1441 lines
174 KiB
XML
1441 lines
174 KiB
XML
<document id="BB2AC7D4F5CF9A85B599B95E5607FD4F" ID-DOI="10.11646/zootaxa.3949.3.7" ID-GBIF-Dataset="47bc52a0-b711-4dc9-8116-305d4ee7137b" ID-ISSN="1175-5326" ID-Zenodo-Dep="243672" ID-ZooBank="92E0842F-1DE0-4E93-9FAD-D524B9339D2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461047110103" checkinUser="plazi" docAuthor="Sideleva, Valentina G., Naseka, Alexander M. & Zhidkov, Zakhar V." docDate="2015" docId="03E81F32FFACFF84DFEB2A46FE11FCAB" docLanguage="en" docName="zt03949p430.pdf" docOrigin="Zootaxa 3949 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cottus gratzianowi Sideleva, Naseka & Zhidkov, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="430" masterDocId="FFD1674AFFADFF8FDF7C2C2AFA34FD7B" masterDocTitle="A new species of Cottus from the Onega River drainage, White Sea basin (Actinopterygii: Scorpaeniformes: Cottidae)" masterLastPageNumber="430" masterPageNumber="419" pageNumber="420" updateTime="1698577459748" updateUser="plazi">
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<mods:mods id="23D48351332880F7CD49ADA5F540B1C8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="58041375DAD8204E8D4A22942E94D7BE">
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<mods:title id="0274616AB959C2C4DF19B208E654D3F9">A new species of Cottus from the Onega River drainage, White Sea basin (Actinopterygii: Scorpaeniformes: Cottidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B339330A65F32A41B11D0C702B278163" type="personal">
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<mods:role id="AAB14B3DB88CD8AE5F956FD8422AA42E">
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<mods:roleTerm id="A46CEBC3EA252DBBD3234905A3CE19C9">Author</mods:roleTerm>
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<mods:namePart id="9B2D8B3B366CACCB76C7DD95A9495167">Sideleva, Valentina G.</mods:namePart>
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</mods:name>
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<mods:name id="BA844C99A1B0A1FFD658FC7C0BB28131" type="personal">
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<mods:role id="1591875EBC1E89870DB958489F26BC78">
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<mods:roleTerm id="45F17A42D81610B5305C5D02E58D8775">Author</mods:roleTerm>
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<mods:namePart id="C8F114282EF4448277E009A42DDCC56B">Naseka, Alexander M.</mods:namePart>
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</mods:name>
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<mods:name id="CC709982DCB2856E57CD9361F5ECE638" type="personal">
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<mods:roleTerm id="2B3C38E80AE32529D4692D34D0495F54">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="192BFDB3B0BD6EB15FA12456F1328519">Zhidkov, Zakhar V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="ECB5DAE974CEF6E0C5D3D1CF1BF69891">text</mods:typeOfResource>
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<mods:title id="FA2274380E4C848DB29ACAEB24915949">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F5075B77F0B9487DCCDD30CDD422B143">
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<mods:date id="706ACFB32BAA9C3E1FE2EC2E5A595EAB">2015</mods:date>
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<mods:detail id="A998F6ED61AD6F61B8524F1B707016F9" type="volume">
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<mods:number id="E263D3072D6059B2F0B42E012981B1A3">3949</mods:number>
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</mods:detail>
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<mods:detail id="7FC195C2E3C90E73E0C30C3ABE513D96" type="issue">
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<mods:number id="A243AD9EA978C700FC20FC8EA4B25EC3">3</mods:number>
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<mods:classification id="350F78B38B6268245680BB243E02E79F">journal article</mods:classification>
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<mods:identifier id="015A90989B49BF745E84A02ECA547FEB" type="DOI">10.11646/zootaxa.3949.3.7</mods:identifier>
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<mods:identifier id="23C4497CE0EEBEDFEA01A58E1B04390F" type="GBIF-Dataset">47bc52a0-b711-4dc9-8116-305d4ee7137b</mods:identifier>
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<mods:identifier id="941D66B85508867E6249F4176B19366B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="13C48CD76A5106D1A74E29ADF82F19F8" type="Zenodo-Dep">243672</mods:identifier>
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<treatment id="03E81F32FFACFF84DFEB2A46FE11FCAB" ID-DOI="http://doi.org/10.5281/zenodo.5615421" ID-GBIF-Taxon="119611126" ID-Zenodo-Dep="5615421" LSID="urn:lsid:plazi:treatment:03E81F32FFACFF84DFEB2A46FE11FCAB" httpUri="http://treatment.plazi.org/id/03E81F32FFACFF84DFEB2A46FE11FCAB" lastPageId="11" lastPageNumber="430" pageId="1" pageNumber="420">
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<subSubSection id="C35BFDAFFFACFF8EDFEB2A46FB77FBD3" pageId="1" pageNumber="420" type="nomenclature">
|
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<paragraph id="8BFEAE24FFACFF8EDFEB2A46FBD2FBFC" blockId="1.[151,486,1644,1704]" box="[151,486,1644,1671]" pageId="1" pageNumber="420">
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<heading id="D0B61948FFACFF8EDFEB2A46FBD2FBFC" bold="true" box="[151,486,1644,1671]" fontSize="11" level="1" pageId="1" pageNumber="420" reason="1">
|
||
<emphasis id="B9357236FFACFF8EDFEB2A46FBD2FBFC" bold="true" box="[151,486,1644,1671]" pageId="1" pageNumber="420">
|
||
<emphasis id="B9357236FFACFF8EDFEB2A46FBB5FBFD" bold="true" box="[151,385,1644,1670]" italics="true" pageId="1" pageNumber="420">
|
||
<taxonomicName id="4C41D5A7FFACFF8EDFEB2A46FB4FFBFD" ID-CoL="YYGY" box="[151,379,1644,1670]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="1" pageNumber="420" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">Cottus gratzianowi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A206CF4DFFACFF8EDEF52A47FBD2FBFC" box="[393,486,1645,1671]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFACFF8EDFEB2AA5FB77FBD3" blockId="1.[151,486,1644,1704]" box="[151,323,1679,1704]" pageId="1" pageNumber="420">
|
||
(
|
||
<figureCitation id="137AB2A1FFACFF8EDFE32AA5FAD3FBD3" box="[159,231,1679,1704]" captionStart="FIGURE 2" captionStartId="2.[151,250,1531,1553]" captionTargetBox="[170,1421,514,1477]" captionTargetId="figure@2.[332,1303,901,1482]" captionTargetPageId="2" captionText="FIGURE 2. Cottus gratzianowi sp. nov., holotype, ZIN 55596, male 63.3 mm SL: lateral view (a), dorsal view of head and anterior body (b), ventral view of head and anterior body (c)." httpUri="https://zenodo.org/record/243674/files/figure.png" pageId="1" pageNumber="420">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="137AB2A1FFACFF8EDF8F2ABAFB37FBD3" box="[243,259,1680,1704]" captionStart="FIGURE 3" captionStartId="5.[151,250,1066,1088]" captionTargetBox="[176,1401,238,987]" captionTargetId="figure@5.[151,1436,194,1044]" captionTargetPageId="5" captionText="FIGURE 3. Photograph of Cottus gratzianowi sp. nov., holotype as in Fig. 2." httpUri="https://zenodo.org/record/243675/files/figure.png" pageId="1" pageNumber="420">3</figureCitation>
|
||
,
|
||
<figureCitation id="137AB2A1FFACFF8EDE732AA5FB2BFBD3" box="[271,287,1679,1704]" captionStart="FIGURE 5" captionStartId="6.[151,250,575,597]" captionTargetBox="[161,1425,196,552]" captionTargetId="figure@6.[151,1436,193,554]" captionTargetPageId="6" captionText="FIGURE 5. Photograph of pectoral fin. Arrows show branched middle pectoral-fin rays in Cottus gratzianowi sp. nov. (a) and unbranched middle pectoral-fin rays in Cottus koshewnikowi (b)." httpUri="https://zenodo.org/record/243677/files/figure.png" pageId="1" pageNumber="420">5</figureCitation>
|
||
,
|
||
<figureCitation id="137AB2A1FFACFF8EDE572AA5FB0EFBD3" box="[299,314,1679,1704]" captionStart="FIGURE 6" captionStartId="7.[151,250,918,940]" captionTargetBox="[151,1436,193,895]" captionTargetId="figure@7.[151,1436,193,896]" captionTargetPageId="7" captionText="FIGURE 6. Radiographs of Cottus gratzianowi sp. nov. (a) with 10 + 23 vertebrae and Cottus koshewnikowi (b) with 11 + 23 vertebrae. Arrow shows first caudal vertebra." httpUri="https://zenodo.org/record/243678/files/figure.png" pageId="1" pageNumber="420">6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BFDAFFFACFF8EDFEB2AFDFE1CFA43" pageId="1" pageNumber="420" type="materials_examined">
|
||
<paragraph id="8BFEAE24FFACFF8EDFEB2AFDFE8BFA6F" blockId="1.[151,1436,1751,2028]" pageId="1" pageNumber="420">
|
||
<emphasis id="B9357236FFACFF8EDFEB2AFDFB3FFB8B" bold="true" box="[151,267,1751,1776]" pageId="1" pageNumber="420">
|
||
<typeStatus id="54FA1086FFACFF8EDFEB2AFDFB32FB8B" box="[151,262,1751,1776]" pageId="1" pageNumber="420" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED5036E1FFACFF8EDE692AF2FB70FB94" box="[277,324,1752,1775]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="420">ZIN</collectionCode>
|
||
55596, 63 mm SL; Ukhtomitsa River village of Korotetskaya (Vologda Province), 60°18′19″N, 38°40′30″E, Onega River drainage, White Sea basin; coll. A. Naseka & Z. Zhidkov,
|
||
<date id="FFFF88E4FFACFF8EDB362AD1FE88FA6F" box="[1098,1212,1787,1812]" pageId="1" pageNumber="420" value="2010-07">July 2010</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFACFF8EDFBB2B35FE1CFA43" blockId="1.[151,1436,1751,2028]" box="[199,1064,1823,1848]" pageId="1" pageNumber="420">
|
||
<typeStatus id="54FA1086FFACFF8EDFBB2B35FB0BFA43" box="[199,319,1823,1848]" pageId="1" pageNumber="420" type="paratype">
|
||
<emphasis id="B9357236FFACFF8EDFBB2B35FB0BFA43" bold="true" box="[199,319,1823,1848]" pageId="1" pageNumber="420">Paratypes</emphasis>
|
||
</typeStatus>
|
||
.
|
||
<collectionCode id="ED5036E1FFACFF8EDE302B0AFB4FFA4C" box="[332,379,1824,1847]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="420">ZIN</collectionCode>
|
||
55597,
|
||
<specimenCount id="9D4765ADFFACFF8EDEAA2B0AF844FA43" box="[470,624,1823,1848]" pageId="1" pageNumber="420" type="generic">17 specimens</specimenCount>
|
||
,
|
||
<quantity id="4CB903C1FFACFF8EDD002B35F8C1FA4C" box="[636,757,1823,1848]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" metricValueMax="6.0" metricValueMin="4.2" pageId="1" pageNumber="420" unit="mm" value="51.0" valueMax="60.0" valueMin="42.0">42–60 mm</quantity>
|
||
SL, same data as
|
||
<typeStatus id="54FA1086FFACFF8EDCC32B35FE11FA43" box="[959,1061,1823,1848]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BFDAFFFACFF8DDFBB2B69F90AFCD6" lastPageId="2" lastPageNumber="421" pageId="1" pageNumber="420" type="diagnosis">
|
||
<paragraph id="8BFEAE24FFACFF8DDFBB2B69F90AFCD6" blockId="1.[151,1436,1751,2028]" lastBlockId="2.[151,1436,151,429]" lastPageId="2" lastPageNumber="421" pageId="1" pageNumber="420">
|
||
<emphasis id="B9357236FFACFF8EDFBB2B69FB75FA27" bold="true" box="[199,321,1859,1884]" pageId="1" pageNumber="420">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C41D5A7FFACFF8EDE352B6EF814FA20" box="[329,544,1860,1883]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="1" pageNumber="420" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFACFF8EDE352B6EF814FA20" box="[329,544,1860,1883]" italics="true" pageId="1" pageNumber="420">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFACFF8EDD5B2B69F8B6FA27" box="[551,642,1859,1884]" pageId="1" pageNumber="420" rank="species">
|
||
<emphasis id="B9357236FFACFF8EDD5B2B69F8B6FA27" bold="true" box="[551,642,1859,1884]" pageId="1" pageNumber="420">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is diagnosed from all other described members of the genus
|
||
<taxonomicName id="4C41D5A7FFACFF8EDA492B6EFF4AFA20" box="[1333,1406,1860,1883]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="1" pageNumber="420" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9357236FFACFF8EDA492B6EFF4AFA20" box="[1333,1406,1860,1883]" italics="true" pageId="1" pageNumber="420">Cottus</emphasis>
|
||
</taxonomicName>
|
||
in Europe east of the
|
||
<collectingCountry id="F356EEB4FFACFF8EDE0E2B42FB89FA04" box="[370,445,1896,1919]" name="Egypt" pageId="1" pageNumber="420">Meuse</collectingCountry>
|
||
(except
|
||
<taxonomicName id="4C41D5A7FFACFF8EDD5E2B42F8D6FA04" box="[546,738,1895,1919]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="1" pageNumber="420" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFACFF8EDD5E2B42F8D6FA04" box="[546,738,1895,1919]" italics="true" pageId="1" pageNumber="420">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
) in having the following combination of character states: no transverse dark bands on the pelvic fin, a single median chin canal pore, an incomplete lateral line not reaching behind the anal-fin insertion, and the position of the lateral line which is located considerably above the mid-line of the flank.
|
||
<taxonomicName id="4C41D5A7FFACFF8EDE752BFEFBD4FA90" box="[265,480,2004,2027]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="1" pageNumber="420" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFACFF8EDE752BFEFBD4FA90" box="[265,480,2004,2027]" italics="true" pageId="1" pageNumber="420">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFACFF8EDE942BF9F876FA97" box="[488,578,2003,2028]" pageId="1" pageNumber="420" rank="species">
|
||
<emphasis id="B9357236FFACFF8EDE942BF9F876FA97" bold="true" box="[488,578,2003,2028]" pageId="1" pageNumber="420">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName id="4C41D5A7FFACFF8EDC3D2BFEF9CBFA97" box="[833,1023,2004,2028]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="1" pageNumber="420" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFACFF8EDC3D2BFEF9CBFA97" box="[833,1023,2004,2028]" italics="true" pageId="1" pageNumber="420">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
by a larger eye (horizontal diameter 23−28% HL, equal to or exceeding snout length vs. 16−25% HL, less than snout length), a rounded caudal fin (vs. commonly truncated), frequent presence of one to three branched rays in median part of the pectoral fin (vs. usual absence), an interrupted supratemporal canal commissure with 4 pores (vs. non-interrupted, with 3 pores), the pelvic fin extending to the anus in both sexes (vs. not reaching the anus), a short lateral-line canal extending to below the fifth to tenth ray of the second dorsal fin, with 15−24 pores, with few interruption (vs. commonly extending to below the 9th to 15th ray of the second dorsal fin, with modally 21−27 pores, with few to numerous interruptions), abdominal vertebrae commonly 10 (vs. 11), and contrasting black blotches on all fins including pelvic and anal fins (vs. no blotches on pelvic and anal fins).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3EFEACFFAFFF8DDFEB29D1F934FB54" httpUri="https://zenodo.org/record/243674/files/figure.png" pageId="2" pageNumber="421" targetBox="[170,1421,514,1477]" targetPageId="2">
|
||
<paragraph id="8BFEAE24FFAFFF8DDFEB29D1F934FB54" blockId="2.[151,1435,1531,1583]" pageId="2" pageNumber="421">
|
||
<emphasis id="B9357236FFAFFF8DDFEB29D1FB3AFB6B" bold="true" box="[151,270,1531,1553]" pageId="2" pageNumber="421">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C41D5A7FFAFFF8DDE6029D1FBEBFB6B" box="[284,479,1531,1552]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="421" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAFFF8DDE6029D1FBEBFB6B" box="[284,479,1531,1552]" italics="true" pageId="2" pageNumber="421">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAFFF8DDE9B29D1F80DFB6B" box="[487,569,1531,1552]" pageId="2" pageNumber="421" rank="species">
|
||
<emphasis id="B9357236FFAFFF8DDE9B29D1F80DFB6B" bold="true" box="[487,569,1531,1552]" pageId="2" pageNumber="421">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, ZIN 55596, male 63.3 mm SL: lateral view (a), dorsal view of head and anterior body (b), ventral view of head and anterior body (c).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35BFDAFFFAFFF89DFBB2A76FEDEFA1D" lastPageId="6" lastPageNumber="425" pageId="2" pageNumber="421" type="description">
|
||
<paragraph id="8BFEAE24FFAFFF8DDFBB2A76F9E2FA52" blockId="2.[151,1436,1628,2013]" pageId="2" pageNumber="421">
|
||
<emphasis id="B9357236FFAFFF8DDFBB2A76FB6DFB0E" bold="true" box="[199,345,1628,1653]" pageId="2" pageNumber="421">Description.</emphasis>
|
||
Proportional measurements of the
|
||
<typeStatus id="54FA1086FFAFFF8DDD932A76F965FB0E" box="[751,849,1628,1653]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54FA1086FFAFFF8DDCF12A77F9CCFB0E" box="[909,1016,1629,1653]" pageId="2" pageNumber="421" type="paratype">paratypes</typeStatus>
|
||
are given in
|
||
<tableCitation id="C6C39B9FFFAFFF8DDBE92A76FED8FB0E" box="[1173,1260,1628,1653]" captionStart="TABLE 1" captionStartId="3.[125,207,513,533]" captionTargetBox="[138,1845,573,1680]" captionTargetPageId="3" captionText="TABLE 1. Prοpοrtiοnal measurements οf Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri="http://table.plazi.org/id/DF3EFEACFFAEFF8CDF012E2BFE18FF6D" pageId="2" pageNumber="421" tableUuid="DF3EFEACFFAEFF8CDF012E2BFE18FF6D">Table 1</tableCitation>
|
||
. Body humped behind head, deep and compressed; maximum body depth 4.3−5.5 times in SL (
|
||
<quantity id="4CB903C1FFAFFF8DDB6B2AAAFE6CFBE2" box="[1047,1112,1664,1689]" metricMagnitude="-1" metricUnit="m" metricValue="1.3716" pageId="2" pageNumber="421" unit="in" value="5.4">5.4 in</quantity>
|
||
<typeStatus id="54FA1086FFAFFF8DDB232AAAFEF7FBE2" box="[1119,1219,1664,1689]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
) and markedly less than HL (
|
||
<figureCitation id="137AB2A1FFAFFF8DDE7A2A8EFB64FBC6" box="[262,336,1700,1725]" captionStart="FIGURE 2" captionStartId="2.[151,250,1531,1553]" captionTargetBox="[170,1421,514,1477]" captionTargetId="figure@2.[332,1303,901,1482]" captionTargetPageId="2" captionText="FIGURE 2. Cottus gratzianowi sp. nov., holotype, ZIN 55596, male 63.3 mm SL: lateral view (a), dorsal view of head and anterior body (b), ventral view of head and anterior body (c)." httpUri="https://zenodo.org/record/243674/files/figure.png" pageId="2" pageNumber="421">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="137AB2A1FFAFFF8DDE222A8FFB59FBC6" box="[350,365,1701,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1066,1088]" captionTargetBox="[176,1401,238,987]" captionTargetId="figure@5.[151,1436,194,1044]" captionTargetPageId="5" captionText="FIGURE 3. Photograph of Cottus gratzianowi sp. nov., holotype as in Fig. 2." httpUri="https://zenodo.org/record/243675/files/figure.png" pageId="2" pageNumber="421">3</figureCitation>
|
||
). Caudal peduncle length 6.8−9 times in SL (
|
||
<quantity id="4CB903C1FFAFFF8DDC052A8EF988FBC6" box="[889,956,1700,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.7271999999999998" pageId="2" pageNumber="421" unit="in" value="6.8">6.8 in</quantity>
|
||
<typeStatus id="54FA1086FFAFFF8DDCB92A8EFE1FFBC6" box="[965,1067,1700,1725]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
), caudal peduncle depth 1.7−2.3 times in its length (
|
||
<quantity id="4CB903C1FFAFFF8DDE112AE2FB9AFB9A" box="[365,430,1736,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.588" pageId="2" pageNumber="421" unit="in" value="2.2">2.2 in</quantity>
|
||
<typeStatus id="54FA1086FFAFFF8DDEC82AE2F82FFB9A" box="[436,539,1736,1761]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
). Prickles (modified scales) numerous, located below and above lateral line; just behind pectoral base, area of prickling extended to back and gradually tapered caudad, terminating below middle of second dorsal fin. Prickle possessing roundish basis and short sharp spine.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFAFFF8BDFBB2B1EF85DFC1E" blockId="2.[151,1436,1628,2013]" lastBlockId="4.[151,1437,151,789]" lastPageId="4" lastPageNumber="423" pageId="2" pageNumber="421">
|
||
Head large, its length, only slightly less than or equal to width, 2.5−3.3 times in SL (
|
||
<quantity id="4CB903C1FFAFFF8DDB132B1FFE9BFA36" box="[1135,1199,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" pageId="2" pageNumber="421" unit="in" value="3.3">3.3 in</quantity>
|
||
<typeStatus id="54FA1086FFAFFF8DDBC92B1EFF2FFA36" box="[1205,1307,1844,1869]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
), depressed (
|
||
<figureCitation id="137AB2A1FFAFFF8DDFE32B72FAD7FA0B" box="[159,227,1880,1905]" captionStart="FIGURE 2" captionStartId="2.[151,250,1531,1553]" captionTargetBox="[170,1421,514,1477]" captionTargetId="figure@2.[332,1303,901,1482]" captionTargetPageId="2" captionText="FIGURE 2. Cottus gratzianowi sp. nov., holotype, ZIN 55596, male 63.3 mm SL: lateral view (a), dorsal view of head and anterior body (b), ventral view of head and anterior body (c)." httpUri="https://zenodo.org/record/243674/files/figure.png" pageId="2" pageNumber="421">Fig. 2</figureCitation>
|
||
a −c). Small and very shallow dermal tubercles on dorsal surface of head. Interbranchial space wide, 2.5−3.8 times in HL (
|
||
<quantity id="4CB903C1FFAFFF8DDE402B57FBB7FAEE" box="[316,387,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" pageId="2" pageNumber="421" unit="in" value="2.7">2.7 in</quantity>
|
||
<typeStatus id="54FA1086FFAFFF8DDEF22B56FBC6FAEE" box="[398,498,1916,1941]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<figureCitation id="137AB2A1FFAFFF8DDD722B56F86CFAEE" box="[526,600,1916,1941]" captionStart="FIGURE 2" captionStartId="2.[151,250,1531,1553]" captionTargetBox="[170,1421,514,1477]" captionTargetId="figure@2.[332,1303,901,1482]" captionTargetPageId="2" captionText="FIGURE 2. Cottus gratzianowi sp. nov., holotype, ZIN 55596, male 63.3 mm SL: lateral view (a), dorsal view of head and anterior body (b), ventral view of head and anterior body (c)." httpUri="https://zenodo.org/record/243674/files/figure.png" pageId="2" pageNumber="421">Fig. 2</figureCitation>
|
||
c). Three preopercular spines; uppermost spine longest, slightly curved upward, its length about 6 times in SL; second spine short, about 3 times in length of uppermost spine; third spine blunt and covered by skin. Snout steep and elongated, its length 3.7−5 times in HL (
|
||
<quantity id="4CB903C1FFAFFF8DDB122BEFFE84FAA6" box="[1134,1200,1988,2013]" metricMagnitude="-1" metricUnit="m" metricValue="1.0668" pageId="2" pageNumber="421" unit="in" value="4.2">4.2 in</quantity>
|
||
<typeStatus id="54FA1086FFAFFF8DDBCB2BEEFF2AFAA6" box="[1207,1310,1988,2013]" pageId="2" pageNumber="421" type="holotype">holotype</typeStatus>
|
||
). Anterior nostril forming short cylindrical pigmented tube. Posterior nostril as slender tube, 3 times as small as anterior nostril. Eye relatively large, eye horizontal diameter 3.5−4.4 times in HL (
|
||
<quantity id="4CB903C1FFA9FF8BDC952C96FE18FDAE" box="[1001,1068,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.0668" pageId="4" pageNumber="423" unit="in" value="4.2">4.2 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDB492C96FEAFFDAE" box="[1077,1179,188,213]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
), equal or greater than snout length; dorsal margin of orbit elevated. Interorbital space narrow; its width 1.2−2 times in eye horizontal diameter (
|
||
<quantity id="4CB903C1FFA9FF8BDE752D2EFB7FFC66" box="[265,331,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="423" unit="in" value="1.5">1.5 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDE2E2D2EFB82FC66" box="[338,438,260,285]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
) about 6.5−9.3 times in HL (
|
||
<quantity id="4CB903C1FFA9FF8BDC7C2D2EF976FC66" box="[768,834,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" pageId="4" pageNumber="423" unit="in" value="6.5">6.5 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDC352D2EF984FC66" box="[841,944,260,285]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
). Mouth horizontal and terminal; upper jaw extending caudad to below anterior margin of pupil. Length of upper jaw 2.8−3.7 times in HL (
|
||
<quantity id="4CB903C1FFA9FF8BDBA12D02FF15FC3B" box="[1245,1313,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" pageId="4" pageNumber="423" unit="in" value="2.8">2.8 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDA552D0DFFA4FC3B" box="[1321,1424,295,320]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
); lower jaw slightly shorter than upper jaw.
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFAEFF8CDF012E2BFE18FF6D" ID-Table-UUID="DF3EFEACFFAEFF8CDF012E2BFE18FF6D" box="[125,1068,513,534]" httpUri="http://table.plazi.org/id/DF3EFEACFFAEFF8CDF012E2BFE18FF6D" pageId="3" pageNumber="422" targetBox="[138,1845,573,1680]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="8BFEAE24FFAEFF8CDF012E2BFE18FF6D" blockId="3.[125,1068,513,534]" box="[125,1068,513,534]" pageId="3" pageNumber="422">
|
||
<emphasis id="B9357236FFAEFF8CDF012E2BFAD5FF6E" bold="true" box="[125,225,513,533]" pageId="3" pageNumber="422">TABLE 1</emphasis>
|
||
. Prοpοrtiοnal measurements οf
|
||
<taxonomicName id="4C41D5A7FFAEFF8CDD652E2BF8E6FF6D" box="[537,722,513,534]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAEFF8CDD652E2BF8E6FF6D" box="[537,722,513,534]" italics="true" pageId="3" pageNumber="422">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAEFF8CDDA42E2BF913FF6E" box="[728,807,513,533]" pageId="3" pageNumber="422" rank="species">
|
||
<emphasis id="B9357236FFAEFF8CDDA42E2BF913FF6E" bold="true" box="[728,807,513,533]" pageId="3" pageNumber="422">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C41D5A7FFAEFF8CDC2B2E2BFE13FF6D" box="[855,1063,513,534]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="koshewnikowi">Cottus koshewnikowi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFAEFF8CDDE22E60FD00FBEB" pageId="3" pageNumber="422">
|
||
<table id="F9415C84FFAE0070DFF62E17FD01FBEB" box="[138,1845,573,1680]" gridcols="6" gridrows="29" pageId="3" pageNumber="422">
|
||
<tr id="3571AC66FFAE0070DFF62E17FD01FF17" box="[138,1845,573,620]" gridrow="0" pageId="3" pageNumber="422" rowspan-0="1" rowspan-2="1">
|
||
<th id="76A0C51AFFAE0070DD552E17F9B1FF17" box="[553,901,573,620]" gridcol="1" gridrow="0" pageId="3" pageNumber="422">
|
||
<emphasis id="B9357236FFAEFF8CDDE22E60F901FF24" box="[670,821,586,607]" italics="true" pageId="3" pageNumber="422">
|
||
<taxonomicName id="4C41D5A7FFAEFF8CDDE22E60F905FF24" box="[670,817,586,607]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="gratzianowi">C. gratzianowi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
n=9
|
||
</th>
|
||
<th id="76A0C51AFFAE0070DC882E17FED3FF17" box="[1012,1255,573,620]" gridcol="3" gridrow="0" pageId="3" pageNumber="422">
|
||
<emphasis id="B9357236FFAEFF8CDB692E17FEF4FF29" box="[1045,1216,573,594]" italics="true" pageId="3" pageNumber="422">
|
||
<taxonomicName id="4C41D5A7FFAEFF8CDB692E17FE89FF29" box="[1045,1213,573,594]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="koshewnikowi">C. koshewnikowi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
vοlga (syntypes), n=21
|
||
</th>
|
||
<th id="76A0C51AFFAE0070DA792E17FC3FFF17" box="[1285,1547,573,620]" gridcol="4" gridrow="0" pageId="3" pageNumber="422">
|
||
<emphasis id="B9357236FFAEFF8CDA482E17FFEBFF29" box="[1332,1503,573,594]" italics="true" pageId="3" pageNumber="422">
|
||
<taxonomicName id="4C41D5A7FFAEFF8CDA482E17FFEFFF29" box="[1332,1499,573,594]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="koshewnikowi">C. koshewnikowi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Pechοra, n=18
|
||
</th>
|
||
<th id="76A0C51AFFAE0070D9482E17FD01FF17" box="[1588,1845,573,620]" gridcol="5" gridrow="0" pageId="3" pageNumber="422">
|
||
<emphasis id="B9357236FFAEFF8CD9272E17FD32FF29" box="[1627,1798,573,594]" italics="true" pageId="3" pageNumber="422">
|
||
<taxonomicName id="4C41D5A7FFAEFF8CD9272E17FD36FF29" box="[1627,1794,573,594]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="koshewnikowi">C. koshewnikowi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Nοrthern Dvina, n=5
|
||
</th>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62E5EFD01FFD8" box="[138,1845,628,675]" gridrow="1" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62E5EF83DFFD8" box="[138,521,628,675]" gridcol="0" gridrow="1" pageId="3" pageNumber="422">Character</th>
|
||
<td id="76A0C51AFFAE0070DD552E5EF9B1FFD8" box="[553,901,628,675]" gridcol="1" gridrow="1" pageId="3" pageNumber="422">hοlοtype, range M mm</td>
|
||
<td id="76A0C51AFFAE0070DCDA2E5EF9E5FFD8" box="[934,977,628,675]" gridcol="2" gridrow="1" pageId="3" pageNumber="422">SD</td>
|
||
<td id="76A0C51AFFAE0070DC882E5EFED3FFD8" box="[1012,1255,628,675]" gridcol="3" gridrow="1" pageId="3" pageNumber="422">range M SD</td>
|
||
<td id="76A0C51AFFAE0070DA792E5EFC3FFFD8" box="[1285,1547,628,675]" gridcol="4" gridrow="1" pageId="3" pageNumber="422">range M SD</td>
|
||
<td id="76A0C51AFFAE0070D9482E5EFD01FFD8" box="[1588,1845,628,675]" gridcol="5" gridrow="1" pageId="3" pageNumber="422">range M SD</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62E84FD01FFB9" box="[138,1845,686,706]" gridrow="2" pageId="3" pageNumber="422" rowspan-2="1">
|
||
<th id="76A0C51AFFAE0070DFF62E84F83DFFB9" box="[138,521,686,706]" gridcol="0" gridrow="2" pageId="3" pageNumber="422">ΤL</th>
|
||
<td id="76A0C51AFFAE0070DD552E84F9B1FFB9" box="[553,901,686,706]" gridcol="1" gridrow="2" pageId="3" pageNumber="422">77.0 55.0‾72.5(77.0)</td>
|
||
<td id="76A0C51AFFAE0070DC882E84FED3FFB9" box="[1012,1255,686,706]" gridcol="3" gridrow="2" pageId="3" pageNumber="422">49.0‾88.0</td>
|
||
<td id="76A0C51AFFAE0070DA792E84FC3FFFB9" box="[1285,1547,686,706]" gridcol="4" gridrow="2" pageId="3" pageNumber="422">66.0‾105.0</td>
|
||
<td id="76A0C51AFFAE0070D9482E84FD01FFB9" box="[1588,1845,686,706]" gridcol="5" gridrow="2" pageId="3" pageNumber="422">73.5‾93.5</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62EE6FD01FF9B" box="[138,1845,716,736]" gridrow="3" pageId="3" pageNumber="422" rowspan-2="1">
|
||
<th id="76A0C51AFFAE0070DFF62EE6F83DFF9B" box="[138,521,716,736]" gridcol="0" gridrow="3" pageId="3" pageNumber="422">SL</th>
|
||
<td id="76A0C51AFFAE0070DD552EE6F9B1FF9B" box="[553,901,716,736]" gridcol="1" gridrow="3" pageId="3" pageNumber="422">63.3 42.5‾58.0(63.3)</td>
|
||
<td id="76A0C51AFFAE0070DC882EE6FED3FF9B" box="[1012,1255,716,736]" gridcol="3" gridrow="3" pageId="3" pageNumber="422">40.0‾76.0</td>
|
||
<td id="76A0C51AFFAE0070DA792EE6FC3FFF9B" box="[1285,1547,716,736]" gridcol="4" gridrow="3" pageId="3" pageNumber="422">52.0‾80.5</td>
|
||
<td id="76A0C51AFFAE0070D9482EE6FD01FF9B" box="[1588,1845,716,736]" gridcol="5" gridrow="3" pageId="3" pageNumber="422">63.5‾77.0</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62EC1FD01FF84" box="[138,1845,747,767]" gridrow="4" pageId="3" pageNumber="422" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76A0C51AFFAE0070DFF62EC1F83DFF84" box="[138,521,747,767]" gridcol="0" gridrow="4" pageId="3" pageNumber="422">%SL</th>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62F2CFD01FE5D" box="[138,1845,774,806]" gridrow="5" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62F2CF83DFE5D" box="[138,521,774,806]" gridcol="0" gridrow="5" pageId="3" pageNumber="422">Predοrsal length</th>
|
||
<td id="76A0C51AFFAE0070DD552F2CF9B1FE5D" box="[553,901,774,806]" gridcol="1" gridrow="5" pageId="3" pageNumber="422">34.8 33.6‾37.7 35.7</td>
|
||
<td id="76A0C51AFFAE0070DCDA2F2CF9E5FE5D" box="[934,977,774,806]" gridcol="2" gridrow="5" pageId="3" pageNumber="422">1.22</td>
|
||
<td id="76A0C51AFFAE0070DC882F2CFED3FE5D" box="[1012,1255,774,806]" gridcol="3" gridrow="5" pageId="3" pageNumber="422">31.9‾38.3 34.7 1.94</td>
|
||
<td id="76A0C51AFFAE0070DA792F2CFC3FFE5D" box="[1285,1547,774,806]" gridcol="4" gridrow="5" pageId="3" pageNumber="422">30.0‾37.8 33.6 2.19</td>
|
||
<td id="76A0C51AFFAE0070D9482F2CFD01FE5D" box="[1588,1845,774,806]" gridcol="5" gridrow="5" pageId="3" pageNumber="422">33.1‾36.8 34.7 1.38</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62F07FD01FE37" box="[138,1845,813,844]" gridrow="6" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62F07F83DFE37" box="[138,521,813,844]" gridcol="0" gridrow="6" pageId="3" pageNumber="422">Preanal length</th>
|
||
<td id="76A0C51AFFAE0070DD552F07F9B1FE37" box="[553,901,813,844]" gridcol="1" gridrow="6" pageId="3" pageNumber="422">55.3 54.3‾59.1 56.5</td>
|
||
<td id="76A0C51AFFAE0070DCDA2F07F9E5FE37" box="[934,977,813,844]" gridcol="2" gridrow="6" pageId="3" pageNumber="422">1.63</td>
|
||
<td id="76A0C51AFFAE0070DC882F07FED3FE37" box="[1012,1255,813,844]" gridcol="3" gridrow="6" pageId="3" pageNumber="422">50.8‾58.6 55.4 2.07</td>
|
||
<td id="76A0C51AFFAE0070DA792F07FC3FFE37" box="[1285,1547,813,844]" gridcol="4" gridrow="6" pageId="3" pageNumber="422">49.8‾58.2 53.5 2.74</td>
|
||
<td id="76A0C51AFFAE0070D9482F07FD01FE37" box="[1588,1845,813,844]" gridcol="5" gridrow="6" pageId="3" pageNumber="422">54.5‾58.4 56.5 1.84</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62F79FD01FE08" box="[138,1845,851,883]" gridrow="7" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62F79F83DFE08" box="[138,521,851,883]" gridcol="0" gridrow="7" pageId="3" pageNumber="422">Maximum bοdy depth</th>
|
||
<td id="76A0C51AFFAE0070DD552F79F9B1FE08" box="[553,901,851,883]" gridcol="1" gridrow="7" pageId="3" pageNumber="422">18.6 18.1‾23.3 19.8</td>
|
||
<td id="76A0C51AFFAE0070DCDA2F79F9E5FE08" box="[934,977,851,883]" gridcol="2" gridrow="7" pageId="3" pageNumber="422">1.60</td>
|
||
<td id="76A0C51AFFAE0070DC882F79FED3FE08" box="[1012,1255,851,883]" gridcol="3" gridrow="7" pageId="3" pageNumber="422">15.9‾23.2 19.2 1.68</td>
|
||
<td id="76A0C51AFFAE0070DA792F79FC3FFE08" box="[1285,1547,851,883]" gridcol="4" gridrow="7" pageId="3" pageNumber="422">16.0‾26.1 17.7 2.44</td>
|
||
<td id="76A0C51AFFAE0070D9482F79FD01FE08" box="[1588,1845,851,883]" gridcol="5" gridrow="7" pageId="3" pageNumber="422">17.5‾21.3 18.6 1.60</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62F50FD01FEE1" box="[138,1845,890,922]" gridrow="8" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62F50F83DFEE1" box="[138,521,890,922]" gridcol="0" gridrow="8" pageId="3" pageNumber="422">Depth οf caudal peduncle</th>
|
||
<td id="76A0C51AFFAE0070DD552F50F9B1FEE1" box="[553,901,890,922]" gridcol="1" gridrow="8" pageId="3" pageNumber="422">6.8 6.2‾7.8 7.0</td>
|
||
<td id="76A0C51AFFAE0070DCDA2F50F9E5FEE1" box="[934,977,890,922]" gridcol="2" gridrow="8" pageId="3" pageNumber="422">0.46</td>
|
||
<td id="76A0C51AFFAE0070DC882F50FED3FEE1" box="[1012,1255,890,922]" gridcol="3" gridrow="8" pageId="3" pageNumber="422">3.5‾7.0 5.1 0.90</td>
|
||
<td id="76A0C51AFFAE0070DA792F50FC3FFEE1" box="[1285,1547,890,922]" gridcol="4" gridrow="8" pageId="3" pageNumber="422">4.7‾7.6 6.1 0.67</td>
|
||
<td id="76A0C51AFFAE0070D9482F50FD01FEE1" box="[1588,1845,890,922]" gridcol="5" gridrow="8" pageId="3" pageNumber="422">5.4‾7.3 6.3 0.76</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62F8AFD01FEBB" box="[138,1845,928,960]" gridrow="9" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62F8AF83DFEBB" box="[138,521,928,960]" gridcol="0" gridrow="9" pageId="3" pageNumber="422">Length οf caudal peduncle</th>
|
||
<td id="76A0C51AFFAE0070DD552F8AF9B1FEBB" box="[553,901,928,960]" gridcol="1" gridrow="9" pageId="3" pageNumber="422">14.7 11.0‾14.7 12.8</td>
|
||
<td id="76A0C51AFFAE0070DCDA2F8AF9E5FEBB" box="[934,977,928,960]" gridcol="2" gridrow="9" pageId="3" pageNumber="422">1.23</td>
|
||
<td id="76A0C51AFFAE0070DC882F8AFED3FEBB" box="[1012,1255,928,960]" gridcol="3" gridrow="9" pageId="3" pageNumber="422">6.6‾11.2 8.9 1.35</td>
|
||
<td id="76A0C51AFFAE0070DA792F8AFC3FFEBB" box="[1285,1547,928,960]" gridcol="4" gridrow="9" pageId="3" pageNumber="422">10.8‾20.4 14.6 2.49</td>
|
||
<td id="76A0C51AFFAE0070D9482F8AFD01FEBB" box="[1588,1845,928,960]" gridcol="5" gridrow="9" pageId="3" pageNumber="422">13.3‾17.3 15.7 1.52</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62FEDFD01FE9C" box="[138,1845,967,999]" gridrow="10" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62FEDF83DFE9C" box="[138,521,967,999]" gridcol="0" gridrow="10" pageId="3" pageNumber="422">Length οf first dοrsal-fin base</th>
|
||
<td id="76A0C51AFFAE0070DD552FEDF9B1FE9C" box="[553,901,967,999]" gridcol="1" gridrow="10" pageId="3" pageNumber="422">15.8 15.8‾20.2 18.3</td>
|
||
<td id="76A0C51AFFAE0070DCDA2FEDF9E5FE9C" box="[934,977,967,999]" gridcol="2" gridrow="10" pageId="3" pageNumber="422">1.50</td>
|
||
<td id="76A0C51AFFAE0070DC882FEDFED3FE9C" box="[1012,1255,967,999]" gridcol="3" gridrow="10" pageId="3" pageNumber="422">13.7‾20.4 17.5 2.03</td>
|
||
<td id="76A0C51AFFAE0070DA792FEDFC3FFE9C" box="[1285,1547,967,999]" gridcol="4" gridrow="10" pageId="3" pageNumber="422">16.0‾20.9 18.6 1.50</td>
|
||
<td id="76A0C51AFFAE0070D9482FEDFD01FE9C" box="[1588,1845,967,999]" gridcol="5" gridrow="10" pageId="3" pageNumber="422">17.3‾21.7 18.6 1.93</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62FC4FD01F976" box="[138,1845,1006,1037]" gridrow="11" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62FC4F83DF976" box="[138,521,1006,1037]" gridcol="0" gridrow="11" pageId="3" pageNumber="422">Length οf secοnd dοrsal-fin base</th>
|
||
<td id="76A0C51AFFAE0070DD552FC4F9B1F976" box="[553,901,1006,1037]" gridcol="1" gridrow="11" pageId="3" pageNumber="422">41.1 35.3‾44.8 41.2</td>
|
||
<td id="76A0C51AFFAE0070DCDA2FC4F9E5F976" box="[934,977,1006,1037]" gridcol="2" gridrow="11" pageId="3" pageNumber="422">2.71</td>
|
||
<td id="76A0C51AFFAE0070DC882FC4FED3F976" box="[1012,1255,1006,1037]" gridcol="3" gridrow="11" pageId="3" pageNumber="422">38.0‾46.3 42.0 2.24</td>
|
||
<td id="76A0C51AFFAE0070DA792FC4FC3FF976" box="[1285,1547,1006,1037]" gridcol="4" gridrow="11" pageId="3" pageNumber="422">39.0‾46.6 42.3 2.14</td>
|
||
<td id="76A0C51AFFAE0070D9482FC4FD01F976" box="[1588,1845,1006,1037]" gridcol="5" gridrow="11" pageId="3" pageNumber="422">38.6‾44.2 42.0 2.24</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF6283EFD01F94F" box="[138,1845,1044,1076]" gridrow="12" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF6283EF83DF94F" box="[138,521,1044,1076]" gridcol="0" gridrow="12" pageId="3" pageNumber="422">Length οf anal-fin base</th>
|
||
<td id="76A0C51AFFAE0070DD55283EF9B1F94F" box="[553,901,1044,1076]" gridcol="1" gridrow="12" pageId="3" pageNumber="422">30.8 30.8‾36.2 32.9</td>
|
||
<td id="76A0C51AFFAE0070DCDA283EF9E5F94F" box="[934,977,1044,1076]" gridcol="2" gridrow="12" pageId="3" pageNumber="422">1.97</td>
|
||
<td id="76A0C51AFFAE0070DC88283EFED3F94F" box="[1012,1255,1044,1076]" gridcol="3" gridrow="12" pageId="3" pageNumber="422">30.9‾40.0 34.5 2.28</td>
|
||
<td id="76A0C51AFFAE0070DA79283EFC3FF94F" box="[1285,1547,1044,1076]" gridcol="4" gridrow="12" pageId="3" pageNumber="422">30.1‾36.8 33.2 2.01</td>
|
||
<td id="76A0C51AFFAE0070D948283EFD01F94F" box="[1588,1845,1044,1076]" gridcol="5" gridrow="12" pageId="3" pageNumber="422">26.8‾33.8 30.5 2.51</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62811FD01F920" box="[138,1845,1083,1115]" gridrow="13" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62811F83DF920" box="[138,521,1083,1115]" gridcol="0" gridrow="13" pageId="3" pageNumber="422">Length οf lοngest first dοrsal-fin ray</th>
|
||
<td id="76A0C51AFFAE0070DD552811F9B1F920" box="[553,901,1083,1115]" gridcol="1" gridrow="13" pageId="3" pageNumber="422">10.4 9.6‾12.4 10.6</td>
|
||
<td id="76A0C51AFFAE0070DCDA2811F9E5F920" box="[934,977,1083,1115]" gridcol="2" gridrow="13" pageId="3" pageNumber="422">0.84</td>
|
||
<td id="76A0C51AFFAE0070DC882811FED3F920" box="[1012,1255,1083,1115]" gridcol="3" gridrow="13" pageId="3" pageNumber="422">6.8‾10.5 8.1 1.09</td>
|
||
<td id="76A0C51AFFAE0070DA792811FC3FF920" box="[1285,1547,1083,1115]" gridcol="4" gridrow="13" pageId="3" pageNumber="422">7.0‾11.8 9.8 1.51</td>
|
||
<td id="76A0C51AFFAE0070D9482811FD01F920" box="[1588,1845,1083,1115]" gridcol="5" gridrow="13" pageId="3" pageNumber="422">6.3‾9.9 8.7 1.44</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF6284BFD01F9FA" box="[138,1845,1121,1153]" gridrow="14" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF6284BF83DF9FA" box="[138,521,1121,1153]" gridcol="0" gridrow="14" pageId="3" pageNumber="422">Length οf lοngest secοnd dοrsal-fin ray</th>
|
||
<td id="76A0C51AFFAE0070DD55284BF9B1F9FA" box="[553,901,1121,1153]" gridcol="1" gridrow="14" pageId="3" pageNumber="422">15.8 13.3‾15.8 14.5</td>
|
||
<td id="76A0C51AFFAE0070DCDA284BF9E5F9FA" box="[934,977,1121,1153]" gridcol="2" gridrow="14" pageId="3" pageNumber="422">0.92</td>
|
||
<td id="76A0C51AFFAE0070DC88284BFED3F9FA" box="[1012,1255,1121,1153]" gridcol="3" gridrow="14" pageId="3" pageNumber="422">9.7‾16.1 12.3 1.26</td>
|
||
<td id="76A0C51AFFAE0070DA79284BFC3FF9FA" box="[1285,1547,1121,1153]" gridcol="4" gridrow="14" pageId="3" pageNumber="422">10.6‾16.0 13.5 1.59</td>
|
||
<td id="76A0C51AFFAE0070D948284BFD01F9FA" box="[1588,1845,1121,1153]" gridcol="5" gridrow="14" pageId="3" pageNumber="422">11.3‾13.0 12.0 0.77</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF628A2FD01F9DC" box="[138,1845,1160,1191]" gridrow="15" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF628A2F83DF9DC" box="[138,521,1160,1191]" gridcol="0" gridrow="15" pageId="3" pageNumber="422">Length οf lοngest anal-fin ray</th>
|
||
<td id="76A0C51AFFAE0070DD5528A2F9B1F9DC" box="[553,901,1160,1191]" gridcol="1" gridrow="15" pageId="3" pageNumber="422">15.5 13.3‾16.6 15.0</td>
|
||
<td id="76A0C51AFFAE0070DCDA28A2F9E5F9DC" box="[934,977,1160,1191]" gridcol="2" gridrow="15" pageId="3" pageNumber="422">1.09</td>
|
||
<td id="76A0C51AFFAE0070DC8828A2FED3F9DC" box="[1012,1255,1160,1191]" gridcol="3" gridrow="15" pageId="3" pageNumber="422">10.2‾15.2 12.8 1.49</td>
|
||
<td id="76A0C51AFFAE0070DA7928A2FC3FF9DC" box="[1285,1547,1160,1191]" gridcol="4" gridrow="15" pageId="3" pageNumber="422">10.6‾17.8 14.3 2.00</td>
|
||
<td id="76A0C51AFFAE0070D94828A2FD01F9DC" box="[1588,1845,1160,1191]" gridcol="5" gridrow="15" pageId="3" pageNumber="422">12.4‾15.8 14.0 1.45</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62884FD01F9B5" box="[138,1845,1198,1230]" gridrow="16" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62884F83DF9B5" box="[138,521,1198,1230]" gridcol="0" gridrow="16" pageId="3" pageNumber="422">Pectοral fin length</th>
|
||
<td id="76A0C51AFFAE0070DD552884F9B1F9B5" box="[553,901,1198,1230]" gridcol="1" gridrow="16" pageId="3" pageNumber="422">29.2 29.2‾33.6 31.1</td>
|
||
<td id="76A0C51AFFAE0070DCDA2884F9E5F9B5" box="[934,977,1198,1230]" gridcol="2" gridrow="16" pageId="3" pageNumber="422">1.54</td>
|
||
<td id="76A0C51AFFAE0070DC882884FED3F9B5" box="[1012,1255,1198,1230]" gridcol="3" gridrow="16" pageId="3" pageNumber="422">24.1‾30.4 27.3 1.76</td>
|
||
<td id="76A0C51AFFAE0070DA792884FC3FF9B5" box="[1285,1547,1198,1230]" gridcol="4" gridrow="16" pageId="3" pageNumber="422">25.6‾32.7 29.0 2.07</td>
|
||
<td id="76A0C51AFFAE0070D9482884FD01F9B5" box="[1588,1845,1198,1230]" gridcol="5" gridrow="16" pageId="3" pageNumber="422">23.5‾32.0 28.0 3.25</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF628FFFD01F98E" box="[138,1845,1237,1269]" gridrow="17" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF628FFF83DF98E" box="[138,521,1237,1269]" gridcol="0" gridrow="17" pageId="3" pageNumber="422">Pelvic fin length</th>
|
||
<td id="76A0C51AFFAE0070DD5528FFF9B1F98E" box="[553,901,1237,1269]" gridcol="1" gridrow="17" pageId="3" pageNumber="422">21.2 18.8‾21.2 20.5</td>
|
||
<td id="76A0C51AFFAE0070DCDA28FFF9E5F98E" box="[934,977,1237,1269]" gridcol="2" gridrow="17" pageId="3" pageNumber="422">0.85</td>
|
||
<td id="76A0C51AFFAE0070DC8828FFFED3F98E" box="[1012,1255,1237,1269]" gridcol="3" gridrow="17" pageId="3" pageNumber="422">17.1‾22.7 19.4 1.66</td>
|
||
<td id="76A0C51AFFAE0070DA7928FFFC3FF98E" box="[1285,1547,1237,1269]" gridcol="4" gridrow="17" pageId="3" pageNumber="422">18.0‾23.6 20.8 1.59</td>
|
||
<td id="76A0C51AFFAE0070D94828FFFD01F98E" box="[1588,1845,1237,1269]" gridcol="5" gridrow="17" pageId="3" pageNumber="422">17.6‾20.9 19.4 1.33</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF628D1FD01F860" box="[138,1845,1275,1307]" gridrow="18" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF628D1F83DF860" box="[138,521,1275,1307]" gridcol="0" gridrow="18" pageId="3" pageNumber="422">Head length (HL)</th>
|
||
<td id="76A0C51AFFAE0070DD5528D1F9B1F860" box="[553,901,1275,1307]" gridcol="1" gridrow="18" pageId="3" pageNumber="422">30.0 30.0‾39.5 33.7</td>
|
||
<td id="76A0C51AFFAE0070DCDA28D1F9E5F860" box="[934,977,1275,1307]" gridcol="2" gridrow="18" pageId="3" pageNumber="422">2.80</td>
|
||
<td id="76A0C51AFFAE0070DC8828D1FED3F860" box="[1012,1255,1275,1307]" gridcol="3" gridrow="18" pageId="3" pageNumber="422">28.7‾33.6 30.9 1.38</td>
|
||
<td id="76A0C51AFFAE0070DA7928D1FC3FF860" box="[1285,1547,1275,1307]" gridcol="4" gridrow="18" pageId="3" pageNumber="422">26.3‾31.2 29.0 1.49</td>
|
||
<td id="76A0C51AFFAE0070D94828D1FD01F860" box="[1588,1845,1275,1307]" gridcol="5" gridrow="18" pageId="3" pageNumber="422">28.2‾32.8 30.1 1.97</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF6290FFD01F842" box="[138,1845,1317,1337]" gridrow="19" pageId="3" pageNumber="422" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76A0C51AFFAE0070DFF6290FF83DF842" box="[138,521,1317,1337]" gridcol="0" gridrow="19" pageId="3" pageNumber="422">%HL</th>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF6296AFD01F824" box="[138,1845,1344,1375]" gridrow="20" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF6296AF83DF824" box="[138,521,1344,1375]" gridcol="0" gridrow="20" pageId="3" pageNumber="422">Head depth</th>
|
||
<td id="76A0C51AFFAE0070DD55296AF9B1F824" box="[553,901,1344,1375]" gridcol="1" gridrow="20" pageId="3" pageNumber="422">60.5 52.0‾60.5 55.1</td>
|
||
<td id="76A0C51AFFAE0070DCDA296AF9E5F824" box="[934,977,1344,1375]" gridcol="2" gridrow="20" pageId="3" pageNumber="422">3.00</td>
|
||
<td id="76A0C51AFFAE0070DC88296AFED3F824" box="[1012,1255,1344,1375]" gridcol="3" gridrow="20" pageId="3" pageNumber="422">46.5‾59.6 52.8 3.37</td>
|
||
<td id="76A0C51AFFAE0070DA79296AFC3FF824" box="[1285,1547,1344,1375]" gridcol="4" gridrow="20" pageId="3" pageNumber="422">44.1‾58.0 50.8 4.34</td>
|
||
<td id="76A0C51AFFAE0070D948296AFD01F824" box="[1588,1845,1344,1375]" gridcol="5" gridrow="20" pageId="3" pageNumber="422">42.2‾58.5 51.6 5.86</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF6294CFD01F8FD" box="[138,1845,1382,1414]" gridrow="21" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF6294CF83DF8FD" box="[138,521,1382,1414]" gridcol="0" gridrow="21" pageId="3" pageNumber="422">Head width</th>
|
||
<td id="76A0C51AFFAE0070DD55294CF9B1F8FD" box="[553,901,1382,1414]" gridcol="1" gridrow="21" pageId="3" pageNumber="422">97.4 83.3‾98.9 90.3</td>
|
||
<td id="76A0C51AFFAE0070DCDA294CF9E5F8FD" box="[934,977,1382,1414]" gridcol="2" gridrow="21" pageId="3" pageNumber="422">5.71</td>
|
||
<td id="76A0C51AFFAE0070DC88294CFED3F8FD" box="[1012,1255,1382,1414]" gridcol="3" gridrow="21" pageId="3" pageNumber="422">74.8‾95.0 83.6 5.70</td>
|
||
<td id="76A0C51AFFAE0070DA79294CFC3FF8FD" box="[1285,1547,1382,1414]" gridcol="4" gridrow="21" pageId="3" pageNumber="422">83.3‾101.0 90.5 4.90</td>
|
||
<td id="76A0C51AFFAE0070D948294CFD01F8FD" box="[1588,1845,1382,1414]" gridcol="5" gridrow="21" pageId="3" pageNumber="422">82.9‾105.0 96.9 9.00</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF629A7FD01F8D0" box="[138,1845,1421,1451]" gridrow="22" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF629A7F83DF8D0" box="[138,521,1421,1451]" gridcol="0" gridrow="22" pageId="3" pageNumber="422">Snοut length</th>
|
||
<td id="76A0C51AFFAE0070DD5529A7F9B1F8D0" box="[553,901,1421,1451]" gridcol="1" gridrow="22" pageId="3" pageNumber="422">23.7 20.0‾26.7 23.8</td>
|
||
<td id="76A0C51AFFAE0070DCDA29A7F9E5F8D0" box="[934,977,1421,1451]" gridcol="2" gridrow="22" pageId="3" pageNumber="422">2.00</td>
|
||
<td id="76A0C51AFFAE0070DC8829A7FED3F8D0" box="[1012,1255,1421,1451]" gridcol="3" gridrow="22" pageId="3" pageNumber="422">21.7‾27.8 25.2 1.74</td>
|
||
<td id="76A0C51AFFAE0070DA7929A7FC3FF8D0" box="[1285,1547,1421,1451]" gridcol="4" gridrow="22" pageId="3" pageNumber="422">25.0‾32.4 28.1 1.99</td>
|
||
<td id="76A0C51AFFAE0070D94829A7FD01F8D0" box="[1588,1845,1421,1451]" gridcol="5" gridrow="22" pageId="3" pageNumber="422">26.1‾29.9 27.4 1.56</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF6299BFD01F8AA" box="[138,1845,1457,1489]" gridrow="23" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF6299BF83DF8AA" box="[138,521,1457,1489]" gridcol="0" gridrow="23" pageId="3" pageNumber="422">Eye hοrizοntal diameter</th>
|
||
<td id="76A0C51AFFAE0070DD55299BF9B1F8AA" box="[553,901,1457,1489]" gridcol="1" gridrow="23" pageId="3" pageNumber="422">23.7 22.5‾28.3 25.2</td>
|
||
<td id="76A0C51AFFAE0070DCDA299BF9E5F8AA" box="[934,977,1457,1489]" gridcol="2" gridrow="23" pageId="3" pageNumber="422">2.16</td>
|
||
<td id="76A0C51AFFAE0070DC88299BFED3F8AA" box="[1012,1255,1457,1489]" gridcol="3" gridrow="23" pageId="3" pageNumber="422">15.9‾22.6 18.9 1.79</td>
|
||
<td id="76A0C51AFFAE0070DA79299BFC3FF8AA" box="[1285,1547,1457,1489]" gridcol="4" gridrow="23" pageId="3" pageNumber="422">17.4‾25.0 21.1 1.76</td>
|
||
<td id="76A0C51AFFAE0070D948299BFD01F8AA" box="[1588,1845,1457,1489]" gridcol="5" gridrow="23" pageId="3" pageNumber="422">18.4‾22.5 21.0 1.89</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF629F2FD01F883" box="[138,1845,1496,1528]" gridrow="24" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF629F2F83DF883" box="[138,521,1496,1528]" gridcol="0" gridrow="24" pageId="3" pageNumber="422">Pοstοrbital length</th>
|
||
<td id="76A0C51AFFAE0070DD5529F2F9B1F883" box="[553,901,1496,1528]" gridcol="1" gridrow="24" pageId="3" pageNumber="422">51.1 42.1‾51.1 46.6</td>
|
||
<td id="76A0C51AFFAE0070DCDA29F2F9E5F883" box="[934,977,1496,1528]" gridcol="2" gridrow="24" pageId="3" pageNumber="422">2.83</td>
|
||
<td id="76A0C51AFFAE0070DC8829F2FED3F883" box="[1012,1255,1496,1528]" gridcol="3" gridrow="24" pageId="3" pageNumber="422">45.2‾55.3 50.3 2.84</td>
|
||
<td id="76A0C51AFFAE0070DA7929F2FC3FF883" box="[1285,1547,1496,1528]" gridcol="4" gridrow="24" pageId="3" pageNumber="422">48.8‾55.9 51.5 2.62</td>
|
||
<td id="76A0C51AFFAE0070D94829F2FD01F883" box="[1588,1845,1496,1528]" gridcol="5" gridrow="24" pageId="3" pageNumber="422">49.0‾55.7 51.4 2.90</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF629D5FD01FB65" box="[138,1845,1535,1566]" gridrow="25" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF629D5F83DFB65" box="[138,521,1535,1566]" gridcol="0" gridrow="25" pageId="3" pageNumber="422">Interοrbital width</th>
|
||
<td id="76A0C51AFFAE0070DD5529D5F9B1FB65" box="[553,901,1535,1566]" gridcol="1" gridrow="25" pageId="3" pageNumber="422">15.3 10.7‾15.3 13.5</td>
|
||
<td id="76A0C51AFFAE0070DCDA29D5F9E5FB65" box="[934,977,1535,1566]" gridcol="2" gridrow="25" pageId="3" pageNumber="422">1.63</td>
|
||
<td id="76A0C51AFFAE0070DC8829D5FED3FB65" box="[1012,1255,1535,1566]" gridcol="3" gridrow="25" pageId="3" pageNumber="422">7.7‾13.6 9.8 2.00</td>
|
||
<td id="76A0C51AFFAE0070DA7929D5FC3FFB65" box="[1285,1547,1535,1566]" gridcol="4" gridrow="25" pageId="3" pageNumber="422">13.0‾20.7 17.0 2.20</td>
|
||
<td id="76A0C51AFFAE0070D94829D5FD01FB65" box="[1588,1845,1535,1566]" gridcol="5" gridrow="25" pageId="3" pageNumber="422">12.2‾15.0 13.6 1.18</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62A0FFD01FB3E" box="[138,1845,1573,1605]" gridrow="26" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62A0FF83DFB3E" box="[138,521,1573,1605]" gridcol="0" gridrow="26" pageId="3" pageNumber="422">Upper ϳaw length</th>
|
||
<td id="76A0C51AFFAE0070DD552A0FF9B1FB3E" box="[553,901,1573,1605]" gridcol="1" gridrow="26" pageId="3" pageNumber="422">35.8 26.8‾35.8 32.1</td>
|
||
<td id="76A0C51AFFAE0070DCDA2A0FF9E5FB3E" box="[934,977,1573,1605]" gridcol="2" gridrow="26" pageId="3" pageNumber="422">2.57</td>
|
||
<td id="76A0C51AFFAE0070DC882A0FFED3FB3E" box="[1012,1255,1573,1605]" gridcol="3" gridrow="26" pageId="3" pageNumber="422">26.1‾34.8 30.5 2.56</td>
|
||
<td id="76A0C51AFFAE0070DA792A0FFC3FFB3E" box="[1285,1547,1573,1605]" gridcol="4" gridrow="26" pageId="3" pageNumber="422">30.9‾40.2 35.4 3.30</td>
|
||
<td id="76A0C51AFFAE0070D9482A0FFD01FB3E" box="[1588,1845,1573,1605]" gridcol="5" gridrow="26" pageId="3" pageNumber="422">34.7‾37.0 35.8 0.94</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62A61FD01FB12" box="[138,1845,1611,1641]" gridrow="27" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62A61F83DFB12" box="[138,521,1611,1641]" gridcol="0" gridrow="27" pageId="3" pageNumber="422">Interbranchial width</th>
|
||
<td id="76A0C51AFFAE0070DD552A61F9B1FB12" box="[553,901,1611,1641]" gridcol="1" gridrow="27" pageId="3" pageNumber="422">36.8 25.8‾40.6 34.3</td>
|
||
<td id="76A0C51AFFAE0070DCDA2A61F9E5FB12" box="[934,977,1611,1641]" gridcol="2" gridrow="27" pageId="3" pageNumber="422">4.74</td>
|
||
<td id="76A0C51AFFAE0070DC882A61FED3FB12" box="[1012,1255,1611,1641]" gridcol="3" gridrow="27" pageId="3" pageNumber="422">26.2‾38.1 32.8 2.60</td>
|
||
<td id="76A0C51AFFAE0070DA792A61FC3FFB12" box="[1285,1547,1611,1641]" gridcol="4" gridrow="27" pageId="3" pageNumber="422">27.8‾40.7 34.2 3.42</td>
|
||
<td id="76A0C51AFFAE0070D9482A61FD01FB12" box="[1588,1845,1611,1641]" gridcol="5" gridrow="27" pageId="3" pageNumber="422">30.6‾35.5 32.7 2.28</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAE0070DFF62A5AFD01FBEB" box="[138,1845,1648,1680]" gridrow="28" pageId="3" pageNumber="422">
|
||
<th id="76A0C51AFFAE0070DFF62A5AF83DFBEB" box="[138,521,1648,1680]" gridcol="0" gridrow="28" pageId="3" pageNumber="422">Length οf gill slit</th>
|
||
<td id="76A0C51AFFAE0070DD552A5AF9B1FBEB" box="[553,901,1648,1680]" gridcol="1" gridrow="28" pageId="3" pageNumber="422">57.9 46.3‾57.9 52.3</td>
|
||
<td id="76A0C51AFFAE0070DCDA2A5AF9E5FBEB" box="[934,977,1648,1680]" gridcol="2" gridrow="28" pageId="3" pageNumber="422">4.31</td>
|
||
<td id="76A0C51AFFAE0070DC882A5AFED3FBEB" box="[1012,1255,1648,1680]" gridcol="3" gridrow="28" pageId="3" pageNumber="422">38.5‾51.1 43.7 4.08</td>
|
||
<td id="76A0C51AFFAE0070DA792A5AFC3FFBEB" box="[1285,1547,1648,1680]" gridcol="4" gridrow="28" pageId="3" pageNumber="422">44.4‾60.5 53.2 4.36</td>
|
||
<td id="76A0C51AFFAE0070D9482A5AFD01FBEB" box="[1588,1845,1648,1680]" gridcol="5" gridrow="28" pageId="3" pageNumber="422">54.8‾59.8 56.5 2.07</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFBB2D45FE14FF63" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="423">Teeth simple, conical, similar in shape and size, grouped in bands on jaws and prevomer; no teeth on palatines. Upper lip thick, lateral lip lobes well developed. Gill slit large (46−58% HL), located completely above pectoralfin base. Outer gill rakers on first gill arch 3 or 4, small and tuberculate, covered with spines at tip. Inner gill rakers on first gill arch 4 or 5, two median gill rakers longer than marginal ones, of trapezoidal shape, laterally compressed and expanded at base, top and sides, armored with tiny sharp spines.</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFBB2E0EFB44FF1B" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="423">Genital papilla of breeding males triangular, short, not reaching base of first anal ray. Three piloric caecae, one long and two short.</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFBB2E46FB81FE6E" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="423">
|
||
Fin-ray counts are given in
|
||
<tableCitation id="C6C39B9FFFA9FF8BDE832E46F862FFFF" box="[511,598,620,645]" captionStart="TABLE 2" captionStartId="4.[151,239,836,859]" captionText="TABLE 2. Frequency of occurrence of fin-ray counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi; counts for holotype are with an asterisk." pageId="4" pageNumber="423">Table 2</tableCitation>
|
||
. First dorsal fin originating behind vertical through dorsal tip of opercular flap; length of first dorsal-fin base about 5−6.3 times in SL and about 2.2−3 times in length of second dorsal fin base (
|
||
<quantity id="4CB903C1FFA9FF8BDFA42E9EFB2FFFB6" box="[216,283,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="4" pageNumber="423" unit="in" value="2.6">2.6 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDE5E2E9EFBBDFFB6" box="[290,393,692,717]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
). No gap between end of terminal membrane of first dorsal fin and second dorsal-fin origin; length of second dorsal-fin base 2.2−2.8 times in SL (
|
||
<quantity id="4CB903C1FFA9FF8BDD8A2EF2F90CFF8B" box="[758,824,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" pageId="4" pageNumber="423" unit="in" value="2.4">2.4 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDC3C2EFDF993FF8B" box="[832,935,727,752]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
); first ray of second dorsal-fin about 3 times as short as its longest ray.
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFA9FF8BDFEB2F6EFBC2FE00" pageId="4" pageNumber="423">
|
||
<paragraph id="8BFEAE24FFA9FF8BDFEB2F6EFBC2FE00" blockId="4.[151,1435,835,891]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9357236FFA9FF8BDFEB2F6EFB30FE20" bold="true" box="[151,260,836,859]" pageId="4" pageNumber="423">TABLE 2</emphasis>
|
||
. Frequency of occurrence of fin-ray counts in
|
||
<taxonomicName id="4C41D5A7FFA9FF8BDC7D2F69F9FAFE20" box="[769,974,835,859]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA9FF8BDC7D2F69F9FAFE20" box="[769,974,835,859]" italics="true" pageId="4" pageNumber="423">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA9FF8BDCAA2F6EFE19FE20" box="[982,1069,836,859]" pageId="4" pageNumber="423" rank="species">
|
||
<emphasis id="B9357236FFA9FF8BDCAA2F6EFE19FE20" bold="true" box="[982,1069,836,859]" pageId="4" pageNumber="423">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C41D5A7FFA9FF8BDB182F69FF72FE20" box="[1124,1350,835,859]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA9FF8BDB182F69FF72FE20" box="[1124,1350,835,859]" italics="true" pageId="4" pageNumber="423">Cottus koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
; counts for holotype are with an asterisk.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFA9FF8BDE032FBCFF7DFED7" blockId="4.[159,1402,918,1300]" box="[383,1353,918,940]" pageId="4" pageNumber="423">First dorsal fin Second dorsal fin Anal fin Pectoral fin</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDE032FEBFAD0F965" blockId="4.[159,1402,918,1300]" pageId="4" pageNumber="423">
|
||
VI VII
|
||
<date id="FFFF88E4FFA9FF8BDD782FEBF888FEAD" box="[516,700,960,982]" pageId="4" pageNumber="423" value="1916-08-15">VIII 15 16</date>
|
||
17 18
|
||
<date id="FFFF88E4FFA9FF8BDC092FEAFE2FFEAD" box="[885,1051,960,982]" pageId="4" pageNumber="423" value="1919-12-20">19 20 12</date>
|
||
13 14 13 14 15
|
||
<taxonomicName id="4C41D5A7FFA9FF8BDFE32FC0FB01FE84" box="[159,309,1002,1023]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="gratzianowi">
|
||
<emphasis id="B9357236FFA9FF8BDFE32FC0FB01FE84" box="[159,309,1002,1023]" italics="true" pageId="4" pageNumber="423">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
2 14*
|
||
<date id="FFFF88E4FFA9FF8BDD782FC3F936FE84" box="[516,770,1001,1023]" pageId="4" pageNumber="423" value="2013-04-02">2 4 13</date>
|
||
* 1 12* 6 4* 12 2 (n=18)
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFE32818FB18F91D" blockId="4.[159,1402,918,1300]" pageId="4" pageNumber="423">
|
||
<taxonomicName id="4C41D5A7FFA9FF8BDFE32818FB7EF93D" box="[159,330,1073,1095]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA9FF8BDFE32818FB7EF93D" box="[159,330,1073,1095]" italics="true" pageId="4" pageNumber="423">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
<date id="FFFF88E4FFA9FF8BDE03281BF825F93C" box="[383,529,1073,1095]" pageId="4" pageNumber="423" value="1919-01-03">1 19 3</date>
|
||
<date id="FFFF88E4FFA9FF8BDDDE281BF973F93C" box="[674,839,1073,1095]" pageId="4" pageNumber="423" value="2011-09-02">2 9 11</date>
|
||
<date id="FFFF88E4FFA9FF8BDC09281BFE54F93C" box="[885,1120,1073,1095]" pageId="4" pageNumber="423" value="2012-08-01">1 8 12</date>
|
||
<date id="FFFF88E4FFA9FF8BDBF2281BFF00F93C" box="[1166,1332,1073,1095]" pageId="4" pageNumber="423" value="1916-04-03">3 4 16</date>
|
||
3 (n=23), Volga
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFE32850FB74F9D5" blockId="4.[159,1402,918,1300]" pageId="4" pageNumber="423">
|
||
<taxonomicName id="4C41D5A7FFA9FF8BDFE32850FB7EF9F5" box="[159,330,1145,1167]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA9FF8BDFE32850FB7EF9F5" box="[159,330,1145,1167]" italics="true" pageId="4" pageNumber="423">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
3 2 2 3 4 1 1 3 1 (n=5), Northern
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFE3289CFAE9F9B7" blockId="4.[159,1402,918,1300]" box="[159,221,1206,1228]" pageId="4" pageNumber="423">Dvina</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFE328CAFB75F86F" blockId="4.[159,1402,918,1300]" pageId="4" pageNumber="423">
|
||
<taxonomicName id="4C41D5A7FFA9FF8BDFE328CAFB7EF98E" box="[159,330,1248,1269]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA9FF8BDFE328CAFB7EF98E" box="[159,330,1248,1269]" italics="true" pageId="4" pageNumber="423">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
5 6 3 5 2 1 3 8 2 8 1 (n=11), Pechora
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFBB297FFB4EFBC9" blockId="4.[151,1437,1365,1966]" pageId="4" pageNumber="423">
|
||
Anal fin originating at short distance from anus, below third ray of second dorsal fin; length of anal-fin base about 1.3 times as short as length of second dorsal fin base, and 2.8−3 times in SL (
|
||
<quantity id="4CB903C1FFA9FF8BDB322950FE4EF8E9" box="[1102,1146,1401,1426]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="423" unit="in" value="3.0">3 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDBFE2953FEDCF8E9" box="[1154,1256,1401,1426]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
), about equal to head length; length of longest anal-fin ray about equal to length of longest second dorsal-fin ray (equal in
|
||
<typeStatus id="54FA1086FFA9FF8BDFEB29EBFACAF8A1" box="[151,254,1473,1498]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
). Caudal fin rounded, with 11 or 12 rays supported by hypural plate, 8 middle rays branched and upper 2 and lower 1−2 rays unbranched, and 12−14 (6−7 dorsal and 6−7 ventral) procurrent rays. Pelvic fin extending to anus; two inner soft rays longer than two outer rays; outer ray with long free tip; narrow pelvic base 7 times in length of longest pelvic-fin ray. Pectoral-fin base oblique, broad, its base about 3 times in HL; pectoral-fin length 3−3.4 times in SL (
|
||
<quantity id="4CB903C1FFA9FF8BDE0E2A78FB81FB11" box="[370,437,1617,1642]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="423" unit="in" value="3.4">3.4 in</quantity>
|
||
<typeStatus id="54FA1086FFA9FF8BDEC12A7BF810FB11" box="[445,548,1617,1642]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
); tip of longest ray reaching below base of third or fourth ray of second dorsal fin; one to three medial (longest) rays clearly branched in 56% of examined specimens (in
|
||
<typeStatus id="54FA1086FFA9FF8BDB952A5FFF7FFBF5" box="[1257,1355,1653,1678]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
and 9
|
||
<typeStatus id="54FA1086FFA9FF8BDFEB2AB0FB30FBC9" box="[151,260,1690,1714]" pageId="4" pageNumber="423" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<figureCitation id="137AB2A1FFA9FF8BDE672AB3FB6AFBC9" box="[283,350,1689,1714]" captionStart="FIGURE 5" captionStartId="6.[151,250,575,597]" captionTargetBox="[161,1425,196,552]" captionTargetId="figure@6.[151,1436,193,554]" captionTargetPageId="6" captionText="FIGURE 5. Photograph of pectoral fin. Arrows show branched middle pectoral-fin rays in Cottus gratzianowi sp. nov. (a) and unbranched middle pectoral-fin rays in Cottus koshewnikowi (b)." httpUri="https://zenodo.org/record/243677/files/figure.png" pageId="4" pageNumber="423">Fig. 5</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA9FF8BDFBB2A97F90CFAD5" blockId="4.[151,1437,1365,1966]" pageId="4" pageNumber="423">
|
||
<emphasis id="B9357236FFA9FF8BDFBB2A97FB97FBAE" box="[199,419,1725,1749]" italics="true" pageId="4" pageNumber="423">Lateral-line system</emphasis>
|
||
. Cephalic sensory canals developed and interconnected except preoperculo-mandibular sensory canal. Supraorbital canal pores 3 (on each side); infraorbital canal pores 8−9 (usually 8); temporal canal pores 3. Temporal canals of left and right side connected via supratemporal commissure interrupted in middle, with two paired pores instead of one; preoperculo-mandibular canal pores 11 on both sides with five paired pores and single large median pore on chin. Lateral-line canal with 15−24 pores (averaging 19.8) (
|
||
<tableCitation id="C6C39B9FFFA9FF8BDB062B67FEE5FA1D" box="[1146,1233,1869,1894]" captionStart="TABLE 3" captionStartId="6.[151,239,667,690]" captionTargetBox="[159,1388,718,1058]" captionTargetPageId="6" captionText="TABLE 3. Frequency of occurrence of lateral-line counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri="http://table.plazi.org/id/DF3EFEACFFABFF89DFEB2EB1FF57FFC9" pageId="4" pageNumber="423" tableUuid="DF3EFEACFFABFF89DFEB2EB1FF57FFC9">Table 3</tableCitation>
|
||
), extending along dorsal profile of body, reaching below fifth to tenth ray of second dorsal fin, sometimes (in
|
||
<specimenCount id="9D4765ADFFA9FF8BDBDA2B58FF05FAF1" box="[1190,1329,1905,1930]" pageId="4" pageNumber="423" type="generic">2 specimens</specimenCount>
|
||
from 18) interrupted at very end with one or two free canal segments.
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFA8FF8ADFEB2800F990F93B" box="[151,932,1066,1088]" httpUri="https://zenodo.org/record/243675/files/figure.png" pageId="5" pageNumber="424" targetBox="[176,1401,238,987]" targetPageId="5">
|
||
<paragraph id="8BFEAE24FFA8FF8ADFEB2800F990F93B" blockId="5.[151,932,1066,1088]" box="[151,932,1066,1088]" pageId="5" pageNumber="424">
|
||
<emphasis id="B9357236FFA8FF8ADFEB2800FB39F944" bold="true" box="[151,269,1066,1088]" pageId="5" pageNumber="424">FIGURE 3</emphasis>
|
||
. Photograph of
|
||
<taxonomicName id="4C41D5A7FFA8FF8ADED32800F845F93B" box="[431,625,1066,1088]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA8FF8ADED32800F845F93B" box="[431,625,1066,1088]" italics="true" pageId="5" pageNumber="424">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA8FF8ADD0B2800F8F6F944" box="[631,706,1066,1087]" pageId="5" pageNumber="424" rank="species">
|
||
<emphasis id="B9357236FFA8FF8ADD0B2800F8F6F944" bold="true" box="[631,706,1066,1087]" pageId="5" pageNumber="424">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype as in Fig. 2.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3EFEACFFA8FF8ADFEB2B77FE29FA08" box="[151,1053,1885,1907]" httpUri="https://zenodo.org/record/243676/files/figure.png" pageId="5" pageNumber="424" targetBox="[151,1436,1137,1863]" targetPageId="5">
|
||
<paragraph id="8BFEAE24FFA8FF8ADFEB2B77FE29FA08" blockId="5.[151,1053,1885,1907]" box="[151,1053,1885,1907]" pageId="5" pageNumber="424">
|
||
<emphasis id="B9357236FFA8FF8ADFEB2B77FB26FA08" bold="true" box="[151,274,1885,1907]" pageId="5" pageNumber="424">FIGURE 4.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="4C41D5A7FFA8FF8ADED32B74F8B2FA08" box="[431,646,1886,1907]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA8FF8ADED32B74F8B2FA08" box="[431,646,1886,1907]" italics="true" pageId="5" pageNumber="424">Cottus koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
, syntype, ZMMU Р-2688, 71.0 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3EFEACFFABFF89DFEB2E15F91CFF0F" httpUri="https://zenodo.org/record/243677/files/figure.png" pageId="6" pageNumber="425" targetBox="[161,1425,196,552]" targetPageId="6">
|
||
<paragraph id="8BFEAE24FFABFF89DFEB2E15F91CFF0F" blockId="6.[151,1435,575,628]" pageId="6" pageNumber="425">
|
||
<emphasis id="B9357236FFABFF89DFEB2E15FB39FF2E" bold="true" box="[151,269,575,597]" pageId="6" pageNumber="425">FIGURE 5</emphasis>
|
||
. Photograph of pectoral fin. Arrows show branched middle pectoral-fin rays in
|
||
<taxonomicName id="4C41D5A7FFABFF89DB412E6AFECBFF2E" box="[1085,1279,576,597]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFABFF89DB412E6AFECBFF2E" box="[1085,1279,576,597]" italics="true" pageId="6" pageNumber="425">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFABFF89DA7A2E6AFF7AFF2E" box="[1286,1358,576,597]" pageId="6" pageNumber="425" rank="species">sp.nov.</taxonomicNameLabel>
|
||
(a) and unbranched middle pectoral-fin rays in
|
||
<taxonomicName id="4C41D5A7FFABFF89DD5B2E74F8C9FF08" box="[551,765,606,627]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DD5B2E74F8C9FF08" box="[551,765,606,627]" italics="true" pageId="6" pageNumber="425">Cottus koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(b).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3EFEACFFABFF89DFEB2EB1FF57FFC9" ID-Table-UUID="DF3EFEACFFABFF89DFEB2EB1FF57FFC9" box="[151,1379,666,691]" httpUri="http://table.plazi.org/id/DF3EFEACFFABFF89DFEB2EB1FF57FFC9" pageId="6" pageNumber="425" targetBox="[159,1388,718,1058]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="8BFEAE24FFABFF89DFEB2EB1FF57FFC9" blockId="6.[151,1379,666,691]" box="[151,1379,666,691]" pageId="6" pageNumber="425">
|
||
<emphasis id="B9357236FFABFF89DFEB2EB1FB37FFC9" bold="true" box="[151,259,667,690]" pageId="6" pageNumber="425">TABLE 3</emphasis>
|
||
. Frequency of occurrence of lateral-line counts in
|
||
<taxonomicName id="4C41D5A7FFABFF89DC632EB1F9DEFFC9" box="[799,1002,667,690]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFABFF89DC632EB1F9DEFFC9" box="[799,1002,667,690]" italics="true" pageId="6" pageNumber="425">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFABFF89DC8D2EB1FE73FFC9" box="[1009,1095,667,690]" pageId="6" pageNumber="425" rank="species">
|
||
<emphasis id="B9357236FFABFF89DC8D2EB1FE73FFC9" bold="true" box="[1009,1095,667,690]" pageId="6" pageNumber="425">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C41D5A7FFABFF89DB062EB1FF6BFFC9" box="[1146,1375,666,690]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">Cottus koshewnikowi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFABFF89DEFE2EE4FF54F97F" pageId="6" pageNumber="425">
|
||
<table id="F9415C84FFAB0070DFE32EE4FF58F959" box="[159,1388,718,1058]" gridcols="14" gridrows="5" pageId="6" pageNumber="425">
|
||
<tr id="3571AC66FFAB0070DFE32EE4FF58FF9F" box="[159,1388,718,740]" gridrow="0" pageId="6" pageNumber="425" rowspan-0="1">
|
||
<th id="76A0C51AFFAB0070DEFE2EE4FBAFFF9F" box="[386,411,718,740]" gridcol="1" gridrow="0" pageId="6" pageNumber="425">15</th>
|
||
<th id="76A0C51AFFAB0070DEA82EE4FBD9FF9F" box="[468,493,718,740]" gridcol="2" gridrow="0" pageId="6" pageNumber="425">16</th>
|
||
<th id="76A0C51AFFAB0070DD592EE4F80AFF9F" box="[549,574,718,740]" gridcol="3" gridrow="0" pageId="6" pageNumber="425">17</th>
|
||
<th id="76A0C51AFFAB0070DD0A2EE4F8BBFF9F" box="[630,655,718,740]" gridcol="4" gridrow="0" pageId="6" pageNumber="425">18</th>
|
||
<th id="76A0C51AFFAB0070DDB42EE4F8D5FF9F" box="[712,737,718,740]" gridcol="5" gridrow="0" pageId="6" pageNumber="425">19</th>
|
||
<th id="76A0C51AFFAB0070DC652EE4F906FF9F" box="[793,818,718,740]" gridcol="6" gridrow="0" pageId="6" pageNumber="425">20</th>
|
||
<th id="76A0C51AFFAB0070DC172EE4F9B0FF9F" box="[875,900,718,740]" gridcol="7" gridrow="0" pageId="6" pageNumber="425">21</th>
|
||
<th id="76A0C51AFFAB0070DCC02EE4F9E1FF9F" box="[956,981,718,740]" gridcol="8" gridrow="0" pageId="6" pageNumber="425">22</th>
|
||
<th id="76A0C51AFFAB0070DB712EE4FE13FF9F" box="[1037,1063,718,740]" gridcol="9" gridrow="0" pageId="6" pageNumber="425">23</th>
|
||
<th id="76A0C51AFFAB0070DB232EE4FE4CFF9F" box="[1119,1144,718,740]" gridcol="10" gridrow="0" pageId="6" pageNumber="425">24</th>
|
||
<th id="76A0C51AFFAB0070DBCC2EE4FEFDFF9F" box="[1200,1225,718,740]" gridcol="11" gridrow="0" pageId="6" pageNumber="425">25</th>
|
||
<th id="76A0C51AFFAB0070DA7E2EE4FF2FFF9F" box="[1282,1307,718,740]" gridcol="12" gridrow="0" pageId="6" pageNumber="425">26</th>
|
||
<th id="76A0C51AFFAB0070DA2F2EE4FF58FF9F" box="[1363,1388,718,740]" gridcol="13" gridrow="0" pageId="6" pageNumber="425">27</th>
|
||
</tr>
|
||
<tr id="3571AC66FFAB0070DFE32EDDFF58FE57" box="[159,1388,759,812]" gridrow="1" pageId="6" pageNumber="425" rowspan-11="1" rowspan-12="1" rowspan-13="1">
|
||
<th id="76A0C51AFFAB0070DFE32EDDFB7EFE57" box="[159,330,759,812]" gridcol="0" gridrow="1" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32ED2FB01FE76" box="[159,309,760,781]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gratzianowi">
|
||
<emphasis id="B9357236FFABFF89DFE32ED2FB01FE76" box="[159,309,760,781]" italics="true" pageId="6" pageNumber="425">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
(n=17)
|
||
</th>
|
||
<td id="76A0C51AFFAB0070DEFE2EDDFBAFFE57" box="[386,411,759,812]" gridcol="1" gridrow="1" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DEA82EDDFBD9FE57" box="[468,493,759,812]" gridcol="2" gridrow="1" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DD592EDDF80AFE57" box="[549,574,759,812]" gridcol="3" gridrow="1" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DD0A2EDDF8BBFE57" box="[630,655,759,812]" gridcol="4" gridrow="1" pageId="6" pageNumber="425">3*</td>
|
||
<td id="76A0C51AFFAB0070DDB42EDDF8D5FE57" box="[712,737,759,812]" gridcol="5" gridrow="1" pageId="6" pageNumber="425">4</td>
|
||
<td id="76A0C51AFFAB0070DC652EDDF906FE57" box="[793,818,759,812]" gridcol="6" gridrow="1" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DC172EDDF9B0FE57" box="[875,900,759,812]" gridcol="7" gridrow="1" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DCC02EDDF9E1FE57" box="[956,981,759,812]" gridcol="8" gridrow="1" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DB712EDDFE13FE57" box="[1037,1063,759,812]" gridcol="9" gridrow="1" pageId="6" pageNumber="425">2</td>
|
||
<td id="76A0C51AFFAB0070DB232EDDFE4CFE57" box="[1119,1144,759,812]" gridcol="10" gridrow="1" pageId="6" pageNumber="425">2</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAB0070DFE32F15FF58FE0F" box="[159,1388,831,884]" gridrow="2" pageId="6" pageNumber="425" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
||
<th id="76A0C51AFFAB0070DFE32F15FB7EFE0F" box="[159,330,831,884]" gridcol="0" gridrow="2" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32F6AFB7EFE2E" box="[159,330,832,853]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DFE32F6AFB7EFE2E" box="[159,330,832,853]" italics="true" pageId="6" pageNumber="425">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(n=31), Volga
|
||
</th>
|
||
<td id="76A0C51AFFAB0070DC172F15F9B0FE0F" box="[875,900,831,884]" gridcol="7" gridrow="2" pageId="6" pageNumber="425">2</td>
|
||
<td id="76A0C51AFFAB0070DCC02F15F9E1FE0F" box="[956,981,831,884]" gridcol="8" gridrow="2" pageId="6" pageNumber="425">2</td>
|
||
<td id="76A0C51AFFAB0070DB712F15FE13FE0F" box="[1037,1063,831,884]" gridcol="9" gridrow="2" pageId="6" pageNumber="425">5</td>
|
||
<td id="76A0C51AFFAB0070DB232F15FE4CFE0F" box="[1119,1144,831,884]" gridcol="10" gridrow="2" pageId="6" pageNumber="425">8</td>
|
||
<td id="76A0C51AFFAB0070DBCC2F15FEFDFE0F" box="[1200,1225,831,884]" gridcol="11" gridrow="2" pageId="6" pageNumber="425">6</td>
|
||
<td id="76A0C51AFFAB0070DA7E2F15FF2FFE0F" box="[1282,1307,831,884]" gridcol="12" gridrow="2" pageId="6" pageNumber="425">4</td>
|
||
<td id="76A0C51AFFAB0070DA2F2F15FF58FE0F" box="[1363,1388,831,884]" gridcol="13" gridrow="2" pageId="6" pageNumber="425">4</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAB0070DFE32FADFF58FEA1" box="[159,1388,903,986]" gridrow="3" pageId="6" pageNumber="425" rowspan-1="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76A0C51AFFAB0070DFE32FADFB7EFEA1" box="[159,330,903,986]" gridcol="0" gridrow="3" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32FA2FB7EFEE6" box="[159,330,904,925]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DFE32FA2FB7EFEE6" box="[159,330,904,925]" italics="true" pageId="6" pageNumber="425">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(n=5), Northern Dvina
|
||
</th>
|
||
<td id="76A0C51AFFAB0070DC652FADF906FEA1" box="[793,818,903,986]" gridcol="6" gridrow="3" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DCC02FADF9E1FEA1" box="[956,981,903,986]" gridcol="8" gridrow="3" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DB232FADFE4CFEA1" box="[1119,1144,903,986]" gridcol="10" gridrow="3" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DBCC2FADFEFDFEA1" box="[1200,1225,903,986]" gridcol="11" gridrow="3" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DA7E2FADFF2FFEA1" box="[1282,1307,903,986]" gridcol="12" gridrow="3" pageId="6" pageNumber="425">1</td>
|
||
</tr>
|
||
<tr id="3571AC66FFAB0070DFE32FC4FF58F959" box="[159,1388,1006,1058]" gridrow="4" pageId="6" pageNumber="425" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-7="1">
|
||
<th id="76A0C51AFFAB0070DFE32FC4FB7EF959" box="[159,330,1006,1058]" gridcol="0" gridrow="4" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32FC4FB7EF978" box="[159,330,1006,1027]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DFE32FC4FB7EF978" box="[159,330,1006,1027]" italics="true" pageId="6" pageNumber="425">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(n=11), Pechora
|
||
</th>
|
||
<td id="76A0C51AFFAB0070DD592FC4F80AF959" box="[549,574,1006,1058]" gridcol="3" gridrow="4" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DD0A2FC4F8BBF959" box="[630,655,1006,1058]" gridcol="4" gridrow="4" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DC652FC4F906F959" box="[793,818,1006,1058]" gridcol="6" gridrow="4" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DCC02FC4F9E1F959" box="[956,981,1006,1058]" gridcol="8" gridrow="4" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DB712FC4FE13F959" box="[1037,1063,1006,1058]" gridcol="9" gridrow="4" pageId="6" pageNumber="425">3</td>
|
||
<td id="76A0C51AFFAB0070DB232FC4FE4CF959" box="[1119,1144,1006,1058]" gridcol="10" gridrow="4" pageId="6" pageNumber="425">3</td>
|
||
<td id="76A0C51AFFAB0070DBCC2FC4FEFDF959" box="[1200,1225,1006,1058]" gridcol="11" gridrow="4" pageId="6" pageNumber="425">5</td>
|
||
<td id="76A0C51AFFAB0070DA7E2FC4FF2FF959" box="[1282,1307,1006,1058]" gridcol="12" gridrow="4" pageId="6" pageNumber="425">1</td>
|
||
<td id="76A0C51AFFAB0070DA2F2FC4FF58F959" box="[1363,1388,1006,1058]" gridcol="13" gridrow="4" pageId="6" pageNumber="425">1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFBB284EFE30F9DB" blockId="6.[151,1436,1123,1184]" pageId="6" pageNumber="425">
|
||
<emphasis id="B9357236FFABFF89DFBB284EFB00F907" box="[199,308,1124,1148]" italics="true" pageId="6" pageNumber="425">Vertebrae</emphasis>
|
||
. Abdominal vertebrae 10 or 11; caudal vertebrae 22 or 23, total 32−34 (
|
||
<tableCitation id="C6C39B9FFFABFF89DB162849FEF7F907" box="[1130,1219,1123,1148]" captionStart="TABLE 4" captionStartId="6.[151,239,1227,1250]" captionText="TABLE 4. Frequency of occurrence of vertebral counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi." pageId="6" pageNumber="425">Table 4</tableCitation>
|
||
); long, thin pleural ribs on the parapophyses of commonly three last abdominal vertebrae (
|
||
<figureCitation id="137AB2A1FFABFF89DCCE28ADF9C3F9DB" box="[946,1015,1159,1184]" captionStart="FIGURE 6" captionStartId="7.[151,250,918,940]" captionTargetBox="[151,1436,193,895]" captionTargetId="figure@7.[151,1436,193,896]" captionTargetPageId="7" captionText="FIGURE 6. Radiographs of Cottus gratzianowi sp. nov. (a) with 10 + 23 vertebrae and Cottus koshewnikowi (b) with 11 + 23 vertebrae. Arrow shows first caudal vertebra." httpUri="https://zenodo.org/record/243678/files/figure.png" pageId="6" pageNumber="425">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFABFF89DFEB28E1FF7AF999" box="[151,1358,1226,1251]" pageId="6" pageNumber="425">
|
||
<paragraph id="8BFEAE24FFABFF89DFEB28E1FF7AF999" blockId="6.[151,1358,1226,1251]" box="[151,1358,1226,1251]" pageId="6" pageNumber="425">
|
||
<emphasis id="B9357236FFABFF89DFEB28E1FB37F999" bold="true" box="[151,259,1227,1250]" pageId="6" pageNumber="425">TABLE 4</emphasis>
|
||
. Frequency of occurrence of vertebral counts in
|
||
<taxonomicName id="4C41D5A7FFABFF89DC7628E1F9E1F999" box="[778,981,1227,1250]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFABFF89DC7628E1F9E1F999" box="[778,981,1227,1250]" italics="true" pageId="6" pageNumber="425">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFABFF89DCA028E1FE06F999" box="[988,1074,1227,1250]" pageId="6" pageNumber="425" rank="species">
|
||
<emphasis id="B9357236FFABFF89DCA028E1FE06F999" bold="true" box="[988,1074,1227,1250]" pageId="6" pageNumber="425">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C41D5A7FFABFF89DB1928E1FF7EF999" box="[1125,1354,1226,1250]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">Cottus koshewnikowi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFABFF89DE0328D4FF3FF86F" blockId="6.[159,1391,1278,1696]" box="[383,1291,1278,1300]" pageId="6" pageNumber="425">Predorsal 2 Abdominal Caudal vertebrae Total vertebrae</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DE032936F88BF849" blockId="6.[159,1391,1278,1696]" box="[383,703,1308,1330]" pageId="6" pageNumber="425">vertebrae vertebrae</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DE03296CFF5BF827" blockId="6.[159,1391,1278,1696]" box="[383,1391,1350,1372]" pageId="6" pageNumber="425">
|
||
<date id="FFFF88E4FFABFF89DE03296CF81BF827" box="[383,559,1350,1372]" pageId="6" pageNumber="425" value="2010-09-08">8 9 10</date>
|
||
<date id="FFFF88E4FFABFF89DD1D296CF926F827" box="[609,786,1350,1372]" pageId="6" pageNumber="425" value="2012-11-10">10 11 12</date>
|
||
21 22 23 24 32 33 34 35
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFE3295FFAD0F8D2" blockId="6.[159,1391,1278,1696]" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE3295FFB01F8F1" box="[159,309,1397,1418]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gratzianowi">
|
||
<emphasis id="B9357236FFABFF89DFE3295FFB01F8F1" box="[159,309,1397,1418]" italics="true" pageId="6" pageNumber="425">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
2 16* 12 6* 6 12*
|
||
<date id="FFFF88E4FFABFF89DB0F295EFF2CF8F1" box="[1139,1304,1396,1418]" pageId="6" pageNumber="425" value="2014-02-02">2 14 2</date>
|
||
* (n=18)
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFE32997FB06F88A" blockId="6.[159,1391,1278,1696]" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32997FB7EF8A9" box="[159,330,1469,1490]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DFE32997FB7EF8A9" box="[159,330,1469,1490]" italics="true" pageId="6" pageNumber="425">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
<date id="FFFF88E4FFABFF89DE032996F817F8A9" box="[383,547,1468,1490]" pageId="6" pageNumber="425" value="2012-09-02">9 12 2</date>
|
||
<date id="FFFF88E4FFABFF89DD1D2996F965F8A9" box="[609,849,1468,1490]" pageId="6" pageNumber="425" value="1922-01-02">1 22 2</date>
|
||
<date id="FFFF88E4FFABFF89DCEC2996FEB4F8A9" box="[912,1152,1468,1490]" pageId="6" pageNumber="425" value="2012-02-09">12 9 2</date>
|
||
13 8 (n= 23), Volga
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFE32A2FFB74FB42" blockId="6.[159,1391,1278,1696]" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32A2FFB7EFB61" box="[159,330,1541,1562]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DFE32A2FFB7EFB61" box="[159,330,1541,1562]" italics="true" pageId="6" pageNumber="425">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
5 5 1 4 1 4 (n=5), Northern
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFE32A68FAE9FB23" blockId="6.[159,1391,1278,1696]" box="[159,221,1602,1624]" pageId="6" pageNumber="425">Dvina</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFE32A46FB75FBDB" blockId="6.[159,1391,1278,1696]" pageId="6" pageNumber="425">
|
||
<taxonomicName id="4C41D5A7FFABFF89DFE32A46FB7EFBFB" box="[159,330,1643,1665]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFABFF89DFE32A46FB7EFBFB" box="[159,330,1643,1665]" italics="true" pageId="6" pageNumber="425">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
1 10
|
||
<date id="FFFF88E4FFABFF89DDD12A41F9A9FBFA" box="[685,925,1643,1665]" pageId="6" pageNumber="425" value="2010-02-01">10 1 2</date>
|
||
8 1 2 7 2 (n=11), Pechora
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFABFF89DFBB2ACAFEDEFA1D" blockId="6.[151,1437,1760,2038]" pageId="6" pageNumber="425">
|
||
<emphasis id="B9357236FFABFF89DFBB2ACAFB6BFB82" bold="true" box="[199,351,1760,1785]" pageId="6" pageNumber="425">Colouration.</emphasis>
|
||
Body background light olive; overall body dark mottled, blotches forming marble pattern (
|
||
<figureCitation id="137AB2A1FFABFF89DA132ACAFA9CFA66" captionStart="FIGURE 2" captionStartId="2.[151,250,1531,1553]" captionTargetBox="[170,1421,514,1477]" captionTargetId="figure@2.[332,1303,901,1482]" captionTargetPageId="2" captionText="FIGURE 2. Cottus gratzianowi sp. nov., holotype, ZIN 55596, male 63.3 mm SL: lateral view (a), dorsal view of head and anterior body (b), ventral view of head and anterior body (c)." httpUri="https://zenodo.org/record/243674/files/figure.png" pageId="6" pageNumber="425">Fig. 2</figureCitation>
|
||
). Head dorsally dark, with dark blotches laterally similar to body. Distinct melanophores and their clusters densely located on ventral surface of head and bases of pectoral and pelvic fins. Dark spots forming transverse bands on both dorsal, pectoral and caudal fins. First dorsal-fin margin in adult males light, whitish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BFDAFFFABFF89DFBB2B5AF941FA8D" pageId="6" pageNumber="425" type="distribution">
|
||
<paragraph id="8BFEAE24FFABFF89DFBB2B5AF941FA8D" blockId="6.[151,1437,1760,2038]" pageId="6" pageNumber="425">
|
||
<emphasis id="B9357236FFABFF89DFBB2B5AFBCFFAF2" bold="true" box="[199,507,1904,1929]" pageId="6" pageNumber="425">Distribution and habitat.</emphasis>
|
||
The species is currently known from the Ukhtomitsa River, a tributary of the Modlona River (a Lake Vozhe tributary) in the Onega River drainage, the White See basin (
|
||
<figureCitation id="137AB2A1FFABFF89DB922BBFFF0CFAD6" box="[1262,1336,1941,1966]" captionStart="FIGURE 8" captionStartId="10.[151,250,1121,1143]" captionTargetBox="[368,1218,193,1099]" captionTargetId="figure@10.[368,1218,193,1100]" captionTargetPageId="10" captionText="FIGURE 8. Map showing the type locality of Cottus gratzianowi sp. nov. in the Ukhtomitsa River, tributary of Modlona River, Lake Vozhe basin in the Onega River system." httpUri="https://zenodo.org/record/243680/files/figure.png" pageId="6" pageNumber="425">Fig. 8</figureCitation>
|
||
).
|
||
<taxonomicName id="4C41D5A7FFABFF89DA2F2BBFFB29FAAB" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFABFF89DA2F2BBFFB29FAAB" italics="true" pageId="6" pageNumber="425">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFABFF89DE592B92FBB4FAAA" box="[293,384,1976,2001]" pageId="6" pageNumber="425" rank="species">
|
||
<emphasis id="B9357236FFABFF89DE592B92FBB4FAAA" bold="true" box="[293,384,1976,2001]" pageId="6" pageNumber="425">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
occurs in a relatively cool in summer small river with moderate to rapid current, gravel and cobble bottom, and not very dense submerged aquatic vegetation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3EFEACFFAAFF88DFEB2FBCF855FEB1" httpUri="https://zenodo.org/record/243678/files/figure.png" pageId="7" pageNumber="426" targetBox="[151,1436,193,895]" targetPageId="7">
|
||
<paragraph id="8BFEAE24FFAAFF88DFEB2FBCF855FEB1" blockId="7.[151,1435,918,970]" pageId="7" pageNumber="426">
|
||
<emphasis id="B9357236FFAAFF88DFEB2FBCFB3BFED0" bold="true" box="[151,271,918,940]" pageId="7" pageNumber="426">FIGURE 6</emphasis>
|
||
. Radiographs of
|
||
<taxonomicName id="4C41D5A7FFAAFF88DEBF2FBCF8B2FED7" box="[451,646,918,940]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DEBF2FBCF8B2FED7" box="[451,646,918,940]" italics="true" pageId="7" pageNumber="426">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DDF22FBCF8D5FED0" box="[654,737,918,939]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DDF22FBCF8D5FED0" bold="true" box="[654,737,918,939]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(a) with 10+23 vertebrae and
|
||
<taxonomicName id="4C41D5A7FFAAFF88DB5C2FBCFECCFED0" box="[1056,1272,918,939]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DB5C2FBCFECCFED0" box="[1056,1272,918,939]" italics="true" pageId="7" pageNumber="426">Cottus koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(b) with 11+23 vertebrae. Arrow shows first caudal vertebra.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35BFDAFFFAAFF85DFBB2FDDFB9FF858" lastPageId="10" lastPageNumber="429" pageId="7" pageNumber="426" type="etymology">
|
||
<paragraph id="8BFEAE24FFAAFF88DFBB2FDDF98FF907" blockId="7.[151,1436,1015,2012]" pageId="7" pageNumber="426">
|
||
<emphasis id="B9357236FFAAFF88DFBB2FDDFB7AF96B" bold="true" box="[199,334,1015,1040]" pageId="7" pageNumber="426">Etymology.</emphasis>
|
||
The new species is named in honour of Valerian Ivanovich Gratzianow [Gratsianov] (
|
||
<date id="FFFF88E4FFAAFF88DA772FDDFFA4F96B" box="[1291,1424,1015,1040]" pageId="7" pageNumber="426" value="1876" valueMax="1932">1876−1932</date>
|
||
), the author of the first taxonomic review of Russian fishes "Versuch einer Übersicht der Fische des Russischen Reiches in systematischer und geographischer Hinsicht" (
|
||
<bibRefCitation id="EFD0D3D5FFAAFF88DC652815F9D8F92C" author="Gratzianow" box="[793,1004,1087,1112]" pageId="7" pageNumber="426" refString="Gratzianow, V. I. (1907 a) Versuch einer Ubersicht der Fische des Russischen Reiches in systematischer und geographischer Hinsicht. Moscow, 567 pp." type="book" year="1907" yearSuffix="a">Gratzianow 1907a</bibRefCitation>
|
||
), who described
|
||
<taxonomicName id="4C41D5A7FFAAFF88DBD4286AFFA1F92C" box="[1192,1429,1087,1111]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DBD4286AFFA1F92C" box="[1192,1429,1087,1111]" italics="true" pageId="7" pageNumber="426">Cottus koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
. We retained the original latinised spelling of the surname (Gratzianow).
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFAAFF88DFBB28ADFB35F803" blockId="7.[151,1436,1015,2012]" pageId="7" pageNumber="426">
|
||
<emphasis id="B9357236FFAAFF88DFBB28ADFB56F9DB" bold="true" box="[199,354,1159,1184]" pageId="7" pageNumber="426">Comparison.</emphasis>
|
||
<taxonomicName id="4C41D5A7FFAAFF88DE1528A2F874F9E4" box="[361,576,1160,1183]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DE1528A2F874F9E4" box="[361,576,1160,1183]" italics="true" pageId="7" pageNumber="426">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DD3B28ADF895F9DB" box="[583,673,1159,1184]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DD3B28ADF895F9DB" bold="true" box="[583,673,1159,1184]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
belongs to
|
||
<taxonomicName id="4C41D5A7FFAAFF88DC5928A2F9B2F9E4" box="[805,902,1159,1183]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gobio">
|
||
<emphasis id="B9357236FFAAFF88DC5928A2F9B2F9E4" box="[805,902,1159,1183]" italics="true" pageId="7" pageNumber="426">C. gobio</emphasis>
|
||
</taxonomicName>
|
||
species group as possessing no transverse dark bands on the pelvic fin, a commonly single median chin canal pore and non-spatulate prickles (see Kottelat & Freyhof 2007). Freshwater sculpins from the White Sea basin have not been distinguished from Volga sculpin and appeared in literature under the name either
|
||
<taxonomicName id="4C41D5A7FFAAFF88DDF928DEF921F870" box="[645,789,1267,1291]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gobio">
|
||
<emphasis id="B9357236FFAAFF88DDF928DEF921F870" box="[645,789,1267,1291]" italics="true" pageId="7" pageNumber="426">Cottus gobio</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<bibRefCitation id="EFD0D3D5FFAAFF88DC2928D9FE25F877" author="Novoselov" box="[853,1041,1267,1292]" pageId="7" pageNumber="426" refString="Novoselov A. P. (Ed.) (2008) Red data book of the Arkhangelsk Region. Pisces. Arkhangelsk, pp. 239 - 247. (In Russian)" type="book chapter" year="2008">Novoselov 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD0D3D5FFAAFF88DB6028D9FF22F877" author="Novoselov" box="[1052,1302,1267,1292]" pageId="7" pageNumber="426" refString="Novoselov, A. P., Studenov, I .. I. & Pavlenko, V. I. (2013) Ecological diversity of freshwater fish fauna of the White Sea-Kuloi Peninsula (Arkhangelsk region). Arctic: Ecology and Economy, 4, 70 - 80. (In Russian)" type="book chapter" year="2013">
|
||
Novoselov
|
||
<emphasis id="B9357236FFAAFF88DBE028DEFEF9F870" box="[1180,1229,1267,1291]" italics="true" pageId="7" pageNumber="426">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD0D3D5FFAAFF88DA5E28D9FAE5F854" author="Parin" pageId="7" pageNumber="426" refString="Parin, N. V., Evseenko, S. A. & Vasil'eva, E. D. (2014) Fishes of Russian Seas. Annotated Catalogue, KMK Scientific Press Ltd. Moskow, 241 - 245." type="book chapter" year="2014">
|
||
Parin
|
||
<emphasis id="B9357236FFAAFF88DA1828DEFFA1F870" box="[1380,1429,1267,1291]" italics="true" pageId="7" pageNumber="426">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
) or
|
||
<taxonomicName id="4C41D5A7FFAAFF88DF812932FB8FF854" box="[253,443,1303,1327]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DF812932FB8FF854" box="[253,443,1303,1327]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(e.g., Kottelat & Freyhof 2007).
|
||
<taxonomicName id="4C41D5A7FFAAFF88DC532932FE31F854" box="[815,1029,1304,1327]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DC532932FE31F854" box="[815,1029,1304,1327]" italics="true" pageId="7" pageNumber="426">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DB70293DFE51F84B" box="[1036,1125,1303,1328]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DB70293DFE51F84B" bold="true" box="[1036,1125,1303,1328]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C41D5A7FFAAFF88DBE02932FF6DF854" box="[1180,1369,1303,1327]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DBE02932FF6DF854" box="[1180,1369,1303,1327]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
share an incomplete lateral line not reaching behind the anal-fin insertion and located considerably above the mid-line of the flank.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFAAFF87DFBB29AEFE20FCD6" blockId="7.[151,1436,1015,2012]" lastBlockId="8.[151,1436,151,429]" lastPageId="8" lastPageNumber="427" pageId="7" pageNumber="426">
|
||
<taxonomicName id="4C41D5A7FFAAFF88DFBB29AEFBAAF8E0" box="[199,414,1412,1435]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DFBB29AEFBAAF8E0" box="[199,414,1412,1435]" italics="true" pageId="7" pageNumber="426">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DEDB29A9F837F8E7" box="[423,515,1411,1436]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DEDB29A9F837F8E7" bold="true" box="[423,515,1411,1436]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
form the Onega River drainage is distinguished from
|
||
<taxonomicName id="4C41D5A7FFAAFF88DB0E29AEFF05F8E0" box="[1138,1329,1411,1435]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DB0E29AEFF05F8E0" box="[1138,1329,1411,1435]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
from the Volga (
|
||
<typeStatus id="54FA1086FFAAFF88DF9B298DFB2DF8C4" box="[231,281,1447,1471]" pageId="7" pageNumber="426">type</typeStatus>
|
||
locality) by a set of the following character states.
|
||
<taxonomicName id="4C41D5A7FFAAFF88DC212982FE00F8C4" box="[861,1076,1448,1471]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DC212982FE00F8C4" box="[861,1076,1448,1471]" italics="true" pageId="7" pageNumber="426">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DB40298DFEA2F8BB" box="[1084,1174,1447,1472]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DB40298DFEA2F8BB" bold="true" box="[1084,1174,1447,1472]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
possesses a larger eye: eye horizontal diameter, 23−28 (mean 25) % HL, is equal to or exceeds snout length, 20−27 (mean 24) % HL (vs. eye diameter less than snout length, 16−23, mean 19, and 22−28, mean 25, respectively) (
|
||
<tableCitation id="C6C39B9FFFAAFF88DBCA29C5FF39FB7C" box="[1206,1293,1519,1544]" captionStart="TABLE 1" captionStartId="3.[125,207,513,533]" captionTargetBox="[138,1845,573,1680]" captionTargetPageId="3" captionText="TABLE 1. Prοpοrtiοnal measurements οf Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri="http://table.plazi.org/id/DF3EFEACFFAEFF8CDF012E2BFE18FF6D" pageId="7" pageNumber="426" tableUuid="DF3EFEACFFAEFF8CDF012E2BFE18FF6D">Table 1</tableCitation>
|
||
), a rounded caudal fin (vs. truncated) (
|
||
<figureCitation id="137AB2A1FFAAFF88DEBD2A39F831FB57" box="[449,517,1555,1580]" captionStart="FIGURE 3" captionStartId="5.[151,250,1066,1088]" captionTargetBox="[176,1401,238,987]" captionTargetId="figure@5.[151,1436,194,1044]" captionTargetPageId="5" captionText="FIGURE 3. Photograph of Cottus gratzianowi sp. nov., holotype as in Fig. 2." httpUri="https://zenodo.org/record/243675/files/figure.png" pageId="7" pageNumber="426">Fig. 3</figureCitation>
|
||
,
|
||
<figureCitation id="137AB2A1FFAAFF88DD6F2A3EF817FB57" box="[531,547,1556,1580]" captionStart="FIGURE 4" captionStartId="5.[151,250,1885,1907]" captionTargetBox="[151,1436,1137,1863]" captionTargetId="figure@5.[151,1436,1137,1863]" captionTargetPageId="5" captionText="FIGURE 4. Photograph of Cottus koshewnikowi, syntype, ZMMU Р- 2688, 71.0 mm SL." httpUri="https://zenodo.org/record/243676/files/figure.png" pageId="7" pageNumber="426">4</figureCitation>
|
||
), the pelvic fin extending to the anus in both sexes (vs. not reaching the anus), frequent presence of one to three branched rays in median part of the pectoral fin (56% of specimens vs. 0 to 2%) (
|
||
<figureCitation id="137AB2A1FFAAFF88DFE32A71FAD2FB0F" box="[159,230,1627,1652]" captionStart="FIGURE 5" captionStartId="6.[151,250,575,597]" captionTargetBox="[161,1425,196,552]" captionTargetId="figure@6.[151,1436,193,554]" captionTargetPageId="6" captionText="FIGURE 5. Photograph of pectoral fin. Arrows show branched middle pectoral-fin rays in Cottus gratzianowi sp. nov. (a) and unbranched middle pectoral-fin rays in Cottus koshewnikowi (b)." httpUri="https://zenodo.org/record/243677/files/figure.png" pageId="7" pageNumber="426">Fig. 5</figureCitation>
|
||
), the supratemporal canal commissure often interrupted in the middle and possessing 4 pores (vs. noninterrupted, with 3 pores), a short lateral-line canal extending to below the fifth to tenth ray of the second dorsal fin, with 15−24 pores (vs. extending to below the ninth to fifteenth ray of the second dorsal fin, with 21−27 pores) (
|
||
<tableCitation id="C6C39B9FFFAAFF88DFE22AEDFACDFBA4" box="[158,249,1735,1760]" captionStart="TABLE 3" captionStartId="6.[151,239,667,690]" captionTargetBox="[159,1388,718,1058]" captionTargetPageId="6" captionText="TABLE 3. Frequency of occurrence of lateral-line counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri="http://table.plazi.org/id/DF3EFEACFFABFF89DFEB2EB1FF57FFC9" pageId="7" pageNumber="426" tableUuid="DF3EFEACFFABFF89DFEB2EB1FF57FFC9">Table 3</tableCitation>
|
||
). Only six from 18 (33%) specimens of
|
||
<taxonomicName id="4C41D5A7FFAAFF88DD942AE2F9A0FBA4" box="[744,916,1736,1759]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DD942AE2F9A0FBA4" box="[744,916,1736,1759]" italics="true" pageId="7" pageNumber="426">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DCDC2AEDF9CBFB9B" box="[928,1023,1735,1760]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DCDC2AEDF9CBFB9B" bold="true" box="[928,1023,1735,1760]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have the lateral-line canal slightly interrupted at its end with one or two free canal segments while
|
||
<taxonomicName id="4C41D5A7FFAAFF88DC0F2AC6FE05FA78" box="[883,1073,1771,1795]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DC0F2AC6FE05FA78" box="[883,1073,1771,1795]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
commonly (94% of specimens) have the lateral line considerably interrupted with two to nine free segments in its posterior section. Post-pectoral prickling in
|
||
<taxonomicName id="4C41D5A7FFAAFF88DE5C2B1EFBF2FA30" box="[288,454,1844,1867]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DE5C2B1EFBF2FA30" box="[288,454,1844,1867]" italics="true" pageId="7" pageNumber="426">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DEB12B19F813FA37" box="[461,551,1843,1868]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DEB12B19F813FA37" bold="true" box="[461,551,1843,1868]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is dense and occupies a wider area than in
|
||
<taxonomicName id="4C41D5A7FFAAFF88DB762B1EFEFCFA30" box="[1034,1224,1843,1867]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DB762B1EFEFCFA30" box="[1034,1224,1843,1867]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
where prickels are sparse and located only under the pectoral fin. The new species has contrasting black blotches on all fins including the pelvic and anal fins while no blotches on either pelvic or anal fin are developed in
|
||
<taxonomicName id="4C41D5A7FFAAFF88DB182B56FF16FAE8" box="[1124,1314,1915,1939]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DB182B56FF16FAE8" box="[1124,1314,1915,1939]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
. There are also some differences in fin-ray counts between
|
||
<taxonomicName id="4C41D5A7FFAAFF88DDAB2B8AF9B6FACC" box="[727,898,1952,1975]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFAAFF88DDAB2B8AF9B6FACC" box="[727,898,1952,1975]" italics="true" pageId="7" pageNumber="426">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFAAFF88DCF22BB5F9D8FAC3" box="[910,1004,1951,1976]" pageId="7" pageNumber="426" rank="species">
|
||
<emphasis id="B9357236FFAAFF88DCF22BB5F9D8FAC3" bold="true" box="[910,1004,1951,1976]" pageId="7" pageNumber="426">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C41D5A7FFAAFF88DB502B8AFEDBFACC" box="[1068,1263,1951,1975]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFAAFF88DB502B8AFEDBFACC" box="[1068,1263,1951,1975]" italics="true" pageId="7" pageNumber="426">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<typeStatus id="54FA1086FFAAFF88DA102B8AFFA8FAC3" box="[1388,1436,1952,1976]" pageId="7" pageNumber="426">type</typeStatus>
|
||
locality (
|
||
<tableCitation id="C6C39B9FFFAAFF88DF862BE9FB66FAA7" box="[250,338,1987,2012]" captionStart="TABLE 2" captionStartId="4.[151,239,836,859]" captionText="TABLE 2. Frequency of occurrence of fin-ray counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi; counts for holotype are with an asterisk." pageId="7" pageNumber="426">Table 2</tableCitation>
|
||
), the former having lower numbers of second dorsal-fin rays (15−18, averaging 16.8, vs. 16−20, averaging 17.5) and anal-fin rays (12−13, averaging 12.3, vs. 12−14, averaging 12.8). In the new species, total vertebrae are 32−34 similar to
|
||
<taxonomicName id="4C41D5A7FFA5FF87DD7F2C96F8F0FDAF" box="[515,708,188,212]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA5FF87DD7F2C96F8F0FDAF" box="[515,708,188,212]" italics="true" pageId="8" pageNumber="427">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
from the Volga but in
|
||
<taxonomicName id="4C41D5A7FFA5FF87DCA52C96FEB0FDAF" box="[985,1156,188,212]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA5FF87DCA52C96FEB0FDAF" box="[985,1156,188,212]" italics="true" pageId="8" pageNumber="427">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA5FF87DBF32C96FED8FDAE" box="[1167,1260,188,213]" pageId="8" pageNumber="427" rank="species">
|
||
<emphasis id="B9357236FFA5FF87DBF32C96FED8FDAE" bold="true" box="[1167,1260,188,213]" pageId="8" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the number of abdominal vertebrae is most often 10 (averaging 10.3) vs. commonly
|
||
<quantity id="4CB903C1FFA5FF87DCE12CCAF9E0FD83" box="[925,980,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="8" pageNumber="427" unit="in" value="11.0">11 in</quantity>
|
||
Volga
|
||
<taxonomicName id="4C41D5A7FFA5FF87DB5D2CCAFEEAFD83" box="[1057,1246,224,248]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA5FF87DB5D2CCAFEEAFD83" box="[1057,1246,224,248]" italics="true" pageId="8" pageNumber="427">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(averaging 11.0) while the number of caudal vertebrae is commonly 23 (averaging 22.7) vs. 21−23, commonly 22 (averaging 22.3). This vertebral feature, a longer caudal region, in combination with a shorter anal fin (commonly 12 rays vs. 13, see
|
||
<tableCitation id="C6C39B9FFFA5FF87DFEB2D66FAD8FC1F" box="[151,236,332,357]" captionStart="TABLE 2" captionStartId="4.[151,239,836,859]" captionText="TABLE 2. Frequency of occurrence of fin-ray counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi; counts for holotype are with an asterisk." pageId="8" pageNumber="427">Table 2</tableCitation>
|
||
) determines a relatively longer caudal peduncle in the new species as shown above (mean 12.8% SL vs. 8.7% SL, see
|
||
<tableCitation id="C6C39B9FFFA5FF87DE442D45FBBFFCF3" box="[312,395,367,392]" captionStart="TABLE 1" captionStartId="3.[125,207,513,533]" captionTargetBox="[138,1845,573,1680]" captionTargetPageId="3" captionText="TABLE 1. Prοpοrtiοnal measurements οf Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri="http://table.plazi.org/id/DF3EFEACFFAEFF8CDF012E2BFE18FF6D" pageId="8" pageNumber="427" tableUuid="DF3EFEACFFAEFF8CDF012E2BFE18FF6D">Table 1</tableCitation>
|
||
). The new species and the Volga
|
||
<taxonomicName id="4C41D5A7FFA5FF87DC6E2D5AF9E5FCF3" box="[786,977,368,392]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA5FF87DC6E2D5AF9E5FCF3" box="[786,977,368,392]" italics="true" pageId="8" pageNumber="427">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
are well discriminated by a complex of morphometric characters as can be seen from results of a DFA presented below.
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFA5FF87DFEB28C5FE59F858" httpUri="https://zenodo.org/record/243679/files/figure.png" pageId="8" pageNumber="427" targetBox="[341,1244,469,1236]" targetPageId="8">
|
||
<paragraph id="8BFEAE24FFA5FF87DFEB28C5FE59F858" blockId="8.[151,1436,1263,1315]" pageId="8" pageNumber="427">
|
||
<emphasis id="B9357236FFA5FF87DFEB28C5FB24F87F" bold="true" box="[151,272,1263,1285]" pageId="8" pageNumber="427">FIGURE 7</emphasis>
|
||
. Result of the DFA carried out on morphometric characters to discriminate
|
||
<taxonomicName id="4C41D5A7FFA5FF87DB4A28C5FECEF87E" box="[1078,1274,1263,1285]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA5FF87DB4A28C5FECEF87E" box="[1078,1274,1263,1285]" italics="true" pageId="8" pageNumber="427">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA5FF87DA7828C5FF6CF87F" box="[1284,1368,1263,1284]" pageId="8" pageNumber="427" rank="species">
|
||
<emphasis id="B9357236FFA5FF87DA7828C5FF6CF87F" bold="true" box="[1284,1368,1263,1284]" pageId="8" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(solid circles) and
|
||
<taxonomicName id="4C41D5A7FFA5FF87DE6F2924F833F858" authority="Volga" authorityName="Volga" box="[275,519,1293,1315]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA5FF87DE6F2924FB8AF858" box="[275,446,1294,1315]" italics="true" pageId="8" pageNumber="427">C. koshewnikowi</emphasis>
|
||
: Volga
|
||
</taxonomicName>
|
||
(circles), Northern Dvina (triangles), and Pechora (squares).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFA5FF87DFBB2966FEC9FA97" blockId="8.[151,1437,1355,2028]" pageId="8" pageNumber="427">
|
||
<collectingCountry id="F356EEB4FFA5FF87DFBB2966FAD2F81F" box="[199,230,1356,1380]" name="American Samoa" pageId="8" pageNumber="427">As</collectingCountry>
|
||
to sculpins from other rivers of the Arctic basin in Europe, an opinion was expressed that the sculpin from the Pechora River may represent a distinct species,
|
||
<taxonomicName id="4C41D5A7FFA5FF87DD84295AFE5AF8FC" authority="Soldatov, 1924" authorityName="Soldatov" authorityYear="1924" box="[760,1134,1391,1416]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="milvensis">
|
||
<emphasis id="B9357236FFA5FF87DD84295AF980F8FC" box="[760,948,1391,1415]" italics="true" pageId="8" pageNumber="427">Cottus milvensis</emphasis>
|
||
<bibRefCitation id="EFD0D3D5FFA5FF87DCC32945FE5AF8FC" author="Soldatov" box="[959,1134,1391,1416]" pageId="8" pageNumber="427" refString="Soldatov, V. K. (1924) Fishes of Pechora River. Proceedings of Northern Scientific Fishing Expedition, 17, 6, 57 - 60. (In Russian)" type="journal article" year="1924">Soldatov, 1924</bibRefCitation>
|
||
</taxonomicName>
|
||
, though no distinguishing characters were given (Kottelat & Freyhof 2007).
|
||
<bibRefCitation id="EFD0D3D5FFA5FF87DDA129B9F9A1F8D7" author="Soldatov" box="[733,917,1427,1452]" pageId="8" pageNumber="427" refString="Soldatov, V. K. (1924) Fishes of Pechora River. Proceedings of Northern Scientific Fishing Expedition, 17, 6, 57 - 60. (In Russian)" type="journal article" year="1924">Soldatov (1924)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C41D5A7FFA5FF87DB7729BEFEAAF8D0" box="[1035,1182,1427,1451]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gobio">
|
||
<emphasis id="B9357236FFA5FF87DB7729BEFEAAF8D0" box="[1035,1182,1427,1451]" italics="true" pageId="8" pageNumber="427">Cottus gobio</emphasis>
|
||
</taxonomicName>
|
||
in the Pechora River, while two juvenile specimens from the Mylva tributary were described by him as a new subspecies
|
||
<taxonomicName id="4C41D5A7FFA5FF87DA4B2992FACBF888" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="subSpecies" species="gobio" subSpecies="milvensis">
|
||
<emphasis id="B9357236FFA5FF87DA4B2992FACBF888" italics="true" pageId="8" pageNumber="427">C. gobio milvensis</emphasis>
|
||
</taxonomicName>
|
||
(whereabouts of the
|
||
<typeStatus id="54FA1086FFA5FF87DE9129F6F87BF88F" box="[493,591,1500,1524]" pageId="8" pageNumber="427" type="syntype">syntypes</typeStatus>
|
||
are unknown). These two specimens differed from the fish identified as
|
||
<taxonomicName id="4C41D5A7FFA5FF87DAFE29F6FB2BFB6C" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="subSpecies" species="gobio" subSpecies="gobio">
|
||
<emphasis id="B9357236FFA5FF87DAFE29F6FB2BFB6C" italics="true" pageId="8" pageNumber="427">C. gobio gobio</emphasis>
|
||
</taxonomicName>
|
||
by a complete lateral line, longer both dorsal and anal fins, and some departure in the prickling. All specimens we examined (
|
||
<quantity id="4CB903C1FFA5FF87DEC62A0EFBC1FB47" box="[442,501,1571,1596]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="8" pageNumber="427" unit="in" value="18.0">18 in</quantity>
|
||
total available at ZIN, see below in the list of material) were tentatively identified by us as
|
||
<emphasis id="B9357236FFA5FF87DE7F2A62FBF2FB24" box="[259,454,1607,1631]" italics="true" pageId="8" pageNumber="427">
|
||
<taxonomicName id="4C41D5A7FFA5FF87DE7F2A62FBF7FB24" box="[259,451,1607,1631]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikovi">C. koshewnikovi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Volga and Pechora specimens share most diagnostic characters (e.g., a smaller eye) mentioned above in the comparison between
|
||
<taxonomicName id="4C41D5A7FFA5FF87DDE32A46F97CFBF8" box="[671,840,1644,1667]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA5FF87DDE32A46F97CFBF8" box="[671,840,1644,1667]" italics="true" pageId="8" pageNumber="427">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA5FF87DC2E2A41F99AFBFF" box="[850,942,1643,1668]" pageId="8" pageNumber="427" rank="species">
|
||
<emphasis id="B9357236FFA5FF87DC2E2A41F99AFBFF" bold="true" box="[850,942,1643,1668]" pageId="8" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and Volga
|
||
<taxonomicName id="4C41D5A7FFA5FF87DB4F2A46FED8FBF8" box="[1075,1260,1643,1667]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikovi">
|
||
<emphasis id="B9357236FFA5FF87DB4F2A46FED8FBF8" box="[1075,1260,1643,1667]" italics="true" pageId="8" pageNumber="427">C. koshewnikovi</emphasis>
|
||
</taxonomicName>
|
||
as can be seen from
|
||
<tableCitation id="C6C39B9FFFA5FF87DFAB2AA5FB6CFBDC" box="[215,344,1679,1704]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="3.[125,207,513,533]" captionStartId-1="4.[151,239,836,859]" captionStartId-2="6.[151,239,667,690]" captionStartId-3="6.[151,239,1227,1250]" captionTargetBox-0="[138,1845,573,1680]" captionTargetBox-2="[159,1388,718,1058]" captionTargetPageId-0="3" captionTargetPageId-2="6" captionText-0="TABLE 1. Prοpοrtiοnal measurements οf Cottus gratzianowi sp. nov. and Cottus koshewnikowi." captionText-1="TABLE 2. Frequency of occurrence of fin-ray counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi; counts for holotype are with an asterisk." captionText-2="TABLE 3. Frequency of occurrence of lateral-line counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi." captionText-3="TABLE 4. Frequency of occurrence of vertebral counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri-0="http://table.plazi.org/id/DF3EFEACFFAEFF8CDF012E2BFE18FF6D" httpUri-2="http://table.plazi.org/id/DF3EFEACFFABFF89DFEB2EB1FF57FFC9" pageId="8" pageNumber="427" tableUuid-0="DF3EFEACFFAEFF8CDF012E2BFE18FF6D" tableUuid-2="DF3EFEACFFABFF89DFEB2EB1FF57FFC9">Tables 1−4</tableCitation>
|
||
. However, the Pechora specimens differ from the Volga specimens in having a slightly longer second dorsal fin, 16−19 rays vs. 17−19, though mode value, 18, is the same (
|
||
<tableCitation id="C6C39B9FFFA5FF87DB732A99FE53FBB7" box="[1039,1127,1715,1740]" captionStart="TABLE 2" captionStartId="4.[151,239,836,859]" captionText="TABLE 2. Frequency of occurrence of fin-ray counts in Cottus gratzianowi sp. nov. and Cottus koshewnikowi; counts for holotype are with an asterisk." pageId="8" pageNumber="427">Table 2</tableCitation>
|
||
), a markedly longer caudal peduncle (11−20 % SL)−longer than in
|
||
<taxonomicName id="4C41D5A7FFA5FF87DD2B2AF2F8CBFB94" box="[599,767,1752,1775]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA5FF87DD2B2AF2F8CBFB94" box="[599,767,1752,1775]" italics="true" pageId="8" pageNumber="427">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA5FF87DC7A2AFDF954FB8B" box="[774,864,1751,1776]" pageId="8" pageNumber="427" rank="species">
|
||
<emphasis id="B9357236FFA5FF87DC7A2AFDF954FB8B" bold="true" box="[774,864,1751,1776]" pageId="8" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and, accordingly, a longer caudal vertebral region, 22−24, modally 23. The number of total vertebrae, 33−35, modally 34, is higher in the Pechora specimens than in the Volga sample. This feature needs a further study based on a more numerous material. However, the number of abdominal vertebrae is
|
||
<quantity id="4CB903C1FFA5FF87DEE32B6EFBECFA27" box="[415,472,1859,1884]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="8" pageNumber="427" unit="in" value="11.0">11 in</quantity>
|
||
the Pechora sample (a character state typical of
|
||
<taxonomicName id="4C41D5A7FFA5FF87DC812B6EFE82FA20" box="[1021,1206,1859,1883]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikovi">
|
||
<emphasis id="B9357236FFA5FF87DC812B6EFE82FA20" box="[1021,1206,1859,1883]" italics="true" pageId="8" pageNumber="427">C. koshewnikovi</emphasis>
|
||
</taxonomicName>
|
||
) in contrast to
|
||
<quantity id="4CB903C1FFA5FF87DA1E2B6EFFA8FA27" box="[1378,1436,1859,1884]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="427" unit="in" value="10.0">10 in</quantity>
|
||
<taxonomicName id="4C41D5A7FFA5FF87DFEB2B42FB09FA04" box="[151,317,1896,1919]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA5FF87DFEB2B42FB09FA04" box="[151,317,1896,1919]" italics="true" pageId="8" pageNumber="427">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA5FF87DE3F2B4DFBA8FAFB" box="[323,412,1895,1920]" pageId="8" pageNumber="427" rank="species">
|
||
<emphasis id="B9357236FFA5FF87DE3F2B4DFBA8FAFB" bold="true" box="[323,412,1895,1920]" pageId="8" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<collectingCountry id="F356EEB4FFA5FF87DEDF2B42FBF6FAFB" box="[419,450,1896,1920]" name="American Samoa" pageId="8" pageNumber="427">As</collectingCountry>
|
||
to the Northern Dvina, a geographically close drainage in the White Sea basin, only five specimens were available for the examination including three old specimens in poor condition. The morphometric characters, fin-ray, lateral-line, and vertebral counts indicate to their probable conspecificity with the Pechora species we tentatively identify as
|
||
<taxonomicName id="4C41D5A7FFA5FF87DD6C2BFEF8F3FA90" box="[528,711,2003,2027]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="koshewnikovi">
|
||
<emphasis id="B9357236FFA5FF87DD6C2BFEF8F3FA90" box="[528,711,2003,2027]" italics="true" pageId="8" pageNumber="427">C. koshewnikovi</emphasis>
|
||
</taxonomicName>
|
||
, but one specimen had branched pectoral-fin rays.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA4FF86DFBB2CB2F8B4FC66" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="428">
|
||
<emphasis id="B9357236FFA4FF86DFBB2CB2F892FDCB" bold="true" box="[199,678,151,176]" pageId="9" pageNumber="428">
|
||
<emphasis id="B9357236FFA4FF86DFBB2CB2F879FDCB" bold="true" box="[199,589,151,176]" italics="true" pageId="9" pageNumber="428">Discriminant Function Analysis</emphasis>
|
||
(
|
||
<emphasis id="B9357236FFA4FF86DD1A2CB2F8A9FDCB" bold="true" box="[614,669,152,176]" italics="true" pageId="9" pageNumber="428">DFA</emphasis>
|
||
)
|
||
</emphasis>
|
||
<emphasis id="B9357236FFA4FF86DDCA2CB2FE37FDCB" box="[694,1027,152,176]" italics="true" pageId="9" pageNumber="428">of morphometric characters</emphasis>
|
||
was performed for 53 examined specimens (as in
|
||
<tableCitation id="C6C39B9FFFA4FF86DE182C96FB88FDAF" box="[356,444,188,213]" captionStart="TABLE 1" captionStartId="3.[125,207,513,533]" captionTargetBox="[138,1845,573,1680]" captionTargetPageId="3" captionText="TABLE 1. Prοpοrtiοnal measurements οf Cottus gratzianowi sp. nov. and Cottus koshewnikowi." httpUri="http://table.plazi.org/id/DF3EFEACFFAEFF8CDF012E2BFE18FF6D" pageId="9" pageNumber="428" tableUuid="DF3EFEACFFAEFF8CDF012E2BFE18FF6D">Table 1</tableCitation>
|
||
):
|
||
<specimenCount id="9D4765ADFFA4FF86DEA42C96F852FDAE" box="[472,614,188,213]" pageId="9" pageNumber="428" type="generic">9 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C41D5A7FFA4FF86DDE82C96F974FDAF" box="[660,832,188,212]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA4FF86DDE82C96F893FDA8" box="[660,679,188,211]" italics="true" pageId="9" pageNumber="428">C</emphasis>
|
||
.
|
||
<emphasis id="B9357236FFA4FF86DDC62C97F974FDAF" box="[698,832,189,212]" italics="true" pageId="9" pageNumber="428">gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA4FF86DC372C96F99DFDAE" box="[843,937,188,213]" pageId="9" pageNumber="428" rank="species">
|
||
<emphasis id="B9357236FFA4FF86DC372C96F99DFDAE" bold="true" box="[843,937,188,213]" pageId="9" pageNumber="428">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<typeStatus id="54FA1086FFA4FF86DCC02C96FE14FDAE" box="[956,1056,188,213]" pageId="9" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
and 8
|
||
<typeStatus id="54FA1086FFA4FF86DB052C96FEDDFDAF" box="[1145,1257,188,212]" pageId="9" pageNumber="428" type="paratype">paratypes</typeStatus>
|
||
),
|
||
<specimenCount id="9D4765ADFFA4FF86DA7C2C96FFA8FDAE" box="[1280,1436,188,213]" pageId="9" pageNumber="428" type="generic">21 specimens</specimenCount>
|
||
including
|
||
<typeStatus id="54FA1086FFA4FF86DE7B2CCAFB5DFD83" box="[263,361,224,248]" pageId="9" pageNumber="428" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C41D5A7FFA4FF86DEF22CCAF87FFD83" box="[398,587,224,248]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA4FF86DEF22CCAF87FFD83" box="[398,587,224,248]" italics="true" pageId="9" pageNumber="428">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
(Volga River drainage),
|
||
<specimenCount id="9D4765ADFFA4FF86DC232CF5F9DDFD83" box="[863,1001,223,248]" pageId="9" pageNumber="428" type="generic">5 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C41D5A7FFA4FF86DB712CCAFEFEFD83" box="[1037,1226,224,248]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA4FF86DB712CCAFEFEFD83" box="[1037,1226,224,248]" italics="true" pageId="9" pageNumber="428">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
from the Northern Dvina, and
|
||
<specimenCount id="9D4765ADFFA4FF86DE652D2EFB85FC66" box="[281,433,260,285]" pageId="9" pageNumber="428" type="generic">18 specimens</specimenCount>
|
||
from the Pechora.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA4FF86DFBB2D0DF816FFFF" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="428">
|
||
The DFA using morphometric measurements revealed significant morphological shape differences between the new species and the other samples (
|
||
<figureCitation id="137AB2A1FFA4FF86DD2C2D66F8A0FC1F" box="[592,660,332,357]" captionStart="FIGURE 7" captionStartId="8.[151,250,1263,1285]" captionTargetBox="[341,1244,469,1236]" captionTargetId="figure@8.[340,1247,463,1242]" captionTargetPageId="8" captionText="FIGURE 7. Result of the DFA carried out on morphometric characters to discriminate Cottus gratzianowi sp. nov. (solid circles) and C. koshewnikowi: Volga (circles), Northern Dvina (triangles), and Pechora (squares)." httpUri="https://zenodo.org/record/243679/files/figure.png" pageId="9" pageNumber="428">Fig. 7</figureCitation>
|
||
,
|
||
<tableCitation id="C6C39B9FFFA4FF86DDDC2D66F8C1FC1E" box="[672,757,332,357]" captionStart="TABLE 5" captionStartId="9.[151,239,832,855]" captionTargetBox="[159,1404,946,1495]" captionTargetPageId="9" captionText="TABLE 5. Statistical results of the DFA (Onega, Volga, Pechora and Northern Dvina samples, n = 53) based on morphometric characters. Characters are listed in order of increasing value of Partial Lambda (e. g. decreasing of particular character contribution to general discrimination)." httpUri="http://table.plazi.org/id/DF3EFEACFFA4FF86DFEB2F6AF924FEEC" pageId="9" pageNumber="428" tableUuid="DF3EFEACFFA4FF86DFEB2F6AF924FEEC">Table 5</tableCitation>
|
||
). This analysis correctly classified all individuals of the new species,
|
||
<taxonomicName id="4C41D5A7FFA4FF86DF852D5AFB97FCFC" box="[249,419,368,391]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA4FF86DF852D5AFB97FCFC" box="[249,419,368,391]" italics="true" pageId="9" pageNumber="428">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA4FF86DED12D45F83EFCF3" box="[429,522,367,392]" pageId="9" pageNumber="428" rank="species">
|
||
<emphasis id="B9357236FFA4FF86DED12D45F83EFCF3" bold="true" box="[429,522,367,392]" pageId="9" pageNumber="428">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, as well as the Volga sculpin (
|
||
<taxonomicName id="4C41D5A7FFA4FF86DC0A2D5AFE0CFCF3" box="[886,1080,368,392]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA4FF86DC0A2D5AFE0CFCF3" box="[886,1080,368,392]" italics="true" pageId="9" pageNumber="428">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
), and the combined sample of individuals from the Northern Dvina and the Pechora tentatively identified as
|
||
<taxonomicName id="4C41D5A7FFA4FF86DB6B2DBEFEE3FCD7" box="[1047,1239,404,428]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA4FF86DB6B2DBEFEE3FCD7" box="[1047,1239,404,428]" italics="true" pageId="9" pageNumber="428">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
. Statistics values are as follows: Wilks’ Lambda 0.00121, approx. F (69, 81) =9.9956, p<0.0000, which indicate almost perfect discrimination.
|
||
<taxonomicName id="4C41D5A7FFA4FF86DE362DF6F810FC8F" box="[330,548,476,500]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA4FF86DE362DF6F810FC8F" box="[330,548,476,500]" italics="true" pageId="9" pageNumber="428">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA4FF86DD522DF6F8B8FC8E" box="[558,652,476,501]" pageId="9" pageNumber="428" rank="species">
|
||
<emphasis id="B9357236FFA4FF86DD522DF6F8B8FC8E" bold="true" box="[558,652,476,501]" pageId="9" pageNumber="428">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from the Onega River drainage is the most distant from the Volga sculpin (Squared Mahalanobis Distance equal 116.5); the Pechora and the Northern Dvina samples are closer (72.4 and 96.1, respectively) but still significantly distant. The most removed, by the morphometric characters considered, are the Volga and the Northern Dvina samples (224.6) and the Volga and the Pechora (167.4). For all mentioned comparisons, p<0.0000.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA4FF86DFBB2EA5FBE1FE6E" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="428">
|
||
Partial Lambdas, demonstrating the unique contribution of the respective variable to the discriminatory power of the whole model, are given in
|
||
<tableCitation id="C6C39B9FFFA4FF86DD622E9EF842FFB6" box="[542,630,692,717]" captionStart="TABLE 5" captionStartId="9.[151,239,832,855]" captionTargetBox="[159,1404,946,1495]" captionTargetPageId="9" captionText="TABLE 5. Statistical results of the DFA (Onega, Volga, Pechora and Northern Dvina samples, n = 53) based on morphometric characters. Characters are listed in order of increasing value of Partial Lambda (e. g. decreasing of particular character contribution to general discrimination)." httpUri="http://table.plazi.org/id/DF3EFEACFFA4FF86DFEB2F6AF924FEEC" pageId="9" pageNumber="428" tableUuid="DF3EFEACFFA4FF86DFEB2F6AF924FEEC">Table 5</tableCitation>
|
||
. The most significant (Partial Lambda <0.7) for discrimination of the samples under consideration are length of caudal peduncle, head depth, snout length, head width, length of gill slit, and horizontal eye diameter.
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFA4FF86DFEB2F6AF924FEEC" ID-Table-UUID="DF3EFEACFFA4FF86DFEB2F6AF924FEEC" httpUri="http://table.plazi.org/id/DF3EFEACFFA4FF86DFEB2F6AF924FEEC" pageId="9" pageNumber="428" targetBox="[159,1404,946,1495]" targetIsTable="true" targetPageId="9">
|
||
<paragraph id="8BFEAE24FFA4FF86DFEB2F6AF924FEEC" blockId="9.[151,1435,831,919]" pageId="9" pageNumber="428">
|
||
<emphasis id="B9357236FFA4FF86DFEB2F6AFB25FE2C" bold="true" box="[151,273,832,855]" pageId="9" pageNumber="428">TABLE 5.</emphasis>
|
||
Statistical results of the DFA (Onega, Volga, Pechora and Northern Dvina samples, n=53) based on morphometric characters. Characters are listed in order of increasing value of Partial Lambda (e.g. decreasing of particular character contribution to general discrimination).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFA4FF86DFE32F98FF56F8AC" blockId="9.[159,1404,946,1950]" pageId="9" pageNumber="428">
|
||
<table id="F9415C84FFA40070DFE32F98FF48F8AC" box="[159,1404,946,1495]" gridcols="6" gridrows="5" pageId="9" pageNumber="428">
|
||
<tr id="3571AC66FFA40070DFE32F98FF48F96B" box="[159,1404,946,1040]" gridrow="0" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE32F98FBF9F96B" box="[159,461,946,1040]" gridcol="0" gridrow="0" pageId="9" pageNumber="428">Character Length of caudal peduncle</th>
|
||
<th id="76A0C51AFFA40070DD312F98F894F96B" box="[589,672,946,1040]" gridcol="1" gridrow="0" pageId="9" pageNumber="428">Wilks' Lambda 0.0029</th>
|
||
<th id="76A0C51AFFA40070DDA82F98F913F96B" box="[724,807,946,1040]" gridcol="2" gridrow="0" pageId="9" pageNumber="428">Partial Lambda 0.4154</th>
|
||
<th id="76A0C51AFFA40070DC1E2F98F9F0F96B" box="[866,964,946,1040]" gridcol="3" gridrow="0" pageId="9" pageNumber="428">F-remove (3.27) 12.6647</th>
|
||
<th id="76A0C51AFFA40070DB7A2F98FE78F96B" box="[1030,1100,946,1040]" gridcol="4" gridrow="0" pageId="9" pageNumber="428">p-level 0.0000</th>
|
||
<th id="76A0C51AFFA40070DBE82F98FF48F96B" box="[1172,1404,946,1040]" gridcol="5" gridrow="0" pageId="9" pageNumber="428">Tolerance R-Square 0.3598 0.6402</th>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE3280EFF48F9F7" box="[159,1404,1060,1164]" gridrow="1" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE3280EFBF9F9F7" box="[159,461,1060,1164]" gridcol="0" gridrow="1" pageId="9" pageNumber="428">Head depth Snout length Head width</th>
|
||
<td id="76A0C51AFFA40070DD31280EF894F9F7" box="[589,672,1060,1164]" gridcol="1" gridrow="1" pageId="9" pageNumber="428">0.0024 0.0020 0.0020</td>
|
||
<td id="76A0C51AFFA40070DDA8280EF913F9F7" box="[724,807,1060,1164]" gridcol="2" gridrow="1" pageId="9" pageNumber="428">0.5046 0.6069 0.6167</td>
|
||
<td id="76A0C51AFFA40070DC1E280EF9F0F9F7" box="[866,964,1060,1164]" gridcol="3" gridrow="1" pageId="9" pageNumber="428">8.8362 5.8283 5.5939</td>
|
||
<td id="76A0C51AFFA40070DB7A280EFE78F9F7" box="[1030,1100,1060,1164]" gridcol="4" gridrow="1" pageId="9" pageNumber="428">0.0003 0.0033 0.0041</td>
|
||
<td id="76A0C51AFFA40070DBE8280EFF48F9F7" box="[1172,1404,1060,1164]" gridcol="5" gridrow="1" pageId="9" pageNumber="428">0.2696 0.7304 0.3929 0.6071 0.2281 0.7719</td>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE3288AFF48F873" box="[159,1404,1184,1288]" gridrow="2" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE3288AFBF9F873" box="[159,461,1184,1288]" gridcol="0" gridrow="2" pageId="9" pageNumber="428">Length of gill slit Horizontal eye diameter Predorsal length</th>
|
||
<td id="76A0C51AFFA40070DD31288AF894F873" box="[589,672,1184,1288]" gridcol="1" gridrow="2" pageId="9" pageNumber="428">0.0019 0.0018 0.0017</td>
|
||
<td id="76A0C51AFFA40070DDA8288AF913F873" box="[724,807,1184,1288]" gridcol="2" gridrow="2" pageId="9" pageNumber="428">0.6551 0.6640 0.7270</td>
|
||
<td id="76A0C51AFFA40070DC1E288AF9F0F873" box="[866,964,1184,1288]" gridcol="3" gridrow="2" pageId="9" pageNumber="428">4.7379 4.5536 3.3803</td>
|
||
<td id="76A0C51AFFA40070DB7A288AFE78F873" box="[1030,1100,1184,1288]" gridcol="4" gridrow="2" pageId="9" pageNumber="428">0.0088 0.0105 0.0326</td>
|
||
<td id="76A0C51AFFA40070DBE8288AFF48F873" box="[1172,1404,1184,1288]" gridcol="5" gridrow="2" pageId="9" pageNumber="428">0.3634 0.6366 0.3853 0.6147 0.2514 0.7486</td>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE32936FF48F8FF" box="[159,1404,1308,1412]" gridrow="3" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE32936FBF9F8FF" box="[159,461,1308,1412]" gridcol="0" gridrow="3" pageId="9" pageNumber="428">Head length Pelvic fin length Length of first dorsal-fin base</th>
|
||
<td id="76A0C51AFFA40070DD312936F894F8FF" box="[589,672,1308,1412]" gridcol="1" gridrow="3" pageId="9" pageNumber="428">0.0016 0.0016 0.0015</td>
|
||
<td id="76A0C51AFFA40070DDA82936F913F8FF" box="[724,807,1308,1412]" gridcol="2" gridrow="3" pageId="9" pageNumber="428">0.7508 0.7755 0.7999</td>
|
||
<td id="76A0C51AFFA40070DC1E2936F9F0F8FF" box="[866,964,1308,1412]" gridcol="3" gridrow="3" pageId="9" pageNumber="428">2.9872 2.6060 2.2518</td>
|
||
<td id="76A0C51AFFA40070DB7A2936FE78F8FF" box="[1030,1100,1308,1412]" gridcol="4" gridrow="3" pageId="9" pageNumber="428">0.0486 0.0723 0.1051</td>
|
||
<td id="76A0C51AFFA40070DBE82936FF48F8FF" box="[1172,1404,1308,1412]" gridcol="5" gridrow="3" pageId="9" pageNumber="428">0.1858 0.8142 0.2136 0.7865 0.3291 0.6709</td>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE329B2FF48F8AC" box="[159,1404,1432,1495]" gridrow="4" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE329B2FBF9F8AC" box="[159,461,1432,1495]" gridcol="0" gridrow="4" pageId="9" pageNumber="428">Interorbital width Length of anal-fin base</th>
|
||
<td id="76A0C51AFFA40070DD3129B2F894F8AC" box="[589,672,1432,1495]" gridcol="1" gridrow="4" pageId="9" pageNumber="428">0.0015 0.0015</td>
|
||
<td id="76A0C51AFFA40070DDA829B2F913F8AC" box="[724,807,1432,1495]" gridcol="2" gridrow="4" pageId="9" pageNumber="428">0.8044 0.8067</td>
|
||
<td id="76A0C51AFFA40070DC1E29B2F9F0F8AC" box="[866,964,1432,1495]" gridcol="3" gridrow="4" pageId="9" pageNumber="428">2.1883 2.1567</td>
|
||
<td id="76A0C51AFFA40070DB7A29B2FE78F8AC" box="[1030,1100,1432,1495]" gridcol="4" gridrow="4" pageId="9" pageNumber="428">0.1125 0.1164</td>
|
||
<td id="76A0C51AFFA40070DBE829B2FF48F8AC" box="[1172,1404,1432,1495]" gridcol="5" gridrow="4" pageId="9" pageNumber="428">0.4320 0.5680 0.3663 0.6337</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA4FF86DFE329C0FF56FAE5" blockId="9.[159,1404,946,1950]" pageId="9" pageNumber="428">
|
||
<table id="F9415C84FFA40070DFE329C0FF48FAE5" box="[159,1404,1514,1950]" gridcols="6" gridrows="5" pageId="9" pageNumber="428">
|
||
<tr id="3571AC66FFA40070DFE329C0FF48FB7B" box="[159,1404,1514,1536]" gridrow="0" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE329C0F8A7FB7B" box="[159,659,1514,1536]" gridcol="0" gridrow="0" pageId="9" pageNumber="428">Length of longest second dorsal-fin ray 0.0015</th>
|
||
<th id="76A0C51AFFA40070DDA829C0F92EFB7B" box="[724,794,1514,1536]" gridcol="1" gridrow="0" pageId="9" pageNumber="428">0.8261</th>
|
||
<th id="76A0C51AFFA40070DC1E29C0F99CFB7B" box="[866,936,1514,1536]" gridcol="2" gridrow="0" pageId="9" pageNumber="428">1.8942</th>
|
||
<th id="76A0C51AFFA40070DB7A29C0FE78FB7B" box="[1030,1100,1514,1536]" gridcol="3" gridrow="0" pageId="9" pageNumber="428">0.1544</th>
|
||
<th id="76A0C51AFFA40070DBE829C0FEEEFB7B" box="[1172,1242,1514,1536]" gridcol="4" gridrow="0" pageId="9" pageNumber="428">0.4893</th>
|
||
<th id="76A0C51AFFA40070DA6029C0FF48FB7B" box="[1308,1404,1514,1536]" gridcol="5" gridrow="0" pageId="9" pageNumber="428">0.5107</th>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE32A3EFF48FB07" box="[159,1404,1556,1660]" gridrow="1" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE32A3EF8A7FB07" box="[159,659,1556,1660]" gridcol="0" gridrow="1" pageId="9" pageNumber="428">
|
||
Interbranchial width 0.0015 Length of second dorsal-fin base 0.0014 Preanal length (
|
||
<emphasis id="B9357236FFA4FF86DE432A4DFB6FFB07" box="[319,347,1639,1660]" italics="true" pageId="9" pageNumber="428">aA</emphasis>
|
||
) 0.0014
|
||
</th>
|
||
<td id="76A0C51AFFA40070DDA82A3EF92EFB07" box="[724,794,1556,1660]" gridcol="1" gridrow="1" pageId="9" pageNumber="428">0.8287 0.8453 0.8497</td>
|
||
<td id="76A0C51AFFA40070DC1E2A3EF99CFB07" box="[866,936,1556,1660]" gridcol="2" gridrow="1" pageId="9" pageNumber="428">1.8599 1.6475 1.5916</td>
|
||
<td id="76A0C51AFFA40070DB7A2A3EFE78FB07" box="[1030,1100,1556,1660]" gridcol="3" gridrow="1" pageId="9" pageNumber="428">0.1602 0.2018 0.2145</td>
|
||
<td id="76A0C51AFFA40070DBE82A3EFEEEFB07" box="[1172,1242,1556,1660]" gridcol="4" gridrow="1" pageId="9" pageNumber="428">0.3954 0.3516 0.3582</td>
|
||
<td id="76A0C51AFFA40070DA602A3EFF48FB07" box="[1308,1404,1556,1660]" gridcol="5" gridrow="1" pageId="9" pageNumber="428">0.6046 0.6484 0.6418</td>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE32ABAFF48FB83" box="[159,1404,1680,1784]" gridrow="2" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE32ABAF8A7FB83" box="[159,659,1680,1784]" gridcol="0" gridrow="2" pageId="9" pageNumber="428">Length of longest first dorsal-fin ray 0.0014 Depth of caudal peduncle 0.0014 Upper jaw length 0.0014</th>
|
||
<td id="76A0C51AFFA40070DDA82ABAF92EFB83" box="[724,794,1680,1784]" gridcol="1" gridrow="2" pageId="9" pageNumber="428">0.8719 0.8777 0.8802</td>
|
||
<td id="76A0C51AFFA40070DC1E2ABAF99CFB83" box="[866,936,1680,1784]" gridcol="2" gridrow="2" pageId="9" pageNumber="428">1.3223 1.2546 1.2252</td>
|
||
<td id="76A0C51AFFA40070DB7A2ABAFE78FB83" box="[1030,1100,1680,1784]" gridcol="3" gridrow="2" pageId="9" pageNumber="428">0.2877 0.3096 0.3197</td>
|
||
<td id="76A0C51AFFA40070DBE82ABAFEEEFB83" box="[1172,1242,1680,1784]" gridcol="4" gridrow="2" pageId="9" pageNumber="428">0.4410 0.2901 0.4721</td>
|
||
<td id="76A0C51AFFA40070DA602ABAFF48FB83" box="[1308,1404,1680,1784]" gridcol="5" gridrow="2" pageId="9" pageNumber="428">0.5590 0.7099 0.5279</td>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE32B26FF48FA0F" box="[159,1404,1804,1908]" gridrow="3" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE32B26F8A7FA0F" box="[159,659,1804,1908]" gridcol="0" gridrow="3" pageId="9" pageNumber="428">Pectoral fin length 0.0014 Length of longest anal-fin ray 0.0014 Maximum body depth 0.0013</th>
|
||
<td id="76A0C51AFFA40070DDA82B26F92EFA0F" box="[724,794,1804,1908]" gridcol="1" gridrow="3" pageId="9" pageNumber="428">0.8961 0.9000 0.9104</td>
|
||
<td id="76A0C51AFFA40070DC1E2B26F99CFA0F" box="[866,936,1804,1908]" gridcol="2" gridrow="3" pageId="9" pageNumber="428">1.0432 0.9998 0.8855</td>
|
||
<td id="76A0C51AFFA40070DB7A2B26FE78FA0F" box="[1030,1100,1804,1908]" gridcol="3" gridrow="3" pageId="9" pageNumber="428">0.3894 0.4080 0.4611</td>
|
||
<td id="76A0C51AFFA40070DBE82B26FEEEFA0F" box="[1172,1242,1804,1908]" gridcol="4" gridrow="3" pageId="9" pageNumber="428">0.2535 0.4021 0.4004</td>
|
||
<td id="76A0C51AFFA40070DA602B26FF48FA0F" box="[1308,1404,1804,1908]" gridcol="5" gridrow="3" pageId="9" pageNumber="428">0.7465 0.5979 0.5996</td>
|
||
</tr>
|
||
<tr id="3571AC66FFA40070DFE32BA2FF48FAE5" box="[159,1404,1928,1950]" gridrow="4" pageId="9" pageNumber="428">
|
||
<th id="76A0C51AFFA40070DFE32BA2F8A7FAE5" box="[159,659,1928,1950]" gridcol="0" gridrow="4" pageId="9" pageNumber="428">Postorbital length 0.0013</th>
|
||
<td id="76A0C51AFFA40070DDA82BA2F92EFAE5" box="[724,794,1928,1950]" gridcol="1" gridrow="4" pageId="9" pageNumber="428">0.9112</td>
|
||
<td id="76A0C51AFFA40070DC1E2BA2F99CFAE5" box="[866,936,1928,1950]" gridcol="2" gridrow="4" pageId="9" pageNumber="428">0.8770</td>
|
||
<td id="76A0C51AFFA40070DB7A2BA2FE78FAE5" box="[1030,1100,1928,1950]" gridcol="3" gridrow="4" pageId="9" pageNumber="428">0.4653</td>
|
||
<td id="76A0C51AFFA40070DBE82BA2FEEEFAE5" box="[1172,1242,1928,1950]" gridcol="4" gridrow="4" pageId="9" pageNumber="428">0.4598</td>
|
||
<td id="76A0C51AFFA40070DA602BA2FF48FAE5" box="[1308,1404,1928,1950]" gridcol="5" gridrow="4" pageId="9" pageNumber="428">0.5402</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF3EFEACFFA7FF85DFEB284BF850F9ED" httpUri="https://zenodo.org/record/243680/files/figure.png" pageId="10" pageNumber="429" targetBox="[368,1218,193,1099]" targetPageId="10">
|
||
<paragraph id="8BFEAE24FFA7FF85DFEB284BF850F9ED" blockId="10.[151,1436,1121,1174]" pageId="10" pageNumber="429">
|
||
<emphasis id="B9357236FFA7FF85DFEB284BFB26F90D" bold="true" box="[151,274,1121,1143]" pageId="10" pageNumber="429">FIGURE 8.</emphasis>
|
||
Map showing the type locality of
|
||
<taxonomicName id="4C41D5A7FFA7FF85DD172848F918F90C" box="[619,812,1122,1143]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA7FF85DD172848F918F90C" box="[619,812,1122,1143]" italics="true" pageId="10" pageNumber="429">Cottus gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA7FF85DC4D2848F9B5F90C" box="[817,897,1122,1143]" pageId="10" pageNumber="429" rank="species">
|
||
<emphasis id="B9357236FFA7FF85DC4D2848F9B5F90C" bold="true" box="[817,897,1122,1143]" pageId="10" pageNumber="429">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the Ukhtomitsa River, tributary of Modlona River, Lake Vozhe basin in the Onega River system.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFEAE24FFA7FF85DFBB28E8FB9FF858" blockId="10.[151,1437,1218,2038]" pageId="10" pageNumber="429">
|
||
Thus, the Pechora and the Northern Dvina combined sample is well discriminated by a complex of morphometric characters from both
|
||
<taxonomicName id="4C41D5A7FFA7FF85DD5528CDF8D2F984" box="[553,742,1255,1279]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA7FF85DD5528CDF8D2F984" box="[553,742,1255,1279]" italics="true" pageId="10" pageNumber="429">C. koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
from the Volga and
|
||
<taxonomicName id="4C41D5A7FFA7FF85DCB428CDFE5AF985" box="[968,1134,1255,1278]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="gratzianowi" status="sp. nov.">
|
||
<emphasis id="B9357236FFA7FF85DCB428CDFE5AF985" box="[968,1134,1255,1278]" italics="true" pageId="10" pageNumber="429">C. gratzianowi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A206CF4DFFA7FF85DB0928CCFEFAF984" box="[1141,1230,1254,1279]" pageId="10" pageNumber="429" rank="species">
|
||
<emphasis id="B9357236FFA7FF85DB0928CCFEFAF984" bold="true" box="[1141,1230,1254,1279]" pageId="10" pageNumber="429">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A further study of these sculpins is needed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35BFDAFFFA7FF84DFBB2904FE11FCAB" lastPageId="11" lastPageNumber="430" pageId="10" pageNumber="429" type="materials_examined">
|
||
<paragraph id="8BFEAE24FFA7FF85DFBB2904FE99FB5B" blockId="10.[151,1437,1218,2038]" pageId="10" pageNumber="429">
|
||
<emphasis id="B9357236FFA7FF85DFBB2904FBEEF83C" bold="true" box="[199,474,1326,1351]" pageId="10" pageNumber="429">Comparative material.</emphasis>
|
||
<taxonomicName id="4C41D5A7FFA7FF85DE982905F8E0F83C" box="[484,724,1327,1351]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="koshewnikowi">
|
||
<emphasis id="B9357236FFA7FF85DE982905F8E0F83C" box="[484,724,1327,1351]" italics="true" pageId="10" pageNumber="429">Cottus koshewnikowi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9357236FFA7FF85DD992905F988F83C" box="[741,956,1327,1351]" italics="true" pageId="10" pageNumber="429">Caspian Sea basin</emphasis>
|
||
:
|
||
<typeStatus id="54FA1086FFA7FF85DCB22905FE0DF83C" box="[974,1081,1327,1351]" pageId="10" pageNumber="429" type="syntype">Syntypes</typeStatus>
|
||
:
|
||
<collectionCode id="ED5036E1FFA7FF85DB3A291AFEA9F83C" box="[1094,1181,1328,1351]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="429">ZMMU</collectionCode>
|
||
Р-2001 (4, SL
|
||
<quantity id="4CB903C1FFA7FF85DA2A2904FAF3F810" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.3" metricValueMin="4.7" pageId="10" pageNumber="429" unit="mm" value="50.0" valueMax="53.0" valueMin="47.0">47–53 mm</quantity>
|
||
), Goredva River at village of Ligachevo;
|
||
<collectionCode id="ED5036E1FFA7FF85DDDC2979F8C3F811" box="[672,759,1363,1386]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="429">ZMMU</collectionCode>
|
||
Р-2658 (20, SL
|
||
<quantity id="4CB903C1FFA7FF85DCC92978FE07F810" box="[949,1075,1362,1387]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.6" metricValueMin="4.4" pageId="10" pageNumber="429" unit="mm" value="60.0" valueMax="76.0" valueMin="44.0">44–76 mm</quantity>
|
||
), same locality;
|
||
<collectionCode id="ED5036E1FFA7FF85DB902979FF77F811" box="[1260,1347,1363,1386]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="429">ZMMU</collectionCode>
|
||
<collectionCode id="ED5036E1FFA7FF85DA372978FF50F810" box="[1355,1380,1362,1387]" pageId="10" pageNumber="429">P-</collectionCode>
|
||
2688 (3, SL
|
||
<quantity id="4CB903C1FFA7FF85DF95295CFB5EF8F4" box="[233,362,1398,1423]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" metricValueMax="7.1" metricValueMin="4.6" pageId="10" pageNumber="429" unit="mm" value="58.5" valueMax="71.0" valueMin="46.0">46−71 mm</quantity>
|
||
), same locality;
|
||
<collectionCode id="ED5036E1FFA7FF85DD5B2952F84AF8F4" box="[551,638,1400,1423]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="429">ZMMU</collectionCode>
|
||
Р-3118 (1, SL
|
||
<quantity id="4CB903C1FFA7FF85DC4F295CF9BCF8F4" box="[819,904,1398,1423]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="429" unit="mm" value="56.0">56 mm</quantity>
|
||
), same locality. Non-type material:
|
||
<collectionCode id="ED5036E1FFA7FF85DA5F2952FF4EF8F4" box="[1315,1402,1400,1423]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="429">ZMMU</collectionCode>
|
||
Р- 3912 (1, SL
|
||
<quantity id="4CB903C1FFA7FF85DE5729B1FB49F8C8" box="[299,381,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="429" unit="mm" value="40.0">40 mm</quantity>
|
||
) Seliger Lake;
|
||
<collectionCode id="ED5036E1FFA7FF85DD4D29B1F86BF8C9" box="[561,607,1435,1458]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
15540 (10, SL
|
||
<quantity id="4CB903C1FFA7FF85DC6629B0F9A1F8C8" box="[794,917,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" metricValueMax="8.4" metricValueMin="4.5" pageId="10" pageNumber="429" unit="mm" value="64.5" valueMax="84.0" valueMin="45.0">45-84 mm</quantity>
|
||
), Kos’va River (left tributary of Kama);
|
||
<collectionCode id="ED5036E1FFA7FF85DA1129B1FFA8F8C9" box="[1389,1436,1435,1458]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
15777 (4, SL
|
||
<quantity id="4CB903C1FFA7FF85DE532994FB9FF8AC" box="[303,427,1470,1495]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="7.8" metricValueMin="6.2" pageId="10" pageNumber="429" unit="mm" value="70.0" valueMax="78.0" valueMin="62.0">62−78 mm</quantity>
|
||
) Kama River at village of Ust-Divia;
|
||
<collectionCode id="ED5036E1FFA7FF85DC2C29EAF94BF8AC" box="[848,895,1472,1495]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
55581 (60, SL
|
||
<quantity id="4CB903C1FFA7FF85DB502994FE9DF8AC" box="[1068,1193,1470,1495]" metricMagnitude="-2" metricUnit="m" metricValue="7.45" metricValueMax="9.8" metricValueMin="5.1" pageId="10" pageNumber="429" unit="mm" value="74.5" valueMax="98.0" valueMin="51.0">51–98 mm</quantity>
|
||
), Kama River at town of Perm';
|
||
<collectionCode id="ED5036E1FFA7FF85DE7B29C9FB02F881" box="[263,310,1507,1530]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
55582 (11, SL
|
||
<quantity id="4CB903C1FFA7FF85DE9629C8F85DF880" box="[490,617,1506,1531]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.6" metricValueMin="3.9" pageId="10" pageNumber="429" unit="mm" value="52.5" valueMax="66.0" valueMin="39.0">39–66 mm</quantity>
|
||
), Oka River at town of Kaluga;
|
||
<collectionCode id="ED5036E1FFA7FF85DCA629C9FE3CF881" box="[986,1032,1507,1530]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
55583 (29, SL
|
||
<quantity id="4CB903C1FFA7FF85DBC229C8FF65F880" box="[1214,1361,1506,1531]" metricMagnitude="-2" metricUnit="m" metricValue="7.175" metricValueMax="9.5" metricValueMin="4.85" pageId="10" pageNumber="429" unit="mm" value="71.75" valueMax="95.0" valueMin="48.5">48.5–95 mm</quantity>
|
||
), Kara River (Kama system);
|
||
<collectionCode id="ED5036E1FFA7FF85DEE92A22FBF7FB64" box="[405,451,1544,1567]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
55685 (5, SL
|
||
<quantity id="4CB903C1FFA7FF85DD192A2CF8D0FB64" box="[613,740,1542,1567]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" metricValueMax="9.7" metricValueMin="4.5" pageId="10" pageNumber="429" unit="mm" value="71.0" valueMax="97.0" valueMin="45.0">45−97 mm</quantity>
|
||
), Kolosleyka River, Kama River system.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA7FF85DFBB2A01FE6AFA18" blockId="10.[151,1437,1218,2038]" pageId="10" pageNumber="429">
|
||
<emphasis id="B9357236FFA7FF85DFBB2A01FBB4FB38" box="[199,384,1579,1603]" italics="true" pageId="10" pageNumber="429">Baltic Sea basin</emphasis>
|
||
:
|
||
<collectionCode id="ED5036E1FFA7FF85DEEE2A01FBF5FB39" box="[402,449,1579,1602]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
8568 (5, SL
|
||
<quantity id="4CB903C1FFA7FF85DD272A00F935FB38" box="[603,769,1578,1603]" metricMagnitude="-2" metricUnit="m" metricValue="6.68" metricValueMax="7.279999999999999" metricValueMin="6.08" pageId="10" pageNumber="429" unit="mm" value="66.8" valueMax="72.8" valueMin="60.8">60.8–72.8 mm</quantity>
|
||
) Neva River at St Petersburg;
|
||
<collectionCode id="ED5036E1FFA7FF85DB1E2A01FEA5FB39" box="[1122,1169,1579,1602]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
1909 (4, SL 54.7–68.0 mm) Neva River;
|
||
<collectionCode id="ED5036E1FFA7FF85DE142A7AFBA2FB1C" box="[360,406,1616,1639]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
3718 (1, SL
|
||
<quantity id="4CB903C1FFA7FF85DD4F2A64F8E8FB1C" box="[563,732,1614,1639]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.65" metricValueMin="3.45" pageId="10" pageNumber="429" unit="mm" value="45.5" valueMax="56.5" valueMin="34.5">34.5–56.5 mm</quantity>
|
||
), Neva River;
|
||
<collectionCode id="ED5036E1FFA7FF85DCF82A7AF987FB1C" box="[900,947,1616,1639]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
14439 (5, SL 57.8–82.0 mm) Neva River, Ostrovki;
|
||
<collectionCode id="ED5036E1FFA7FF85DE712A59FB0FFBF1" box="[269,315,1651,1674]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
14493 (2, SL 60.8–100.0 mm), same locality;
|
||
<collectionCode id="ED5036E1FFA7FF85DC0F2A59F996FBF1" box="[883,930,1651,1674]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
23480 (5, SL
|
||
<quantity id="4CB903C1FFA7FF85DB252A58FF36FBF0" box="[1113,1282,1650,1675]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.05" metricValueMin="1.95" pageId="10" pageNumber="429" unit="mm" value="25.0" valueMax="30.5" valueMin="19.5">19.5–30.5 mm</quantity>
|
||
) Neva River downstream of Mga River mouth;
|
||
<collectionCode id="ED5036E1FFA7FF85DD662AB2F87DFBD4" box="[538,585,1688,1711]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
36171 (3, SL 37.0–
|
||
<quantity id="4CB903C1FFA7FF85DC572ABCF94FFBD4" box="[811,891,1686,1711]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="429" unit="mm" value="51.0">51 mm</quantity>
|
||
), Neva River at St Petersburg;
|
||
<collectionCode id="ED5036E1FFA7FF85DBA82AB2FF37FBD4" box="[1236,1283,1688,1711]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
40739 (2, SL 40.5–41.0 mm), Ladoga Lake;
|
||
<collectionCode id="ED5036E1FFA7FF85DE8D2A91F814FBA9" box="[497,544,1723,1746]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
41251 (2, SL 61.0–73.0 mm), Neva River at Ostrovky;
|
||
<collectionCode id="ED5036E1FFA7FF85DBF32A91FE8AFBA9" box="[1167,1214,1723,1746]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
41253 (2, SL
|
||
<quantity id="4CB903C1FFA7FF85DA212A90FB35FB8C" metricMagnitude="-2" metricUnit="m" metricValue="6.85" metricValueMax="6.95" metricValueMin="6.75" pageId="10" pageNumber="429" unit="mm" value="68.5" valueMax="69.5" valueMin="67.5">67.5– 69.5 mm</quantity>
|
||
), Neva River at St Petersburg;
|
||
<collectionCode id="ED5036E1FFA7FF85DD142ACAF8A2FB8C" box="[616,662,1760,1783]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
51904 (6, SL 41.0–
|
||
<quantity id="4CB903C1FFA7FF85DCFD2AF4F9DEFB8C" box="[897,1002,1758,1783]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" pageId="10" pageNumber="429" unit="mm" value="51.5">51.5 mm</quantity>
|
||
), Ladoga Lake,
|
||
<quantity id="4CB903C1FFA7FF85DBD92AF4FED4FB8C" box="[1189,1248,1758,1783]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="429" unit="km" value="5.0">5 кm</quantity>
|
||
from Kraskovo, depth
|
||
<quantity id="4CB903C1FFA7FF85DFA12B28FB16FA61" box="[221,290,1794,1819]" metricMagnitude="0" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="429" unit="m" value="4.5">4.5 m</quantity>
|
||
;
|
||
<collectionCode id="ED5036E1FFA7FF85DE522B29FB69FA61" box="[302,349,1795,1818]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
52249 (1, SL 78.0 mm), Lute River at Sikovitsy, Pskov Province;
|
||
<collectionCode id="ED5036E1FFA7FF85DB2B2B29FEB2FA61" box="[1111,1158,1795,1818]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
52861 (2, SL 40.0–52.0 mm), Ladoga Lake;
|
||
<collectionCode id="ED5036E1FFA7FF85DE072B02FB9DFA44" box="[379,425,1832,1855]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
54243 (5, SL 66.2–73.0 mm), Neva River delta;
|
||
<collectionCode id="ED5036E1FFA7FF85DCB22B02F9C9FA44" box="[974,1021,1832,1855]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
54247 (1, SL 71.0 mm), Neva River, depth
|
||
<quantity id="4CB903C1FFA7FF85DFA02B60FB76FA19" box="[220,322,1866,1891]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="10" pageNumber="429" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">15–20 m</quantity>
|
||
;
|
||
<collectionCode id="ED5036E1FFA7FF85DE312B61FB48FA19" box="[333,380,1867,1890]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
55686 (75, SL 48.0–
|
||
<quantity id="4CB903C1FFA7FF85DD172B60F8E6FA18" box="[619,722,1866,1891]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" pageId="10" pageNumber="429" unit="mm" value="57.5">57.5 mm</quantity>
|
||
), Izhora River at town of Gatchina.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA7FF85DFBB2B5AFB08FA8D" blockId="10.[151,1437,1218,2038]" pageId="10" pageNumber="429">
|
||
<emphasis id="B9357236FFA7FF85DFBB2B5AFBA4FAFC" box="[199,400,1903,1927]" italics="true" pageId="10" pageNumber="429">Barentz Sea basin</emphasis>
|
||
:
|
||
<collectionCode id="ED5036E1FFA7FF85DEE22B5AFBF8FAFC" box="[414,460,1904,1927]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
11331 (7, SL
|
||
<quantity id="4CB903C1FFA7FF85DD102B44F927FAFC" box="[620,787,1902,1927]" metricMagnitude="-2" metricUnit="m" metricValue="7.284999999999999" metricValueMax="8.85" metricValueMin="5.72" pageId="10" pageNumber="429" unit="mm" value="72.85" valueMax="88.5" valueMin="57.2">57.2–88.5 mm</quantity>
|
||
), Pechora River;
|
||
<collectionCode id="ED5036E1FFA7FF85DCA82B5AFE37FAFC" box="[980,1027,1904,1927]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
37102 (2, SL 52.0–58.0 mm), upper Pechora River, 62о N,
|
||
<geoCoordinate id="EE75C8E3FFA7FF85DEF62BBFFBD3FAD5" box="[394,487,1938,1966]" direction="east" orientation="longitude" pageId="10" pageNumber="429" precision="925" value="58.733334">58о44’E</geoCoordinate>
|
||
;
|
||
<collectionCode id="ED5036E1FFA7FF85DE8F2BBCF816FAD6" box="[499,546,1942,1965]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
41252 (1, SL
|
||
<quantity id="4CB903C1FFA7FF85DDBF2BBCF922FAD5" box="[707,790,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="429" unit="mm" value="70.0">70 mm</quantity>
|
||
), Pechora River;
|
||
<collectionCode id="ED5036E1FFA7FF85DCA42BBCFE32FAD6" box="[984,1030,1942,1965]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
50221 (3, SL
|
||
<quantity id="4CB903C1FFA7FF85DBD42BBCFF13FAD5" box="[1192,1319,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" metricValueMax="7.9" metricValueMin="7.7" pageId="10" pageNumber="429" unit="mm" value="78.0" valueMax="79.0" valueMin="77.0">77−79 mm</quantity>
|
||
), Ydzhyd- Lyaga River, tributary of Ilych, Pechora drainage;
|
||
<collectionCode id="ED5036E1FFA7FF85DD9B2B90F921FAAA" box="[743,789,1978,2001]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="429">ZIN</collectionCode>
|
||
55687 (6, SL
|
||
<quantity id="4CB903C1FFA7FF85DCBB2B93FE7DFAA9" box="[967,1097,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="6.85" metricValueMax="7.4" metricValueMin="6.3" pageId="10" pageNumber="429" unit="mm" value="68.5" valueMax="74.0" valueMin="63.0">63−74 mm</quantity>
|
||
), Koshym River, tributary of Pechora River.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA6FF84DFBB2CB2F81FFD83" blockId="11.[151,1436,151,464]" pageId="11" pageNumber="430">
|
||
<emphasis id="B9357236FFA6FF84DFBB2CB2FB4AFDCB" box="[199,382,152,176]" italics="true" pageId="11" pageNumber="430">White Sea basin</emphasis>
|
||
:
|
||
<collectionCode id="ED5036E1FFA6FF84DEF12CB2FB88FDD4" box="[397,444,152,175]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
41256 (1, SL 77.0 mm), Northern Dvina River at town of Arkhangelsk;
|
||
<collectionCode id="ED5036E1FFA6FF84DB852CB2FF1CFDD4" box="[1273,1320,152,175]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
52124 (2, SL 75, 74.6), Yavzora River, tributary of Pinega, Northern Dvina drainage;
|
||
<collectionCode id="ED5036E1FFA6FF84DC9D2C97FE24FDAF" box="[993,1040,189,212]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
55688 (2, SL 63, 71 mm), Dvinitsa River, Northern Dvina river system.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA6FF84DFBB2D2EFB91FC3B" blockId="11.[151,1436,151,464]" pageId="11" pageNumber="430">
|
||
<emphasis id="B9357236FFA6FF84DFBB2D2EFB6AFC67" box="[199,350,260,284]" italics="true" pageId="11" pageNumber="430">
|
||
<taxonomicName id="4C41D5A7FFA6FF84DFBB2D2EFB6EFC67" box="[199,346,260,284]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="430" phylum="Chordata" rank="species" species="gobio">Cottus gobio</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED5036E1FFA6FF84DE182D2FFBA6FC67" box="[356,402,261,284]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
13510 (1, SL
|
||
<quantity id="4CB903C1FFA6FF84DD4F2D2EF8AFFC67" box="[563,667,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" pageId="11" pageNumber="430" unit="mm" value="58.5">58.5 mm</quantity>
|
||
), Sillamae;
|
||
<collectionCode id="ED5036E1FFA6FF84DC622D2FF979FC67" box="[798,845,261,284]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
41255 (1, SL 62.0 mm), Baltic Sea, Svybiriken Bay,
|
||
<collectingCountry id="F356EEB4FFA6FF84DFEB2D0DFB02FC3B" box="[151,310,295,320]" name="Aland Islands" pageId="11" pageNumber="430">Aland Islands</collectingCountry>
|
||
: Gradeso.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA6FF84DFBB2D66F8C8FCD6" blockId="11.[151,1436,151,464]" pageId="11" pageNumber="430">
|
||
<taxonomicName id="4C41D5A7FFA6FF84DFBB2D66FBA4FC1F" box="[199,400,332,356]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="430" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B9357236FFA6FF84DFBB2D66FBA4FC1F" box="[199,400,332,356]" italics="true" pageId="11" pageNumber="430">Cottus poecilopus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<typeStatus id="54FA1086FFA6FF84DEE32D66F83EFC1F" box="[415,522,332,356]" pageId="11" pageNumber="430" type="syntype">Syntypes</typeStatus>
|
||
:
|
||
<collectionCode id="ED5036E1FFA6FF84DD692D67F868FC1F" box="[533,604,333,356]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430">NMW</collectionCode>
|
||
78816 (1, SL
|
||
<quantity id="4CB903C1FFA6FF84DC7C2D66F952FC1F" box="[768,870,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="8.45" pageId="11" pageNumber="430" unit="mm" value="84.5">84.5 mm</quantity>
|
||
) Ober-Ungarn, Karpaten;
|
||
<collectionCode id="ED5036E1FFA6FF84DBF22D67FEE2FC1F" box="[1166,1238,333,356]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430">NMW</collectionCode>
|
||
78816 (2, SL 88, 92 mm) same locality;
|
||
<collectionCode id="ED5036E1FFA6FF84DEEB2D5AFBEAFCFC" box="[407,478,368,391]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430">NMW</collectionCode>
|
||
6729 (3, SL
|
||
<quantity id="4CB903C1FFA6FF84DD122D5AF8D8FCF3" box="[622,748,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" metricValueMax="9.2" metricValueMin="8.2" pageId="11" pageNumber="430" unit="mm" value="87.0" valueMax="92.0" valueMin="82.0">82−92 mm</quantity>
|
||
); non
|
||
<typeStatus id="54FA1086FFA6FF84DC4C2D5AF951FCF3" box="[816,869,368,392]" pageId="11" pageNumber="430">type</typeStatus>
|
||
:
|
||
<collectionCode id="ED5036E1FFA6FF84DC122D5AF9A9FCFC" box="[878,925,368,391]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
315 (1, 90 mm), Vistula River,
|
||
<collectionCode id="ED5036E1FFA6FF84DB802D5AFF1FFCFC" box="[1276,1323,368,391]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="430">ZIN</collectionCode>
|
||
37661 (1, SL
|
||
<quantity id="4CB903C1FFA6FF84DFC22DBEFB26FCD7" box="[190,274,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="11" pageNumber="430" unit="mm" value="66.0">66 mm</quantity>
|
||
), Teresva River, Danube drainage,
|
||
<collectingCountry id="F356EEB4FFA6FF84DDE62DBEF8CCFCD6" box="[666,760,404,429]" name="Ukraine" pageId="11" pageNumber="430">Ukraine</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFEAE24FFA6FF84DFBB2D92FE11FCAB" blockId="11.[151,1436,151,464]" box="[199,1061,439,464]" pageId="11" pageNumber="430">
|
||
<taxonomicName id="4C41D5A7FFA6FF84DFBB2D92FB6EFCAB" box="[199,346,440,464]" class="Actinopterygii" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="430" phylum="Chordata" rank="species" species="metae">
|
||
<emphasis id="B9357236FFA6FF84DFBB2D92FB6EFCAB" box="[199,346,440,464]" italics="true" pageId="11" pageNumber="430">Cottus metae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<collectionCode id="ED5036E1FFA6FF84DE1B2D92FB9BFCB4" box="[359,431,440,463]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="430">NMW</collectionCode>
|
||
(3, SL
|
||
<quantity id="4CB903C1FFA6FF84DD7F2D9DF8B6FCAB" box="[515,642,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" metricValueMax="7.9" metricValueMin="7.5" pageId="11" pageNumber="430" unit="mm" value="77.0" valueMax="79.0" valueMin="75.0">75−79 mm</quantity>
|
||
), Krupa River, Sava system, Danube.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |