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<document ID-DOI="10.11646/zootaxa.5263.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7835794" ID-ZooBank="0C4BE979-F298-400E-BFB8-485F244A2D3B" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1681724172769" checkinUser="plazi" docAuthor="Li, Li-Fen, Liu, Ping, Li, Bing &amp; Peng, Xian-Jin" docDate="2023" docId="8B2387910359FFA7FF176CC3074708AE" docLanguage="en" docName="zootaxa.5263.4.3.pdf" docOrigin="Zootaxa 5263 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clubiona flammaforma Li &amp; Liu &amp; Li &amp; Peng 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="523" masterDocId="771AFFE90358FFA4FF806F60040A0F2D" masterDocTitle="Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China" masterLastPageNumber="530" masterPageNumber="520" pageNumber="521" updateTime="1681724637021" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Li-Fen</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4581-6794</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China. &amp; llfyjzy 2020 @ 126. com; https: // orcid. org / 0000 - 0003 - 4581 - 6794</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Ping</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-4959-2735</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China. &amp; pingzi 129 @ 126. com; https: // orcid. org / 0000 - 0002 - 4959 - 2735</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Bing</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7106-4680</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Langfang Normal University, Langfang, Hebei 065000, China. &amp; libing @ lfnu. edu. cn; https: // orcid. org / 0000 - 0002 - 7106 - 4680</mods:affiliation>
<mods:nameIdentifier type="email">libing@lfnu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peng, Xian-Jin</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2614-3910</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China. &amp; xjpeng @ 126. com; https: // orcid. org / 0000 - 0002 - 2614 - 3910</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5263</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>520</mods:start>
<mods:end>530</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.4.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5263.4.3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7835794</mods:identifier>
<mods:identifier type="ZooBank">0C4BE979-F298-400E-BFB8-485F244A2D3B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7835788" ID-Zenodo-Dep="7835788" LSID="urn:lsid:plazi:treatment:8B2387910359FFA7FF176CC3074708AE" httpUri="http://treatment.plazi.org/id/8B2387910359FFA7FF176CC3074708AE" lastPageId="3" lastPageNumber="523" pageId="1" pageNumber="521">
<subSubSection box="[151,538,931,958]" pageId="1" pageNumber="521" type="nomenclature">
<paragraph blockId="1.[151,538,931,995]" box="[151,538,931,958]" pageId="1" pageNumber="521">
<heading bold="true" box="[151,538,931,958]" fontSize="11" level="1" pageId="1" pageNumber="521" reason="1">
<emphasis bold="true" box="[151,538,931,958]" pageId="1" pageNumber="521">
<taxonomicName authority="Li &amp; Liu &amp; Li &amp; Peng, 2023" authorityName="Li &amp; Liu &amp; Li &amp; Peng" authorityYear="2023" box="[151,437,931,958]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="flammaforma" status="sp. nov.">
<emphasis bold="true" box="[151,437,931,958]" italics="true" pageId="1" pageNumber="521">Clubiona flammaforma</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[445,538,932,958]" pageId="1" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,276,968,995]" pageId="1" pageNumber="521" type="description">
<paragraph blockId="1.[151,538,931,995]" box="[151,276,968,995]" pageId="1" pageNumber="521">
<figureCitation box="[151,220,968,994]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1413,417,1876]" captionTargetId="figure-124@2.[151,1435,417,1876]" captionTargetPageId="2" captionText="FIGURE 1. Clubiona flammaforma sp. nov., male holotype: A habitus, dorsal view; B ditto, ventral view; C left palp, prolateral view; D ditto, ventral view; E ditto, retrolateral view; F ditto, dorsal view; G RTA, dorsal view. Abbreviations: C = conductor; E = embolus; EB = embolar base; RTA = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7835796" httpUri="https://zenodo.org/record/7835796/files/figure.png" pageId="1" pageNumber="521">Figs 1</figureCitation>
,
<figureCitation box="[234,250,968,994]" captionStart="FIGURE 2" captionStartId="3.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-25@3.[151,1435,181,1622]" captionTargetPageId="3" captionText="FIGURE 2. Clubiona flammaforma sp. nov., female paratype: A, habitus, dorsal view; B ditto, ventral view. C epigyne, ventral view; D vulva, dorsal view.Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilization ducts." figureDoi="http://doi.org/10.5281/zenodo.7835798" httpUri="https://zenodo.org/record/7835798/files/figure.png" pageId="1" pageNumber="521">2</figureCitation>
,
<figureCitation box="[262,276,969,995]" captionStart="FIGURE 7" captionStartId="9.[151,250,1284,1308]" captionTargetBox="[156,1434,185,1247]" captionTargetId="figure-25@9.[151,1435,181,1260]" captionTargetPageId="9" captionText="FIGURE 7. Collection localities of Clubiona flammaforma sp. nov., Clubiona subcylindrata Wang, Chen &amp; Zhang, 2018, and Clubiona tianpingshan sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7835808" httpUri="https://zenodo.org/record/7835808/files/figure.png" pageId="1" pageNumber="521">7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="521" type="materials_examined">
<paragraph blockId="1.[151,1437,1034,2033]" pageId="1" pageNumber="521">
<emphasis bold="true" box="[151,327,1034,1061]" pageId="1" pageNumber="521">Type material.</emphasis>
<materialsCitation collectingDate="2016-04-27" collectionCode="T" collectorName="W. Liu &amp; Tian &amp; C. Zeng" country="China" county="Enshi City" elevation="1836" latitude="31.35279" location="Yanduhe Town" longLatPrecision="1" longitude="110.39937" municipality="Badong county" pageId="1" pageNumber="521" specimenCount="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis bold="true" box="[336,454,1035,1061]" pageId="1" pageNumber="521">
<typeStatus box="[336,449,1035,1061]" pageId="1" pageNumber="521">Holotype</typeStatus>
:
</emphasis>
♁,
<collectingCountry box="[500,593,1035,1061]" name="China" pageId="1" pageNumber="521">CHINA</collectingCountry>
:
<collectingRegion box="[605,784,1035,1061]" country="China" name="Hubei" pageId="1" pageNumber="521">Hubei Province</collectingRegion>
,
<collectingCounty box="[798,919,1035,1061]" pageId="1" pageNumber="521">Enshi City</collectingCounty>
,
<collectingMunicipality box="[933,1106,1035,1061]" pageId="1" pageNumber="521">Badong county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:8B2387910359FFA7FF176CC3074708AE:0655605C0359FFA5FBDE6B6B01010B08" box="[1118,1291,1035,1061]" country="China" county="Enshi City" latitude="31.35279" longLatPrecision="1" longitude="110.39937" municipality="Badong county" name="Yanduhe Town" pageId="1" pageNumber="521" stateProvince="Hubei">Yanduhe Town</location>
,
<location LSID="urn:lsid:plazi:treatment:8B2387910359FFA7FF176CC3074708AE:0655605C0359FFA5FA986B6B04E10B64" country="China" county="Enshi City" latitude="31.35279" longLatPrecision="1" longitude="110.39937" municipality="Badong county" name="Songziyuan Village" pageId="1" pageNumber="521" stateProvince="Hubei">Songziyuan Village</location>
,
<geoCoordinate box="[250,389,1070,1097]" degrees="31.35279" direction="north" orientation="latitude" pageId="1" pageNumber="521" precision="1" value="31.35279">31.35279°N</geoCoordinate>
,
<geoCoordinate box="[404,554,1071,1097]" degrees="110.39937" direction="east" orientation="longitude" pageId="1" pageNumber="521" precision="1" value="110.39937">110.39937°E</geoCoordinate>
,
<quantity box="[569,660,1071,1097]" metricMagnitude="3" metricUnit="m" metricValue="1.8359999999999999" pageId="1" pageNumber="521" unit="m" value="1836.0">
<elevation box="[569,660,1071,1097]" metricMagnitude="3" metricUnit="m" metricValue="1.8359999999999999" pageId="1" pageNumber="521" unit="m" value="1836.0">1836 m</elevation>
</quantity>
,
<date box="[676,802,1071,1097]" pageId="1" pageNumber="521" value="2016-04-27">
<collectingDate box="[676,802,1071,1097]" pageId="1" pageNumber="521" value="2016-04-27">27.IV.2016</collectingDate>
</date>
,
<collectorName box="[817,901,1071,1097]" pageId="1" pageNumber="521">W. Liu</collectorName>
,
<collectionCode box="[916,934,1071,1097]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="521">T</collectionCode>
.
<collectorName box="[948,998,1071,1097]" pageId="1" pageNumber="521">Tian</collectorName>
&amp;
<collectorName box="[1042,1137,1071,1097]" pageId="1" pageNumber="521">C. Zeng</collectorName>
leg. (HNU-HB-IV-1607)
</materialsCitation>
.
<materialsCitation collectingDate="2016-04-28" collectionCode="T" collectorName="W. Liu" country="China" county="Enshi City" elevation="1340" latitude="31.35067" location="Yanduhe Town" longLatPrecision="1" longitude="110.42625" municipality="Badong county" pageId="1" pageNumber="521" specimenCount="2" specimenCount-female="2" stateProvince="Hubei" typeStatus="paratype">
<emphasis bold="true" box="[151,280,1107,1133]" pageId="1" pageNumber="521">
<typeStatus box="[151,275,1107,1133]" pageId="1" pageNumber="521">Paratypes</typeStatus>
:
</emphasis>
1♁, same data as holotype (HNU-HB-IV-1607);
<specimenCount box="[821,858,1107,1133]" pageId="1" pageNumber="521" type="female">2♀</specimenCount>
, same place as
<typeStatus box="[1024,1125,1107,1133]" pageId="1" pageNumber="521">holotype</typeStatus>
,
<geoCoordinate box="[1134,1274,1106,1133]" degrees="31.35067" direction="north" orientation="latitude" pageId="1" pageNumber="521" precision="1" value="31.35067">31.35067°N</geoCoordinate>
,
<geoCoordinate box="[1283,1432,1106,1133]" degrees="110.42625" direction="east" orientation="longitude" pageId="1" pageNumber="521" precision="1" value="110.42625">110.42625°E</geoCoordinate>
,
<quantity box="[151,238,1143,1169]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="1" pageNumber="521" unit="m" value="1340.0">
<elevation box="[151,238,1143,1169]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="1" pageNumber="521" unit="m" value="1340.0">1340 m</elevation>
</quantity>
,
<date box="[250,376,1143,1169]" pageId="1" pageNumber="521" value="2016-04-28">
<collectingDate box="[250,376,1143,1169]" pageId="1" pageNumber="521" value="2016-04-28">28.IV.2016</collectingDate>
</date>
,
<collectorName box="[387,464,1143,1169]" pageId="1" pageNumber="521">W. Liu</collectorName>
<emphasis box="[471,527,1143,1169]" italics="true" pageId="1" pageNumber="521">et al.</emphasis>
leg. (HNU-HB-IV-1608)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="521" type="etymology">
<paragraph blockId="1.[151,1437,1034,2033]" pageId="1" pageNumber="521">
<emphasis bold="true" box="[199,334,1179,1205]" pageId="1" pageNumber="521">Etymology.</emphasis>
The specific epithet is the combination of the Latin
<emphasis box="[908,992,1179,1205]" italics="true" pageId="1" pageNumber="521">flamma</emphasis>
(fire) and
<emphasis box="[1107,1179,1179,1205]" italics="true" pageId="1" pageNumber="521">formis</emphasis>
(form), referring to the flame-shaped RTA, adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="521" type="diagnosis">
<paragraph blockId="1.[151,1437,1034,2033]" pageId="1" pageNumber="521">
<emphasis bold="true" box="[199,322,1250,1276]" pageId="1" pageNumber="521">Diagnosis.</emphasis>
<taxonomicName authority="Li &amp; Liu &amp; Li &amp; Peng, 2023" authorityName="Li &amp; Liu &amp; Li &amp; Peng" authorityYear="2023" box="[332,597,1251,1277]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="flammaforma" status="sp. nov.">
<emphasis box="[332,597,1251,1277]" italics="true" pageId="1" pageNumber="521">Clubiona flammaforma</emphasis>
</taxonomicName>
<emphasis bold="true" box="[607,700,1251,1277]" pageId="1" pageNumber="521">
<taxonomicNameLabel box="[607,700,1251,1277]" pageId="1" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName authority="Paik, 1985" authorityName="Paik" authorityYear="1985" box="[832,1091,1250,1277]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[832,957,1251,1277]" italics="true" pageId="1" pageNumber="521">C. rostrata</emphasis>
Paik, 1985
</taxonomicName>
(see
<bibRefCitation author="Zhu M. S. &amp; Zhang B. S." box="[1156,1384,1251,1277]" pageId="1" pageNumber="521" refId="ref5432" refString="Zhu M. S. &amp; Zhang B. S. (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, 558 pp." type="book" year="2011">Zhu &amp; Zhang 2011</bibRefCitation>
: fig. 266A–E) in having similar course of both embolus in male and copulatory ducts in female, but can be distinguished by: (1) the RTA trifurcate (
<figureCitation box="[465,543,1323,1349]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1413,417,1876]" captionTargetId="figure-124@2.[151,1435,417,1876]" captionTargetPageId="2" captionText="FIGURE 1. Clubiona flammaforma sp. nov., male holotype: A habitus, dorsal view; B ditto, ventral view; C left palp, prolateral view; D ditto, ventral view; E ditto, retrolateral view; F ditto, dorsal view; G RTA, dorsal view. Abbreviations: C = conductor; E = embolus; EB = embolar base; RTA = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7835796" httpUri="https://zenodo.org/record/7835796/files/figure.png" pageId="1" pageNumber="521">Fig 1F</figureCitation>
) (vs. uniramous, with a ventral protuberance in
<taxonomicName authorityName="Paik" authorityYear="1985" box="[1107,1232,1323,1349]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[1107,1232,1323,1349]" italics="true" pageId="1" pageNumber="521">C. rostrata</emphasis>
</taxonomicName>
); (2) the embolic base with a stronger dentiform process extended upwards distally in ventral view (
<figureCitation box="[1086,1167,1359,1385]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1413,417,1876]" captionTargetId="figure-124@2.[151,1435,417,1876]" captionTargetPageId="2" captionText="FIGURE 1. Clubiona flammaforma sp. nov., male holotype: A habitus, dorsal view; B ditto, ventral view; C left palp, prolateral view; D ditto, ventral view; E ditto, retrolateral view; F ditto, dorsal view; G RTA, dorsal view. Abbreviations: C = conductor; E = embolus; EB = embolar base; RTA = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7835796" httpUri="https://zenodo.org/record/7835796/files/figure.png" pageId="1" pageNumber="521">Fig 1D</figureCitation>
) (vs. with a weak tooth extended horizontally in
<taxonomicName authorityName="Paik" authorityYear="1985" box="[432,554,1395,1421]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[432,554,1395,1421]" italics="true" pageId="1" pageNumber="521">C. rostrata</emphasis>
</taxonomicName>
); (3) the posterior margin of epigyne recurve, without median excavation (
<figureCitation captionStart="FIGURE 2" captionStartId="3.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-25@3.[151,1435,181,1622]" captionTargetPageId="3" captionText="FIGURE 2. Clubiona flammaforma sp. nov., female paratype: A, habitus, dorsal view; B ditto, ventral view. C epigyne, ventral view; D vulva, dorsal view.Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilization ducts." figureDoi="http://doi.org/10.5281/zenodo.7835798" httpUri="https://zenodo.org/record/7835798/files/figure.png" pageId="1" pageNumber="521">Fig 2C</figureCitation>
) (vs. margine procurve, with oblong excavation in
<taxonomicName authorityName="Paik" authorityYear="1985" box="[778,902,1431,1457]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[778,902,1431,1457]" italics="true" pageId="1" pageNumber="521">C. rostrata</emphasis>
</taxonomicName>
); (4) the proximal portion of copulatory ducts separated from each other and arc-shaped (
<figureCitation box="[632,713,1467,1493]" captionStart="FIGURE 2" captionStartId="3.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-25@3.[151,1435,181,1622]" captionTargetPageId="3" captionText="FIGURE 2. Clubiona flammaforma sp. nov., female paratype: A, habitus, dorsal view; B ditto, ventral view. C epigyne, ventral view; D vulva, dorsal view.Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilization ducts." figureDoi="http://doi.org/10.5281/zenodo.7835798" httpUri="https://zenodo.org/record/7835798/files/figure.png" pageId="1" pageNumber="521">Fig 2D</figureCitation>
) (vs. close to each other and straight in
<taxonomicName authorityName="Paik" authorityYear="1985" box="[1160,1281,1467,1493]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="521" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[1160,1281,1467,1493]" italics="true" pageId="1" pageNumber="521">C. rostrata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="523" pageId="1" pageNumber="521" type="description">
<paragraph blockId="1.[151,1437,1034,2033]" pageId="1" pageNumber="521">
<emphasis bold="true" box="[199,415,1502,1529]" pageId="1" pageNumber="521">Description. Male</emphasis>
(
<typeStatus box="[431,532,1503,1529]" pageId="1" pageNumber="521">holotype</typeStatus>
) (
<figureCitation box="[557,683,1503,1529]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1413,417,1876]" captionTargetId="figure-124@2.[151,1435,417,1876]" captionTargetPageId="2" captionText="FIGURE 1. Clubiona flammaforma sp. nov., male holotype: A habitus, dorsal view; B ditto, ventral view; C left palp, prolateral view; D ditto, ventral view; E ditto, retrolateral view; F ditto, dorsal view; G RTA, dorsal view. Abbreviations: C = conductor; E = embolus; EB = embolar base; RTA = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7835796" httpUri="https://zenodo.org/record/7835796/files/figure.png" pageId="1" pageNumber="521">Fig. 1A, B</figureCitation>
). Total length 3.91; carapace 1.98 long, 1.35 wide; abdomen 2.15 long, 1.12 wide. Carapace dark yellow; fovea reddish and longitudinal, cervical groove inconspicuous, radial grooves visible. Eyes: AER slightly recurved, PER slightly precurved in dorsal view. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.12, PLE 0.10, AME–AME 0.04, AME–ALE 0.03, PME–PME 0.17, PME–PLE 0.10, MOQL 0.20, MOQA 0.26, MOQP 0.42. Chelicerae yellowish, with five promarginal teeth and seven retromarginal teeth. Endites and labium yellow. Sternum yellow, almost oval. Legs yellow; tibiae I and II with two pairs of ventral spines, metatarsus I and II with a pair of ventral spines. Leg measurements: I 4.04 (1.13, 1.76, 0.77, 0.45), II 4.22 (1.19, 1.77, 0.81, 0.45), III 3.59 (1.14, 1.25, 0.78, 0.42), IV 5.84 (1.91, 1.88, 1.59, 0.46). Abdomen elongate oval, dorsum yellowish white, with brown pinnate pattern posteriorly and two pairs of muscular depressions; venter yellowish white. Spinneret yellowish white.
</paragraph>
<paragraph blockId="1.[151,1437,1034,2033]" lastBlockId="2.[151,1437,151,393]" lastPageId="2" lastPageNumber="522" pageId="1" pageNumber="521">
Palp (
<figureCitation box="[262,383,1863,1889]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1413,417,1876]" captionTargetId="figure-124@2.[151,1435,417,1876]" captionTargetPageId="2" captionText="FIGURE 1. Clubiona flammaforma sp. nov., male holotype: A habitus, dorsal view; B ditto, ventral view; C left palp, prolateral view; D ditto, ventral view; E ditto, retrolateral view; F ditto, dorsal view; G RTA, dorsal view. Abbreviations: C = conductor; E = embolus; EB = embolar base; RTA = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7835796" httpUri="https://zenodo.org/record/7835796/files/figure.png" pageId="1" pageNumber="521">Fig. 1C–G</figureCitation>
). Femur and patella without apophysis. Tibia as long as patella and about 1/3 length of cymbium, RTA longer than tibia, sclerotized and trifurcated, flame-shaped, ventral branch tuberculate with blunt tip, lateral branch strongest with pointed tip, dorsal branch spiniform with pointed tip. Cymbium longer than wide. Tegulum relatively flattened, longer than wide. Conductor groovelike, membranous, located distally on retrolateral side of tegulum. Embolus slender, angled across the distal end of tegulum and extended to about 2/3 length of tegulum; embolic base broad with a stronger dentiform process extended upwards distally. Sperm duct slender and twisted in ventral view.
</paragraph>
<paragraph blockId="2.[151,1437,151,393]" pageId="2" pageNumber="522">
<emphasis bold="true" box="[199,286,223,249]" pageId="2" pageNumber="522">Female</emphasis>
(one of HNU-HB-IV-1608) (
<figureCitation box="[620,745,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-25@3.[151,1435,181,1622]" captionTargetPageId="3" captionText="FIGURE 2. Clubiona flammaforma sp. nov., female paratype: A, habitus, dorsal view; B ditto, ventral view. C epigyne, ventral view; D vulva, dorsal view.Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilization ducts." figureDoi="http://doi.org/10.5281/zenodo.7835798" httpUri="https://zenodo.org/record/7835798/files/figure.png" pageId="2" pageNumber="522">Fig. 2A–B</figureCitation>
). Total length 4.82. Carapace 2.09 long, 1.50 wide; abdomen 2.71 long, 1.58 wide. Eye sizes and interdistances: AME 0.13, ALE 0.14, PME 0.14, PLE 0.12, AME–AME 0.08, AME–ALE 0.06, PME–PME 0.22, PME–PLE 0.13, ALE–PLE 0.07, MOQL 0.27, MOQA 0.32, MOQP 0.47. Leg measurements: I 3.76 (1.14, 1.54, 0.70, 0.38), II 4.06 (1.26, 1.66, 0.72, 0.42), III 3.73 (1.14, 1.30, 0.90, 0.39), IV 5.77 (1.88, 1.77, 1.63, 0.49). Darker than male, patterns same as in male.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7835796" ID-Zenodo-Dep="7835796" httpUri="https://zenodo.org/record/7835796/files/figure.png" pageId="2" pageNumber="522" startId="2.[151,250,1899,1923]" targetBox="[151,1413,417,1876]" targetPageId="2">
<paragraph blockId="2.[151,1437,1899,1996]" pageId="2" pageNumber="522">
<emphasis bold="true" box="[151,274,1899,1923]" pageId="2" pageNumber="522">FIGURE 1.</emphasis>
<taxonomicName authority="Li &amp; Liu &amp; Li &amp; Peng, 2023" authorityName="Li &amp; Liu &amp; Li &amp; Peng" authorityYear="2023" box="[279,516,1900,1924]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="522" phylum="Arthropoda" rank="species" species="flammaforma" status="sp. nov.">
<emphasis box="[279,516,1900,1924]" italics="true" pageId="2" pageNumber="522">Clubiona flammaforma</emphasis>
</taxonomicName>
<emphasis bold="true" box="[521,601,1900,1924]" pageId="2" pageNumber="522">
<taxonomicNameLabel box="[521,601,1900,1924]" pageId="2" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype: A habitus, dorsal view; B ditto, ventral view; C left palp, prolateral view; D ditto, ventral view; E ditto, retrolateral view; F ditto, dorsal view; G RTA, dorsal view. Abbreviations: C = conductor; E = embolus; EB = embolar base; RTA = retrolateral tibial apophysis.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7835798" ID-Zenodo-Dep="7835798" httpUri="https://zenodo.org/record/7835798/files/figure.png" pageId="3" pageNumber="523" startId="3.[151,250,1646,1670]" targetBox="[151,1435,181,1622]" targetPageId="3">
<paragraph blockId="3.[151,1436,1646,1706]" pageId="3" pageNumber="523">
<emphasis bold="true" box="[151,274,1646,1670]" pageId="3" pageNumber="523">FIGURE 2.</emphasis>
<taxonomicName authority="Li &amp; Liu &amp; Li &amp; Peng, 2023" authorityName="Li &amp; Liu &amp; Li &amp; Peng" authorityYear="2023" box="[280,517,1646,1670]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="523" phylum="Arthropoda" rank="species" species="flammaforma" status="sp. nov.">
<emphasis box="[280,517,1646,1670]" italics="true" pageId="3" pageNumber="523">Clubiona flammaforma</emphasis>
</taxonomicName>
<emphasis bold="true" box="[522,602,1646,1670]" pageId="3" pageNumber="523">
<taxonomicNameLabel box="[522,602,1646,1670]" pageId="3" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype: A, habitus, dorsal view; B ditto, ventral view. C epigyne, ventral view; D vulva, dorsal view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilization ducts.
</paragraph>
</caption>
<paragraph blockId="3.[151,1437,1752,1923]" pageId="3" pageNumber="523">
Epigyne (
<figureCitation box="[305,422,1752,1779]" captionStart="FIGURE 2" captionStartId="3.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-25@3.[151,1435,181,1622]" captionTargetPageId="3" captionText="FIGURE 2. Clubiona flammaforma sp. nov., female paratype: A, habitus, dorsal view; B ditto, ventral view. C epigyne, ventral view; D vulva, dorsal view.Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilization ducts." figureDoi="http://doi.org/10.5281/zenodo.7835798" httpUri="https://zenodo.org/record/7835798/files/figure.png" pageId="3" pageNumber="523">Fig.2C, D</figureCitation>
).With trapezoid epigynal plate.Copulatory openings united at postmedian potion of epigyne. Copulatory ducts longitudinally arc-shaped and extended upwards, then twisted into oval loops. Spermathecae with two oval chambers respectively, interior chambers smaller than lateral chambers. Fertilization ducts located anteriorly inserted on spermathecal interior chambers.
</paragraph>
</subSubSection>
<subSubSection box="[199,845,1896,1923]" pageId="3" pageNumber="523" type="distribution">
<paragraph blockId="3.[151,1437,1752,1923]" box="[199,845,1896,1923]" pageId="3" pageNumber="523">
<emphasis bold="true" box="[199,353,1896,1922]" pageId="3" pageNumber="523">Distribution.</emphasis>
Known only from the
<typeStatus box="[608,656,1897,1923]" pageId="3" pageNumber="523">type</typeStatus>
locality (
<figureCitation box="[763,831,1896,1923]" captionStart="FIGURE 7" captionStartId="9.[151,250,1284,1308]" captionTargetBox="[156,1434,185,1247]" captionTargetId="figure-25@9.[151,1435,181,1260]" captionTargetPageId="9" captionText="FIGURE 7. Collection localities of Clubiona flammaforma sp. nov., Clubiona subcylindrata Wang, Chen &amp; Zhang, 2018, and Clubiona tianpingshan sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7835808" httpUri="https://zenodo.org/record/7835808/files/figure.png" pageId="3" pageNumber="523">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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