinidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="family">Paracortinidae</taxonomicName>
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(
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<bibRefCitation id="814F0ABE880885A61E7C60E727E129D4" DOI="https://doi.org/10.2476/asjaa.53.93" author="Stoev, P" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="341" pagination="93 - 103" refId="B21" refString="Stoev, P, Geoffroy, JJ, 2004. Review of the millipede family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida). Acta Arachnologica 53 (2): 93 - 103, DOI: https://doi.org/10.2476/asjaa.53.93" title="Review of the millipede family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida)." url="https://doi.org/10.2476/asjaa.53.93" volume="53" year="2004">Stoev and Geoffroy 2004</bibRefCitation>
|
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;
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<bibRefCitation id="9397655A8C6844C5C8DBDC3B2BF8ACE8" DOI="https://doi.org/10.3897/zookeys.517.9949" author="Liu, WX" journalOrPublisher="ZooKeys" pageId="0" pageNumber="341" pagination="123 - 140" refId="B9" refString="Liu, WX, Tian, MY, 2015. Two new cave-dwelling species of the millipede genus Paracortina Wang & Zhang, 1993 from southern China (Diplopoda, Callipodida, Paracortinidae). ZooKeys 517: 123 - 140, DOI: https://doi.org/10.3897/zookeys.517.9949" title="Two new cave-dwelling species of the millipede genus Paracortina Wang & Zhang, 1993 from southern China (Diplopoda, Callipodida, Paracortinidae)." url="https://doi.org/10.3897/zookeys.517.9949" volume="517" year="2015">Liu and Tian 2015</bibRefCitation>
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) mention eight or nine
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<normalizedToken id="0CB7C5CB8B37DA224CBA658C2D923308" originalValue="“macrosetae”">"macrosetae"</normalizedToken>
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|
on the small prefemoroidal process and use this character to separate the species from its congener
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<taxonomicName id="00C87A3F0EFBD3EFE2AB8337AA34BB42" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. leptoclada" order="Callipodida" pageId="0" pageNumber="341" phylum="Arthropoda" rank="species" species="leptoclada">
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<emphasis id="32F6294AF06987B3E1F981F623E389E9" italics="true" pageId="0" pageNumber="341">P. leptoclada</emphasis>
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</taxonomicName>
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. However, in the specimens we examined, this number greatly exceeds that (see Figs
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<figureCitation id="872C76088242C1FD79F81D2EA25783FF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM) A head, anterior pleurotergites and legs, frontal view B-D gonopods B anterior view C posterior view D telopodite distal part. Abbreviations: b = falcate mesal process of coxa 2; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; ps = parasolenomere; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure21" httpUri="https://binary.pensoft.net/fig/958796" pageId="0" pageNumber="341">21</figureCitation>
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|
,
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<figureCitation id="F616A9D39688ADE909684536FCD650D0" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Paracortina voluta Wang & Zhang, 1993, male (Rd 5346 ZMUM), left gonopod A anterior view B posterior view C lateral view D mesal view. Abbreviations: b = falcate mesal process of coxa; Cl = lateral lobe of coxa; dl = distal lamella of telopodite; pf 1 = prefemoroidal process 1; pf 2 = prefemoroidal process 2; s = solenomere; T = telopodite." figureDoi="10.3897/zookeys.1187.113473.figure22" httpUri="https://binary.pensoft.net/fig/958797" pageId="0" pageNumber="341">22</figureCitation>
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), proving that this character is not reliable for species discrimination.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |