163 lines
17 KiB
XML
163 lines
17 KiB
XML
<document id="017BE6F8DF265EFD75683C463AE0CE3E" ID-DOI="10.5281/zenodo.158466" ID-GBIF-Dataset="d5d8bbf6-677e-4341-923a-60deb8e2fa0d" ID-ISSN="11755326" ID-Zenodo-Dep="158466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461037551613" checkinUser="plazi" docAuthor="Greathead, David J. & Evenhuis, Neal L." docDate="2004" docId="8B54AF764C6AFF9CFED9FA456C39F9AA" docLanguage="en" docName="zt00773.pdf" docOrigin="Zootaxa 773" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Empidideicus completus Bezzi 1926" docType="treatment" docVersion="11" lastPageNumber="14" masterDocId="776DD70E4C66FF91FFD1FFE26F70FFBC" masterDocTitle="New species of Bombylioidea in Mario Bezzi’s Unpublished Hungarian Museum Manuscript" masterLastPageNumber="56" masterPageNumber="1" pageNumber="13" updateTime="1698574864920" updateUser="plazi">
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<mods:titleInfo id="7CB00FD6B8F532C763CA436A7DF622A2">
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<mods:title id="E0F9F80B95C213FA51963DAA9A88A885">New species of Bombylioidea in Mario Bezzi’s Unpublished Hungarian Museum Manuscript</mods:title>
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</mods:titleInfo>
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<mods:name id="1BBCA0A95954BD39BD0D1793CE836B66" type="personal">
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<mods:roleTerm id="A1A54BFD81293552209059FEDC05B90F">Author</mods:roleTerm>
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<mods:namePart id="6B3492DF272FA678965DA67A705E3055">Greathead, David J.</mods:namePart>
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<mods:name id="7958A9C032A57A4ED2E3A47DB9B44FFF" type="personal">
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<mods:roleTerm id="DE8B33591928895D55291234CCF11637">Author</mods:roleTerm>
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<mods:namePart id="2B3B1B8E4533C13E159BC9475D2C05A2">Evenhuis, Neal L.</mods:namePart>
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<mods:typeOfResource id="388FBCA9F63D14E7D9AFAF7C7E1036DE">text</mods:typeOfResource>
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<mods:title id="3FCA11F919B8AC6E8408D010CB499336">Zootaxa</mods:title>
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<mods:part id="28D96A88627EF30FD872195E06EAE359">
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<mods:date id="587794BF35B0A07FAD14DD07A8135C5E">2004</mods:date>
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<mods:detail id="F870E4AA436F4EDF433B5B81199703AA" type="volume">
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<mods:number id="5A088E5F8CAD51C607C7C36252407CED">773</mods:number>
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<mods:start id="30182A59D7C1A948A22EFFAF4F580EC3">1</mods:start>
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<mods:end id="7DCFA68ECEE592684CA4054A91FEC664">56</mods:end>
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<mods:classification id="77340ABE0EB9A9D3A2AD47D333286E0A">journal article</mods:classification>
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<mods:identifier id="3CF826380587BABC07484C220C8EF754" type="DOI">10.5281/zenodo.158466</mods:identifier>
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<mods:identifier id="B748853648F5AA1F898F448E09FB7888" type="Zenodo-Dep">158466</mods:identifier>
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<treatment id="8B54AF764C6AFF9CFED9FA456C39F9AA" ID-DOI="http://doi.org/10.5281/zenodo.6270378" ID-GBIF-Taxon="119609310" ID-Zenodo-Dep="6270378" LSID="urn:lsid:plazi:treatment:8B54AF764C6AFF9CFED9FA456C39F9AA" httpUri="http://treatment.plazi.org/id/8B54AF764C6AFF9CFED9FA456C39F9AA" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<subSubSection id="4BE74DEB4C6AFF9DFED9FA456BF5F9BB" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="03421E604C6AFF9DFED9FA456DB2FA7D" blockId="12.[264,706,1447,1473]" box="[264,706,1447,1473]" pageId="12" pageNumber="13">
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<heading id="580AA90C4C6AFF9DFED9FA456DB2FA7D" bold="true" box="[264,706,1447,1473]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<taxonomicName id="C4FD65E34C6AFF9DFED9FA456DB2FA7D" authority="Bezzi, 1926" authorityName="Bezzi" authorityYear="1926" box="[264,706,1447,1473]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="completus">
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<emphasis id="3189C2724C6AFF9DFED9FA456DB2FA7D" bold="true" box="[264,706,1447,1473]" pageId="12" pageNumber="13">
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<emphasis id="3189C2724C6AFF9DFED9FA456D5DFA7D" bold="true" box="[264,557,1447,1473]" italics="true" pageId="12" pageNumber="13">Empidideicus completus</emphasis>
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Bezzi, 1926
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="03421E604C6AFF9DFED9FA0F6BF5F9BB" blockId="12.[264,1324,1516,1793]" box="[264,1157,1516,1543]" pageId="12" pageNumber="13">
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26.
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<taxonomicName id="C4FD65E34C6AFF9DFEE5FA0F6D22F9BA" box="[308,594,1517,1542]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="completus">
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<emphasis id="3189C2724C6AFF9DFEE5FA0F6D22F9BA" box="[308,594,1517,1542]" italics="true" pageId="12" pageNumber="13">Empidideicus completus</emphasis>
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</taxonomicName>
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1Ψ
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<collectingCountry id="7BEA5EF04C6AFF9DFD52FA0F6DA7F9BB" box="[643,727,1517,1543]" name="Eritrea" pageId="12" pageNumber="13">Eritrea</collectingCountry>
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: Assab, 1907 (Katona). MS page 23.
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</paragraph>
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</subSubSection>
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<subSubSection id="4BE74DEB4C6AFF9DFEE7F9F96B14F8BD" pageId="12" pageNumber="13" type="description">
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<paragraph id="03421E604C6AFF9DFEE7F9F96C65F9D1" blockId="12.[264,1324,1516,1793]" pageId="12" pageNumber="13">
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<emphasis id="3189C2724C6AFF9DFEE7F9F96EC6F98F" bold="true" box="[310,438,1563,1587]" pageId="12" pageNumber="13">Bezzi, 1926</emphasis>
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: 254 – included in a key to species of
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<taxonomicName id="C4FD65E34C6AFF9DFC87F9F96CA1F98E" box="[854,977,1563,1586]" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Cyrtosiinae">Cyrtosiinae</taxonomicName>
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[=
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<taxonomicName id="C4FD65E34C6AFF9DFC26F9F96BDBF98E" box="[1015,1195,1563,1586]" class="Insecta" family="Mythicomyiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Mythicomyiidae</taxonomicName>
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] and a footnote: ‘espèce d’Assab, appartennant au Musée de Budapest’, validating the species and indicating the
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<typeStatus id="DC46A0C24C6AFF9DFE2EF9B46D5DF9D1" box="[511,557,1622,1645]" pageId="12" pageNumber="13">type</typeStatus>
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deposited in HNHM.
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</paragraph>
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<paragraph id="03421E604C6AFF9DFEE7F9966EA3F914" blockId="12.[264,1324,1516,1793]" pageId="12" pageNumber="13">
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Hesse, 1967: 108 – placed it (calling it an Egyptian [sic] species) in the subgenus
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<taxonomicName id="C4FD65E34C6AFF9DFB74F9966EF5F914" class="Insecta" family="Mythicomyiidae" genus="Anomaloptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="3189C2724C6AFF9DFB74F9966EF5F914" italics="true" pageId="12" pageNumber="13">Anomaloptilus</emphasis>
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</taxonomicName>
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Hesse.
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</paragraph>
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<paragraph id="03421E604C6AFF9DFEE9F94C6CE7F95F" blockId="12.[264,1324,1516,1793]" pageId="12" pageNumber="13">
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Bowden, 1980: 383 – treated
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<taxonomicName id="C4FD65E34C6AFF9DFDA5F94D6C7FF97A" box="[628,783,1711,1734]" class="Insecta" family="Mythicomyiidae" genus="Anomaloptilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="3189C2724C6AFF9DFDA5F94D6C7FF97A" box="[628,783,1711,1734]" italics="true" pageId="12" pageNumber="13">Anomaloptilus</emphasis>
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</taxonomicName>
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as a junior synonym of
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<taxonomicName id="C4FD65E34C6AFF9DFBC0F94D6BD2F97A" box="[1041,1186,1711,1734]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="3189C2724C6AFF9DFBC0F94D6BD2F97A" box="[1041,1186,1711,1734]" italics="true" pageId="12" pageNumber="13">Empidideicus</emphasis>
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</taxonomicName>
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. Evenhuis, 2002: 32 –
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<typeStatus id="DC46A0C24C6AFF9DFDF0F92E6D3FF95F" box="[545,591,1740,1763]" pageId="12" pageNumber="13">type</typeStatus>
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in HNHM noted as destroyed.
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</paragraph>
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<paragraph id="03421E604C6AFF9DFEE9F90B6B14F8BD" blockId="12.[264,1324,1516,1793]" box="[312,1124,1769,1793]" pageId="12" pageNumber="13">
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<emphasis id="3189C2724C6AFF9DFEE9F90B6E08F8BD" bold="true" box="[312,376,1769,1793]" pageId="12" pageNumber="13">
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<typeStatus id="DC46A0C24C6AFF9DFEE9F90B6E08F8BD" box="[312,376,1769,1793]" pageId="12" pageNumber="13">Types</typeStatus>
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</emphasis>
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: No specimens found.
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<typeStatus id="DC46A0C24C6AFF9DFDA0F9086DA4F8BD" box="[625,724,1770,1793]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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female in HNHM destroyed in 1956.
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</paragraph>
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</subSubSection>
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<subSubSection id="4BE74DEB4C6AFF9CFED9F8C66C39F9AA" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
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<paragraph id="03421E604C6AFF9CFED9F8C66CB5FEE2" blockId="12.[264,1323,1828,1894]" lastBlockId="13.[264,1323,284,550]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<emphasis id="3189C2724C6AFF9DFED9F8C66E0AF882" bold="true" box="[264,378,1828,1854]" pageId="12" pageNumber="13">Remarks</emphasis>
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. The species was based on a single female in HNHM that was destroyed in 1956. No further specimens have been found from
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<collectingCountry id="7BEA5EF04C6AFF9DFCA2F8AE6CB3F8DA" box="[883,963,1868,1894]" name="Eritrea" pageId="12" pageNumber="13">Eritrea</collectingCountry>
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or nearby localities including
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<collectingCountry id="7BEA5EF04C6BFF9CFED9FEFE6E29FE8A" box="[264,345,284,310]" name="Yemen" pageId="13" pageNumber="14">Yemen</collectingCountry>
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that fit the characters of this species. It is hereby relegated to
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<emphasis id="3189C2724C6BFF9CFBEFFEFE6B9CFE89" box="[1086,1260,284,309]" italics="true" pageId="13" pageNumber="14">nomen dubium</emphasis>
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until such time as specimens fitting the description can be found.
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</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FE8E6D9FFE3A" blockId="13.[264,1323,284,550]" box="[312,751,364,390]" pageId="13" pageNumber="14">Bezzi’s Latin diagnosis is given here:</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FE766D27FE42" blockId="13.[264,1323,284,550]" pageId="13" pageNumber="14">“Pallide luteus, pedibus concoloribus, proboscide antennisque nigris, thoracis disco abdominisque segmentorum basi nigris opacis, alis albidohyalinis, nervis decolaribus, cellula discoidali completa”.</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FDEE6CFDFD9A" blockId="13.[264,1323,284,550]" box="[312,909,524,550]" pageId="13" pageNumber="14">Bezzi’s full description (translated from the Latin):</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FDBE6DBFFDCA" blockId="13.[312,719,604,630]" box="[312,719,604,630]" pageId="13" pageNumber="14">Length of body and wing. 1.3 mm.</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FD4F6C30FBF8" blockId="13.[312,1274,685,1092]" pageId="13" pageNumber="14">“Head pale yellow, occiput opaque black, frons bare, parallel laterally; third of head lateral part equal; face bare horizontal [= prognathus], eyes round, large. Antennae inserted in a straight line between the vertex and oral margin, all black, bases approximate, short, basal segments minute, third short, ovate, truncated, style strong placed terminal. Proboscis strong, thick, sharp, black, directed backward, underside longer. Thorax and remainder of body bare, opaque black dorsally; humeri, lateral line, crescentshaped spot above humeri, and other small sutures yellow; pleurae all ochraceous, blackish markings; below black. Scutellum ochraceous, dark black basomedially, black below. Halter large pale yellow, club white. Squama indistinct. Abdomen white, first four segments with black basal fasciae. Legs all yellowishwhite, unarmed and bare; femora well thickened; claws and pulvilli small but distinct. Wing whitish hyaline, without spots, veins pale; bases of first, third and transverse veins thick, remainder very tenuous difficult to distinguish. Inferior branch of fourth vein with upper branch much from transverse scarcely connected so that discoidal cell from second posterior distinctly placed.”</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FB8E6A5BFA9A" blockId="13.[264,1324,1132,1558]" pageId="13" pageNumber="14">
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Bezzi manuscript has a key to three species from the “Ethiopian Region” that he placed in
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<taxonomicName id="C4FD65E34C6BFF9CFEABFB776D69FB12" box="[378,537,1173,1198]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="3189C2724C6BFF9CFEABFB776D69FB12" box="[378,537,1173,1198]" italics="true" pageId="13" pageNumber="14">Empidideicus</emphasis>
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</taxonomicName>
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:
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<taxonomicName id="C4FD65E34C6BFF9CFDF8FB776D9AFB12" authority="Bezzi" authorityName="Bezzi" box="[553,746,1172,1198]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="beckeri">
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<emphasis id="3189C2724C6BFF9CFDF8FB776DD0FB12" box="[553,672,1173,1198]" italics="true" pageId="13" pageNumber="14">E. beckeri</emphasis>
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Bezzi
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</taxonomicName>
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(now in
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<taxonomicName id="C4FD65E34C6BFF9CFC82FB776B7AFB12" box="[851,1034,1173,1198]" class="Insecta" family="Bombyliidae" genus="Cephalodromia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="3189C2724C6BFF9CFC82FB776B7AFB12" box="[851,1034,1173,1198]" italics="true" pageId="13" pageNumber="14">Cephalodromia</emphasis>
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</taxonomicName>
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),
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<taxonomicName id="C4FD65E34C6BFF9CFBF2FB776B97FB12" authority="Bezzi" authorityName="Bezzi" box="[1059,1255,1172,1198]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="melleus">
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<emphasis id="3189C2724C6BFF9CFBF2FB776B45FB12" box="[1059,1077,1173,1198]" italics="true" pageId="13" pageNumber="14">E</emphasis>
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.
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<emphasis id="3189C2724C6BFF9CFB92FB776BEDFB12" box="[1091,1181,1173,1198]" italics="true" pageId="13" pageNumber="14">melleus</emphasis>
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Bezzi
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</taxonomicName>
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(now in
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<taxonomicName id="C4FD65E34C6BFF9CFEFBFB5F6ECBFB6A" box="[298,443,1213,1238]" class="Insecta" family="Bombyliidae" genus="Mnemomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="3189C2724C6BFF9CFEFBFB5F6ECBFB6A" box="[298,443,1213,1238]" italics="true" pageId="13" pageNumber="14">Mnemomyia</emphasis>
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</taxonomicName>
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), and
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<taxonomicName id="C4FD65E34C6BFF9CFDDCFB5F6D8EFB6A" authority="Bezzi." authorityName="Bezzi." box="[525,766,1212,1238]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="completus">
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<emphasis id="3189C2724C6BFF9CFDDCFB5F6DD8FB6A" box="[525,680,1213,1238]" italics="true" pageId="13" pageNumber="14">E. completus</emphasis>
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Bezzi.
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</taxonomicName>
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<taxonomicName id="C4FD65E34C6BFF9CFCD9FB5F6B5AFB6A" box="[776,1066,1213,1238]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="completus">
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<emphasis id="3189C2724C6BFF9CFCD9FB5F6B5AFB6A" box="[776,1066,1213,1238]" italics="true" pageId="13" pageNumber="14">Empidideicus completus</emphasis>
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</taxonomicName>
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was characterised in that key by having a complete discoidal cell (cell d), a yellow thorax with matte black dorsally, and an abdomen that was white with black spots on the basal portions of the tergites.
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</paragraph>
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<paragraph id="03421E604C6BFF9CFEE9FAD66C39F9AA" blockId="13.[264,1324,1132,1558]" pageId="13" pageNumber="14">
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These combinations of characters are found in a number of specimens from the area near the
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<typeStatus id="DC46A0C24C6BFF9CFEBEFABE6ED2FACA" box="[367,418,1372,1398]" pageId="13" pageNumber="14">type</typeStatus>
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locality. However, the specific characters mentioned in the description, specifically, the scutellum being yellowish [Bezzi’s “
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<emphasis id="3189C2724C6BFF9CFC8EFA676CCFFA22" box="[863,959,1413,1438]" italics="true" pageId="13" pageNumber="14">luteum”</emphasis>
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= ochraceous or mudyellow] with matte black basomedially in combination with a yellow mesonotum with opaque black dorsally cannot be found in any described species nor in any undetermined material from Africa or the Middle East currently at hand.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |