881 lines
105 KiB
XML
881 lines
105 KiB
XML
<document id="121FB889D71D8DF370CAAC99DFCAFF2E" ID-DOI="10.5281/zenodo.186100" ID-GBIF-Dataset="06b79f4e-5e07-4315-9636-eb8a036e3172" ID-ISSN="1175-5326" ID-Zenodo-Dep="186100" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045997293" checkinUser="plazi" docAuthor="Bartels, Paul J., Pilato, Giovanni, Lisi, Oscar & Nelson, Diane R." docDate="2009" docId="03974A7FFFEDFFF833AF30306562BE8D" docLanguage="en" docName="zt02022p057.pdf" docOrigin="Zootaxa 2022" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Macrobiotus halei Bartels, Pilato, Lisi & Nelson, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FFAE3207FFE8FFF333383550606CBE55" masterDocTitle="Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee / North Carolina, USA (North America): two new species and six new records" masterLastPageNumber="57" masterPageNumber="45" pageNumber="50" updateTime="1698577029529" updateUser="plazi">
|
||
<mods:mods id="CCB4A6120A8A978B0A4EE3B8E4525233" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="24D80E12A96D1D3D27FEA336320F6217">
|
||
<mods:title id="F74983AF98939F0DD05DFB6B48AFBF4B">Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee / North Carolina, USA (North America): two new species and six new records</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="010A2A93B4C7DA22CB1FBEFD7C62FE11" type="personal">
|
||
<mods:role id="C13D59B1CA1AACB25853037A1918BA41">
|
||
<mods:roleTerm id="FCE999353FDAAA54A89AE3A178CA33A7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E69108366623E32C0CD678158F4F7A83">Bartels, Paul J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7ED96907375C7C9A5B34117967B0C405" type="personal">
|
||
<mods:role id="7E8B4D59AD001E63367E5D48EFFFE6D4">
|
||
<mods:roleTerm id="FDBC1D7F7C81E38BBC2244D93A222FF6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="16E600822E37724C9C5C0451090EB89A">Pilato, Giovanni</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F839FCE0058D1AD5C7552D9D1B7125B6" type="personal">
|
||
<mods:role id="3071041B1EFB60D6A949099025C14267">
|
||
<mods:roleTerm id="87F6FD7D7D711FFC66F40FCDD721367F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="621D4EA97E378FCCAAD6FC2395B432CB">Lisi, Oscar</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="75C1D2E74F83C7A75B115E6BD8274CDA" type="personal">
|
||
<mods:role id="A78DAC47932D07C1FEFA602623A8EC68">
|
||
<mods:roleTerm id="F3B9DBFA4C4CF86DB290E161BE849CB4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A00A198B93AC45024578FC9619254C4D">Nelson, Diane R.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="625AF2D2545D742A8A3F7799C82F063D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="C2F5D8C97ED3A385E9EBA5D1ABECF78D" type="host">
|
||
<mods:titleInfo id="C58E5FA1B649D8CC0726C6308A512483">
|
||
<mods:title id="CEAAA2BB7973EEE79B93F5C6F2418926">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CEA86F692CA6C1B5C10965D6B7DD80D8">
|
||
<mods:date id="F6DA939138FB1F6387D1A2C2BB083921">2009</mods:date>
|
||
<mods:detail id="A0526F9EA9283296121E5F4CF271A3F4" type="volume">
|
||
<mods:number id="9750B55E49FE8CD26C5865E0E6B8D5FE">2022</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B1780FA659D1B4CE00F90808E296FDE2" unit="page">
|
||
<mods:start id="703948F03F4620AFD60EA2211E1A295D">45</mods:start>
|
||
<mods:end id="741B6942952FD052B9A68FFF144CDDB1">57</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="B0CE2E0B1686502021A93D2303E820F0">journal article</mods:classification>
|
||
<mods:identifier id="A52630D89AB4C916CAFFD19054EDF1E2" type="DOI">10.5281/zenodo.186100</mods:identifier>
|
||
<mods:identifier id="55820BFF5F98E6FF12F55A843E3654B2" type="GBIF-Dataset">06b79f4e-5e07-4315-9636-eb8a036e3172</mods:identifier>
|
||
<mods:identifier id="6A27CEA25420C9FD9D3434928F630AA1" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="9E9A1BB1F34EECED17DCEAE1F4B6C35B" type="Zenodo-Dep">186100</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03974A7FFFEDFFF833AF30306562BE8D" ID-DOI="http://doi.org/10.5281/zenodo.5626064" ID-GBIF-Taxon="119610561" ID-Zenodo-Dep="5626064" LSID="urn:lsid:plazi:treatment:03974A7FFFEDFFF833AF30306562BE8D" httpUri="http://treatment.plazi.org/id/03974A7FFFEDFFF833AF30306562BE8D" lastPageId="11" lastPageNumber="55" pageId="5" pageNumber="50">
|
||
<subSubSection id="C324A8E2FFEDFFF633AF30306168BBC8" pageId="5" pageNumber="50" type="nomenclature">
|
||
<paragraph id="8B81FB69FFEDFFF633AF303061B4BB2F" blockId="5.[151,472,1376,1437]" box="[151,472,1376,1402]" pageId="5" pageNumber="50">
|
||
<heading id="D0C94C05FFEDFFF633AF303061B4BB2F" bold="true" box="[151,472,1376,1402]" fontSize="11" level="1" pageId="5" pageNumber="50" reason="1">
|
||
<emphasis id="B94A277BFFEDFFF633AF303061B4BB2F" bold="true" box="[151,472,1376,1402]" pageId="5" pageNumber="50">
|
||
<taxonomicName id="4C3E80EAFFEDFFF633AF3030611FBB2F" box="[151,371,1376,1402]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="50" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFEDFFF633AF3030611FBB2F" bold="true" box="[151,371,1376,1402]" italics="true" pageId="5" pageNumber="50">Macrobiotus halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFEDFFF63242303061B4BB2F" box="[378,472,1376,1402]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFEDFFF633AF30D36168BBC8" blockId="5.[151,472,1376,1437]" box="[151,260,1411,1437]" pageId="5" pageNumber="50">
|
||
<figureCitation id="1305E7ECFFEDFFF633AF30D36168BBC8" box="[151,260,1411,1437]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6. A" captionStart-3="FIGURE 7. A" captionStartId-0="7.[151,257,1660,1684]" captionStartId-1="8.[151,256,931,955]" captionStartId-2="9.[151,256,1463,1487]" captionStartId-3="10.[151,255,1547,1571]" captionTargetBox-0="[161,1427,205,1631]" captionTargetBox-1="[155,1428,198,902]" captionTargetBox-2="[164,1426,281,1447]" captionTargetBox-3="[160,1427,205,1511]" captionTargetId-0="figure@7.[151,1436,194,1637]" captionTargetId-1="figure@8.[151,1436,194,907]" captionTargetId-2="figure@9.[151,1436,271,1455]" captionTargetId-3="figure@10.[151,1436,194,1523]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" captionText-1="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" captionText-2="FIGURE 6. A, buccal-pharyngeal apparatus of Macrobiotus vanescens. B, claws of Macrobiotus priviterae. C, egg of Macrobiotus danielisae. D, egg of Macrobiotus lorenae. (Scale bars = 10.0 µm)" captionText-3="FIGURE 7. A, claws on the second and third pairs of legs of Macrobiotus gerlachae. B, egg of Macrobiotus gerlachae. C, claws on the first and second pairs of legs of Macrobiotus corgatensis. D, egg of Macrobiotus corgatensis. (Scale bars = 10.0 µm)" httpUri-0="https://zenodo.org/record/186104/files/figure.png" httpUri-1="https://zenodo.org/record/186105/files/figure.png" httpUri-2="https://zenodo.org/record/186106/files/figure.png" httpUri-3="https://zenodo.org/record/186107/files/figure.png" pageId="5" pageNumber="50">Figs. 4–7</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324A8E2FFEDFFF633AF30836119B925" pageId="5" pageNumber="50" type="materials_examined">
|
||
<paragraph id="8B81FB69FFEDFFF633AF30836261B86E" blockId="5.[151,1437,1488,2020]" pageId="5" pageNumber="50">
|
||
<emphasis id="B94A277BFFEDFFF633AF30836128BBB8" bold="true" box="[151,324,1491,1517]" pageId="5" pageNumber="50">
|
||
<typeStatus id="548545CBFFEDFFF633AF308360B9BBB8" box="[151,213,1491,1517]" pageId="5" pageNumber="50">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
Brushy Mountain
|
||
<collectionCode id="ED2F63ACFFEDFFF6311F30836205BBB8" box="[551,617,1491,1517]" pageId="5" pageNumber="50">ATBI</collectionCode>
|
||
Plot (
|
||
<geoCoordinate id="EE0A9DAEFFEDFFF631893083631CBBB8" box="[689,880,1488,1517]" direction="west" orientation="longitude" pageId="5" pageNumber="50" precision="1" value="-83.432175">W 83o 25.9307’</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0A9DAEFFEDFFF630443083645FBBB8" box="[892,1075,1488,1517]" direction="north" orientation="latitude" pageId="5" pageNumber="50" precision="1" value="35.675995">N 35o 40.5596’</geoCoordinate>
|
||
,
|
||
<quantity id="4CC6568CFFEDFFF63707308364F6BBB8" box="[1087,1178,1491,1517]" metricMagnitude="3" metricUnit="m" metricValue="1.468" pageId="5" pageNumber="50" unit="m" value="1468.0">1468 m</quantity>
|
||
asl),
|
||
<collectionCode id="ED2F63ACFFEDFFF637E53083652FBBB8" LSID="urn:lsid:biocol.org:col:13136" box="[1245,1347,1491,1517]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="5" pageNumber="50">GSMNP</collectionCode>
|
||
, Sevier County, Tennessee,
|
||
<collectingCountry id="F329BBF9FFEDFFF632BC30AA61AFB841" box="[388,451,1530,1556]" name="United States of America" pageId="5" pageNumber="50">USA</collectingCountry>
|
||
. In soil and leaf litter in a heath bald community (dominant species
|
||
<taxonomicName id="4C3E80EAFFEDFFF637D630AA6165B86F" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="50" phylum="Tracheophyta" rank="species" species="maximum">
|
||
<emphasis id="B94A277BFFEDFFF637D630AA6165B86F" italics="true" pageId="5" pageNumber="50">Rhododendron maximum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3E80EAFFEDFFF6327A33716190B86F" box="[322,508,1569,1594]" class="Magnoliopsida" family="Ericaceae" genus="Kalmia" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="50" phylum="Tracheophyta" rank="species" species="latifolia">
|
||
<emphasis id="B94A277BFFEDFFF6327A33716190B86F" box="[322,508,1569,1594]" italics="true" pageId="5" pageNumber="50">Kalmia latifolia</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFEDFFF633FD3317655DB8DD" blockId="5.[151,1437,1488,2020]" pageId="5" pageNumber="50">
|
||
<emphasis id="B94A277BFFEDFFF633FD331761ACB834" bold="true" box="[197,448,1607,1633]" pageId="5" pageNumber="50">Material examined:</emphasis>
|
||
<typeStatus id="548545CBFFEDFFF632F133176251B834" box="[457,573,1607,1633]" pageId="5" pageNumber="50" type="holotype">Holotype</typeStatus>
|
||
: Brushy Mountain.
|
||
<typeStatus id="548545CBFFEDFFF63013331763C8B834" box="[811,932,1607,1633]" pageId="5" pageNumber="50" type="paratype">Paratypes</typeStatus>
|
||
: Brushy Mountain:
|
||
<specimenCount id="9D3830E0FFEDFFF637AA3317649DB834" box="[1170,1265,1607,1633]" pageId="5" pageNumber="50" type="adult">2 adults</specimenCount>
|
||
and
|
||
<specimenCount id="9D3830E0FFEDFFF6361533176517B834" box="[1325,1403,1607,1633]" pageId="5" pageNumber="50" type="egg">3 eggs</specimenCount>
|
||
(2 with fully developed embryos); Cataloochee:
|
||
<specimenCount id="9D3830E0FFEDFFF63197333E6353B8DD" box="[687,831,1646,1672]" pageId="5" pageNumber="50" type="generic">7 specimens</specimenCount>
|
||
and
|
||
<specimenCount id="9D3830E0FFEDFFF63040333E63D4B8DD" box="[888,952,1646,1672]" pageId="5" pageNumber="50" type="egg">1 egg</specimenCount>
|
||
(with fully-developed embryo).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFEDFFF633FD33C46119B925" blockId="5.[151,1437,1488,2020]" pageId="5" pageNumber="50">
|
||
<emphasis id="B94A277BFFEDFFF633FD33C461CBB8FB" bold="true" box="[197,423,1684,1710]" pageId="5" pageNumber="50">Species diagnosis:</emphasis>
|
||
Colorless; cuticle with very small tubercles, quite difficult to see; very fine granules present on all legs, more evident on legs IV; eyes absent. Buccal armature in oral cavity of
|
||
<taxonomicName id="4C3E80EAFFEDFFF637D733EB653AB881" box="[1263,1366,1723,1748]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="50" phylum="Tardigrada" rank="species" species="richtersi">
|
||
<emphasis id="B94A277BFFEDFFF637D733EB653AB881" box="[1263,1366,1723,1748]" italics="true" pageId="5" pageNumber="50">richtersi</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="548545CBFFEDFFF6366633EB65FBB880" box="[1374,1431,1723,1749]" pageId="5" pageNumber="50">type</typeStatus>
|
||
; buccal tube wide; stylet supports inserted on the buccal tube at 81.0–82.7% of its length; three macroplacoids and a microplacoid (far away from the third macroplacoid) present. Claws of the
|
||
<taxonomicName id="4C3E80EAFFEDFFF63752325964B7B977" box="[1130,1243,1801,1826]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="50" phylum="Tardigrada" rank="species" species="hufelandi">
|
||
<emphasis id="B94A277BFFEDFFF63752325964B7B977" box="[1130,1243,1801,1826]" italics="true" pageId="5" pageNumber="50">hufelandi</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="548545CBFFEDFFF637DB32596570B976" box="[1251,1308,1801,1827]" pageId="5" pageNumber="50">type</typeStatus>
|
||
; accessory points and lunules present. Eggs with truncated conical processes with a reticular design; egg shell areolated; areolae sculptured.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324A8E2FFEDFFF533FD322C6098BA51" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="description">
|
||
<paragraph id="8B81FB69FFEDFFF533FD322C64A8BFB2" blockId="5.[151,1437,1488,2020]" lastBlockId="6.[151,1437,152,1028]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
|
||
<emphasis id="B94A277BFFEDFFF633FD322C6258B9C3" bold="true" box="[197,564,1916,1942]" pageId="5" pageNumber="50">
|
||
Description of the
|
||
<typeStatus id="548545CBFFEDFFF63280322C6241B9C3" box="[440,557,1916,1942]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Body length 504.0 µm; colorless; eyes absent; cuticle with very small circular or elongated tubercles, more visible on the dorsal and lateral body wall, but very difficult to see (
|
||
<figureCitation id="1305E7ECFFEDFFF6365432F360CBB9B1" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="5" pageNumber="50">Fig. 4</figureCitation>
|
||
A, B); smaller granules present on all legs (
|
||
<figureCitation id="1305E7ECFFEDFFF63185329A6364B9B1" box="[701,776,1994,2020]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="5" pageNumber="50">Fig. 4</figureCitation>
|
||
B), but more evident on hind legs, especially in larger animals. Mouth terminal with 10 peribuccal lamellae; buccal armature in oral cavity well-developed (
|
||
<figureCitation id="1305E7ECFFEEFFF5360135C86513BEE7" box="[1337,1407,152,178]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="6" pageNumber="51">Fig. 4</figureCitation>
|
||
C) with anterior band of small teeth, posterior band of triangular teeth (some of them fused), and three dorsal and three ventral transverse ridges. Some supplementary teeth present between the ring of triangular teeth and the transverse ridges on the ventral wall of the buccal cavity. Buccal tube 62.3 µm long and 17 µm wide (
|
||
<emphasis id="B94A277BFFEEFFF53654345C65E8BF70" box="[1388,1412,268,293]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 27.3); stylet supports inserted on the buccal tube at 81.6% of its length (
|
||
<emphasis id="B94A277BFFEEFFF537363462644ABF1E" box="[1038,1062,306,331]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 81.6). Pharyngeal bulb with apophyses, three rod-shaped macroplacoids and a small microplacoid (with antero-lateral wings) far away from the third macroplacoid, a characteristic typical of species in the
|
||
<taxonomicName id="4C3E80EAFFEEFFF530FE34D06445BFCC" box="[966,1065,384,409]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="51" phylum="Tardigrada" rank="species" species="richtersi">
|
||
<emphasis id="B94A277BFFEEFFF530FE34D06445BFCC" box="[966,1065,384,409]" italics="true" pageId="6" pageNumber="51">richtersi</emphasis>
|
||
</taxonomicName>
|
||
group. First macroplacoid 10.9 µm long (
|
||
<emphasis id="B94A277BFFEEFFF5323334F7614DBF95" box="[267,289,423,448]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 17.5), the second 7.8 µm (
|
||
<emphasis id="B94A277BFFEEFFF5314C34F762E7BF95" box="[628,651,423,448]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 12.5), the third 12.1 µm (
|
||
<emphasis id="B94A277BFFEEFFF530EA34F76385BF95" box="[978,1001,423,448]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 19.4), the microplacoid 3.9 µm (
|
||
<emphasis id="B94A277BFFEEFFF536BD34F765F0BF95" box="[1413,1436,423,448]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 6.3); entire placoid row 47.9 µm (
|
||
<emphasis id="B94A277BFFEEFFF53104349D623EBFB3" box="[572,594,461,486]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 76.9); macroplacoid row 37.1 µm long (
|
||
<emphasis id="B94A277BFFEEFFF53771349D640CBFB3" box="[1097,1120,461,486]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 59.6).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFEEFFF533FD34A464DDBCD7" blockId="6.[151,1437,152,1028]" pageId="6" pageNumber="51">
|
||
Claws, of the
|
||
<taxonomicName id="4C3E80EAFFEEFFF5324834A4618FBC58" box="[368,483,500,525]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="51" phylum="Tardigrada" rank="species" species="hufelandi">
|
||
<emphasis id="B94A277BFFEEFFF5324834A4618FBC58" box="[368,483,500,525]" italics="true" pageId="6" pageNumber="51">hufelandi</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="548545CBFFEEFFF532D334A46248BC5B" box="[491,548,500,526]" pageId="6" pageNumber="51">type</typeStatus>
|
||
, well-developed (
|
||
<figureCitation id="1305E7ECFFEEFFF531C634A46324BC5B" box="[766,840,500,526]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="6" pageNumber="51">Fig. 4</figureCitation>
|
||
D, E), with evident accessory points on the main branches. Internal and external claws on the third pair of legs 13.5 µm long (
|
||
<emphasis id="B94A277BFFEEFFF5372D374A6440BC66" box="[1045,1068,538,563]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 21.7) and 14.7 µm long (
|
||
<emphasis id="B94A277BFFEEFFF53655374A65E8BC66" box="[1389,1412,538,563]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 23.6), respectively; anterior and posterior claws on the hind legs 16.3 µm long (
|
||
<emphasis id="B94A277BFFEEFFF537743711640FBC0F" box="[1100,1123,577,602]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 26.2) and 17.4 µm long (
|
||
<emphasis id="B94A277BFFEEFFF53399373860D4BCD4" box="[161,184,616,641]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
= 27.9), respectively. Other claws not measurable due to their unfavorable orientation.
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFEEFFF533FD37DE641EBDC5" blockId="6.[151,1437,152,1028]" pageId="6" pageNumber="51">
|
||
Eggs, laid freely, spherical with truncated conical processes (
|
||
<figureCitation id="1305E7ECFFEEFFF530B537DE63B8BCFD" box="[909,980,654,680]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="6" pageNumber="51">Fig. 5</figureCitation>
|
||
). In two measured eggs the diameter is 91.0 and 96.0 µm, excluding the processes, and 117.0 and 121.0 µm including them; 11 processes present around the circumference,
|
||
<quantity id="4CC6568CFFEEFFF532DE378C623FBCA3" box="[486,595,732,758]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.858" metricValueMin="6.096" pageId="6" pageNumber="51" unit="in" value="25.5" valueMax="27.0" valueMin="24.0">24–27 in</quantity>
|
||
the hemisphere. Processes 11.0–14.0 µm high, with basal diameter 22.0–23.5 µm. Processes with dense reticular design with elongated meshes having sinuous margins on the basal portion, almost isodiametric on the distal portion (
|
||
<figureCitation id="1305E7ECFFEEFFF530123679631CBD16" box="[810,880,809,835]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="6" pageNumber="51">Fig. 5</figureCitation>
|
||
B). Egg shell areolated between processes. The large areolae, with thick margins, have a more or less evident thickened central portion with circular light spots forming an irregular reticular design (
|
||
<figureCitation id="1305E7ECFFEEFFF531AD362662B6BDC5" box="[661,730,886,912]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="6" pageNumber="51">Fig. 5</figureCitation>
|
||
C) (similar to pits in a strawberry).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFEEFFF533FD36CD6098BA51" blockId="6.[151,1437,152,1028]" pageId="6" pageNumber="51">
|
||
The
|
||
<typeStatus id="548545CBFFEEFFF533C336CD6100BDE2" box="[251,364,925,951]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="548545CBFFEEFFF5317C36CD62C7BDE2" box="[580,683,925,951]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
in both qualitative and metric characters. Measurements and
|
||
<emphasis id="B94A277BFFEEFFF536BD36CD65F0BDE3" box="[1413,1436,925,950]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
values of selected morphological structures for the smallest and largest examined specimens are given in
|
||
<tableCitation id="C6BCCED2FFEEFFF533AF36BA609DBA51" box="[151,241,1002,1028]" captionStart="TABLE 2" captionStartId="6.[151,239,1079,1103]" captionTargetBox="[151,1179,1130,1836]" captionTargetPageId="6" captionText="TABLE 2. Macrobiotus halei sp. nov. metric characters (all values in µm, pt ratios in brackets)." httpUri="http://table.plazi.org/id/DF41ABE1FFEEFFF533AF316764FBBA1A" pageId="6" pageNumber="51" tableUuid="DF41ABE1FFEEFFF533AF316764FBBA1A">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF41ABE1FFEEFFF533AF316764FBBA1A" ID-Table-UUID="DF41ABE1FFEEFFF533AF316764FBBA1A" box="[151,1175,1079,1103]" httpUri="http://table.plazi.org/id/DF41ABE1FFEEFFF533AF316764FBBA1A" pageId="6" pageNumber="51" targetBox="[151,1179,1130,1836]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="8B81FB69FFEEFFF533AF316764FBBA1A" blockId="6.[151,1175,1079,1103]" box="[151,1175,1079,1103]" pageId="6" pageNumber="51">
|
||
<emphasis id="B94A277BFFEEFFF533AF31676164BA1A" bold="true" box="[151,264,1079,1103]" pageId="6" pageNumber="51">TABLE 2.</emphasis>
|
||
<taxonomicName id="4C3E80EAFFEEFFF53228316861BFBA1A" box="[272,467,1080,1103]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="51" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFEEFFF53228316861BFBA1A" box="[272,467,1080,1103]" italics="true" pageId="6" pageNumber="51">Macrobiotus halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFEEFFF532E23167625CBA1A" box="[474,560,1079,1103]" pageId="6" pageNumber="51" rank="species">
|
||
<emphasis id="B94A277BFFEEFFF532E23167625CBA1A" bold="true" box="[474,560,1079,1103]" pageId="6" pageNumber="51">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
metric characters (all values in µm,
|
||
<emphasis id="B94A277BFFEEFFF5308D316863A6BA1A" box="[949,970,1080,1103]" italics="true" pageId="6" pageNumber="51">pt</emphasis>
|
||
ratios in brackets).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81FB69FFEEFFF53175313A6432B979" pageId="6" pageNumber="51">
|
||
<table id="F93E09C9FFEE000C33AF313A64F7B979" box="[151,1179,1130,1836]" gridcols="3" gridrows="17" pageId="6" pageNumber="51">
|
||
<tr id="350EF92BFFEE000C33AF313A64F7BAD4" box="[151,1179,1130,1153]" gridrow="0" pageId="6" pageNumber="51" rowspan-0="1">
|
||
<th id="76DF9057FFEE000C3175313A637CBAD4" box="[589,784,1130,1153]" gridcol="1" gridrow="0" pageId="6" pageNumber="51">smallest specimen</th>
|
||
<th id="76DF9057FFEE000C30D1313A64F7BAD4" box="[1001,1179,1130,1153]" gridcol="2" gridrow="0" pageId="6" pageNumber="51">largest specimen</th>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF31C564F7BAF9" box="[151,1179,1173,1196]" gridrow="1" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF31C561E9BAF9" box="[151,389,1173,1196]" gridcol="0" gridrow="1" pageId="6" pageNumber="51">Body length</th>
|
||
<td id="76DF9057FFEE000C317531C5637CBAF9" box="[589,784,1173,1196]" gridcol="1" gridrow="1" pageId="6" pageNumber="51">405.0</td>
|
||
<td id="76DF9057FFEE000C30D131C564F7BAF9" box="[1001,1179,1173,1196]" gridcol="2" gridrow="1" pageId="6" pageNumber="51">630.0</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF319064F7BA82" box="[151,1179,1216,1239]" gridrow="2" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF319061E9BA82" box="[151,389,1216,1239]" gridcol="0" gridrow="2" pageId="6" pageNumber="51">Buccal tube length</th>
|
||
<td id="76DF9057FFEE000C31753190637CBA82" box="[589,784,1216,1239]" gridcol="1" gridrow="2" pageId="6" pageNumber="51">53.5</td>
|
||
<td id="76DF9057FFEE000C30D1319064F7BA82" box="[1001,1179,1216,1239]" gridcol="2" gridrow="2" pageId="6" pageNumber="51">66.8</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF31BA64F7BB54" box="[151,1179,1258,1281]" gridrow="3" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF31BA61E9BB54" box="[151,389,1258,1281]" gridcol="0" gridrow="3" pageId="6" pageNumber="51">Buccal tube width</th>
|
||
<td id="76DF9057FFEE000C317531BA637CBB54" box="[589,784,1258,1281]" gridcol="1" gridrow="3" pageId="6" pageNumber="51">14.2 [26.5]</td>
|
||
<td id="76DF9057FFEE000C30D131BA64F7BB54" box="[1001,1179,1258,1281]" gridcol="2" gridrow="3" pageId="6" pageNumber="51">19.1 [28.6]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF304564F7BB79" box="[151,1179,1301,1324]" gridrow="4" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF304561E9BB79" box="[151,389,1301,1324]" gridcol="0" gridrow="4" pageId="6" pageNumber="51">Stylet supports</th>
|
||
<td id="76DF9057FFEE000C31753045637CBB79" box="[589,784,1301,1324]" gridcol="1" gridrow="4" pageId="6" pageNumber="51">43.6 [81.5]</td>
|
||
<td id="76DF9057FFEE000C30D1304564F7BB79" box="[1001,1179,1301,1324]" gridcol="2" gridrow="4" pageId="6" pageNumber="51">54.4 [81.5]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF301064F7BB02" box="[151,1179,1344,1367]" gridrow="5" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF301061E9BB02" box="[151,389,1344,1367]" gridcol="0" gridrow="5" pageId="6" pageNumber="51">Placoid row</th>
|
||
<td id="76DF9057FFEE000C31753010637CBB02" box="[589,784,1344,1367]" gridcol="1" gridrow="5" pageId="6" pageNumber="51">36.9 [69.0]</td>
|
||
<td id="76DF9057FFEE000C30D1301064F7BB02" box="[1001,1179,1344,1367]" gridcol="2" gridrow="5" pageId="6" pageNumber="51">49.6 [74.3]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF303A64F7BBD4" box="[151,1179,1386,1409]" gridrow="6" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF303A61E9BBD4" box="[151,389,1386,1409]" gridcol="0" gridrow="6" pageId="6" pageNumber="51">Macroplacoid row</th>
|
||
<td id="76DF9057FFEE000C3175303A637CBBD4" box="[589,784,1386,1409]" gridcol="1" gridrow="6" pageId="6" pageNumber="51">27.8 [52.0]</td>
|
||
<td id="76DF9057FFEE000C30D1303A64F7BBD4" box="[1001,1179,1386,1409]" gridcol="2" gridrow="6" pageId="6" pageNumber="51">37.0 [55.4]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF30C564F7BBF9" box="[151,1179,1429,1452]" gridrow="7" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF30C561E9BBF9" box="[151,389,1429,1452]" gridcol="0" gridrow="7" pageId="6" pageNumber="51">First macroplacoid</th>
|
||
<td id="76DF9057FFEE000C317530C5637CBBF9" box="[589,784,1429,1452]" gridcol="1" gridrow="7" pageId="6" pageNumber="51">9.2 [17.2]</td>
|
||
<td id="76DF9057FFEE000C30D130C564F7BBF9" box="[1001,1179,1429,1452]" gridcol="2" gridrow="7" pageId="6" pageNumber="51">12.4 [18.6]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF309064F7BB82" box="[151,1179,1472,1495]" gridrow="8" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF309061E9BB82" box="[151,389,1472,1495]" gridcol="0" gridrow="8" pageId="6" pageNumber="51">Second macroplacoid</th>
|
||
<td id="76DF9057FFEE000C31753090637CBB82" box="[589,784,1472,1495]" gridcol="1" gridrow="8" pageId="6" pageNumber="51">6.4 [12.0]</td>
|
||
<td id="76DF9057FFEE000C30D1309064F7BB82" box="[1001,1179,1472,1495]" gridcol="2" gridrow="8" pageId="6" pageNumber="51">8.8 [13.2]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF30BA64F7B854" box="[151,1179,1514,1537]" gridrow="9" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF30BA61E9B854" box="[151,389,1514,1537]" gridcol="0" gridrow="9" pageId="6" pageNumber="51">Third macroplacoid</th>
|
||
<td id="76DF9057FFEE000C317530BA637CB854" box="[589,784,1514,1537]" gridcol="1" gridrow="9" pageId="6" pageNumber="51">9.8 [18.3]</td>
|
||
<td id="76DF9057FFEE000C30D130BA64F7B854" box="[1001,1179,1514,1537]" gridcol="2" gridrow="9" pageId="6" pageNumber="51">12.7 [19.0]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF334564F7B879" box="[151,1179,1557,1580]" gridrow="10" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF334561E9B879" box="[151,389,1557,1580]" gridcol="0" gridrow="10" pageId="6" pageNumber="51">Microplacoid</th>
|
||
<td id="76DF9057FFEE000C31753345637CB879" box="[589,784,1557,1580]" gridcol="1" gridrow="10" pageId="6" pageNumber="51">3.4 [6.4]</td>
|
||
<td id="76DF9057FFEE000C30D1334564F7B879" box="[1001,1179,1557,1580]" gridcol="2" gridrow="10" pageId="6" pageNumber="51">4.0 [6.0]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF331064F7B802" box="[151,1179,1600,1623]" gridrow="11" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF331061E9B802" box="[151,389,1600,1623]" gridcol="0" gridrow="11" pageId="6" pageNumber="51">Internal claw II</th>
|
||
<td id="76DF9057FFEE000C31753310637CB802" box="[589,784,1600,1623]" gridcol="1" gridrow="11" pageId="6" pageNumber="51">10.9 [20.4]</td>
|
||
<td id="76DF9057FFEE000C30D1331064F7B802" box="[1001,1179,1600,1623]" gridcol="2" gridrow="11" pageId="6" pageNumber="51">14.1 [21.1]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF333A64F7B8D4" box="[151,1179,1642,1665]" gridrow="12" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF333A61E9B8D4" box="[151,389,1642,1665]" gridcol="0" gridrow="12" pageId="6" pageNumber="51">External claw II</th>
|
||
<td id="76DF9057FFEE000C3175333A637CB8D4" box="[589,784,1642,1665]" gridcol="1" gridrow="12" pageId="6" pageNumber="51">12.2 [22.8]</td>
|
||
<td id="76DF9057FFEE000C30D1333A64F7B8D4" box="[1001,1179,1642,1665]" gridcol="2" gridrow="12" pageId="6" pageNumber="51">15.5 [23.2]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF33C564F7B8F9" box="[151,1179,1685,1708]" gridrow="13" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF33C561E9B8F9" box="[151,389,1685,1708]" gridcol="0" gridrow="13" pageId="6" pageNumber="51">Internal claw III</th>
|
||
<td id="76DF9057FFEE000C317533C5637CB8F9" box="[589,784,1685,1708]" gridcol="1" gridrow="13" pageId="6" pageNumber="51">11.6 [21.7]</td>
|
||
<td id="76DF9057FFEE000C30D133C564F7B8F9" box="[1001,1179,1685,1708]" gridcol="2" gridrow="13" pageId="6" pageNumber="51">14.6 [21.9]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF339064F7B882" box="[151,1179,1728,1751]" gridrow="14" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF339061E9B882" box="[151,389,1728,1751]" gridcol="0" gridrow="14" pageId="6" pageNumber="51">External claw III</th>
|
||
<td id="76DF9057FFEE000C31753390637CB882" box="[589,784,1728,1751]" gridcol="1" gridrow="14" pageId="6" pageNumber="51">12.6 [23.6]</td>
|
||
<td id="76DF9057FFEE000C30D1339064F7B882" box="[1001,1179,1728,1751]" gridcol="2" gridrow="14" pageId="6" pageNumber="51">16.3 [24.4]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF33BA64F7B954" box="[151,1179,1770,1793]" gridrow="15" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF33BA61E9B954" box="[151,389,1770,1793]" gridcol="0" gridrow="15" pageId="6" pageNumber="51">Anterior claw IV</th>
|
||
<td id="76DF9057FFEE000C317533BA637CB954" box="[589,784,1770,1793]" gridcol="1" gridrow="15" pageId="6" pageNumber="51">13.0 [24.3]</td>
|
||
<td id="76DF9057FFEE000C30D133BA64F7B954" box="[1001,1179,1770,1793]" gridcol="2" gridrow="15" pageId="6" pageNumber="51">17.5 [26.2]</td>
|
||
</tr>
|
||
<tr id="350EF92BFFEE000C33AF324564F7B979" box="[151,1179,1813,1836]" gridrow="16" pageId="6" pageNumber="51">
|
||
<th id="76DF9057FFEE000C33AF324561E9B979" box="[151,389,1813,1836]" gridcol="0" gridrow="16" pageId="6" pageNumber="51">Posterior claw IV</th>
|
||
<td id="76DF9057FFEE000C31753245637CB979" box="[589,784,1813,1836]" gridcol="1" gridrow="16" pageId="6" pageNumber="51">15.0 [28.0]</td>
|
||
<td id="76DF9057FFEE000C30D1324564F7B979" box="[1001,1179,1813,1836]" gridcol="2" gridrow="16" pageId="6" pageNumber="51">19.1 [28.6]</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C324A8E2FFEEFFF533FD32266447B98B" pageId="6" pageNumber="51" type="etymology">
|
||
<paragraph id="8B81FB69FFEEFFF533FD32266447B98B" blockId="6.[151,1436,1910,2014]" pageId="6" pageNumber="51">
|
||
<emphasis id="B94A277BFFEEFFF533FD3226613AB9C5" bold="true" box="[197,342,1910,1936]" pageId="6" pageNumber="51">Etymology:</emphasis>
|
||
The name ‘
|
||
<emphasis id="B94A277BFFEEFFF532DD3227625CB9C5" box="[485,560,1911,1936]" italics="true" pageId="6" pageNumber="51">
|
||
<taxonomicName id="4C3E80EAFFEEFFF532DD32276249B9C5" box="[485,549,1911,1936]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="51" phylum="Tardigrada" rank="species" species="halei">halei</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
is in honor of Mr. Gilbert Hale in appreciation of his long-time service as lab assistant for Dr. Diane Nelson. He has produced thousands of slides for the tardigrade inventory of the ATBI, and he is exceptional at finding eggs. Gilbert is a “mighty bear hunter”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF41ABE1FFEFFFF433AF332C646CB88C" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="7" pageNumber="52" targetBox="[161,1427,205,1631]" targetPageId="7">
|
||
<paragraph id="8B81FB69FFEFFFF433AF332C646CB88C" blockId="7.[151,1437,1660,1753]" pageId="7" pageNumber="52">
|
||
<emphasis id="B94A277BFFEFFFF433AF332C6171B8C1" bold="true" box="[151,285,1660,1684]" pageId="7" pageNumber="52">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C3E80EAFFEFFFF4321C332D6182B8C1" box="[292,494,1661,1684]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="52" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFEFFFF4321C332D6182B8C1" box="[292,494,1661,1684]" italics="true" pageId="7" pageNumber="52">Macrobiotus halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFEFFFF432CE332C6223B8C1" box="[502,591,1660,1684]" pageId="7" pageNumber="52" rank="species">
|
||
<emphasis id="B94A277BFFEFFFF432CE332C6223B8C1" bold="true" box="[502,591,1660,1684]" pageId="7" pageNumber="52">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C324A8E2FFEFFFFB33FD325B6380BABE" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52" type="materials_examined">
|
||
<paragraph id="8B81FB69FFEFFFF433FD325B61CFB9B2" blockId="7.[151,1438,1803,2023]" pageId="7" pageNumber="52">
|
||
<emphasis id="B94A277BFFEFFFF433FD325B61DBB970" bold="true" box="[197,439,1803,1829]" pageId="7" pageNumber="52">
|
||
<typeStatus id="548545CBFFEFFFF433FD325B6169B970" box="[197,261,1803,1829]" pageId="7" pageNumber="52">Type</typeStatus>
|
||
depositories:
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="548545CBFFEFFFF432C1325B6209B970" box="[505,613,1803,1829]" pageId="7" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED2F63ACFFEFFFF4314E325B62BCB970" LSID="urn:lsid:biocol.org:col:34871" box="[630,720,1803,1829]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="52">USNM</collectionCode>
|
||
# 1120692) and 3
|
||
<typeStatus id="548545CBFFEFFFF4308C325B6447B970" box="[948,1067,1803,1829]" pageId="7" pageNumber="52" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="ED2F63ACFFEFFFF43704325B64FAB970" LSID="urn:lsid:biocol.org:col:34871" box="[1084,1174,1803,1829]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="52">USNM</collectionCode>
|
||
# 1120693, 1120694, 1120695 (egg)) are deposited in the National Museum of Natural History, Smithsonian Institution, Washington, DC,
|
||
<collectingCountry id="F329BBF9FFEFFFF43242320961D0B926" box="[378,444,1881,1907]" name="United States of America" pageId="7" pageNumber="52">USA</collectingCountry>
|
||
. The remaining
|
||
<typeStatus id="548545CBFFEFFFF431B432096369B926" box="[652,773,1881,1907]" pageId="7" pageNumber="52" type="paratype">paratypes</typeStatus>
|
||
are deposited in the
|
||
<collectionCode id="ED2F63ACFFEFFFF4372A32096410B926" LSID="urn:lsid:biocol.org:col:13136" box="[1042,1148,1881,1907]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13136" name="Great Smoky Mountains National Park" pageId="7" pageNumber="52">GSMNP</collectionCode>
|
||
collection; the Nelson collection in the Department of Biological Sciences, East Tennessee State University, Johnson City, Tennessee,
|
||
<collectingCountry id="F329BBF9FFEFFFF4321A32F6610EB995" box="[290,354,1958,1984]" name="United States of America" pageId="7" pageNumber="52">USA</collectingCountry>
|
||
; and the Binda and Pilato collection in the Department of Animal Biology, University of Catania, Catania,
|
||
<collectingCountry id="F329BBF9FFEFFFF4325F329D61F3B9B2" box="[359,415,1997,2023]" name="Italy" pageId="7" pageNumber="52">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF41ABE1FFE0FFFB33AF36F3618ABD88" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="8" pageNumber="53" targetBox="[155,1428,198,902]" targetPageId="8">
|
||
<paragraph id="8B81FB69FFE0FFFB33AF36F3618ABD88" blockId="8.[151,1436,931,989]" pageId="8" pageNumber="53">
|
||
<emphasis id="B94A277BFFE0FFFB33AF36F36170BDEE" bold="true" box="[151,284,931,955]" pageId="8" pageNumber="53">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C3E80EAFFE0FFFB321B36F36186BDEF" box="[291,490,931,954]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB321B36F36186BDEF" box="[291,490,931,954]" italics="true" pageId="8" pageNumber="53">Macrobiotus halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB32CA36F36226BDEE" box="[498,586,931,955]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB32CA36F36226BDEE" bold="true" box="[498,586,931,955]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81FB69FFE0FFFB33FD315F6380BABE" blockId="8.[151,1437,1039,2032]" pageId="8" pageNumber="53">
|
||
<emphasis id="B94A277BFFE0FFFB33FD315F61B3BA7C" bold="true" box="[197,479,1039,1065]" pageId="8" pageNumber="53">Differential Diagnosis:</emphasis>
|
||
<emphasis id="B94A277BFFE0FFFB32DE3140634CBA7C" box="[486,800,1039,1065]" italics="true" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C3E80EAFFE0FFFB32DE314062D2BA7C" box="[486,702,1040,1065]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">Macrobiotus halei</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB31FC315F634CBA7C" box="[708,800,1039,1065]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB31FC315F634CBA7C" bold="true" box="[708,800,1039,1065]" italics="true" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C3E80EAFFE0FFFB3081314060B8BA05" authority="Pilato, Binda & Catanzaro, 1991" authorityName="Pilato, Binda & Catanzaro" authorityYear="1991" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE0FFFB30813140643BBA7C" box="[953,1111,1040,1065]" italics="true" pageId="8" pageNumber="53">M. vanescens</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB3767315F60B8BA05" author="Pilato" pageId="8" pageNumber="57" refString="Pilato, G., Binda, M. G. & Catanzaro, R. (1991) Remarks on some tardigrades of the African fauna with description of three new species of Macrobiotus Schultze 1894. Tropical Zoology, 4, 167 - 178." type="journal article" year="1991">Pilato, Binda & Catanzaro, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33D8316663E4BA05" authority="Binda, Pilato, Moncada & Napolitano, 2001" authorityName="Binda, Pilato, Moncada & Napolitano" authorityYear="2001" box="[224,904,1078,1104]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="priviterae">
|
||
<emphasis id="B94A277BFFE0FFFB33D831666115BA1A" box="[224,377,1078,1103]" italics="true" pageId="8" pageNumber="53">M. priviterae</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB32B9316663E4BA05" author="Binda" box="[385,904,1078,1104]" pageId="8" pageNumber="56" refString="Binda, M. G., Pilato, G., Moncada, E. & Napolitano, G. (2001) Some tardigrades from Central Africa with the description of two new species: Macrobiotus ragonesei and M. priviterae (Eutardigrada Macrobiotidae). Tropical Zoology, 14, 233 - 242." type="journal article" year="2001">Binda, Pilato, Moncada & Napolitano, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB30AC3166651DBA05" authority="Pilato, Binda & Lisi, 2006" authorityName="Pilato, Binda & Lisi" authorityYear="2006" box="[916,1393,1078,1104]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB30AC3166645EBA1A" box="[916,1074,1078,1103]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB37023166651DBA05" author="Pilato" box="[1082,1393,1078,1104]" pageId="8" pageNumber="57" refString="Pilato, G., Binda, M. G. & Lisi, O. (2006) Three new species of eutardigrades from the Seychelles. New Zealand Journal of Zoology, 33, 39 - 48." type="journal article" year="2006">Pilato, Binda & Lisi, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB3644316661F3BA22" authority="Biserov, 1996" authorityName="Biserov" authorityYear="1996" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="lorenae">
|
||
<emphasis id="B94A277BFFE0FFFB36443166609CBA23" italics="true" pageId="8" pageNumber="53">M. lorenae</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB33C0310D61F3BA22" author="Biserov" box="[248,415,1117,1143]" pageId="8" pageNumber="56" refString="Biserov, V. (1996) Macrobiotus lorenae sp. n., a new species of Tardigrada (Eutardigrada, Macrobiotidae) from the Russian Far East. Arthropoda Selecta, 5 (1 - 2), 145 - 149." type="journal article" year="1996">Biserov, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB3293310D63FBBA22" authority="Pilato, Binda & Lisi , 2004" authorityName="Pilato, Binda & Lisi" authorityYear="2004" box="[427,919,1117,1143]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="gerlachae">
|
||
<emphasis id="B94A277BFFE0FFFB3293310D6225BA23" box="[427,585,1117,1142]" italics="true" pageId="8" pageNumber="53">M. gerlachae</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB3169310D63FBBA22" author="Pilato" box="[593,919,1117,1143]" pageId="8" pageNumber="57" refString="Pilato, G., Binda, M. G. & Lisi, O. (2004) Notes on tardigrades of the Seychelles with the description of three new species. Italian Journal of Zoology, 71, 171 - 178." type="journal article" year="2004">Pilato, Binda & Lisi, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB309B310D65FBBA22" authority="Pilato, Binda & Lisi, 2002" authorityName="Pilato, Binda & Lisi" authorityYear="2002" box="[931,1431,1117,1143]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="corgatensis">
|
||
<emphasis id="B94A277BFFE0FFFB309B310D643EBA23" box="[931,1106,1117,1142]" italics="true" pageId="8" pageNumber="53">M. corgatensis</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB3762310D65FBBA22" author="Pilato" box="[1114,1431,1117,1143]" pageId="8" pageNumber="57" refString="Pilato, G., Binda, M. G. & Lisi, O. (2002) Notes on tardigrades of the Seychelles with description of two new species. Bollettino dell'Accademia Gioenia di Scienze Naturali, Catania 35, 361, 503 - 517." type="journal article" year="2002">Pilato, Binda & Lisi, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33AF31D463EABAC8" authority="Michalczyk, Kaczmarek & Weglarska, 2006" authorityName="Michalczyk, Kaczmarek & Weglarska" authorityYear="2006" box="[151,902,1155,1181]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="sklodowskae">
|
||
<emphasis id="B94A277BFFE0FFFB33AF31D46136BAC8" box="[151,346,1156,1181]" italics="true" pageId="8" pageNumber="53">M. sklodowskae</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB325B31D363EABAC8" author="Michalczyk" box="[355,902,1155,1181]" pageId="8" pageNumber="57" refString="Michalczyk, L., Kaczmarek, L. & Weglarska, B. (2006) Macrobiotus sklodowskae sp. nov. (Tardigrada: Eutardigrada: Macrobiotidae, richtersi - group) from Cyprus. Zootaxa, 1371, 45 - 56." type="journal article" year="2006">Michalczyk, Kaczmarek & Weglarska, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB30AC31D4649BBAC8" authority="Tumanov, 2005" authorityName="Tumanov" authorityYear="2005" box="[916,1271,1155,1181]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="alekseevi">
|
||
<emphasis id="B94A277BFFE0FFFB30AC31D463C0BAC8" box="[916,940,1156,1181]" italics="true" pageId="8" pageNumber="53">M</emphasis>
|
||
.
|
||
<emphasis id="B94A277BFFE0FFFB308531D46441BAC8" box="[957,1069,1156,1181]" italics="true" pageId="8" pageNumber="53">alekseevi</emphasis>
|
||
<bibRefCitation id="EFAF8698FFE0FFFB370E31D3649BBAC8" author="Tumanov" box="[1078,1271,1155,1181]" pageId="8" pageNumber="57" refString="Tumanov, D. V. (2005) Notes on the Tardigrada of Thailand, with a description of Macrobiotus alekseevi sp. nov. (Eutardigrada, Macrobiotidae). Zootaxa, 999, 1 - 16." type="journal article" year="2005">Tumanov, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E80EAFFE0FFFB363D31D462F3BA91" authority="Kaczmarek, Michalczyk & Diduszko, 2005" authorityName="Kaczmarek, Michalczyk & Diduszko" authorityYear="2005" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="garynhai">
|
||
<emphasis id="B94A277BFFE0FFFB363D31D465F0BAC8" box="[1285,1436,1156,1181]" italics="true" pageId="8" pageNumber="53">M. garynhai</emphasis>
|
||
Kaczmarek, Michalczyk &
|
||
<bibRefCitation id="EFAF8698FFE0FFFB32D831FA62F3BA91" author="Diduszko" box="[480,671,1194,1220]" pageId="8" pageNumber="56" refString="Kaczmarek, L., Michalczk, L. & Diduszko, D. (2005) Some tardigrades from Siberia (Russia Baikal region) with a description of Macrobiotus garynhai sp. nov. (Eutardigrada: Macrobiotidae: richtersi group). Zootaxa, 1053, 35 - 45." type="journal article" year="2005">Diduszko, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3E80EAFFE0FFFB31E731FA6548BA91" authority="Michalczyk & Kaczmarek, 2006" authorityName="Michalczyk & Kaczmarek" authorityYear="2006" box="[735,1316,1194,1220]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="magdalenae">
|
||
<emphasis id="B94A277BFFE0FFFB31E731FA63FBBA96" box="[735,919,1194,1219]" italics="true" pageId="8" pageNumber="53">M. magdalenae</emphasis>
|
||
Michalczyk & Kaczmarek,
|
||
<quantity id="4CC6568CFFE0FFFB37D131FA6548BA91" box="[1257,1316,1194,1220]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="8" pageNumber="53" unit="in" value="2006.0">2006</quantity>
|
||
</taxonomicName>
|
||
in having sculptured egg areolae, but it differs from them in some other characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324A8E2FFE0FFF933FD31A8615BB920" lastPageId="10" lastPageNumber="55" pageId="8" pageNumber="53" type="description">
|
||
<paragraph id="8B81FB69FFE0FFFB33FD31A863D4BB86" blockId="8.[151,1437,1039,2032]" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33FD31A8614BBB44" box="[197,295,1272,1297]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33FD31A8614BBB44" box="[197,295,1272,1297]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB321531A761E6BB44" box="[301,394,1271,1297]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB321531A761E6BB44" bold="true" box="[301,394,1271,1297]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE0FFFB311C31A862ADBB44" box="[548,705,1272,1297]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE0FFFB311C31A862ADBB44" box="[548,705,1272,1297]" italics="true" pageId="8" pageNumber="53">M. vanescens</emphasis>
|
||
</taxonomicName>
|
||
by having small cuticular tubercles (difficult to see) instead of a very fine punctation; stylet supports inserted on the buccal tube in a more caudal position (
|
||
<emphasis id="B94A277BFFE0FFFB37EB304E6486BB62" box="[1235,1258,1310,1335]" italics="true" pageId="8" pageNumber="53">pt</emphasis>
|
||
= 81.0–
|
||
<quantity id="4CC6568CFFE0FFFB3673304E65F0BB6D" box="[1355,1436,1310,1336]" metricMagnitude="0" metricUnit="m" metricValue="2.10058" pageId="8" pageNumber="53" unit="in" value="82.7">82.7 in</quantity>
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33AF30156090BB0B" box="[151,252,1349,1375]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33AF301560C3BB0B" box="[151,175,1349,1374]" italics="true" pageId="8" pageNumber="53">M</emphasis>
|
||
.
|
||
<emphasis id="B94A277BFFE0FFFB33F830156090BB0B" box="[192,252,1349,1374]" italics="true" pageId="8" pageNumber="53">halei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94A277BFFE0FFFB323C3014610ABB0B" bold="true" box="[260,358,1348,1374]" pageId="8" pageNumber="53">
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB323C3014610ABB0B" box="[260,358,1348,1374]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<quantity id="4CC6568CFFE0FFFB324E30156263BB0A" box="[374,527,1349,1375]" metricMagnitude="0" metricUnit="m" metricValue="1.9558" metricValueMax="1.9685" metricValueMin="1.9431" pageId="8" pageNumber="53" unit="in" value="77.0" valueMax="77.5" valueMin="76.5">76.5–77.5 in</quantity>
|
||
<taxonomicName id="4C3E80EAFFE0FFFB312F301562D6BB0B" box="[535,698,1349,1374]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE0FFFB312F301562D6BB0B" box="[535,698,1349,1374]" italics="true" pageId="8" pageNumber="53">M. vanescens</emphasis>
|
||
</taxonomicName>
|
||
); a longer microplacoid (
|
||
<emphasis id="B94A277BFFE0FFFB30CA30156465BB0B" box="[1010,1033,1349,1374]" italics="true" pageId="8" pageNumber="53">pt</emphasis>
|
||
= about 6.0 in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB37FD30156546BB0B" box="[1221,1322,1349,1374]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB37FD30156546BB0B" box="[1221,1322,1349,1374]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB360B301465F8BB0B" box="[1331,1428,1348,1374]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB360B301465F8BB0B" bold="true" box="[1331,1428,1348,1374]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, about 4.0 in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB3214303C61A0BBD0" box="[300,460,1388,1413]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE0FFFB3214303C61A0BBD0" box="[300,460,1388,1413]" italics="true" pageId="8" pageNumber="53">M. vanescens</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="1305E7ECFFE0FFFB32DF303B6256BBD0" box="[487,570,1387,1413]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="8" pageNumber="53">Figs. 4</figureCitation>
|
||
C and 6A); and a different shape of the egg processes, which are slightly shorter (11.0–14.0 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB32E230C2622FBBFE" box="[474,579,1426,1451]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei">
|
||
<emphasis id="B94A277BFFE0FFFB32E230C2622FBBFE" box="[474,579,1426,1451]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
, 16.0–17.0 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB301630C263BABBFE" box="[814,982,1426,1451]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE0FFFB301630C263BABBFE" box="[814,982,1426,1451]" italics="true" pageId="8" pageNumber="53">M. vanescens</emphasis>
|
||
</taxonomicName>
|
||
) while the basal diameter is similar (22.0–23.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB325230E961A7BB87" box="[362,459,1465,1490]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB325230E961A7BB87" box="[362,459,1465,1490]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB32EB30E8625CBB87" box="[467,560,1464,1490]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB32EB30E8625CBB87" bold="true" box="[467,560,1464,1490]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 24.0–25.0 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB303130E963CBBB87" box="[777,935,1465,1490]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE0FFFB303130E963CBBB87" box="[777,935,1465,1490]" italics="true" pageId="8" pageNumber="53">M. vanescens</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE0FFFB33FD30B06102B838" blockId="8.[151,1437,1039,2032]" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33FD30B0614BBBAC" box="[197,295,1504,1529]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33FD30B0614BBBAC" box="[197,295,1504,1529]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB3215308F61E7BBAC" box="[301,395,1503,1529]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB3215308F61E7BBAC" bold="true" box="[301,395,1503,1529]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE0FFFB311D30B062D3BBAC" box="[549,703,1504,1529]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="priviterae">
|
||
<emphasis id="B94A277BFFE0FFFB311D30B062D3BBAC" box="[549,703,1504,1529]" italics="true" pageId="8" pageNumber="53">M. priviterae</emphasis>
|
||
</taxonomicName>
|
||
by lacking eyes and by having a sculptured cuticle, a different shape of the claws (
|
||
<figureCitation id="1305E7ECFFE0FFFB3245335661A2B875" box="[381,462,1542,1568]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="8" pageNumber="53">Figs. 4</figureCitation>
|
||
C and 6B), egg processes with similar height but a larger basal diameter (22.0–23.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33D1337D613DB813" box="[233,337,1581,1606]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33D1337D613DB813" box="[233,337,1581,1606]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB3262337C61D1B813" box="[346,445,1580,1606]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB3262337C61D1B813" bold="true" box="[346,445,1580,1606]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 13.0–16.3 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB319D337D6326B813" box="[677,842,1581,1606]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="priviterae">
|
||
<emphasis id="B94A277BFFE0FFFB319D337D6326B813" box="[677,842,1581,1606]" italics="true" pageId="8" pageNumber="53">M. priviterae</emphasis>
|
||
</taxonomicName>
|
||
) and a more evident reticular design with more elongated meshes.
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE0FFFB33FD332A6454B900" blockId="8.[151,1437,1039,2032]" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33FD332A614BB8C6" box="[197,295,1658,1683]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33FD332A614BB8C6" box="[197,295,1658,1683]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB3216332961E7B8C6" box="[302,395,1657,1683]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB3216332961E7B8C6" bold="true" box="[302,395,1657,1683]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE0FFFB311E332A62A9B8C6" box="[550,709,1658,1683]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB311E332A62A9B8C6" box="[550,709,1658,1683]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
</taxonomicName>
|
||
by having different cuticular ornamentation (small circular or elongate tubercles in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB32AE33F16196B8EF" box="[406,506,1697,1722]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB32AE33F16196B8EF" box="[406,506,1697,1722]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB313A33F0620DB8EF" box="[514,609,1696,1722]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB313A33F0620DB8EF" bold="true" box="[514,609,1696,1722]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, small polygons of variable and irregular shape in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB37FB33F16509B8EF" box="[1219,1381,1697,1722]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB37FB33F16509B8EF" box="[1219,1381,1697,1722]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
</taxonomicName>
|
||
), by having shorter egg processes (11.0–14 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB318933986378B8B4" box="[689,788,1736,1761]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB318933986378B8B4" box="[689,788,1736,1761]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB302433976316B8B4" box="[796,890,1735,1761]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB302433976316B8B4" bold="true" box="[796,890,1735,1761]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 17.2 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB372B339864DFB8B4" box="[1043,1203,1736,1761]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB372B339864DFB8B4" box="[1043,1203,1736,1761]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
</taxonomicName>
|
||
), of different shape (truncated cones in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB32BF33BE6181B952" box="[391,493,1774,1799]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB32BF33BE6181B952" box="[391,493,1774,1799]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB32CC33BE6239B95D" box="[500,597,1774,1800]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB32CC33BE6239B95D" bold="true" box="[500,597,1774,1800]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, cones in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB31EB33BE631BB952" box="[723,887,1774,1799]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB31EB33BE631BB952" box="[723,887,1774,1799]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1305E7ECFFE0FFFB30B133BE63B1B95D" box="[905,989,1774,1800]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="8" pageNumber="53">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="1305E7ECFFE0FFFB372233BE6446B95D" box="[1050,1066,1774,1800]" captionStart="FIGURE 6. A" captionStartId="9.[151,256,1463,1487]" captionTargetBox="[164,1426,281,1447]" captionTargetId="figure@9.[151,1436,271,1455]" captionTargetPageId="9" captionText="FIGURE 6. A, buccal-pharyngeal apparatus of Macrobiotus vanescens. B, claws of Macrobiotus priviterae. C, egg of Macrobiotus danielisae. D, egg of Macrobiotus lorenae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186106/files/figure.png" pageId="8" pageNumber="53">6</figureCitation>
|
||
C), and with the height clearly shorter than the basal diameter while in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB31463245634EB97B" box="[638,802,1813,1838]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB31463245634EB97B" box="[638,802,1813,1838]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
</taxonomicName>
|
||
the height is similar to the basal diameter (the basal diameter is 22.0–23.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB32D1326C6226B900" box="[489,586,1852,1877]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB32D1326C6226B900" box="[489,586,1852,1877]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB316A326B62C3B900" box="[594,687,1851,1877]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB316A326B62C3B900" bold="true" box="[594,687,1851,1877]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 17.5–20.0 µm in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB30B0326C644BB900" box="[904,1063,1852,1877]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE0FFFB30B0326C644BB900" box="[904,1063,1852,1877]" italics="true" pageId="8" pageNumber="53">M. danielisae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE0FFFB33FD3232630EB99C" blockId="8.[151,1437,1039,2032]" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33FD32326141B92E" box="[197,301,1890,1915]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33FD32326141B92E" box="[197,301,1890,1915]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB320E323161F6B92E" box="[310,410,1889,1915]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB320E323161F6B92E" bold="true" box="[310,410,1889,1915]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE0FFFB317B323262A7B92E" box="[579,715,1890,1915]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="lorenae">
|
||
<emphasis id="B94A277BFFE0FFFB317B323262A7B92E" box="[579,715,1890,1915]" italics="true" pageId="8" pageNumber="53">M. lorenae</emphasis>
|
||
</taxonomicName>
|
||
by having very small cuticular tubercles and by having a different shape of the egg processes (
|
||
<figureCitation id="1305E7ECFFE0FFFB315732D962ABB9F6" box="[623,711,1929,1955]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="8" pageNumber="53">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="1305E7ECFFE0FFFB303D32D9637AB9F6" box="[773,790,1929,1955]" captionStart="FIGURE 6. A" captionStartId="9.[151,256,1463,1487]" captionTargetBox="[164,1426,281,1447]" captionTargetId="figure@9.[151,1436,271,1455]" captionTargetPageId="9" captionText="FIGURE 6. A, buccal-pharyngeal apparatus of Macrobiotus vanescens. B, claws of Macrobiotus priviterae. C, egg of Macrobiotus danielisae. D, egg of Macrobiotus lorenae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186106/files/figure.png" pageId="8" pageNumber="53">6</figureCitation>
|
||
D): truncated cones in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB370F32D964CCB9F7" box="[1079,1184,1929,1954]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB370F32D964CCB9F7" box="[1079,1184,1929,1954]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB379132D86562B9F7" box="[1193,1294,1928,1954]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB379132D86562B9F7" bold="true" box="[1193,1294,1928,1954]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and cones terminating with a long, thin, flexible filament in
|
||
<taxonomicName id="4C3E80EAFFE0FFFB31E432E06337B99C" box="[732,859,1968,1993]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="lorenae">
|
||
<emphasis id="B94A277BFFE0FFFB31E432E06337B99C" box="[732,859,1968,1993]" italics="true" pageId="8" pageNumber="53">M. lorenae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE0FFFA33FD3286645ABE8D" blockId="8.[151,1437,1039,2032]" lastBlockId="9.[151,1436,152,216]" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53">
|
||
<taxonomicName id="4C3E80EAFFE0FFFB33FD3286614AB9BA" box="[197,294,2006,2031]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE0FFFB33FD3286614AB9BA" box="[197,294,2006,2031]" italics="true" pageId="8" pageNumber="53">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE0FFFB3214328661E5B9A5" box="[300,393,2006,2032]" pageId="8" pageNumber="53" rank="species">
|
||
<emphasis id="B94A277BFFE0FFFB3214328661E5B9A5" bold="true" box="[300,393,2006,2032]" pageId="8" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE0FFFB311B328662D2B9BA" box="[547,702,2006,2031]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="53" phylum="Tardigrada" rank="species" species="gerlachae">
|
||
<emphasis id="B94A277BFFE0FFFB311B328662D2B9BA" box="[547,702,2006,2031]" italics="true" pageId="8" pageNumber="53">M. gerlachae</emphasis>
|
||
</taxonomicName>
|
||
by having a cuticle ornamented with tubercles, slightly shorter and stouter claws (
|
||
<figureCitation id="1305E7ECFFE1FFFA324A35C861A8BEE7" box="[370,452,152,178]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="9" pageNumber="54">Figs. 4</figureCitation>
|
||
D, E and 7A), and larger eggs, with processes having a larger basal diameter (22.0 – 23.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA322635EF61ECBE8D" box="[286,384,191,216]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA322635EF61ECBE8D" box="[286,384,191,216]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA32BF35EE6189BE8D" box="[391,485,190,216]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA32BF35EE6189BE8D" bold="true" box="[391,485,190,216]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 11.8–14.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA318435EF6334BE8D" box="[700,856,191,216]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="gerlachae">
|
||
<emphasis id="B94A277BFFE1FFFA318435EF6334BE8D" box="[700,856,191,216]" italics="true" pageId="9" pageNumber="54">M. gerlachae</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="1305E7ECFFE1FFFA304235EE63A7BE8D" box="[890,971,190,216]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="9" pageNumber="54">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="1305E7ECFFE1FFFA373B35EE647FBE8D" box="[1027,1043,190,216]" captionStart="FIGURE 7. A" captionStartId="10.[151,255,1547,1571]" captionTargetBox="[160,1427,205,1511]" captionTargetId="figure@10.[151,1436,194,1523]" captionTargetPageId="10" captionText="FIGURE 7. A, claws on the second and third pairs of legs of Macrobiotus gerlachae. B, egg of Macrobiotus gerlachae. C, claws on the first and second pairs of legs of Macrobiotus corgatensis. D, egg of Macrobiotus corgatensis. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186107/files/figure.png" pageId="9" pageNumber="54">7</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<caption id="DF41ABE1FFE1FFFA33AF30E76386BBA4" httpUri="https://zenodo.org/record/186106/files/figure.png" pageId="9" pageNumber="54" targetBox="[164,1426,281,1447]" targetPageId="9">
|
||
<paragraph id="8B81FB69FFE1FFFA33AF30E76386BBA4" blockId="9.[151,1435,1463,1521]" pageId="9" pageNumber="54">
|
||
<emphasis id="B94A277BFFE1FFFA33AF30E76177BB9A" bold="true" box="[151,283,1463,1487]" pageId="9" pageNumber="54">FIGURE 6.</emphasis>
|
||
A, buccal-pharyngeal apparatus of
|
||
<taxonomicName id="4C3E80EAFFE1FFFA31A730E863F0BB9A" box="[671,924,1464,1487]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="vanescens">
|
||
<emphasis id="B94A277BFFE1FFFA31A730E863F0BB9A" box="[671,924,1464,1487]" italics="true" pageId="9" pageNumber="54">Macrobiotus vanescens</emphasis>
|
||
</taxonomicName>
|
||
. B, claws of
|
||
<emphasis id="B94A277BFFE1FFFA371630E8655CBB9A" box="[1070,1328,1464,1487]" italics="true" pageId="9" pageNumber="54">
|
||
<taxonomicName id="4C3E80EAFFE1FFFA371630E86540BB9A" box="[1070,1324,1464,1487]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="priviterae">Macrobiotus priviterae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
C, egg of
|
||
<taxonomicName id="4C3E80EAFFE1FFFA33AF308A61FEBBA4" box="[151,402,1498,1521]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="danielisae">
|
||
<emphasis id="B94A277BFFE1FFFA33AF308A61FEBBA4" box="[151,402,1498,1521]" italics="true" pageId="9" pageNumber="54">Macrobiotus danielisae</emphasis>
|
||
</taxonomicName>
|
||
. D, egg of
|
||
<taxonomicName id="4C3E80EAFFE1FFFA3132308A6284BBA4" box="[522,744,1498,1521]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="lorenae">
|
||
<emphasis id="B94A277BFFE1FFFA3132308A6284BBA4" box="[522,744,1498,1521]" italics="true" pageId="9" pageNumber="54">Macrobiotus lorenae</emphasis>
|
||
</taxonomicName>
|
||
. (Scale bars = 10.0 µm)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81FB69FFE1FFFA33FD33746528B8DE" blockId="9.[151,1436,1571,2023]" pageId="9" pageNumber="54">
|
||
<taxonomicName id="4C3E80EAFFE1FFFA33FD33746146B868" box="[197,298,1572,1597]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA33FD33746146B868" box="[197,298,1572,1597]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA320B337361F8B868" box="[307,404,1571,1597]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA320B337361F8B868" bold="true" box="[307,404,1571,1597]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE1FFFA310E33746285B868" box="[566,745,1572,1597]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="corgatensis">
|
||
<emphasis id="B94A277BFFE1FFFA310E33746285B868" box="[566,745,1572,1597]" italics="true" pageId="9" pageNumber="54">M. corgatensis</emphasis>
|
||
</taxonomicName>
|
||
by lacking eyes, and by having small cuticular tubercles instead of a very fine punctation, stouter claws (
|
||
<figureCitation id="1305E7ECFFE1FFFA31E6331A635FB831" box="[734,819,1610,1636]" captionStart="FIGURE 4" captionStartId="7.[151,257,1660,1684]" captionTargetBox="[161,1427,205,1631]" captionTargetId="figure@7.[151,1436,194,1637]" captionTargetPageId="7" captionText="FIGURE 4. Macrobiotus halei sp. nov. (holotype). A, detail of the cuticular ornamentation (arrow). B, detail of the ornamentation on a leg where small tubercles and very small dots are visible. C, buccal-pharyngeal apparatus. D, claws of the third pair of legs. E, claws of the fourth pair of legs. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186104/files/figure.png" pageId="9" pageNumber="54">Figs. 4</figureCitation>
|
||
D, E and 7C), and truncated, conical egg processes (
|
||
<figureCitation id="1305E7ECFFE1FFFA33A73321609DB8DE" box="[159,241,1649,1675]" captionStart="FIGURE 5" captionStartId="8.[151,256,931,955]" captionTargetBox="[155,1428,198,902]" captionTargetId="figure@8.[151,1436,194,907]" captionTargetPageId="8" captionText="FIGURE 5. Macrobiotus halei sp. nov. A, egg. B, reticular design on the egg processes. C, ornamentation of the egg areolae. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186105/files/figure.png" pageId="9" pageNumber="54">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="1305E7ECFFE1FFFA32113321615BB8DE" box="[297,311,1649,1675]" captionStart="FIGURE 7. A" captionStartId="10.[151,255,1547,1571]" captionTargetBox="[160,1427,205,1511]" captionTargetId="figure@10.[151,1436,194,1523]" captionTargetPageId="10" captionText="FIGURE 7. A, claws on the second and third pairs of legs of Macrobiotus gerlachae. B, egg of Macrobiotus gerlachae. C, claws on the first and second pairs of legs of Macrobiotus corgatensis. D, egg of Macrobiotus corgatensis. (Scale bars = 10.0 µm)" httpUri="https://zenodo.org/record/186107/files/figure.png" pageId="9" pageNumber="54">7</figureCitation>
|
||
D) that are shorter (11.0–14.0 µm in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA31DF33216326B8DF" box="[743,842,1649,1674]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA31DF33216326B8DF" box="[743,842,1649,1674]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA3068332063C1B8DF" box="[848,941,1648,1674]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA3068332063C1B8DF" bold="true" box="[848,941,1648,1674]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 20.0–25.0 µm in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA37BE33216558B8DF" box="[1158,1332,1649,1674]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="corgatensis">
|
||
<emphasis id="B94A277BFFE1FFFA37BE33216558B8DF" box="[1158,1332,1649,1674]" italics="true" pageId="9" pageNumber="54">M. corgatensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE1FFFA33FD33C864CCB919" blockId="9.[151,1436,1571,2023]" pageId="9" pageNumber="54">
|
||
<taxonomicName id="4C3E80EAFFE1FFFA33FD33C86144B8E4" box="[197,296,1688,1713]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA33FD33C86144B8E4" box="[197,296,1688,1713]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA320833C761E2B8E4" box="[304,398,1687,1713]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA320833C761E2B8E4" bold="true" box="[304,398,1687,1713]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE1FFFA311533C86286B8E4" box="[557,746,1688,1713]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="sklodowskae">
|
||
<emphasis id="B94A277BFFE1FFFA311533C86286B8E4" box="[557,746,1688,1713]" italics="true" pageId="9" pageNumber="54">M. sklodowskae</emphasis>
|
||
</taxonomicName>
|
||
by lacking eyes, having a sculptured cuticle, a longer and wider buccal tube (the buccal tube width is 14.2–19.1 µm (
|
||
<emphasis id="B94A277BFFE1FFFA306C33EF6307B88D" box="[852,875,1727,1752]" italics="true" pageId="9" pageNumber="54">pt</emphasis>
|
||
= 26.5–28.6) in
|
||
<emphasis id="B94A277BFFE1FFFA370933EF6483B88D" box="[1073,1263,1726,1752]" italics="true" pageId="9" pageNumber="54">
|
||
<taxonomicName id="4C3E80EAFFE1FFFA370933EF64F8B88D" box="[1073,1172,1727,1752]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">M. halei</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA37A433EE6483B88D" box="[1180,1263,1726,1752]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA37A433EE6483B88D" bold="true" box="[1180,1263,1726,1752]" pageId="9" pageNumber="54">sp. nov</emphasis>
|
||
</taxonomicNameLabel>
|
||
</emphasis>
|
||
., 7.1–11.9 µm (
|
||
<emphasis id="B94A277BFFE1FFFA339933B560D5B8AB" box="[161,185,1765,1790]" italics="true" pageId="9" pageNumber="54">pt</emphasis>
|
||
= 17.0–20.5) in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA32BF33B56226B8AB" box="[391,586,1765,1790]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="sklodowskae">
|
||
<emphasis id="B94A277BFFE1FFFA32BF33B56226B8AB" box="[391,586,1765,1790]" italics="true" pageId="9" pageNumber="54">M. sklodowskae</emphasis>
|
||
</taxonomicName>
|
||
), more slender claws, and less slender egg processes with their basal diameter larger with respect to the process height (height 11.0–14.0 µm and basal diameter 22.0–23.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA33AF32626095B91E" box="[151,249,1842,1867]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA33AF32626095B91E" box="[151,249,1842,1867]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA323832626131B919" box="[256,349,1842,1868]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA323832626131B919" bold="true" box="[256,349,1842,1868]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; height 15.2–19.0 µm and basal diameter 29.5 µm in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA30EC326264E3B91E" box="[980,1167,1842,1867]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="sklodowskae">
|
||
<emphasis id="B94A277BFFE1FFFA30EC326264E3B91E" box="[980,1167,1842,1867]" italics="true" pageId="9" pageNumber="54">M. sklodowskae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE1FFFA33FD32096538B9B3" blockId="9.[151,1436,1571,2023]" pageId="9" pageNumber="54">
|
||
<taxonomicName id="4C3E80EAFFE1FFFA33FD3209614BB927" box="[197,295,1881,1906]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA33FD3209614BB927" box="[197,295,1881,1906]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA3217320861E2B927" box="[303,398,1880,1906]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA3217320861E2B927" bold="true" box="[303,398,1880,1906]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE1FFFA3113320962D3B927" box="[555,703,1881,1906]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="alekseevi">
|
||
<emphasis id="B94A277BFFE1FFFA3113320962D3B927" box="[555,703,1881,1906]" italics="true" pageId="9" pageNumber="54">M. alekseevi</emphasis>
|
||
</taxonomicName>
|
||
by having a sculptured cuticle, longer stylet supports inserted on the buccal tube in a more caudal position (
|
||
<emphasis id="B94A277BFFE1FFFA319132D062D3B9CC" box="[681,703,1920,1945]" italics="true" pageId="9" pageNumber="54">pt</emphasis>
|
||
= 81.0–
|
||
<quantity id="4CC6568CFFE1FFFA301932D0631DB9CF" box="[801,881,1920,1946]" metricMagnitude="0" metricUnit="m" metricValue="2.10058" pageId="9" pageNumber="54" unit="in" value="82.7">82.7 in</quantity>
|
||
<taxonomicName id="4C3E80EAFFE1FFFA304032D063B5B9CC" box="[888,985,1920,1945]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA304032D063B5B9CC" box="[888,985,1920,1945]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA30D8322F6451B9CC" box="[992,1085,1919,1945]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA30D8322F6451B9CC" bold="true" box="[992,1085,1919,1945]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 78.1–81.0 in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA37DD32D0651AB9CC" box="[1253,1398,1920,1945]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="alekseevi">
|
||
<emphasis id="B94A277BFFE1FFFA37DD32D0651AB9CC" box="[1253,1398,1920,1945]" italics="true" pageId="9" pageNumber="54">M. alekseevi</emphasis>
|
||
</taxonomicName>
|
||
), a wider buccal tube (
|
||
<emphasis id="B94A277BFFE1FFFA324132F761FCB995" box="[377,400,1959,1984]" italics="true" pageId="9" pageNumber="54">pt</emphasis>
|
||
=
|
||
<quantity id="4CC6568CFFE1FFFA328832F66228B995" box="[432,580,1958,1984]" metricMagnitude="-1" metricUnit="m" metricValue="6.9977" metricValueMax="7.2644" metricValueMin="6.731" pageId="9" pageNumber="54" unit="in" value="27.55" valueMax="28.6" valueMin="26.5">26.5–28.6 in</quantity>
|
||
<taxonomicName id="4C3E80EAFFE1FFFA317332F762C2B995" box="[587,686,1959,1984]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA317332F762C2B995" box="[587,686,1959,1984]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94A277BFFE1FFFA318D32F66376B995" bold="true" box="[693,794,1958,1984]" pageId="9" pageNumber="54">
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA318D32F66378B995" box="[693,788,1958,1984]" pageId="9" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
<quantity id="4CC6568CFFE1FFFA301A32F663DAB995" box="[802,950,1958,1984]" metricMagnitude="-1" metricUnit="m" metricValue="4.6863" metricValueMax="5.1562" metricValueMin="4.2164" pageId="9" pageNumber="54" unit="in" value="18.45" valueMax="20.3" valueMin="16.6">16.6–20.3 in</quantity>
|
||
<taxonomicName id="4C3E80EAFFE1FFFA308532F7643CB995" box="[957,1104,1959,1984]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="alekseevi">
|
||
<emphasis id="B94A277BFFE1FFFA308532F7643CB995" box="[957,1104,1959,1984]" italics="true" pageId="9" pageNumber="54">M. alekseevi</emphasis>
|
||
</taxonomicName>
|
||
) and differently-shaped egg processes (in
|
||
<taxonomicName id="4C3E80EAFFE1FFFA320E329D61F4B9B3" box="[310,408,1997,2022]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="54" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE1FFFA320E329D61F4B9B3" box="[310,408,1997,2022]" italics="true" pageId="9" pageNumber="54">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE1FFFA32A7329D6191B9B2" box="[415,509,1997,2023]" pageId="9" pageNumber="54" rank="species">
|
||
<emphasis id="B94A277BFFE1FFFA32A7329D6191B9B2" bold="true" box="[415,509,1997,2023]" pageId="9" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
they are truncated cones without a terminal cap-like vesicular structure).
|
||
</paragraph>
|
||
<caption id="DF41ABE1FFE2FFF933AF335B6163B832" httpUri="https://zenodo.org/record/186107/files/figure.png" pageId="10" pageNumber="55" targetBox="[160,1427,205,1511]" targetPageId="10">
|
||
<paragraph id="8B81FB69FFE2FFF933AF335B6163B832" blockId="10.[151,1436,1547,1639]" pageId="10" pageNumber="55">
|
||
<emphasis id="B94A277BFFE2FFF933AF335B6175B876" bold="true" box="[151,281,1547,1571]" pageId="10" pageNumber="55">FIGURE 7.</emphasis>
|
||
A, claws on the second and third pairs of legs of
|
||
<taxonomicName id="4C3E80EAFFE2FFF93014335B6449B877" box="[812,1061,1547,1570]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="gerlachae">
|
||
<emphasis id="B94A277BFFE2FFF93014335B6449B877" box="[812,1061,1547,1570]" italics="true" pageId="10" pageNumber="55">Macrobiotus gerlachae</emphasis>
|
||
</taxonomicName>
|
||
. B, egg of
|
||
<taxonomicName id="4C3E80EAFFE2FFF937A4335B65FAB877" box="[1180,1430,1547,1570]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="gerlachae">
|
||
<emphasis id="B94A277BFFE2FFF937A4335B65FAB877" box="[1180,1430,1547,1570]" italics="true" pageId="10" pageNumber="55">Macrobiotus gerlachae</emphasis>
|
||
</taxonomicName>
|
||
. C, claws on the first and second pairs of legs of
|
||
<taxonomicName id="4C3E80EAFFE2FFF931AC337E63F7B810" box="[660,923,1582,1605]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="corgatensis">
|
||
<emphasis id="B94A277BFFE2FFF931AC337E63F7B810" box="[660,923,1582,1605]" italics="true" pageId="10" pageNumber="55">Macrobiotus corgatensis</emphasis>
|
||
</taxonomicName>
|
||
. D, egg of
|
||
<taxonomicName id="4C3E80EAFFE2FFF9372A337E6576B810" box="[1042,1306,1582,1605]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="corgatensis">
|
||
<emphasis id="B94A277BFFE2FFF9372A337E6576B810" box="[1042,1306,1582,1605]" italics="true" pageId="10" pageNumber="55">Macrobiotus corgatensis</emphasis>
|
||
</taxonomicName>
|
||
. (Scale bars = 10.0 µm)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81FB69FFE2FFF933FD33CA652BB954" blockId="10.[151,1436,1689,2025]" pageId="10" pageNumber="55">
|
||
<taxonomicName id="4C3E80EAFFE2FFF933FD33CA6144B8E6" box="[197,296,1690,1715]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE2FFF933FD33CA6144B8E6" box="[197,296,1690,1715]" italics="true" pageId="10" pageNumber="55">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE2FFF9321733C961E2B8E6" box="[303,398,1689,1715]" pageId="10" pageNumber="55" rank="species">
|
||
<emphasis id="B94A277BFFE2FFF9321733C961E2B8E6" bold="true" box="[303,398,1689,1715]" pageId="10" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE2FFF9311333CA62D1B8E6" box="[555,701,1690,1715]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="garynhai">
|
||
<emphasis id="B94A277BFFE2FFF9311333CA62D1B8E6" box="[555,701,1690,1715]" italics="true" pageId="10" pageNumber="55">M. garynhai</emphasis>
|
||
</taxonomicName>
|
||
by having cuticular tubercles and lacking elliptical pores, and by having egg processes that are truncated cones without a cap-like terminal portion, and by the egg shell areolae with a thickened central portion with circular light spots forming an irregular reticular design).
|
||
</paragraph>
|
||
<paragraph id="8B81FB69FFE2FFF933FD325E615BB920" blockId="10.[151,1436,1689,2025]" pageId="10" pageNumber="55">
|
||
<taxonomicName id="4C3E80EAFFE2FFF933FD325E6145B972" box="[197,297,1806,1831]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE2FFF933FD325E6145B972" box="[197,297,1806,1831]" italics="true" pageId="10" pageNumber="55">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE2FFF9320A325D61FFB972" box="[306,403,1805,1831]" pageId="10" pageNumber="55" rank="species">
|
||
<emphasis id="B94A277BFFE2FFF9320A325D61FFB972" bold="true" box="[306,403,1805,1831]" pageId="10" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3E80EAFFE2FFF9310D325E629DB972" box="[565,753,1806,1831]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="magdalenae">
|
||
<emphasis id="B94A277BFFE2FFF9310D325E629DB972" box="[565,753,1806,1831]" italics="true" pageId="10" pageNumber="55">M. magdalenae</emphasis>
|
||
</taxonomicName>
|
||
by lacking eyes, having a sculptured cuticle, and by the shape of the egg processes (truncated cones in
|
||
<taxonomicName id="4C3E80EAFFE2FFF931FE32656345B91B" box="[710,809,1845,1870]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE2FFF931FE32656345B91B" box="[710,809,1845,1870]" italics="true" pageId="10" pageNumber="55">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE2FFF93009326463FCB91B" box="[817,912,1844,1870]" pageId="10" pageNumber="55" rank="species">
|
||
<emphasis id="B94A277BFFE2FFF93009326463FCB91B" bold="true" box="[817,912,1844,1870]" pageId="10" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, cones with elongate terminal portion in
|
||
<taxonomicName id="4C3E80EAFFE2FFF936443265614AB921" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="magdalenae">
|
||
<emphasis id="B94A277BFFE2FFF936443265614AB921" italics="true" pageId="10" pageNumber="55">M. magdalenae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324A8E2FFE2FFF833FD32D16562BE8D" lastPageId="11" lastPageNumber="56" pageId="10" pageNumber="55" type="discussion">
|
||
<paragraph id="8B81FB69FFE2FFF833FD32D16562BE8D" blockId="10.[151,1436,1689,2025]" lastBlockId="11.[151,1435,151,216]" lastPageId="11" lastPageNumber="56" pageId="10" pageNumber="55">
|
||
<emphasis id="B94A277BFFE2FFF933FD32D16136B9CE" bold="true" box="[197,346,1921,1947]" pageId="10" pageNumber="55">Comments:</emphasis>
|
||
Besides the
|
||
<typeStatus id="548545CBFFE2FFF932C132D26208B9C9" box="[505,612,1922,1948]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="548545CBFFE2FFF9319932D2637AB9C9" box="[673,790,1922,1948]" pageId="10" pageNumber="55" type="paratype">paratypes</typeStatus>
|
||
used for the description of the species in this paper, additional specimens of
|
||
<taxonomicName id="4C3E80EAFFE2FFF9328C32F962E6B997" box="[436,650,1961,1986]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="55" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE2FFF9328C32F962E6B997" box="[436,650,1961,1986]" italics="true" pageId="10" pageNumber="55">Macrobiotus halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE2FFF931AA32F86283B997" box="[658,751,1960,1986]" pageId="10" pageNumber="55" rank="species">
|
||
<emphasis id="B94A277BFFE2FFF931AA32F86283B997" bold="true" box="[658,751,1960,1986]" pageId="10" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have been identified within the park. The total is now
|
||
<specimenCount id="9D3830E0FFE2FFF9364832F86179B9BC" pageId="10" pageNumber="55" type="generic">361 specimens</specimenCount>
|
||
, including
|
||
<specimenCount id="9D3830E0FFE2FFF932A0329F619BB9BC" box="[408,503,1999,2025]" pageId="10" pageNumber="55" type="egg">13 eggs</specimenCount>
|
||
, although we expect to find many more as we sort through additional slides of the
|
||
<taxonomicName id="4C3E80EAFFE3FFF833FA35C86120BEE4" box="[194,332,152,177]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="56" phylum="Tardigrada" rank="species" species="richtersi">
|
||
<emphasis id="B94A277BFFE3FFF833FA35C86120BEE4" box="[194,332,152,177]" italics="true" pageId="11" pageNumber="56">M. richtersi</emphasis>
|
||
</taxonomicName>
|
||
group. So far, all
|
||
<taxonomicName id="4C3E80EAFFE3FFF8311835C862EEBEE4" box="[544,642,152,177]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="56" phylum="Tardigrada" rank="species" species="halei" status="sp. nov.">
|
||
<emphasis id="B94A277BFFE3FFF8311835C862EEBEE4" box="[544,642,152,177]" italics="true" pageId="11" pageNumber="56">M. halei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2799A00FFE3FFF831B135C7628BBEE4" box="[649,743,151,177]" pageId="11" pageNumber="56" rank="species">
|
||
<emphasis id="B94A277BFFE3FFF831B135C7628BBEE4" bold="true" box="[649,743,151,177]" pageId="11" pageNumber="56">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
specimens have been found in mosses and lichens on trees and rocks and also frequently in soil and leaf litter. None have been found in our aquatic samples.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |