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<document id="4CFCEBD1E93F7F1A9B5553C5FBBABF19" ID-CLB-Dataset="264974" ID-DOI="10.5852/ejt.2023.890.2261" ID-GBIF-Dataset="ea7ad0da-111f-4709-819b-6a84832ceec5" ID-ISSN="2118-9773" ID-Zenodo-Dep="8305633" ID-ZooBank="5BA69DED-87A2-4041-886F-E11A6D38D52D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1693494219877" checkinUser="felipe" docAuthor="Calcetas, Orlando A." docDate="2023" docId="8B4587A1FFFA59748F4B21389E8C2337" docLanguage="en" docName="ejt-890_184-203.pdf" docOrigin="European Journal of Taxonomy 890" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucopholis ratcliffei Calcetas 2023, sp. nov." docType="treatment" docUuid="43230871-FB2C-4A6F-AF35-EE5F12830927" docUuidSource="ZooBank" docVersion="4" lastPageNumber="199" masterDocId="777CFFD9FFF7597B8D2323739F57247C" masterDocTitle="Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896" masterLastPageNumber="203" masterPageNumber="184" pageNumber="197" updateTime="1699623951618" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D86F780B75BF8875995FEDB03DF084E0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="57B074418714F490841E1E8CC5454EA1">
<mods:title id="51D0F435D4B9478B7159AFC945CF3294">Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896</mods:title>
</mods:titleInfo>
<mods:name id="FDA424363F04BFDE1E4157D0B3D557A4" type="personal">
<mods:role id="A0F8EB281F8D6EF07204AC9698714AD4">
<mods:roleTerm id="35149784039A8F10C3E5155DCF957448">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57865F2AC45B933A19F0E739CE6F450D">Calcetas, Orlando A.</mods:namePart>
<mods:nameIdentifier id="AC8BDB1BD91DD05657C90D04BD570DE4" type="LSID">5A1D3F2A-DB38-4170-987C-783749F2E1BD</mods:nameIdentifier>
<mods:affiliation id="7E7BF1876FB32ACE1F1538BB24BB6E97">Department of Agriculture, IVA-CALABARZON, Regional Crop Protection Center (DA-RCPC) Lipa Agricultural Research and Experiment Station (LARES) Brgy. Maraouy, Lipa City, Batangas 4217, Philippines</mods:affiliation>
<mods:nameIdentifier id="9417D1AE4FB1E6B4F580A189A8BBF4AA" type="email">orlando.calcetas@calabarzon.da.gov.ph &amp; orly.calcetas@yahoo.com.ph</mods:nameIdentifier>
</mods:name>
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<mods:title id="B7D6A97910D1F3495BB3291F7F412A36">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D2F033DF84C8B84AEC95E552DB5F0822">
<mods:date id="FD6A36D0562EC6B0B05947EDF5FCE37F">2023</mods:date>
<mods:detail id="0B14C2308A64B55A02039A9C9F759477" type="pubDate">
<mods:number id="25B57E934669CB8430E96B7B176D85BB">2023-08-31</mods:number>
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<mods:detail id="3F296267DA45356C12AA486698232303" type="volume">
<mods:number id="4BA8D1B77AB66BD9AD40A7F7172EA97E">890</mods:number>
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<mods:classification id="2CB3FCFF2C58DE1894C3BAC669C7F289">journal article</mods:classification>
<mods:identifier id="F8CE48426A1ADF53B6CEFDF8031EE61C" type="CLB-Dataset">264974</mods:identifier>
<mods:identifier id="BDBAE267906E6DDBAA18A844FF551718" type="DOI">10.5852/ejt.2023.890.2261</mods:identifier>
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<mods:identifier id="C591A22C09EDB14FF532FF38D0A69D81" type="ISSN">2118-9773</mods:identifier>
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<treatment id="8B4587A1FFFA59748F4B21389E8C2337" ID-DOI="http://doi.org/10.5281/zenodo.8306350" ID-GBIF-Taxon="212742583" ID-Zenodo-Dep="8306350" LSID="urn:lsid:zoobank.org:act:43230871-FB2C-4A6F-AF35-EE5F12830927" httpUri="http://treatment.plazi.org/id/8B4587A1FFFA59748F4B21389E8C2337" lastPageId="15" lastPageNumber="199" pageId="13" pageNumber="197">
<subSubSection id="4BF6653CFFFA59768F4B21389C9B261A" box="[616,972,587,614]" pageId="13" pageNumber="197" type="nomenclature">
<paragraph id="035336B7FFFA59768F4B21389C9B261A" blockId="13.[375,1213,587,681]" box="[616,972,587,614]" pageId="13" pageNumber="197">
<heading id="581B81DBFFFA59768F4B21389C9B261A" box="[616,972,587,614]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="197" reason="2">
<taxonomicName id="C4EC4D34FFFA59768F4B21389C3D261A" ID-CoL="BSCTG" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[616,874,587,614]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
<emphasis id="3198EAA5FFFA59768F4B21389C3D261A" bold="true" box="[616,874,587,614]" italics="true" pageId="13" pageNumber="197">Leucopholis ratcliffei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFFA59768E52213F9C9B261A" box="[881,972,588,614]" pageId="13" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFFA59768C54211E9C0C26D5" pageId="13" pageNumber="197" type="description">
<paragraph id="035336B7FFFA59768C54211E9BEA26F4" blockId="13.[375,1213,587,681]" box="[375,1213,621,648]" pageId="13" pageNumber="197">
<uri id="777D3AB5FFFA59768C54211E9BEA26F4" box="[375,1213,621,648]" pageId="13" pageNumber="197">
urn:lsid:zoobank.org:act:
<uuid id="774A0C62FFFA59768FB0211E9BEA26F4" box="[659,1213,621,648]" pageId="13" pageNumber="197">43230871-FB2C-4A6F-AF35-EE5F12830927</uuid>
</uri>
</paragraph>
<paragraph id="035336B7FFFA59768FFB21FD9C0C26D5" blockId="13.[375,1213,587,681]" box="[728,859,654,681]" pageId="13" pageNumber="197">
<figureCitation id="9BD72A32FFFA59768FFB21FD9C0C26D5" box="[728,859,654,681]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="13" pageNumber="197">Figs 17–20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFFA59768D9E21BD9B41201A" pageId="13" pageNumber="197" type="diagnosis">
<paragraph id="035336B7FFFA59768D9E21BD9E912694" blockId="13.[189,454,718,744]" box="[189,454,718,744]" pageId="13" pageNumber="197">
<heading id="581B81DBFFFA59768D9E21BD9E912694" bold="true" box="[189,454,718,744]" fontSize="11" level="3" pageId="13" pageNumber="197" reason="3">
<emphasis id="3198EAA5FFFA59768D9E21BD9E912694" bold="true" box="[189,454,718,744]" pageId="13" pageNumber="197">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFFA59768D9E21889B41201A" blockId="13.[187,1400,763,1126]" pageId="13" pageNumber="197">
The new species can be distinguished from all other Philippine
<taxonomicName id="C4EC4D34FFFA59768EEB21889B0D2769" authorityName="Dejean" authorityYear="1833" box="[968,1114,763,789]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFFA59768EEB21889B0D2769" box="[968,1114,763,789]" italics="true" pageId="13" pageNumber="197">Leucopholis</emphasis>
</taxonomicName>
by the paramere of the apical process of the male genitalia, which is distinctly elongate, curved downward and broadly spatulate apically. The apical process of the male genitalia of
<taxonomicName id="C4EC4D34FFFA59768E2F204C9CDA2725" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[780,909,831,857]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
<emphasis id="3198EAA5FFFA59768E2F204C9CDA2725" box="[780,909,831,857]" italics="true" pageId="13" pageNumber="197">L. ratcliffei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFFA59768EB6204C9CB92725" box="[917,1006,831,857]" pageId="13" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
is nearly as long as the phallobase while all other species of Philippine
<taxonomicName id="C4EC4D34FFFA59768F4920139DAE2706" authorityName="Dejean" authorityYear="1833" box="[618,761,864,890]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFFA59768F4920139DAE2706" box="[618,761,864,890]" italics="true" pageId="13" pageNumber="197">Leucopholis</emphasis>
</taxonomicName>
with the metaventral process extending in front of the prosternal process have the apical proces either short or distinctly shorter than the phallobase. The anterior margin of the phallobase of the male genitalia of both
<taxonomicName id="C4EC4D34FFFA59768E0920D79CFB27C2" authorityName="Calcetas" authorityYear="2023" box="[810,940,932,958]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="ratcliffei">
<emphasis id="3198EAA5FFFA59768E0920D79CFB27C2" box="[810,940,932,958]" italics="true" pageId="13" pageNumber="197">L. ratcliffei</emphasis>
</taxonomicName>
and
<taxonomicName id="C4EC4D34FFFA59768EC720D79B0B27C1" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[996,1116,932,957]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
<emphasis id="3198EAA5FFFA59768EC720D79B0B27C1" box="[996,1116,932,957]" italics="true" pageId="13" pageNumber="197">L. stainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFFA5976894320D79BEE27C2" box="[1120,1209,932,958]" pageId="13" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
is bowl-shaped, but its medial margin is slightly convex in
<taxonomicName id="C4EC4D34FFFA59768F8020B59C7127A3" authorityName="Calcetas" authorityYear="2023" box="[675,806,965,991]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="ratcliffei">
<emphasis id="3198EAA5FFFA59768F8020B59C7127A3" box="[675,806,965,991]" italics="true" pageId="13" pageNumber="197">L. ratcliffei</emphasis>
</taxonomicName>
while it is distinctly convex medially in
<taxonomicName id="C4EC4D34FFFA597689D420B59A3A27A3" authorityName="Calcetas" authorityYear="2023" box="[1271,1389,966,991]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="stainesi">
<emphasis id="3198EAA5FFFA597689D420B59A3A27A3" box="[1271,1389,966,991]" italics="true" pageId="13" pageNumber="197">L. stainesi</emphasis>
</taxonomicName>
. Also, the apical process apex is spatulate in
<taxonomicName id="C4EC4D34FFFA59768FE4209B9C1B207D" authorityName="Calcetas" authorityYear="2023" box="[711,844,999,1025]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="ratcliffei">
<emphasis id="3198EAA5FFFA59768FE4209B9C1B207D" box="[711,844,999,1025]" italics="true" pageId="13" pageNumber="197">L. ratcliffei</emphasis>
</taxonomicName>
while it is short and rounded in
<taxonomicName id="C4EC4D34FFFA597689EF209B9A12207D" authorityName="Calcetas" authorityYear="2023" box="[1228,1349,1000,1025]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="stainesi">
<emphasis id="3198EAA5FFFA597689EF209B9A12207D" box="[1228,1349,1000,1025]" italics="true" pageId="13" pageNumber="197">L. stainesi</emphasis>
</taxonomicName>
and
<taxonomicName id="C4EC4D34FFFA59768D9D277A9E6E205E" authorityName="Brenske" authorityYear="1896" box="[190,313,1033,1058]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="semperi">
<emphasis id="3198EAA5FFFA59768D9D277A9E6E205E" box="[190,313,1033,1058]" italics="true" pageId="13" pageNumber="197">L. semperi</emphasis>
</taxonomicName>
, and long and tapered in
<taxonomicName id="C4EC4D34FFFA59768F49277A9DB5205F" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[618,738,1033,1059]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="3198EAA5FFFA59768F49277A9DB5205F" box="[618,738,1033,1059]" italics="true" pageId="13" pageNumber="197">L. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFFA59768FCA277A9C12205F" box="[745,837,1033,1059]" pageId="13" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
The medial anterior margin of the pronotum in
<taxonomicName id="C4EC4D34FFFA59768D9D27589E132038" authorityName="Calcetas" authorityYear="2023" box="[190,324,1066,1092]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="ratcliffei">
<emphasis id="3198EAA5FFFA59768D9D27589E132038" box="[190,324,1066,1092]" italics="true" pageId="13" pageNumber="197">L. ratcliffei</emphasis>
</taxonomicName>
is slightly convex while it is nearly straight in all other species of Philippine
<taxonomicName id="C4EC4D34FFFA597689CB27599A2F2038" authorityName="Dejean" authorityYear="1833" box="[1256,1400,1066,1092]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFFA597689CB27599A2F2038" box="[1256,1400,1066,1092]" italics="true" pageId="13" pageNumber="197">Leucopholis</emphasis>
</taxonomicName>
with the metaventral process extending in front of the prosternal process.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFFA59768D9E27FF9B3F216A" pageId="13" pageNumber="197" type="etymology">
<paragraph id="035336B7FFFA59768D9E27FF9E1320DA" blockId="13.[189,324,1164,1190]" box="[189,324,1164,1190]" pageId="13" pageNumber="197">
<heading id="581B81DBFFFA59768D9E27FF9E1320DA" bold="true" box="[189,324,1164,1190]" fontSize="11" level="3" pageId="13" pageNumber="197" reason="3">
<emphasis id="3198EAA5FFFA59768D9E27FF9E1320DA" bold="true" box="[189,324,1164,1190]" pageId="13" pageNumber="197">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFFA59768D9E27CA9B3F216A" blockId="13.[189,1399,1209,1302]" pageId="13" pageNumber="197">
This new species is named after Dr Brett Ratcliffe, Curator of Insects at the University of
<collectingRegion id="C128F855FFFA597689E427CA9A2020AF" box="[1223,1399,1209,1235]" country="United States of America" name="Nebraska" pageId="13" pageNumber="197">Nebraska State</collectingRegion>
Museum and a Professor in the Department of Entomology. He is a specialist in the taxonomy, biology, ecology, and biogeography of scarab beetles, especially those of the Neotropics.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFFA59768D9E264F9BF721D9" pageId="13" pageNumber="197" type="materials_examined">
<paragraph id="035336B7FFFA59768D9E264F9EFD212A" blockId="13.[189,426,1340,1366]" box="[189,426,1340,1366]" pageId="13" pageNumber="197">
<heading id="581B81DBFFFA59768D9E264F9EFD212A" bold="true" box="[189,426,1340,1366]" fontSize="11" level="3" pageId="13" pageNumber="197" reason="3">
<emphasis id="3198EAA5FFFA59768D9E264F9EFD212A" bold="true" box="[189,426,1340,1366]" pageId="13" pageNumber="197">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFFA59768DC9261A9BF721D9" blockId="13.[189,1184,1385,1445]" pageId="13" pageNumber="197">
<materialsCitation id="B3843CEAFFFA59768DC9261A9BF721D9" ID-GBIF-Occurrence="4402534307" collectingDate="1932-12-05" collectionCode="UNSM" collectorName="H. L. Philipps" country="Philippines" location="Cagayan de Oro City" pageId="13" pageNumber="197" specimenCount="1" specimenCount-male="1" stateProvince="Cagayan de Oro City" typeStatus="holotype">
<emphasis id="3198EAA5FFFA59768DC9261A9E0B21FF" bold="true" box="[234,348,1385,1411]" pageId="13" pageNumber="197">
<typeStatus id="DC578815FFFA59768DC9261A9E0B21FF" box="[234,348,1385,1411]" pageId="13" pageNumber="197" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7BFB7627FFFA59768D9E26F89E3D21D9" box="[189,362,1419,1445]" name="Philippines" pageId="13" pageNumber="197">PHILIPPINES</collectingCountry>
•
<specimenCount id="15EAFD3EFFFA59768CA026F89EE521D9" box="[387,434,1419,1445]" pageId="13" pageNumber="197" type="male">1 ♂</specimenCount>
;
<collectingRegion id="C128F855FFFA59768C9D26F89DED21D9" box="[446,698,1419,1445]" country="Philippines" name="Cagayan de Oro" pageId="13" pageNumber="197">Cagayan de Oro City</collectingRegion>
;
<date id="77521077FFFA59768FE526F99C0F21D9" box="[710,856,1418,1445]" pageId="13" pageNumber="197" value="1932-12-05">
<collectingDate id="6716E99FFFFA59768FE526F99C0F21D9" box="[710,856,1418,1445]" pageId="13" pageNumber="197" value="1932-12-05">5 Dec. 1932</collectingDate>
</date>
;
<collectorName id="AE195361FFFA59768E4726F89B5621D9" box="[868,1025,1419,1445]" pageId="13" pageNumber="197">H.L. Philipps</collectorName>
leg.;
<collectionCode id="65FDAE72FFFA5976896026F89BCB21D9" box="[1091,1180,1419,1445]" collectionName="USA, Nebraska, Lincoln, University of Nebraska State Museum" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="13" pageNumber="197">UNSM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFFA59748D9E26BE9BE622AB" lastPageId="15" lastPageNumber="199" pageId="13" pageNumber="197" type="description">
<paragraph id="035336B7FFFA59768D9E26BE9E0A219A" blockId="13.[189,349,1484,1511]" box="[189,349,1484,1511]" pageId="13" pageNumber="197">
<heading id="581B81DBFFFA59768D9E26BE9E0A219A" bold="true" box="[189,349,1484,1511]" fontSize="11" level="3" pageId="13" pageNumber="197" reason="3">
<emphasis id="3198EAA5FFFA59768D9E26BE9E0A219A" bold="true" box="[189,349,1484,1511]" pageId="13" pageNumber="197">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFFA59768D9E268A9CE22268" blockId="13.[189,949,1529,1556]" box="[189,949,1529,1556]" pageId="13" pageNumber="197">
<collectingCountry id="7BFB7627FFFA59768D9E268A9E16226F" box="[189,321,1529,1555]" name="Philippines" pageId="13" pageNumber="197">Philippines</collectingCountry>
(Mindanao,
<collectingRegion id="C128F855FFFA59768CFB268A9DF0226F" box="[472,679,1529,1555]" country="Philippines" name="Misamis Oriental" pageId="13" pageNumber="197">Misamis Oriental</collectingRegion>
,
<collectingRegion id="C128F855FFFA59768F9026899CFA2268" box="[691,941,1529,1556]" country="Philippines" name="Cagayan de Oro" pageId="13" pageNumber="197">Cagayan de Oro City</collectingRegion>
)
</paragraph>
<paragraph id="035336B7FFFA59768D9E254A9E19222F" blockId="13.[189,334,1593,1619]" box="[189,334,1593,1619]" pageId="13" pageNumber="197">
<heading id="581B81DBFFFA59768D9E254A9E19222F" bold="true" box="[189,334,1593,1619]" fontSize="11" level="3" pageId="13" pageNumber="197" reason="3">
<emphasis id="3198EAA5FFFA59768D9E254A9E19222F" bold="true" box="[189,334,1593,1619]" pageId="13" pageNumber="197">Description</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFFA59768D9E25159EB322FD" blockId="13.[189,484,1638,1665]" box="[189,484,1638,1665]" pageId="13" pageNumber="197">
BODY LENGTH.
<quantity id="C4149B52FFFA59768C5025159E8822FD" box="[371,479,1638,1665]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="13" pageNumber="197" unit="mm" value="42.5">42.5 mm</quantity>
.
</paragraph>
<paragraph id="035336B7FFFA59768D9E25D59E8122BD" blockId="13.[189,470,1702,1729]" box="[189,470,1702,1729]" pageId="13" pageNumber="197">
BODY WIDTH.
<quantity id="C4149B52FFFA59768C4525D59E8522BD" box="[358,466,1702,1729]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="13" pageNumber="197" unit="mm" value="22.5">22.5 mm</quantity>
.
</paragraph>
<paragraph id="035336B7FFFA59768D9E25959BDF235E" blockId="13.[189,1398,1766,1826]" pageId="13" pageNumber="197">
COLOUR. Dorsum dichromatic, head, pronotum, scutellum and legs black to blackish brown; elytra and abdomen dark reddish brown to brownish. Venter monochromatic brown (
<figureCitation id="9BD72A32FFFA59768900247B9B2D235E" box="[1059,1146,1800,1826]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="13" pageNumber="197">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="035336B7FFFA59758D9E243B9D7D25D7" blockId="13.[189,1400,1864,2024]" lastBlockId="14.[189,1400,265,427]" lastPageId="14" lastPageNumber="198" pageId="13" pageNumber="197">HEAD. Clypeus with medial anterior margin slightly convex anteriorly; slightly cleft, slanted at 60° angle laterally; with row of irregular rugose punctures adjacent to posterior margin; each puncture with stiff brownish hair or seta; surface directly above clypeo-labral suture, lustrous, glabrous, impunctate; anterior and lateral margins carinate dorsally; lateral margin thickly carinate; nearly straight medially; anterior angle widely rounded dorsally; dorsal surface covered with yellowish white scales; scales short or long, narrow, elongate, parallel-sided, rounded apically; scales apex slightly bent downward; base of scales with rounded punctures.Apical maXillary palpomere with oval flattened area. Mentum pot-shaped, lustrous, glabrous; with very few hair brushes on each side of medial cleft subanteriorly; with nearly flattened surface, slightly depressed medially. Antennal lamellae length 3.0 mm, nearly as long as entire length of antennomeres II–VII.</paragraph>
<caption id="5793663FFFF959758D9E24D89B8D2395" ID-DOI="http://doi.org/10.5281/zenodo.8306348" ID-Zenodo-Dep="8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="14" pageNumber="198" startId="14.[189,231,1963,1989]" targetBox="[228,1398,506,1905]" targetPageId="14">
<paragraph id="035336B7FFF959758D9E24D89B8D2395" blockId="14.[187,1398,1963,2025]" pageId="14" pageNumber="198">
<emphasis id="3198EAA5FFF959758D9E24D89E6823B9" bold="true" box="[189,319,1963,1989]" pageId="14" pageNumber="198">Fig. 17–20.</emphasis>
<taxonomicName id="C4EC4D34FFF959758C6724D89D6123B9" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[324,566,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="198" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
<emphasis id="3198EAA5FFF959758C6724D89D6123B9" box="[324,566,1963,1989]" italics="true" pageId="14" pageNumber="198">Leucopholis ratcliffei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF959758F1824DF9DC323BA" box="[571,660,1964,1990]" pageId="14" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="DC578815FFF959758FBE24D89C5323B9" box="[669,772,1963,1989]" pageId="14" pageNumber="198" type="holotype">holotype</typeStatus>
,
<specimenCount id="15EAFD3EFFF959758E2E24DF9C7423BA" box="[781,803,1964,1990]" pageId="14" pageNumber="198" type="male">♂</specimenCount>
(UNSM).
<emphasis id="3198EAA5FFF959758EBA24D89CE123B9" bold="true" box="[921,950,1963,1989]" pageId="14" pageNumber="198">17</emphasis>
. Habitus, dorsal aspect.
<emphasis id="3198EAA5FFF9597589E224D89B8923B9" bold="true" box="[1217,1246,1963,1989]" pageId="14" pageNumber="198">18</emphasis>
. Metaventral process, dorsal aspect.
<emphasis id="3198EAA5FFF959758CE824BD9EBF2394" bold="true" box="[459,488,1998,2024]" pageId="14" pageNumber="198">19</emphasis>
. Male genitalia, dorsal aspect.
<emphasis id="3198EAA5FFF959758E7624BD9C252394" bold="true" box="[853,882,1998,2024]" pageId="14" pageNumber="198">20</emphasis>
. Male genitalia, lateral aspect.
</paragraph>
</caption>
<paragraph id="035336B7FFF859748D9E227A9C5E2635" blockId="15.[187,1400,265,585]" pageId="15" pageNumber="199">PRONOTUM. Anterior margin widely concave, slightly convex medially; anterior angle obtuse, widely rounded; with lustrous, glabrous, impunctate callosity apically; lateral margin widely convex medially; anterolateral margin slightly concave, nearly at 45° angle, crenulate; postero lateral margin wedge-shaped, slightly crenulate; posterior angle obtuse, strongly sinuate, rounded apically; posterior margin bisinuate, widely concave; deeply rounded, margin distinctly extended posteriorly; each side slightly carinate except medially; surface covered with scales of variable shapes and sizes; covered mostly with elongate, spindle-shaped, tapered apically scales medio-subanteriorly and medio subposteriorly; covered mostly with narrow, lanceolate, tapered apically scales submedially; covered mostly with elongate, spindle-shaped, rounded apically scales adjacent to lateral margin.</paragraph>
<paragraph id="035336B7FFF859748D9E21019DD82714" blockId="15.[187,1401,625,872]" pageId="15" pageNumber="199">
VENTRAL SIDE OF THORAX. Prosternal process nearly isosceles triangulate, widely rounded apically; surface with flattened impunctate area dorsally. Metaventral process length
<quantity id="C4149B52FFF85974897121E59BE626CD" box="[1106,1201,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="199" unit="mm" value="4.5">4.5 mm</quantity>
, distinctly wide, triangulate basally; elongate towards apex; not constricted subapically; subapical lateral margin distinctly long, nearly parallel-sided; widely rounded apically; lustrous, glabrous, impunctate dorsally; distinctly wide, triangulate basally; elongate, parallel-sided towards apex; rounded apically without subapical constriction; lustrous, glabrous, impunctate, with faint dark line dividing metasternum (
<figureCitation id="9BD72A32FFF859748998205A9A46273F" box="[1211,1297,809,835]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="15" pageNumber="199">Fig. 18</figureCitation>
); ventral and sides covered with hairs and scales.
</paragraph>
<paragraph id="035336B7FFF859748D9E20E29B8027D7" blockId="15.[189,1239,913,939]" box="[189,1239,913,939]" pageId="15" pageNumber="199">LEG. Foretibia bidentate, narrowly rounded apically; posterior metatibia with 23 spicules.</paragraph>
<paragraph id="035336B7FFF859748D9E20A79D7B206E" blockId="15.[188,1399,980,1042]" pageId="15" pageNumber="199">SCUTELLUM. Covered with large and small, elongate, parallel-sided scales; lustrous, impunctate, without scales around posterior margin.</paragraph>
<paragraph id="035336B7FFF859748D9E27489DA8214D" blockId="15.[187,1400,1083,1330]" pageId="15" pageNumber="199">ELYTRON. Anterior margin distinctly wedge-shaped; anterior angle rounded, widely explanate, thickly carinate; upper lateral margin widely explanate, thickly carinate towards anterior angle; lower lateral margin towards posterior angle not eXplanate; posterolateral margin distinctly infleXed towards posterior angle; posterior angle widely rounded; posterior margin slightly sinuate to nearly straight; sutural angle approximately at 90° angle; sutural margin medially and towards sutural angle distinctly carinate; towards scutellum not carinate; each side near sutural angle covered with fringed hairs; surface covered with same scales in pronotum; most scales truncate apically.</paragraph>
<paragraph id="035336B7FFF859748D9E26289A6F21E5" blockId="15.[189,1398,1370,1434]" pageId="15" pageNumber="199">ABDOMEN. Abdominal sternites I–IV covered with short, ovoid lanceolate, rounded or truncate basally, scales tapered apically; abdominal sternite V covered mostly with elongate spindle-shaped scales.</paragraph>
<paragraph id="035336B7FFF859748D9E26B19CAD225A" blockId="15.[189,1400,1474,1574]" pageId="15" pageNumber="199">PYGIDIUM. Posterior margin bisinuate, concave medially; shortly concave medially; posterior margin slightly concave, narrowly explanate; distinctly depressed medially; subposterior margin vertically flattened laterally; surface covered with minute, elongate ovoid scales.</paragraph>
<paragraph id="035336B7FFF859748D9E253C9BE622AB" blockId="15.[188,1399,1614,1751]" pageId="15" pageNumber="199">
MALE GENITALIA. Genitalia length
<quantity id="C4149B52FFF859748F68253D9DE22215" box="[587,693,1614,1641]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="15" pageNumber="199" unit="mm" value="18.5">18.5 mm</quantity>
. Phallobase dorsal apical margin wide, bowl-shaped, slightly convex medially (
<figureCitation id="9BD72A32FFF859748CAD25009EB222F1" box="[398,485,1651,1677]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="15" pageNumber="199">Fig. 19</figureCitation>
). Paramere with dorsal basal margin bisinuate. Apical process length
<quantity id="C4149B52FFF85974883725009A2622F2" box="[1300,1393,1651,1678]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="15" pageNumber="199" unit="mm" value="6.9">6.9 mm</quantity>
, nearly as long as phallobase; distinctly elongate, flattened; curved downward, spatulate apically. Lateral margin of paramere with wide, shallow longitudinal depression on each side (
<figureCitation id="9BD72A32FFF85974896F25CE9BF322AB" box="[1100,1188,1725,1751]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="15" pageNumber="199">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF859748D9E258C9E8C2337" box="[189,475,1791,1867]" pageId="15" pageNumber="199" type="distribution">
<paragraph id="035336B7FFF859748D9E258C9E012365" blockId="15.[189,342,1791,1817]" box="[189,342,1791,1817]" pageId="15" pageNumber="199">
<heading id="581B81DBFFF859748D9E258C9E012365" bold="true" box="[189,342,1791,1817]" fontSize="11" level="3" pageId="15" pageNumber="199" reason="3">
<emphasis id="3198EAA5FFF859748D9E258C9E012365" bold="true" box="[189,342,1791,1817]" pageId="15" pageNumber="199">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF859748D9E24439E8C2337" blockId="15.[189,475,1840,1867]" box="[189,475,1840,1867]" pageId="15" pageNumber="199">
<collectingCountry id="7BFB7627FFF859748D9E24439E162336" box="[189,321,1840,1866]" name="Philippines" pageId="15" pageNumber="199">Philippines</collectingCountry>
(Mindanao).
</paragraph>
</subSubSection>
</treatment>
</document>
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