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<document id="9D429925312B9A2FEFA4BE2C05133B02" ID-CLB-Dataset="3500" ID-DOI="10.11646/zootaxa.5068.1.1" ID-GBIF-Dataset="726fdd32-9199-4ca9-9dcd-2892bc30b2e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5702007" ID-ZooBank="F0D7412B-5968-4459-9BA4-F1A314BA8EC7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636961366237" checkinUser="plazi" docAuthor="Cruz-Gómez, Christopher" docDate="2021" docId="044C3804717DEF23D7D3FD2B21C09462" docLanguage="en" docName="zootaxa.5068.1.1.pdf" docOrigin="Zootaxa 5068 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bhawatsonia fusa Cruz-Gómez 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="F875407C7160EF3CD744FFAA2036932F" masterDocTitle="A new genus and seven new species of chrysopetalids (Annelida, Chrysopetalidae) from the Tropical Eastern Pacific" masterLastPageNumber="59" masterPageNumber="1" pageNumber="30" updateTime="1699468561083" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="096B7B8E2578EC647897CEEA311FDF34" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5052F90D68093D990604073F9C145539">
<mods:title id="DF321CDBCEB5C3D767E4703FEF3E2729">A new genus and seven new species of chrysopetalids (Annelida, Chrysopetalidae) from the Tropical Eastern Pacific</mods:title>
</mods:titleInfo>
<mods:name id="41F8BF0C7B82A919860C987CB0CDDE98" type="personal">
<mods:role id="33B513751D4923D6C401FD42966A3443">
<mods:roleTerm id="ABFBFF853FCBAA7A780CB4B88BBA4B9C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="826BC42E1B53256C21005D878C122E73">Cruz-Gómez, Christopher</mods:namePart>
<mods:nameIdentifier id="67F35236843B0EA0504B761562C93C4A" type="ORCID">0000-0002-2618-9230</mods:nameIdentifier>
<mods:affiliation id="A387508692D2A0CB902CC285A1D7682A">Laboratorio de Sistemática de Invertebrados Marinos (LABSIM), Universidad del Mar (UMAR), campus Puerto Ángel, Oaxaca, México Departamento de Sistemática y Ecología Acuática, El Colegio de la Frontera Sur (ECOSUR), Unidad Chetumal, Quintana Roo, México christopher. cruz-gomez @ hotmail. com; https: // orcid. org / 0000 - 0002 - 2618 - 9230</mods:affiliation>
<mods:nameIdentifier id="364A75575C9693AAE84255F4A2BCEEED" type="email">christopher.cruz-gomez@hotmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="C97254539FAF660C04CC28B831D1A75A">text</mods:typeOfResource>
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<mods:title id="08F51C0DB55A0DDFF3262EE983E38434">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="158A07F78DE98607527AAA22AD511FBA">
<mods:date id="B55A92F389BF70D26348CADB5B76E999">2021</mods:date>
<mods:detail id="39A1B38644DBC241CB92B6B63B04800C" type="pubDate">
<mods:number id="EB45926DC3FCCEA32CB44F06CC890CEB">2021-11-15</mods:number>
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<mods:number id="696B7E19541E1C3DD29ACAD73E62FFFF">5068</mods:number>
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<mods:classification id="AA78411184C8032C388F5F3EC49B65A1">journal article</mods:classification>
<mods:identifier id="83E98C1A59362F794CD56FE135867267" type="CLB-Dataset">3500</mods:identifier>
<mods:identifier id="EFD8827DD954B49DC95C3CC418986790" type="DOI">10.11646/zootaxa.5068.1.1</mods:identifier>
<mods:identifier id="569CDDF1AF3584AB64946A47F653EF93" type="GBIF-Dataset">726fdd32-9199-4ca9-9dcd-2892bc30b2e3</mods:identifier>
<mods:identifier id="E92A1AFD96418329DAC36E27583DF978" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9C8AC09E2EF4925C248010D65747D083" type="Zenodo-Dep">5702007</mods:identifier>
<mods:identifier id="822F81B4C52F6309ABB20CDA5E885B78" type="ZooBank">F0D7412B-5968-4459-9BA4-F1A314BA8EC7</mods:identifier>
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<treatment id="044C3804717DEF23D7D3FD2B21C09462" ID-DOI="http://doi.org/10.5281/zenodo.5706147" ID-GBIF-Taxon="190304398" ID-Zenodo-Dep="5706147" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462" httpUri="http://treatment.plazi.org/id/044C3804717DEF23D7D3FD2B21C09462" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="C4FFDA99717DEF21D7D3FD2B218D91B4" box="[151,443,641,668]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8C5A8912717DEF21D7D3FD2B218D91B4" blockId="29.[151,443,641,704]" box="[151,443,641,668]" pageId="29" pageNumber="30">
<heading id="D7123E7E717DEF21D7D3FD2B218D91B4" bold="true" box="[151,443,641,668]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="BE915500717DEF21D7D3FD2B218D91B4" bold="true" box="[151,443,641,668]" pageId="29" pageNumber="30">
<taxonomicName id="4BE5F291717DEF21D7D3FD2B214791B4" ID-CoL="8SM2X" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[151,369,641,668]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717DEF21D7D3FD2B214791B4" bold="true" box="[151,369,641,668]" italics="true" pageId="29" pageNumber="30">Bhawatsonia fusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5A2E87B717DEF21D63CFD2B218D91B4" box="[376,443,641,667]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717DEF21D7D3FD0F243F9027" pageId="29" pageNumber="30" type="description">
<paragraph id="8C5A8912717DEF21D7D3FD0F21089190" blockId="29.[151,443,641,704]" box="[151,318,677,704]" pageId="29" pageNumber="30">
<figureCitation id="14DE9597717DEF21D7D3FD0F212091EF" box="[151,278,677,704]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="30.[151,250,1845,1869]" captionStartId-1="31.[151,250,1340,1364]" captionTargetBox-0="[236,1352,181,1805]" captionTargetBox-1="[171,1435,185,1316]" captionTargetId-0="figure-26@30.[236,1352,181,1821]" captionTargetId-1="figure-15@31.[151,1435,181,1316]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." captionText-1="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi-0="http://doi.org/10.5281/zenodo.5702063" figureDoi-1="http://doi.org/10.5281/zenodo.5702065" httpUri-0="https://zenodo.org/record/5702063/files/figure.png" httpUri-1="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Figs 14–15</figureCitation>
,
<figureCitation id="14DE9597717DEF21D666FD0F21089190" box="[290,318,677,703]" captionStart="FIGURE 26" captionStartId="49.[151,250,1898,1922]" captionTargetBox="[172,1414,917,1868]" captionTargetId="figure-379@49.[166,1422,913,1874]" captionTargetPageId="49" captionText="FIGURE 26. Previous and new records of chrysopetalids in the Tropical Eastern Pacific. Empty symbols indicate historic records." figureDoi="http://doi.org/10.5281/zenodo.5702090" httpUri="https://zenodo.org/record/5702090/files/figure.png" pageId="29" pageNumber="30">26</figureCitation>
</paragraph>
<paragraph id="8C5A8912717DEF21D7D3FD47243F9027" blockId="29.[151,1033,749,776]" box="[151,1033,749,776]" pageId="29" pageNumber="30">
LSID:
<uri id="F8748510717DEF21D7A0FD47243F9027" box="[228,1033,749,776]" pageId="29" pageNumber="30">
urn:lsid:zoobank.org:act:
<uuid id="F843B3C7717DEF21D6B0FD47243F9027" box="[500,1033,749,776]" pageId="29" pageNumber="30">2F7115F8-50BE-4897-A849-4ADC85C2860F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717DEF21D7D3FC81248C906C" box="[151,1210,811,835]" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="8C5A8912717DEF21D7D3FC81248C906C" blockId="29.[151,1210,811,835]" box="[151,1210,811,835]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="ACF5AE3C717DEF21D7D3FC81248C906C" box="[151,1210,811,835]" pageId="29" pageNumber="30">
<taxonomicName id="4BE5F291717DEF21D7D3FC812107906C" box="[151,305,811,835]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawania" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="undefined-3">
<emphasis id="BE915500717DEF21D7D3FC8120C8906C" box="[151,254,811,835]" italics="true" pageId="29" pageNumber="30">Bhawania</emphasis>
sp. 3
</taxonomicName>
.—
<treatmentCitation id="0D44AF03717DEF21D60BFC812248906C" author="Cruz-Gomez, C. &amp; Salazar-Vallejo, S. &amp; Mora-Vallin, Z. G." box="[335,638,811,835]" page="180" pageId="29" pageNumber="30" year="2021">
<bibRefCitation id="E874F4E3717DEF21D60BFC812248906C" author="Cruz-Gomez, C. &amp; Salazar-Vallejo, S. &amp; Mora-Vallin, Z. G." box="[335,638,811,835]" pageId="29" pageNumber="30" pagination="167 - 186" refId="ref35599" refString="Cruz-Gomez, C., Salazar-Vallejo, S. &amp; Mora-Vallin, Z. G. (2021) Chrysopetalidae Ehlers, 1864. In: de Leon-Gonzalez J. A., Bastida-Zavala J. R., Carrera-Parra L. F., Garcia-Garza M. E., Salazar-Vallejo S. I., Solis-Weiss V. &amp; Tovar-Hernandez M. A. (Eds.), Anelidos Marinos de Mexico y America Tropical. Universidad Autonoma de Nuevo Leon, Nuevo Leon, pp. 167 - 186." type="book chapter" year="2021">
Cruz-Gómez
<emphasis id="BE915500717DEF21D699FC862226906C" box="[477,528,811,835]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2021: 180
</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="14DE9597717DEF21D5CDFC8122E3906C" box="[649,725,811,835]" captionStart="FIGURE 5" captionStartId="11.[151,250,1910,1934]" captionTargetBox="[167,1414,181,1881]" captionTargetId="figure-15@11.[167,1420,181,1886]" captionTargetPageId="11" captionText="FIGURE 5. Paleanotus karlyae n. sp., holotype (ECOSUR-258). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of pygidium, ventral view. D) Parapodium from segment 23. E) Lateral paleae. F) Subunit 1. G) Main palea. H) Apex of G. I–J) Median paleae. K) Unit 1. L) Unit 2. M) Unit 3. N) Unit 4. Scales: 10 µm (KLMN), 20 µm (CEFH), 50 µm (IJ), 100 µm (B), 200 µm (D), 500 µm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702038" httpUri="https://zenodo.org/record/5702038/files/figure.png" pageId="29" pageNumber="30">Fig. 5A</figureCitation>
(key of chrysopetalids from Tropical America).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717DEF21D7D3FCDB233B974C" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8C5A8912717DEF21D7D3FCDB225A90A4" blockId="29.[151,1437,881,2024]" box="[151,620,881,907]" pageId="29" pageNumber="30">
<materialsCitation id="3C8D834F717DEF21D7D3FCDB225E90A4" ID-GBIF-Occurrence="3406812405" box="[151,616,881,907]" country="Mexico" location="Aticama" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Nayarit" typeStatus="holotype">
<emphasis id="BE915500717DEF21D7D3FCDB210F90A4" bold="true" box="[151,313,881,907]" pageId="29" pageNumber="30">
<typeStatus id="535E37B0717DEF21D7D3FCDB20E490A4" box="[151,210,881,907]" pageId="29" pageNumber="30">Type</typeStatus>
locality:
</emphasis>
<location id="893ADFC9717DEF21D604FCDB219490A4" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462:893ADFC9717DEF21D604FCDB219490A4" box="[320,418,881,907]" country="Mexico" name="Aticama" pageId="29" pageNumber="30" stateProvince="Nayarit">Aticama</location>
,
<collectingRegion id="4E2147F0717DEF21D6E9FCDB223290A4" box="[429,516,881,907]" country="Mexico" name="Nayarit" pageId="29" pageNumber="30">Nayarit</collectingRegion>
,
<collectingCountry id="F4F2C982717DEF21D54BFCDB225E90A4" box="[527,616,881,907]" name="Mexico" pageId="29" pageNumber="30">México</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8C5A8912717DEF21D783FC3F236490FC" blockId="29.[151,1437,881,2024]" pageId="29" pageNumber="30">
<emphasis id="BE915500717DEF21D783FC3F21419080" bold="true" box="[199,375,917,943]" pageId="29" pageNumber="30">Type material.</emphasis>
<materialsCitation id="3C8D834F717DEF21D6C4FC3F215B90FC" ID-GBIF-Occurrence="3406812336" collectingDate="2004-11-25" collectionCode="ECOSUR" country="Mexico" elevation="2" location="Aticama" municipality="northern Mexican Pacific" pageId="29" pageNumber="30" specimenCode="ECOSUR-263" specimenCount="1" stateProvince="Nayarit" typeStatus="holotype">
<emphasis id="BE915500717DEF21D6C4FC3F21C09080" bold="true" box="[384,502,917,943]" pageId="29" pageNumber="30">
<typeStatus id="535E37B0717DEF21D6C4FC3F21C79080" box="[384,497,917,943]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DC432169717DEF21D6BBFC3F22AA9080" box="[511,668,917,943]" collectionCode="ECOSUR-" pageId="29" pageNumber="30">ECOSUR-263</specimenCode>
,
<location id="893ADFC9717DEF21D5E3FC3F23359080" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462:893ADFC9717DEF21D5E3FC3F23359080" box="[679,771,917,943]" country="Mexico" municipality="northern Mexican Pacific" name="Aticama" pageId="29" pageNumber="30" stateProvince="Nayarit">Aticama</location>
,
<collectingRegion id="4E2147F0717DEF21D454FC3F23549080" box="[784,866,917,943]" country="Mexico" name="Nayarit" pageId="29" pageNumber="30">Nayarit</collectingRegion>
,
<collectingCountry id="F4F2C982717DEF21D429FC3F23F79080" box="[877,961,917,943]" name="Mexico" pageId="29" pageNumber="30">México</collectingCountry>
,
<collectingMunicipality id="6C3E1368717DEF21D489FC3F24E99080" box="[973,1247,917,943]" pageId="29" pageNumber="30">northern Mexican Pacific</collectingMunicipality>
, on bivalve,
<quantity id="4B1D24F7717DEF21D22FFC3F25AE909F" box="[1387,1432,917,944]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="m" value="2.0">
<elevation id="07C86E21717DEF21D22FFC3F25AE909F" box="[1387,1432,917,944]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="m" value="2.0">2 m</elevation>
</quantity>
,
<date id="F85BAFD2717DEF21D7D3FC13215B90FC" box="[151,365,953,980]" pageId="29" pageNumber="30" value="2004-11-25">
<collectingDate id="E81F563A717DEF21D7D3FC13215B90FC" box="[151,365,953,980]" pageId="29" pageNumber="30" value="2004-11-25">November 25, 2004</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3C8D834F717DEF21D633FC13237890FC" ID-GBIF-Occurrence="3406812373" box="[375,846,953,980]" collectionCode="ECOSUR-" country="Mexico" pageId="29" pageNumber="30" specimenCode="ECOSUR-264" specimenCount="2" stateProvince="Nayarit" typeStatus="paratype">
<emphasis id="BE915500717DEF21D633FC1321DE90FC" bold="true" box="[375,488,953,979]" pageId="29" pageNumber="30">
<typeStatus id="535E37B0717DEF21D633FC1321D590FC" box="[375,483,953,979]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCode id="DC432169717DEF21D6A9FC1322BC90FC" box="[493,650,953,979]" collectionCode="ECOSUR-" pageId="29" pageNumber="30">ECOSUR-264</specimenCode>
, same as
<typeStatus id="535E37B0717DEF21D5AAFC13237890FC" box="[750,846,953,979]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8C5A8912717DEF21D783FC77233B974C" blockId="29.[151,1437,881,2024]" pageId="29" pageNumber="30">
<emphasis id="BE915500717DEF21D783FC7721DA90D8" bold="true" box="[199,492,989,1015]" pageId="29" pageNumber="30">Other material examined.</emphasis>
<materialsCitation id="3C8D834F717DEF21D6B5FC77246490D7" ID-GBIF-Occurrence="3406812420" box="[497,1106,989,1016]" collectionCode="ECOSUR" collectorName="Six" location="Baja California" pageId="29" pageNumber="30" specimenCode="ECOSUR-3219" specimenCount="2" stateProvince="Baja California">
<collectorName id="2110ECC4717DEF21D6B5FC77222290D8" box="[497,532,989,1015]" pageId="29" pageNumber="30">Six</collectorName>
specimens.
<emphasis id="BE915500717DEF21D5D1FC77237890D8" bold="true" box="[661,846,989,1015]" pageId="29" pageNumber="30">
<collectingRegion id="4E2147F0717DEF21D5D1FC77237C90D8" box="[661,842,989,1015]" country="Mexico" name="Baja California" pageId="29" pageNumber="30">Baja California</collectingRegion>
:
</emphasis>
<specimenCode id="DC432169717DEF21D410FC7723C990D8" box="[852,1023,989,1015]" collectionCode="ECOSUR-" pageId="29" pageNumber="30">ECOSUR-3219</specimenCode>
,
<specimenCount id="9AE3429B717DEF21D34CFC77246490D7" box="[1032,1106,989,1016]" pageId="29" pageNumber="30" type="generic">2 spec.</specimenCount>
</materialsCitation>
<materialsCitation id="3C8D834F717DEF21D313FC7721779734" ID-GBIF-Occurrence="3406812374" collectingDate="1986-05-25" collectionCode="SSV" collectorName="Bahia" location="Angeles" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Baja California">
<collectorName id="2110ECC4717DEF21D313FC7724A390D8" box="[1111,1173,989,1015]" pageId="29" pageNumber="30">Bahía</collectorName>
de los
<location id="893ADFC9717DEF21D3A4FC75250F90D5" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462:893ADFC9717DEF21D3A4FC75250F90D5" box="[1248,1337,991,1018]" name="Angeles" pageId="29" pageNumber="30" stateProvince="Baja California">Ángeles</location>
,
<date id="F85BAFD2717DEF21D207FC7420F89734" pageId="29" pageNumber="30" value="1986-05-25">
<collectingDate id="E81F563A717DEF21D207FC7420F89734" pageId="29" pageNumber="30" value="1986-05-25">May 25, 1986</collectingDate>
</date>
, coll.
<collectionCode id="EAF411D7717DEF21D654FBAB21779734" box="[272,321,1025,1051]" pageId="29" pageNumber="30">SSV</collectionCode>
</materialsCitation>
.
<materialsCitation id="3C8D834F717DEF21D60FFBAB22DF9734" ID-GBIF-Occurrence="3406812357" box="[331,745,1025,1051]" collectionCode="ECOSUR" location="Baja California Sur" pageId="29" pageNumber="30" specimenCode="ECOSUR-3220" specimenCount="1" stateProvince="Baja California Sur">
<emphasis id="BE915500717DEF21D60FFBAB22019734" bold="true" box="[331,567,1025,1051]" pageId="29" pageNumber="30">
<collectingRegion id="4E2147F0717DEF21D60FFBAB22049734" box="[331,562,1025,1051]" country="Mexico" name="Baja California Sur" pageId="29" pageNumber="30">Baja California Sur</collectingRegion>
:
</emphasis>
<specimenCode id="DC432169717DEF21D57AFBAB22DF9734" box="[574,745,1025,1051]" collectionCode="ECOSUR-" pageId="29" pageNumber="30">ECOSUR-3220</specimenCode>
</materialsCitation>
,
<materialsCitation id="3C8D834F717DEF21D5B0FBAB21A3976F" ID-GBIF-Occurrence="3406812361" collectingDate="1987-10-04" collectionCode="SSV, ALG" collectorName="El Presidente" location="Bay" pageId="29" pageNumber="30" specimenCode="ECOSUR-3221" specimenCount="2" stateProvince="La Paz">
<collectorName id="2110ECC4717DEF21D5B0FBAB23B59734" box="[756,899,1025,1051]" pageId="29" pageNumber="30">El Presidente</collectorName>
,
<collectingRegion id="4E2147F0717DEF21D4CBFBA823EF9733" box="[911,985,1026,1052]" country="Honduras" name="La Paz" pageId="29" pageNumber="30">La Paz</collectingRegion>
<location id="893ADFC9717DEF21D4A5FBA8243A9733" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462:893ADFC9717DEF21D4A5FBA8243A9733" box="[993,1036,1026,1052]" name="Bay" pageId="29" pageNumber="30" stateProvince="La Paz">Bay</location>
,
<date id="F85BAFD2717DEF21D35CFBAB24FE9734" box="[1048,1224,1025,1051]" pageId="29" pageNumber="30" value="1987-10-04">
<collectingDate id="E81F563A717DEF21D35CFBAB24FE9734" box="[1048,1224,1025,1051]" pageId="29" pageNumber="30" value="1987-10-04">October 4, 1987</collectingDate>
</date>
, coll.
<collectionCode id="EAF411D7717DEF21D24DFBAB250C9734" box="[1289,1338,1025,1051]" pageId="29" pageNumber="30">SSV</collectionCode>
&amp;
<collectionCode id="EAF411D7717DEF21D219FBAB25AE9734" box="[1373,1432,1025,1051]" pageId="29" pageNumber="30">ALG</collectionCode>
;
<specimenCode id="DC432169717DEF21D7D3FB8F21779710" box="[151,321,1061,1087]" collectionCode="ECOSUR-" pageId="29" pageNumber="30">ECOSUR-3221</specimenCode>
,
<specimenCount id="9AE3429B717DEF21D60FFB8F21A3976F" box="[331,405,1061,1088]" pageId="29" pageNumber="30" type="generic">2 spec.</specimenCount>
</materialsCitation>
<materialsCitation id="3C8D834F717DEF21D6DEFB8F23EA9710" ID-GBIF-Occurrence="3406812395" box="[410,988,1061,1088]" collectingDate="1987-10-10" collectionCode="SSV" collectorName="El Presidente" location="Bay" pageId="29" pageNumber="30" specimenCount="1" stateProvince="La Paz">
<collectorName id="2110ECC4717DEF21D6DEFB8F22119710" box="[410,551,1061,1087]" pageId="29" pageNumber="30">El Presidente</collectorName>
,
<collectingRegion id="4E2147F0717DEF21D575FB8C224F976F" box="[561,633,1062,1088]" country="Honduras" name="La Paz" pageId="29" pageNumber="30">La Paz</collectingRegion>
<location id="893ADFC9717DEF21D53BFB8C229C976F" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462:893ADFC9717DEF21D53BFB8C229C976F" box="[639,682,1062,1088]" name="Bay" pageId="29" pageNumber="30" stateProvince="La Paz">Bay</location>
,
<date id="F85BAFD2717DEF21D5F1FB8F23589710" box="[693,878,1061,1087]" pageId="29" pageNumber="30" value="1987-10-10">
<collectingDate id="E81F563A717DEF21D5F1FB8F23589710" box="[693,878,1061,1087]" pageId="29" pageNumber="30" value="1987-10-10">October 10, 1987</collectingDate>
</date>
, coll.
<collectionCode id="EAF411D7717DEF21D4EFFB8F23EA9710" box="[939,988,1061,1087]" pageId="29" pageNumber="30">SSV</collectionCode>
</materialsCitation>
.
<materialsCitation id="3C8D834F717DEF21D4A1FB8F250B976F" ID-GBIF-Occurrence="3406812389" box="[997,1341,1061,1088]" collectionCode="ECOSUR" location="Jalisco" pageId="29" pageNumber="30" specimenCode="ECOSUR-3222" specimenCount="2" stateProvince="Jalisco">
<emphasis id="BE915500717DEF21D4A1FB8F240D9710" bold="true" box="[997,1083,1061,1087]" pageId="29" pageNumber="30">
<collectingRegion id="4E2147F0717DEF21D4A1FB8F24009710" box="[997,1078,1061,1087]" country="Mexico" name="Jalisco" pageId="29" pageNumber="30">Jalisco</collectingRegion>
:
</emphasis>
<specimenCode id="DC432169717DEF21D37BFB8F24DC9710" box="[1087,1258,1061,1087]" collectionCode="ECOSUR-" pageId="29" pageNumber="30">ECOSUR-3222</specimenCode>
,
<specimenCount id="9AE3429B717DEF21D3B0FB8F250B976F" box="[1268,1341,1061,1088]" pageId="29" pageNumber="30" type="generic">2 spec.</specimenCount>
</materialsCitation>
<materialsCitation id="3C8D834F717DEF21D206FB8C233F974C" ID-GBIF-Occurrence="3406812324" collectingDate="2004-12-03" collectionCode="BY, PSS" elevation="4" location="Barra de Navidad" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Jalisco">
<location id="893ADFC9717DEF21D206FB8C20C5974C" LSID="urn:lsid:plazi:treatment:044C3804717DEF23D7D3FD2B21C09462:893ADFC9717DEF21D206FB8C20C5974C" name="Barra de Navidad" pageId="29" pageNumber="30" stateProvince="Jalisco">Barra de Navidad</location>
, on rock,
<quantity id="4B1D24F7717DEF21D61EFBE321B3974B" box="[346,389,1097,1124]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="m" value="4.0">
<elevation id="07C86E21717DEF21D61EFBE321B3974B" box="[346,389,1097,1124]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="m" value="4.0">4 m</elevation>
</quantity>
,
<date id="F85BAFD2717DEF21D6CBFBE32263974C" box="[399,597,1097,1123]" pageId="29" pageNumber="30" value="2004-12-03">
<collectingDate id="E81F563A717DEF21D6CBFBE32263974C" box="[399,597,1097,1123]" pageId="29" pageNumber="30" value="2004-12-03">December 3, 2004</collectingDate>
</date>
, coll.
<collectionCode id="EAF411D7717DEF21D5D7FBE0228F974B" box="[659,697,1098,1124]" pageId="29" pageNumber="30">BY</collectionCode>
&amp;
<collectionCode id="EAF411D7717DEF21D59DFBE3233F974C" box="[729,777,1097,1123]" country="0" httpUri="http://grbio.org/cool/vv09-ccp1" name="Paleontology and Stratigraphic Section of the Geological Institute of the Mongolian Academy of Sciences" pageId="29" pageNumber="30">PSS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717DEF21D783FBC72548941C" pageId="29" pageNumber="30" type="description">
<paragraph id="8C5A8912717DEF21D783FBC7225697E0" blockId="29.[151,1437,881,2024]" pageId="29" pageNumber="30">
<emphasis id="BE915500717DEF21D783FBC7216797A8" bold="true" box="[199,337,1133,1159]" pageId="29" pageNumber="30">Description.</emphasis>
Based on the
<typeStatus id="535E37B0717DEF21D6ACFBC7227E97A8" box="[488,584,1133,1159]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
: complete with 74 segments. TL=
<quantity id="4B1D24F7717DEF21D4FDFBC723CC97A7" box="[953,1018,1133,1160]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="29" pageNumber="30" unit="mm" value="8.0">8 mm</quantity>
, TW= 1.8 mm. Body stiffed, long, narrowed posteriorly (
<figureCitation id="14DE9597717DEF21D620FB3B21FE9784" box="[356,456,1169,1195]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14A</figureCitation>
). Body orange to brown, dorsal and ventral cirri bases darker (
<figureCitation id="14DE9597717DEF21D32AFB3B24E49784" box="[1134,1234,1169,1195]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14G</figureCitation>
). Paleae fan bright maroon to golden, non-imbricated dorsally.
</paragraph>
<paragraph id="8C5A8912717DEF21D783FB7325689670" blockId="29.[151,1437,881,2024]" pageId="29" pageNumber="30">
Prostomium retracted among the first four segments. Lateral antennae long, inserted on antero-ventral prostomial margin, median antenna slightly shorter than lateral ones, inserted in front of anterior eyes. Eyes red-violet, two pairs. Nuchal organ, semicircular nuchal fold, partially covering prostomium (
<figureCitation id="14DE9597717DEF21D40CFA8B23FA9613" box="[840,972,1313,1340]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14B, D</figureCitation>
). Palps short, ovoid, visible only in ventral view. Mouth fold small, placed between segment 3 and 4 (
<figureCitation id="14DE9597717DEF21D5BAFAEF234D964F" box="[766,891,1349,1376]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14C, E</figureCitation>
). Pharynx eversible, not exposed, stylet thick.
</paragraph>
<paragraph id="8C5A8912717DEF21D783FAC320C29574" blockId="29.[151,1437,881,2024]" pageId="29" pageNumber="30">
Parapodium from segment 49, notochaetae in three main groups (
<figureCitation id="14DE9597717DEF21D430FAC323E396AC" box="[884,981,1385,1411]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14H</figureCitation>
). Notochaetae: lateral group inserted below notaciculum, 1 paleae, very slender and symmetrical, with 1–2 (3) internal ribs (
<figureCitation id="14DE9597717DEF21D4A9FA2724739688" box="[1005,1093,1421,1447]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14I</figureCitation>
); subunit 1, 1 palea, slender and symmetrical, with 9–10 (11) internal ribs (
<figureCitation id="14DE9597717DEF21D520FA1B228996E4" box="[612,703,1457,1483]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14J</figureCitation>
). Main group, 7–9 paleae, broad and symmetrical, with (14) 15–18 (20) internal ribs and 4–5 finely raised ribs (
<figureCitation id="14DE9597717DEF21D53EFA7F22E896C0" box="[634,734,1493,1519]" captionStart="FIGURE 14" captionStartId="30.[151,250,1845,1869]" captionTargetBox="[236,1352,181,1805]" captionTargetId="figure-26@30.[236,1352,181,1821]" captionTargetPageId="30" captionText="FIGURE 14. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="29" pageNumber="30">Fig. 14K</figureCitation>
). Median group, 3 paleae, shorter, slender and asymmetrical; the distal-most palea with 10–11 internal ribs and 3 raised ribs; the next proximal palea smaller with (8) 10 internal ribs and 1 raised rib, and the proximal-most and smallest with 8 internal ribs and 1 raised rib (
<figureCitation id="14DE9597717DEF21D348F9B724B99517" box="[1036,1167,1565,1592]" captionStart="FIGURE 15" captionStartId="31.[151,250,1340,1364]" captionTargetBox="[171,1435,185,1316]" captionTargetId="figure-15@31.[151,1435,181,1316]" captionTargetPageId="31" captionText="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Fig. 15A–B</figureCitation>
). All paleae with serrated margins.
</paragraph>
<paragraph id="8C5A8912717DEF21D783F9CF22A0943F" blockId="29.[151,1437,881,2024]" pageId="29" pageNumber="30">
Neuropodium conical, smaller than notopodium. Neurochaetae: unit 1, 2 superior spinigers, blades straight and long, 15–16 times longer than wide (
<figureCitation id="14DE9597717DEF21D6ACF923227C958B" box="[488,586,1673,1700]" captionStart="FIGURE 15" captionStartId="31.[151,250,1340,1364]" captionTargetBox="[171,1435,185,1316]" captionTargetId="figure-15@31.[151,1435,181,1316]" captionTargetPageId="31" captionText="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Fig. 15C</figureCitation>
). Unit 2, 6 falcigers with blades straight and short, 1–2 times longer than wide (
<figureCitation id="14DE9597717DEF21D7DBF907213695E7" box="[159,256,1709,1736]" captionStart="FIGURE 15" captionStartId="31.[151,250,1340,1364]" captionTargetBox="[171,1435,185,1316]" captionTargetId="figure-15@31.[151,1435,181,1316]" captionTargetPageId="31" captionText="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Fig. 15D</figureCitation>
). Unit 3, 4 falcigers with blades straight and short, 2 times longer than wide (
<figureCitation id="14DE9597717DEF21D364F90724AC95E7" box="[1056,1178,1709,1736]" captionStart="FIGURE 15" captionStartId="31.[151,250,1340,1364]" captionTargetBox="[171,1435,185,1316]" captionTargetId="figure-15@31.[151,1435,181,1316]" captionTargetPageId="31" captionText="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Fig. 15E–F</figureCitation>
). Unit 4, 2 falcigers with blades straight and medium-sized, 5–6 times longer than wide (
<figureCitation id="14DE9597717DEF21D468F97B23BB95C3" box="[812,909,1745,1772]" captionStart="FIGURE 15" captionStartId="31.[151,250,1340,1364]" captionTargetBox="[171,1435,185,1316]" captionTargetId="figure-15@31.[151,1435,181,1316]" captionTargetPageId="31" captionText="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Fig. 15G</figureCitation>
). Blades bidentate, bent distally. Pygidium conical with two anal cirri (
<figureCitation id="14DE9597717DEF21D622F95F21FE943F" box="[358,456,1781,1808]" captionStart="FIGURE 15" captionStartId="31.[151,250,1340,1364]" captionTargetBox="[171,1435,185,1316]" captionTargetId="figure-15@31.[151,1435,181,1316]" captionTargetPageId="31" captionText="FIGURE 15. Bhawatsonia fusa n. sp., holotype (ECOSUR-263). A–B) Median paleae. C) Unit 1. D) Unit 2. E–F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="29" pageNumber="30">Fig. 15H</figureCitation>
). Oocytes not seen.
</paragraph>
<paragraph id="8C5A8912717DEF21D783F8B32548941C" blockId="29.[151,1437,881,2024]" box="[199,1406,1817,1844]" pageId="29" pageNumber="30">
<emphasis id="BE915500717DEF21D783F8B32113941C" bold="true" box="[199,293,1817,1843]" pageId="29" pageNumber="30">Habitat.</emphasis>
Subtidal (
<quantity id="4B1D24F7717DEF21D6D4F8B321EE941B" box="[400,472,1817,1844]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
). Specimens of this species were collected mainly on rocks, and as epibionts of bivalves.
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717DEF21D783F89724F29477" box="[199,1220,1853,1882]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8C5A8912717DEF21D783F89724F29477" blockId="29.[151,1437,881,2024]" box="[199,1220,1853,1882]" pageId="29" pageNumber="30">
<emphasis id="BE915500717DEF21D783F89721619478" bold="true" box="[199,343,1853,1879]" pageId="29" pageNumber="30">Distribution.</emphasis>
From Bahía de los Ángeles,
<collectingRegion id="4E2147F0717DEF21D5CEF897231D9478" box="[650,811,1853,1879]" country="Mexico" name="Baja California" pageId="29" pageNumber="30">Baja California</collectingRegion>
to Barra de Navidad,
<collectingRegion id="4E2147F0717DEF21D357F897246C9478" box="[1043,1114,1853,1879]" country="Mexico" name="Jalisco" pageId="29" pageNumber="30">Jalisco</collectingRegion>
(
<figureCitation id="14DE9597717DEF21D32CF89724819477" box="[1128,1207,1853,1880]" captionStart="FIGURE 26" captionStartId="49.[151,250,1898,1922]" captionTargetBox="[172,1414,917,1868]" captionTargetId="figure-379@49.[166,1422,913,1874]" captionTargetPageId="49" captionText="FIGURE 26. Previous and new records of chrysopetalids in the Tropical Eastern Pacific. Empty symbols indicate historic records." figureDoi="http://doi.org/10.5281/zenodo.5702090" httpUri="https://zenodo.org/record/5702090/files/figure.png" pageId="29" pageNumber="30">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717DEF23D783F8CB23C395CF" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8C5A8912717DEF23D783F8CB21939502" blockId="29.[151,1437,881,2024]" lastBlockId="31.[151,1437,1482,1869]" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<emphasis id="BE915500717DEF21D783F8CB21009454" bold="true" box="[199,310,1889,1915]" pageId="29" pageNumber="30">Remarks.</emphasis>
<taxonomicName id="4BE5F291717DEF21D605F8CB22359454" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[321,515,1889,1915]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717DEF21D605F8CB22359454" box="[321,515,1889,1915]" italics="true" pageId="29" pageNumber="30">Bhawatsonia fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717DEF21D549F8CB227B9454" bold="true" box="[525,589,1889,1915]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A5A2E87B717DEF21D549F8CB227B9454" box="[525,589,1889,1915]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, along with
<taxonomicName id="4BE5F291717DEF21D59AF8CB23A79454" box="[734,913,1889,1916]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="BE915500717DEF21D59AF8CB235F9454" box="[734,873,1889,1915]" italics="true" pageId="29" pageNumber="30">Bhawatsonia</emphasis>
sp.
</taxonomicName>
, belongs to the
<taxonomicName id="4BE5F291717DEF21D30FF8CB24E09454" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[1099,1238,1889,1915]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="BE915500717DEF21D30FF8CB24E09454" box="[1099,1238,1889,1915]" italics="true" pageId="29" pageNumber="30">Bhawatsonia</emphasis>
</taxonomicName>
<emphasis id="BE915500717DEF21D3A4F8CB25199454" bold="true" box="[1248,1327,1889,1915]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A5A2E87B717DEF21D3A4F8CB25199454" box="[1248,1327,1889,1915]" pageId="29" pageNumber="30" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
members with broad main paleae and slender median paleae. Specimens of those species can be differentiated by the form of the main paleae; in
<taxonomicName id="4BE5F291717DEF21D607F80021B994EC" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[323,399,1961,1987]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717DEF21D607F80021B994EC" box="[323,399,1961,1987]" italics="true" pageId="29" pageNumber="30">B. fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717DEF21D6D2F80321E594EC" bold="true" box="[406,467,1961,1987]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A5A2E87B717DEF21D6D2F80321E594EC" box="[406,467,1961,1987]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
specimens have broader main paleae, whereas specimens of
<taxonomicName id="4BE5F291717DEF21D322F803252194EB" box="[1126,1303,1961,1988]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="BE915500717DEF21D322F80324C794EC" box="[1126,1265,1961,1987]" italics="true" pageId="29" pageNumber="30">Bhawatsonia</emphasis>
sp.
</taxonomicName>
have slightly slender main paleae. In addition, subunit 1 appears to be taxonomically relevant in the members of the new genus. Specimens of
<taxonomicName id="4BE5F291717FEF23D67CFA6121B196CA" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[312,391,1483,1509]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717FEF23D67CFA6121B196CA" box="[312,391,1483,1509]" italics="true" pageId="31" pageNumber="32">B. fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D6CAFA6121F896CA" bold="true" box="[398,462,1483,1509]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D6CAFA6121F896CA" box="[398,462,1483,1509]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
have the subunit 1 with a rounded tip, and a range of internal ribs of 9–10, maximum of 12; whereas, specimens of
<taxonomicName id="4BE5F291717FEF23D6ACFA4422949526" box="[488,674,1518,1545]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="BE915500717FEF23D6ACFA44224C9527" box="[488,634,1518,1544]" italics="true" pageId="31" pageNumber="32">Bhawatsonia</emphasis>
sp.
</taxonomicName>
have the subunit 1 with an acute tip, and a range of internal ribs of 12–13, maximum of 14.
</paragraph>
<caption id="D89AD99A717EEF22D7D3F89F239794F1" ID-DOI="http://doi.org/10.5281/zenodo.5702063" ID-Zenodo-Dep="5702063" httpUri="https://zenodo.org/record/5702063/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1845,1869]" targetBox="[236,1352,181,1805]" targetPageId="30">
<paragraph id="8C5A8912717EEF22D7D3F89F239794F1" blockId="30.[151,1437,1845,2014]" pageId="30" pageNumber="31">
<emphasis id="BE915500717EEF22D7D3F89F21179462" bold="true" box="[151,289,1845,1869]" pageId="30" pageNumber="31">FIGURE 14.</emphasis>
<taxonomicName id="4BE5F291717EEF22D66CF89F21EB9461" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[296,477,1845,1870]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717EEF22D66CF89F21EB9461" box="[296,477,1845,1870]" italics="true" pageId="30" pageNumber="31">Bhawatsonia fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717EEF22D6A0F89C22289461" bold="true" box="[484,542,1846,1870]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A5A2E87B717EEF22D6A0F89C22289461" box="[484,542,1846,1870]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (ECOSUR-263). A) Complete fragmented body. B) Drawing of prostomium, dorsal view. C) Drawing of prostomium, ventral view. D) Prostomium, dorsal view. E) Prostomium, ventral view; arrow indicates Mo. F) Right side of middle segments, arrow indicates dorsal cirrus. G) Dorsal side of the specimen, showing the middorsal line paleae free. H) Parapodium from segment 49. I) Lateral paleae. J) Subunit 1. K) Main palea, arrow indicates finely raised rib. Scales: 20 µm (IJK), 100 µm (BCDEFH), 500 µm (G), 1 mm (A).
</paragraph>
</caption>
<caption id="D89AD99A717FEF23D7D3FA96215C96B3" ID-DOI="http://doi.org/10.5281/zenodo.5702065" ID-Zenodo-Dep="5702065" httpUri="https://zenodo.org/record/5702065/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1340,1364]" targetBox="[171,1435,185,1316]" targetPageId="31">
<paragraph id="8C5A8912717FEF23D7D3FA96215C96B3" blockId="31.[151,1436,1340,1437]" pageId="31" pageNumber="32">
<emphasis id="BE915500717FEF23D7D3FA962116967B" bold="true" box="[151,288,1340,1364]" pageId="31" pageNumber="32">FIGURE 15.</emphasis>
<taxonomicName id="4BE5F291717FEF23D663FA9621EB967B" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[295,477,1340,1364]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717FEF23D663FA9621EB967B" box="[295,477,1340,1364]" italics="true" pageId="31" pageNumber="32">Bhawatsonia fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D6A0FA962228967B" bold="true" box="[484,542,1340,1364]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D6A0FA962228967B" box="[484,542,1340,1364]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (ECOSUR-263). A
<emphasis id="BE915500717FEF23D408FA96236F967B" bold="true" box="[844,857,1340,1364]" pageId="31" pageNumber="32">–</emphasis>
B) Median paleae. C) Unit 1. D) Unit 2. E
<emphasis id="BE915500717FEF23D24DFA962520967B" bold="true" box="[1289,1302,1340,1364]" pageId="31" pageNumber="32">–</emphasis>
F) Unit 3. G) Unit 4. H) Pygidium, ventral view. I) Complete body, largest specimen (ECOSUR-3219). Scales: 10 µm (CDEFG), 20 µm (AB), 50 µm (H),1 mm (I).
</paragraph>
</caption>
<paragraph id="8C5A8912717FEF23D783F99D23C395CF" blockId="31.[151,1437,1482,1869]" pageId="31" pageNumber="32">
Occurrence with specimens of
<taxonomicName id="4BE5F291717FEF23D566F99D229F957F" authorityName="Cruz-Gómez" authorityYear="2021" baseAuthorityName="Cruz-Gómez" baseAuthorityYear="2021" box="[546,681,1591,1616]" class="Polychaeta" family="Chrysopetalidae" genus="Paleanotus" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="purpurea" status="comb. nov.">
<emphasis id="BE915500717FEF23D566F99D229F957F" box="[546,681,1591,1616]" italics="true" pageId="31" pageNumber="32">B. purpurea</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D5F4F99D2320957E" bold="true" box="[688,790,1591,1617]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D5F4F99D2320957E" box="[688,790,1591,1617]" pageId="31" pageNumber="32" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
is common; however, both species are different.
<taxonomicName id="4BE5F291717FEF23D279F99C213F955A" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717FEF23D279F99C213F955A" italics="true" pageId="31" pageNumber="32">Bhawatsonia fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D64BF9F1217B955A" bold="true" box="[271,333,1627,1653]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D64BF9F1217B955A" box="[271,333,1627,1653]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
possess subunit 1 broad, with slightly higher rib number, 9–10, maximum 11; while
<taxonomicName id="4BE5F291717FEF23D3BEF9F12549955B" authorityName="Cruz-Gómez" authorityYear="2021" baseAuthorityName="Cruz-Gómez" baseAuthorityYear="2021" box="[1274,1407,1627,1652]" class="Polychaeta" family="Chrysopetalidae" genus="Paleanotus" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="purpurea" status="comb. nov.">
<emphasis id="BE915500717FEF23D3BEF9F12549955B" box="[1274,1407,1627,1652]" italics="true" pageId="31" pageNumber="32">B. purpurea</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D2C2F9F120E995B6" bold="true" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D2C2F9F120E995B6" pageId="31" pageNumber="32" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
possess subunit 1 slender, with 8–10 internal ribs. Further, the main paleae of
<taxonomicName id="4BE5F291717FEF23D300F9D524A795B6" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" box="[1092,1169,1663,1689]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="fusa" status="sp. nov.">
<emphasis id="BE915500717FEF23D300F9D524A795B6" box="[1092,1169,1663,1689]" italics="true" pageId="31" pageNumber="32">B. fusa</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D3D3F9D524E395B6" bold="true" box="[1175,1237,1663,1689]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D3D3F9D524E395B6" box="[1175,1237,1663,1689]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is clearly broader and shorter with slightly reduced rib number, 15–18, maximum 20 internal ribs; meanwhile,
<taxonomicName id="4BE5F291717FEF23D3D8F90925179593" authority="Cruz-Gómez, 2021" authorityName="Cruz-Gómez" authorityYear="2021" baseAuthorityName="Cruz-Gómez" baseAuthorityYear="2021" box="[1180,1313,1699,1724]" class="Polychaeta" family="Chrysopetalidae" genus="Paleanotus" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="purpurea" status="sp. nov.">
<emphasis id="BE915500717FEF23D3D8F90925179593" box="[1180,1313,1699,1724]" italics="true" pageId="31" pageNumber="32">B. purpurea</emphasis>
</taxonomicName>
<emphasis id="BE915500717FEF23D263F90925539592" bold="true" box="[1319,1381,1699,1725]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A5A2E87B717FEF23D263F90925539592" box="[1319,1381,1699,1725]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
possesses slender and longer main paleae with 17–18, maximum 23 interna ribs.
</paragraph>
</subSubSection>
<subSubSection id="C4FFDA99717FEF23D783F94121C09462" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8C5A8912717FEF23D783F94121C09462" blockId="31.[151,1437,1482,1869]" pageId="31" pageNumber="32">
<emphasis id="BE915500717FEF23D783F9412178942A" bold="true" box="[199,334,1771,1797]" pageId="31" pageNumber="32">Etymology.</emphasis>
The specific epithet is made after the Latin singular adjective
<taxonomicName id="4BE5F291717FEF23D346F9412407942A" authorityName="Cruz-Gómez" authorityYear="2021" box="[1026,1073,1771,1797]" class="Polychaeta" family="Chrysopetalidae" genus="Bhawatsonia" kingdom="Animalia" order="Phyllodocida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="fusa">
<emphasis id="BE915500717FEF23D346F9412407942A" box="[1026,1073,1771,1797]" italics="true" pageId="31" pageNumber="32">fusa</emphasis>
</taxonomicName>
(-
<emphasis id="BE915500717FEF23D30DF9412454942B" box="[1097,1122,1771,1796]" italics="true" pageId="31" pageNumber="32">us</emphasis>
,
<emphasis id="BE915500717FEF23D32BF94124AD942B" box="[1135,1179,1771,1796]" italics="true" pageId="31" pageNumber="32">-um</emphasis>
) meaning ‘broad’. The name indicates the broad shape of upper middle section of the main paleae, and it is feminine for matching the genus gender (
<bibRefCitation id="E874F4E3717FEF23D7B6F89921419462" author="ICZN" box="[242,375,1843,1869]" pageId="31" pageNumber="32" refId="ref37026" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature (The Natural History Museum), London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 32.2).
</paragraph>
</subSubSection>
</treatment>
</document>
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