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<document id="AC5B5C68DE957CFE066C96D05A54EAE2" ID-ISSN="0024-4082" ID-ZooBank="DB1E4C0F-C529-4F51-973E-D8ED6D84DDFD" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630098224661" checkinUser="felipe" docAuthor="Criscione, Francesco, Hallan, Anders, Puillandre, Nicolas &amp; Fedosov, Alexander" docDate="2021" docId="8B48E757FFB3F84DFC50FCCBF93439E3" docLanguage="en" docName="zlaa088.pdf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Austrobela rufa Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov 2021, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="984" masterDocId="77719F2FFFA5F85AFFE7FF91FD293C5B" masterDocTitle="Where the snails have no name: a molecular phylogeny of Raphitomidae (Neogastropoda: Conoidea) uncovers vast unexplored diversity in the deep seas of temperate southern and eastern Australia" masterLastPageNumber="1000" masterPageNumber="961" pageNumber="983" updateTime="1699144688566" updateUser="plazi">
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<mods:title id="884919282E60D2B31B6E8E22C64CE799">Where the snails have no name: a molecular phylogeny of Raphitomidae (Neogastropoda: Conoidea) uncovers vast unexplored diversity in the deep seas of temperate southern and eastern Australia</mods:title>
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<mods:namePart id="1105CD050851526233A588A931503BCD">Criscione, Francesco</mods:namePart>
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<mods:namePart id="FC1149266E4011615FCD2DA35D36242C">Hallan, Anders</mods:namePart>
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<mods:namePart id="4032BA0CE12B75AD9CF8F1B83418683B">Puillandre, Nicolas</mods:namePart>
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<mods:namePart id="06D8CC89AE14101A29D9E65D619F6EFC">Fedosov, Alexander</mods:namePart>
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<mods:title id="EEA1FE7A9C994FEC27F114AE113E59A5">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="F08FFAB63CF331E10361C43A0BD479A5">2021</mods:date>
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<treatment id="8B48E757FFB3F84DFC50FCCBF93439E3" LSID="urn:lsid:plazi:treatment:8B48E757FFB3F84DFC50FCCBF93439E3" httpUri="http://treatment.plazi.org/id/8B48E757FFB3F84DFC50FCCBF93439E3" lastPageId="23" lastPageNumber="984" pageId="22" pageNumber="983">
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<paragraph id="035E5641FFB3F84CFC50FCCBF82B3F29" blockId="22.[951,1282,858,882]" box="[951,1282,858,882]" pageId="22" pageNumber="983">
<heading id="5816E12DFFB3F84CFC50FCCBF82B3F29" box="[951,1282,858,882]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="983" reason="2">
<taxonomicName id="C4E12DC2FFB3F84CFC50FCCBF9B13F2A" ID-CoL="7Q977" authority="Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov, 2021" authorityName="Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov" authorityYear="2021" box="[951,1176,858,882]" class="Gastropoda" family="Raphitomidae" genus="Austrobela" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="986" phylum="Mollusca" rank="species" species="rufa" status="sp. nov.">AUSTROBELA RUFA</taxonomicName>
<emphasis id="31958A53FFB3F84CFB78FCCCF82B3F29" bold="true" box="[1183,1282,858,882]" pageId="22" pageNumber="983">
<taxonomicNameLabel id="2AA63728FFB3F84CFB78FCCCF82B3F29" box="[1183,1282,858,882]" pageId="22" pageNumber="983" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="4BFB05CAFFB3F84DFC1DFC18F93439E3" lastPageId="23" lastPageNumber="984" pageId="22" pageNumber="983" type="description">
<paragraph id="035E5641FFB3F84CFC1DFC18F9963FF9" blockId="22.[1018,1215,905,930]" box="[1018,1215,905,930]" pageId="22" pageNumber="983">
(
<figureCitation id="9BDA4AC4FFB3F84CFBE5FC18F9723FF9" box="[1026,1115,905,930]" captionStart="Figure 3" captionStartId="15.[163,241,1570,1592]" captionTargetBox="[163,1443,195,1530]" captionTargetId="figure-87@15.[163,1443,195,1530]" captionTargetPageId="15" captionText="Figure 3. Shells of species described herein. A, Notothauma magnifica, holotype AMS C.532707; B, Fusobela parvioculata, holotype TMAG E59231; C, Aplotoma brevitentaculata, holotype AMS C.571635; D, Biconitoma cretosa, holotype AMS C.482313. E, Pagodibela maia, holotype AMS C.571678; F, Austrobela rufa, holotype AMS C.571709; G, Glaciotomella investigator, holotypeAMSC.571621;H, Austrotheta crassidentata, holotype AMSC.519302.Scale bar20 mm (A), 10 mm (B–H)." pageId="22" pageNumber="983">FIGS 3F</figureCitation>
,
<figureCitation id="9BDA4AC4FFB3F84CFB8FFC18F9A13FF9" box="[1128,1160,905,930]" captionStart="Figure 5" captionStartId="17.[163,243,1339,1361]" captionTargetBox="[163,1443,195,1299]" captionTargetId="figure-221@17.[163,1443,195,1299]" captionTargetPageId="17" captionText="Figure 5. Protoconchs of species described herein. A, Trochodaphne cuprosa, holotype AMS C.571611; B, Globodaphne pomum, holotype AMS C.482283; C, Pagodibela maia, holotype AMS C.571678; D, Pueridaphne cirrisulcata, paratype AMS C.563103; E, Austrobela rufa, paratype AMS C. 571681. Scale bar = 500 μm." pageId="22" pageNumber="983">5E</figureCitation>
,
<figureCitation id="9BDA4AC4FFB3F84CFB72FC18F9913FF9" box="[1173,1208,905,930]" captionStart="Figure 6" captionStartId="18.[145,227,1378,1400]" captionTargetBox="[145,1425,195,1338]" captionTargetId="figure-24@18.[145,1425,195,1338]" captionTargetPageId="18" captionText="Figure 6. Hypodermic teeth of species described herein. A, Aplotoma brevitentaculata, holotype AMS C.571635; B, Gladiobela angulata, paratype AMS C.571737; C, Pueridaphne cirrisulcata, paratype AMS C.571609; D: Austrobela rufa, holotype AMS C.571709 (left), paratype AMS C.571679 (right); E, Pagodibela maia, holotype AMS C.571678; F, Fusobela parvioculata, holotype TMAG E59231; G, Biconitoma cretosa, holotype AMS C.482313; H, Austrotheta crassidentata, holotype AMS C.519302. Scale bars = 50 μm." pageId="22" pageNumber="983">6D</figureCitation>
)
</paragraph>
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<emphasis id="31958A53FFB3F84CFCCEFC43F94C3FBC" box="[809,1125,977,999]" italics="true" pageId="22" pageNumber="983">Z o o B a n k r e g i s t r a t i o n</emphasis>
: u r n: l s i d: z o o b a n k. org:act:
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, IN2015_C02_134, (
<geoCoordinate id="66D53086FFB3F84CFAD7FA2FF8A4398F" box="[1328,1421,1470,1492]" degrees="35.345" direction="south" orientation="latitude" pageId="22" pageNumber="983" precision="55" value="-35.345">–35.345</geoCoordinate>
,
<geoCoordinate id="66D53086FFB3F84CFCCEFA4CFEA239A8" box="[809,907,1501,1523]" degrees="134.045" direction="east" orientation="longitude" pageId="22" pageNumber="983" precision="55" value="134.045">134.045</geoCoordinate>
), one wet (
<collectionCode id="65F0CE84FFB3F84CFBF5FA4CF96739A9" box="[1042,1102,1501,1522]" pageId="22" pageNumber="983">AMS</collectionCode>
C.532691); one wet (
<collectionCode id="65F0CE84FFB3F84CFAB3FA4CF8A639A9" box="[1364,1423,1501,1522]" pageId="22" pageNumber="983">AMS</collectionCode>
C.571699)
</materialsCitation>
.
<materialsCitation id="B3895C1CFFB3F84CFC4CFA6DF9ED3A14" collectionCode="AMS" country="Australia" elevation="1127" latitude="-41.209" location="St Helens" longLatPrecision="69" longitude="148.797" pageId="22" pageNumber="983" specimenCode="V06" specimenCount="1" stateProvince="Tasmania" typeStatus="paratype">
<collectingCountry id="7BF616D1FFB3F84CFC4CFA6DF9323A49" box="[939,1051,1532,1554]" name="Australia" pageId="22" pageNumber="983">Australia</collectingCountry>
,
<collectingRegion id="C12598A3FFB3F84CFBC0FA6DF9B53A49" box="[1063,1180,1532,1554]" country="Australia" name="Tasmania" pageId="22" pageNumber="983">Tasmania</collectingRegion>
,
<location id="063E009AFFB3F84CFB4FFA6DF8333A49" LSID="urn:lsid:plazi:treatment:8B48E757FFB3F84DFC50FCCBF93439E3:063E009AFFB3F84CFB4FFA6DF8333A49" box="[1192,1306,1532,1554]" country="Australia" latitude="-41.209" longLatPrecision="69" longitude="148.797" name="St Helens" pageId="22" pageNumber="983" stateProvince="Tasmania">St Helens</location>
flat,
<quantity id="C419FBA4FFB3F84CFABEFA6DFE683A6B" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="22" pageNumber="983" unit="m" value="1127.0">
<elevation id="88CCB172FFB3F84CFABEFA6DFE683A6B" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="22" pageNumber="983" unit="m" value="1127.0">1127 m</elevation>
</quantity>
, IN2018_
<specimenCode id="5347FE3AFFB3F84CFC4AF98BFEF23A6B" box="[941,987,1562,1584]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="983" type="Museum">V06</specimenCode>
_184, (
<geoCoordinate id="66D53086FFB3F84CFBC6F98AF9573A6B" box="[1057,1150,1563,1584]" degrees="41.209" direction="south" orientation="latitude" pageId="22" pageNumber="983" precision="55" value="-41.209">–41.209</geoCoordinate>
,
<geoCoordinate id="66D53086FFB3F84CFB60F98AF9CA3A6A" box="[1159,1251,1563,1585]" degrees="148.797" direction="east" orientation="longitude" pageId="22" pageNumber="983" precision="55" value="148.797">148.797</geoCoordinate>
), one wet (
<collectionCode id="65F0CE84FFB3F84CFAB0F98AF8B83A6B" box="[1367,1425,1563,1584]" pageId="22" pageNumber="983">AMS</collectionCode>
C.574588); one wet (
<collectionCode id="65F0CE84FFB3F84CFBE9F9A8F9613A15" box="[1038,1096,1593,1614]" pageId="22" pageNumber="983">AMS</collectionCode>
C.271201)
</materialsCitation>
.
</paragraph>
<paragraph id="035E5641FFB3F84CFCCEF9E9FE943A90" blockId="22.[809,1425,1656,1739]" pageId="22" pageNumber="983">
<emphasis id="31958A53FFB3F84CFCCEF9E9FE923AD6" box="[809,955,1656,1677]" italics="true" pageId="22" pageNumber="983">Distribution:</emphasis>
Known from the Great Australian Bight, northern
<collectingRegion id="C12598A3FFB3F84CFC7BF907F93B3AF7" box="[924,1042,1686,1708]" country="Australia" name="Tasmania" pageId="22" pageNumber="983">Tasmania</collectingRegion>
and the southern coast of
<collectingRegion id="C12598A3FFB3F84CFABCF906FE903A90" country="Australia" name="New South Wales" pageId="22" pageNumber="983">New South Wales</collectingRegion>
.
</paragraph>
<paragraph id="035E5641FFB3F84CFCCEF965FEA33B1C" blockId="22.[809,1425,1780,1863]" pageId="22" pageNumber="983">
<emphasis id="31958A53FFB3F84CFCCEF965FE863B52" box="[809,943,1780,1801]" italics="true" pageId="22" pageNumber="983">Etymology:</emphasis>
The epithet is derived from the Latin adjective
<emphasis id="31958A53FFB3F84CFC7DF883FEF13B7C" box="[922,984,1810,1831]" italics="true" pageId="22" pageNumber="983">rufus</emphasis>
, red, in reference to the coloration of its shell.
</paragraph>
<paragraph id="035E5641FFB2F84DFF44FF57FC063C86" blockId="23.[163,303,198,221]" box="[163,303,198,221]" pageId="23" pageNumber="984">
<emphasis id="31958A53FFB2F84DFF44FF57FC063C86" box="[163,303,198,221]" italics="true" pageId="23" pageNumber="984">Description</emphasis>
</paragraph>
<paragraph id="035E5641FFB2F84DFF44FF7CFF043FB8" blockId="23.[163,780,237,1884]" pageId="23" pageNumber="984">
Shell (
<figureCitation id="9BDA4AC4FFB2F84DFF13FF7CFC623D59" box="[244,331,237,259]" captionStart="Figure 3" captionStartId="15.[163,241,1570,1592]" captionTargetBox="[163,1443,195,1530]" captionTargetId="figure-87@15.[163,1443,195,1530]" captionTargetPageId="15" captionText="Figure 3. Shells of species described herein. A, Notothauma magnifica, holotype AMS C.532707; B, Fusobela parvioculata, holotype TMAG E59231; C, Aplotoma brevitentaculata, holotype AMS C.571635; D, Biconitoma cretosa, holotype AMS C.482313. E, Pagodibela maia, holotype AMS C.571678; F, Austrobela rufa, holotype AMS C.571709; G, Glaciotomella investigator, holotypeAMSC.571621;H, Austrotheta crassidentata, holotype AMSC.519302.Scale bar20 mm (A), 10 mm (B–H)." pageId="23" pageNumber="984">Fig. 3F</figureCitation>
) (SL = 35.4, SW = 14.6) thin-walled, fusiform, semitranslucent to opaque. Protoconch (
<figureCitation id="9BDA4AC4FFB2F84DFF4CFEBAFDD63D1B" box="[171,255,298,321]" captionStart="Figure 5" captionStartId="17.[163,243,1339,1361]" captionTargetBox="[163,1443,195,1299]" captionTargetId="figure-221@17.[163,1443,195,1299]" captionTargetPageId="17" captionText="Figure 5. Protoconchs of species described herein. A, Trochodaphne cuprosa, holotype AMS C.571611; B, Globodaphne pomum, holotype AMS C.482283; C, Pagodibela maia, holotype AMS C.571678; D, Pueridaphne cirrisulcata, paratype AMS C.563103; E, Austrobela rufa, paratype AMS C. 571681. Scale bar = 500 μm." pageId="23" pageNumber="984">Fig. 5E</figureCitation>
) (based on
<typeStatus id="DC5AE8E3FFB2F84DFE6FFEBAFCD93D1B" box="[392,496,299,320]" pageId="23" pageNumber="984" type="paratype">paratype</typeStatus>
AMS C.571681) orange, cyrthoconoid, multispiral, with 4.5 whorls, first whorl with punctate sculpture (
<figureCitation id="9BDA4AC4FFB2F84DFE28FEF9FF0A3D25" box="[463,547,360,382]" captionStart="Figure 5" captionStartId="17.[163,243,1339,1361]" captionTargetBox="[163,1443,195,1299]" captionTargetId="figure-221@17.[163,1443,195,1299]" captionTargetPageId="17" captionText="Figure 5. Protoconchs of species described herein. A, Trochodaphne cuprosa, holotype AMS C.571611; B, Globodaphne pomum, holotype AMS C.482283; C, Pagodibela maia, holotype AMS C.571678; D, Pueridaphne cirrisulcata, paratype AMS C.563103; E, Austrobela rufa, paratype AMS C. 571681. Scale bar = 500 μm." pageId="23" pageNumber="984">Fig. 5E</figureCitation>
), remaining whorls with fine, evenly distanced arcuate riblets (about 35 on last whorl). Protoconch–teleoconch transition sharply delineated, broadly sinuate. Teleoconch of about six whorls with broad, slightly concave subsutural ramp. Whorl lower portion with axial sculpture of strong, sharp opisthocline ribs, well-pronounced at shoulder of two earliest and last teleoconch whorls; whorls subcylindrical below shoulder. Third and penultimate whorls without pronounced axials. Microsculpture of dense growth lines on subsutural ramp and fine spiral cords on whorl lower portion. Last adult whorl with about 16 axials, vanishing shortly below shoulder. Shell base evenly convex, clearly demarcated from long, slender tapering siphonal canal. Aperture elongate, about half of length of shell, rounded below shoulder and narrowing toward siphonal canal. Outer lip thin, inner lip smooth, with narrow callus on long, straight columella bearing vertical orange stain. Sinus wide, moderately deep, L-shaped. Shell surface glossy, early teleoconch whorls pale orange or cream, subsequent whorls of orange colour.
</paragraph>
<paragraph id="035E5641FFB2F84DFF5CFC7DFF0239B7" blockId="23.[163,780,237,1884]" pageId="23" pageNumber="984">Anatomy based on AMS C.27120, female, except penial characters based on AMS C.571679, male. Animal uniform whitish/cream. Cephalic tentacles large, muscular, long, subcylindrical; large eyes on outer lower base of tentacles. Rhynchostome subcircular, lined with numerous epithelial cells; large, thin-walled introvert with numerous longitudinal folds. Oviduct large; intestine lightly curved along length of oviduct, posterior to it. Penis large, coiling clockwise, simple. Rectal gland greenish, filamentous/bristly in appearance, parallel to anterior of intestine. Rhynchocoel with dark red content. Venom apparatus extremely large, occupying most of rhynchoocoel; proboscis large, elongate; radular sac extremely large; venom gland large, thick, whitish, long and convoluted; muscular bulb lustrous, yellow, extremely large, elongate, with indentation posteriorly where pressed against oesophagus.</paragraph>
<paragraph id="035E5641FFB2F84DFF5CFA64F94B3CA2" blockId="23.[163,780,237,1884]" lastBlockId="23.[827,1442,197,249]" pageId="23" pageNumber="984">
Radula (
<figureCitation id="9BDA4AC4FFB2F84DFEC4FA64FC523A50" box="[291,379,1525,1547]" captionStart="Figure 6" captionStartId="18.[145,227,1378,1400]" captionTargetBox="[145,1425,195,1338]" captionTargetId="figure-24@18.[145,1425,195,1338]" captionTargetPageId="18" captionText="Figure 6. Hypodermic teeth of species described herein. A, Aplotoma brevitentaculata, holotype AMS C.571635; B, Gladiobela angulata, paratype AMS C.571737; C, Pueridaphne cirrisulcata, paratype AMS C.571609; D: Austrobela rufa, holotype AMS C.571709 (left), paratype AMS C.571679 (right); E, Pagodibela maia, holotype AMS C.571678; F, Fusobela parvioculata, holotype TMAG E59231; G, Biconitoma cretosa, holotype AMS C.482313; H, Austrotheta crassidentata, holotype AMS C.519302. Scale bars = 50 μm." pageId="23" pageNumber="984">Fig. 6D</figureCitation>
) (based on: AMS C.571679; AMS C.571709; AMS C.574588) of straight to gently curved, somewhat loosely rolled hypodermic teeth of up to 350 µm in length; lower half of shaft cylindrical, somewhat inflated, distal half weakly tapering toward prominent dorsal and ventral distal barbs, of which ventral barb more distal from tip; dorsal barb sharper than ventral barb. Adapical opening elongate, narrow, boundaries not clearly defined due to loosely overlapping tooth margins, up to one-fifth of shaft length. Basal third of shaft with weakly rugose texture. Base broad, with medium coarse texture on exterior. Basal opening large. Ligament about half width of base, solid, thick.
</paragraph>
<paragraph id="035E5641FFB2F84DFCDCFEBBFE813D19" blockId="23.[827,936,298,322]" box="[827,936,298,322]" pageId="23" pageNumber="984">
<emphasis id="31958A53FFB2F84DFCDCFEBBFE813D19" box="[827,936,298,322]" italics="true" pageId="23" pageNumber="984">Remarks</emphasis>
</paragraph>
<paragraph id="035E5641FFB2F84DFCDCFEC3FE873F6F" blockId="23.[827,1444,338,1464]" pageId="23" pageNumber="984">
This species can be differentiated from other raphitomids by the following combination of characters: a glossy shell with weakly developed sculpture, clearly pronounced but rounded shoulder and yellowish orange to reddish brown coloration; a hypodermic radula with two large, sharply hooked barbs; an extremely large venom apparatus occupying almost the entire rhynchocoel; a thin-walled, strongly folded introvert and the presence of well-developed, large eyes. Although DNA sequence data for the
<typeStatus id="DC5AE8E3FFB2F84DFCDCFD14FEB43EC0" box="[827,925,645,667]" pageId="23" pageNumber="984" type="holotype">holotype</typeStatus>
were not included in the analysis of
<figureCitation id="9BDA4AC4FFB2F84DFADBFD14F8B43EC1" box="[1340,1437,645,667]" captionStart="Figure 2" captionStartId="13.[163,240,1776,1798]" captionTargetBox="[280,1321,198,1689]" captionTargetId="graphics-44@13.[274,955,203,1674]" captionTargetPageId="13" captionText="Figure 2. Bayesian consensus phylogram based on analyses of the concatenated COI, 16S, 12S, 28S and H3 sequences. Thick lines mark branches consistent with the topology of the ML tree.Numbers above branches indicate nodal support (%) by Bayesian posterior probabilities (BPP). Numbers below branches indicate nodal support (%) by ML bootstrap (BTSP). BPP and BTSP values of 100% are both represented by asterisks. Outgroup taxa are not shown. Names of species described herein are in bold." pageId="23" pageNumber="984">Figure 2</figureCitation>
, a
<emphasis id="31958A53FFB2F84DFCB5FD35FEAA3EE2" box="[850,899,676,697]" italics="true" pageId="23" pageNumber="984">COI</emphasis>
sequence was generated and is available on Genbank (AN
<accessionNumber id="1CB2CBA2FFB2F84DFC0FFD52F9463E82" box="[1000,1135,707,729]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN983272" pageId="23" pageNumber="984">MN983272</accessionNumber>
). This is a variable taxon based on shell morphology, with some specimens examined exhibiting shells with little to no distinct sculpture.
</paragraph>
<paragraph id="035E5641FFB2F84DFCB4FCACF93439E3" blockId="23.[827,1444,338,1464]" pageId="23" pageNumber="984">
Based on its shell morphology, this new taxon bears considerable similarity to the north-east Atlantic
<taxonomicName id="C4E12DC2FFB2F84DFCDCFCEBF8B63FCB" authority="(Dautzenberg &amp; Fischer, 1996)" baseAuthorityName="Dautzenberg &amp; Fischer" baseAuthorityYear="1996" box="[827,1439,890,912]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="fulvotincta">
<emphasis id="31958A53FFB2F84DFCDCFCEBF9123FD4" box="[827,1083,890,911]" italics="true" pageId="23" pageNumber="984">Gymnobela fulvotincta</emphasis>
(Dautzenberg &amp; Fischer, 1996)
</taxonomicName>
. However, when compared to the
<typeStatus id="DC5AE8E3FFB2F84DFB48FC08F8393FF4" box="[1199,1296,921,943]" pageId="23" pageNumber="984" type="holotype">holotype</typeStatus>
of the latter, the following characters observed in
<emphasis id="31958A53FFB2F84DFB05FC29F88A3F96" box="[1250,1443,952,973]" italics="true" pageId="23" pageNumber="984">Gym. fulvocincta</emphasis>
distinguishes this species from
<taxonomicName id="C4E12DC2FFB2F84DFB70FC46F9CB3FB0" authorityName="Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov" authorityYear="2021" box="[1175,1250,982,1004]" class="Gastropoda" family="Raphitomidae" genus="Austrobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="rufa">
<emphasis id="31958A53FFB2F84DFB70FC46F9CB3FB0" box="[1175,1250,982,1004]" italics="true" pageId="23" pageNumber="984">A. rufa</emphasis>
</taxonomicName>
: more prominent axial ribs, producing a more angulate whorl profile; a shoulder situated lower on adult whorls; a less cylindrical whorl periphery and a columella that is distinctly curved when observed in apertural view as opposed to the straight columella in
<taxonomicName id="C4E12DC2FFB2F84DFB06FBE1F81838DE" authorityName="Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov" authorityYear="2021" box="[1249,1329,1136,1157]" class="Gastropoda" family="Raphitomidae" genus="Austrobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="rufa">
<emphasis id="31958A53FFB2F84DFB06FBE1F81838DE" box="[1249,1329,1136,1157]" italics="true" pageId="23" pageNumber="984">A. rufa</emphasis>
</taxonomicName>
. In terms of radular morphology, the hypodermic tooth figured in
<bibRefCitation id="67702BB0FFB2F84DFCBDFB3CF95B3899" author="Bouchet P &amp; Waren A" box="[858,1138,1197,1219]" pageId="23" pageNumber="984" pagination="1 - 119" refId="ref27671" refString="Bouchet P, Waren A. 1980. Revision of the north east Atlantic bathyal and abyssal Turridae (Mollusca, Gastropoda). Journal of Molluscan Studies Supplement 8: 1 - 119." type="journal article" year="1980">Bouchet &amp; Warén (1980</bibRefCitation>
: fig. 24) shows barbs that appear distinctly less prominent in
<taxonomicName id="C4E12DC2FFB2F84DFB3CFB5DF8B538BA" baseAuthorityName="Dautzenberg &amp; Fischer" baseAuthorityYear="1996" box="[1243,1436,1228,1249]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="fulvotincta">
<emphasis id="31958A53FFB2F84DFB3CFB5DF8B538BA" box="[1243,1436,1228,1249]" italics="true" pageId="23" pageNumber="984">Gym. fulvotincta</emphasis>
</taxonomicName>
, and a shaft that tapers gradually as opposed to the somewhat inflated lower half of the shaft in
<taxonomicName id="C4E12DC2FFB2F84DFAB7FA98F88A3945" authorityName="Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov" authorityYear="2021" box="[1360,1443,1289,1310]" class="Gastropoda" family="Raphitomidae" genus="Austrobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="rufa">
<emphasis id="31958A53FFB2F84DFAB7FA98F88A3945" box="[1360,1443,1289,1310]" italics="true" pageId="23" pageNumber="984">A. rufa</emphasis>
</taxonomicName>
(
<figureCitation id="9BDA4AC4FFB2F84DFCA4FAB9FEBE3965" box="[835,919,1320,1342]" captionStart="Figure 6" captionStartId="18.[145,227,1378,1400]" captionTargetBox="[145,1425,195,1338]" captionTargetId="figure-24@18.[145,1425,195,1338]" captionTargetPageId="18" captionText="Figure 6. Hypodermic teeth of species described herein. A, Aplotoma brevitentaculata, holotype AMS C.571635; B, Gladiobela angulata, paratype AMS C.571737; C, Pueridaphne cirrisulcata, paratype AMS C.571609; D: Austrobela rufa, holotype AMS C.571709 (left), paratype AMS C.571679 (right); E, Pagodibela maia, holotype AMS C.571678; F, Fusobela parvioculata, holotype TMAG E59231; G, Biconitoma cretosa, holotype AMS C.482313; H, Austrotheta crassidentata, holotype AMS C.519302. Scale bars = 50 μm." pageId="23" pageNumber="984">Fig. 6D</figureCitation>
). When compared to the superficially similar
<taxonomicName id="C4E12DC2FFB2F84DFCDCFAD7F86D3907" authority="(Kuroda &amp; Habe, 1961)" baseAuthorityName="Kuroda &amp; Habe" baseAuthorityYear="1961" box="[827,1348,1350,1372]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="yoshidai">
<emphasis id="31958A53FFB2F84DFCDCFAD7F9003900" box="[827,1065,1350,1371]" italics="true" pageId="23" pageNumber="984">Gymnobela yoshidai</emphasis>
(Kuroda &amp;
<bibRefCitation id="67702BB0FFB2F84DFB51FAD7F8153907" author="Habe T" box="[1206,1340,1350,1372]" pageId="23" pageNumber="984" refId="ref28572" refString="Habe T. 1961. Coloured illustrations of the shells of Japan (II). Hoikusha, Osaka. xii + 183 + 42 pp., 66 pls." type="book" year="1961">Habe, 1961</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="C4E12DC2FFB2F84DFAB6FAD6F88A3900" authorityName="Criscione &amp; Hallan &amp; Puillandre &amp; Fedosov" authorityYear="2021" box="[1361,1443,1350,1372]" class="Gastropoda" family="Raphitomidae" genus="Austrobela" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="984" phylum="Mollusca" rank="species" species="rufa">
<emphasis id="31958A53FFB2F84DFAB6FAD6F88A3900" box="[1361,1443,1350,1372]" italics="true" pageId="23" pageNumber="984">A. rufa</emphasis>
</taxonomicName>
possesses a significantly broader shell and a more sharply inclined subsutural ramp (
<bibRefCitation id="67702BB0FFB2F84DFB29FA15FE5B39EC" author="Bouchet P &amp; Sysoev A" pageId="23" pageNumber="984" pagination="1693 - 1715" refId="ref27640" refString="Bouchet P, Sysoev A. 2001. Typhlosyrinx-like tropical deep-water turriform gastropods (Mollusca, Gastropoda, Conoidea). Journal of Natural History 35: 1693 - 1715." type="journal article" year="2001">Sysoev &amp; Bouchet, 2001</bibRefCitation>
: figs 134–144).
</paragraph>
</subSubSection>
</treatment>
</document>
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