499 lines
70 KiB
XML
499 lines
70 KiB
XML
<document id="D2677CF744EF7F111664BFA6DC5ACB40" ID-CLB-Dataset="264223" ID-DOI="10.5852/ejt.2023.888.2237" ID-GBIF-Dataset="c0a2f738-d7c6-4c71-b178-058487b3b26a" ID-ISSN="2118-9773" ID-Zenodo-Dep="8269539" ID-ZooBank="56507FD3-29FE-4CF7-B205-A339EC9D471A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692626437556" checkinUser="felipe" docAuthor="Frau, Camille" docDate="2023" docId="03E23D6D4B1CFFB9394EFD28EEE01F0B" docLanguage="en" docName="ejt-888_137-158.pdf" docOrigin="European Journal of Taxonomy 888 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.888.2237" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acanthohoplites aschiltaensis" docType="treatment" docVersion="7" lastPageNumber="147" masterDocId="FFDB45154B19FFB33B4BFF93EC2C1E7E" masterDocTitle="The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900)" masterLastPageNumber="158" masterPageNumber="137" pageNumber="142" updateTime="1699595454097" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="94C977F3F68A0AF4A32C852F2336457F">The systematic affinities between the Lower Cretaceous Ammonoidea Protacanthoplites abichi (Anthula, 1900) and Acanthohoplites aschiltaensis (Anthula, 1900)</mods:title>
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<mods:namePart id="B1849EDC9C73E8AF186944E00C115481">Frau, Camille</mods:namePart>
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<mods:affiliation id="C50B3DE15A4685B705045096EE8BED01">Groupement d’Intérêt Paléontologique, Science et Exposition, 60 bd Georges Richard, 83000 Toulon, France</mods:affiliation>
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<treatment id="03E23D6D4B1CFFB9394EFD28EEE01F0B" ID-DOI="http://doi.org/10.5281/zenodo.8270492" ID-GBIF-Taxon="212448185" ID-Zenodo-Dep="8270492" LSID="urn:lsid:plazi:treatment:03E23D6D4B1CFFB9394EFD28EEE01F0B" httpUri="http://treatment.plazi.org/id/03E23D6D4B1CFFB9394EFD28EEE01F0B" lastPageId="10" lastPageNumber="147" pageId="5" pageNumber="142">
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<subSubSection id="C351DFF04B1CFFB6394EFD28E8031CAB" box="[517,1071,699,726]" pageId="5" pageNumber="142" type="nomenclature">
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<paragraph id="8BF48C7B4B1CFFB6394EFD28E8031CAB" blockId="5.[517,1071,699,761]" box="[517,1071,699,726]" pageId="5" pageNumber="142">
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<heading id="D0BC3B174B1CFFB6394EFD28E8031CAB" box="[517,1071,699,726]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="142" reason="2">
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<taxonomicName id="4C4BF7F84B1CFFB6394EFD28E8031CAB" ID-CoL="64F2W" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[517,1071,699,726]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="5" pageNumber="142" phylum="Mollusca" rank="species" species="aschiltaensis">
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<emphasis id="B93F50694B1CFFB6394EFD28EF401CA8" bold="true" box="[517,876,699,726]" italics="true" pageId="5" pageNumber="142">Acanthohoplites aschiltaensis</emphasis>
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(
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<bibRefCitation id="EFDAF18A4B1CFFB63830FD28E80A1CAB" author="Anthula" box="[891,1062,699,725]" firstAuthor="Anthula" pageId="5" pageNumber="142" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351DFF04B1CFFB63933FD4DEE211D44" pageId="5" pageNumber="142" type="description">
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<paragraph id="8BF48C7B4B1CFFB63933FD4DEF901C86" blockId="5.[517,1071,699,761]" box="[632,956,734,761]" pageId="5" pageNumber="142">
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<figureCitation id="137090FE4B1CFFB63933FD4DEED41C87" box="[632,760,734,761]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="5" pageNumber="142">Figs 2L–R</figureCitation>
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,
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<figureCitation id="137090FE4B1CFFB63848FD4CEFA81C87" box="[771,900,735,761]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[223,1364,265,1748]" captionTargetId="figure-81@6.[215,929,275,1535]" captionTargetPageId="6" captionText="Fig. 3. Illustration of the ontogenetic stages observed after the ammonitella (not seen), including Royerianum (green), Abichi (yellow), Crassicostatum-like (orange), Tobleri (violet), and Aschiltaensis (blue). A. Protacanthoplites abichi (Anthula, 1900), lectotype (IPUW 1900-3a). B. Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105). C. A. aschiltaensis, paralectotype (PMU.24107). D–J. Schematic evolution of the whorl section of the plexus P. abichi – A. aschiltaensis. D–E. After Bogdanova & Mikhailova (2016: fig. 57), not to scale. F. After IPUW 1900-3a, not to scale. G. After paralectotype (PMU.24108), at scale. H–J. After paralectotype (PMU.24107), at scale." figureDoi="http://doi.org/10.5281/zenodo.8269546" httpUri="https://zenodo.org/record/8269546/files/figure.png" pageId="5" pageNumber="142">3B–C, F–J</figureCitation>
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,
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<figureCitation id="137090FE4B1CFFB638C4FD4DEF901C86" box="[911,956,734,760]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="7.[189,232,1998,2024]" captionStartId-1="9.[189,232,1998,2024]" captionStartId-2="10.[189,232,1963,1989]" captionTargetBox-0="[189,1398,961,1952]" captionTargetBox-1="[192,1398,605,1956]" captionTargetBox-2="[189,1398,429,1917]" captionTargetId-0="figure-268@7.[-381,1239,803,2018]" captionTargetId-1="figure-117@9.[-49,1050,555,2021]" captionTargetId-2="figure-44@10.[-351,971,289,2052]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Fig. 4. Acanthohoplites aschiltaensis (Anthula, 1900), paralectotype (PMU.24106). Scale bar = 10 mm." captionText-1="Fig. 5. Acanthohoplites aschiltaensis (Anthula, 1900), paralectotype (PMU.24107). Scale bar = 10 mm." captionText-2="Fig. 6. Acanthohoplites aschiltaensis (Anthula, 1900). A–C. Paralectotype (PMU.24107). D–H. Paralectotype (PMU.24108). Scale bars = 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8269548" figureDoi-1="http://doi.org/10.5281/zenodo.8269552" figureDoi-2="http://doi.org/10.5281/zenodo.8269554" httpUri-0="https://zenodo.org/record/8269548/files/figure.png" httpUri-1="https://zenodo.org/record/8269552/files/figure.png" httpUri-2="https://zenodo.org/record/8269554/files/figure.png" pageId="5" pageNumber="142">4–6</figureCitation>
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</paragraph>
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<paragraph id="8BF48C7B4B1CFFB63BF6FCB3EE211D44" blockId="5.[189,525,800,826]" box="[189,525,800,826]" pageId="5" pageNumber="142">
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<heading id="D0BC3B174B1CFFB63BF6FCB3EE211D44" bold="true" box="[189,525,800,826]" fontSize="11" level="3" pageId="5" pageNumber="142" reason="3">
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<emphasis id="B93F50694B1CFFB63BF6FCB3EE211D44" bold="true" box="[189,525,800,826]" pageId="5" pageNumber="142">Conch shape and ontogeny</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351DFF04B1CFFB63BA1FCDCEE2D1D16" box="[234,513,846,873]" pageId="5" pageNumber="142" type="materials_examined">
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<paragraph id="8BF48C7B4B1CFFB63BA1FCDCEE2D1D16" blockId="5.[189,1401,846,1084]" box="[234,513,846,873]" pageId="5" pageNumber="142">
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<materialsCitation id="3B2386264B1CFFB63BA1FCDCEE2D1D16" ID-GBIF-Occurrence="4398934301" box="[234,513,846,873]" collectionCode="PMU" pageId="5" pageNumber="142" specimenCount="1" typeStatus="lectotype">
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<emphasis id="B93F50694B1CFFB63BA1FCDCEE2D1D16" bold="true" box="[234,513,846,873]" pageId="5" pageNumber="142">
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||
<typeStatus id="54F032D94B1CFFB63BA1FCDCED4A1D17" box="[234,358,847,873]" pageId="5" pageNumber="142" type="lectotype">Lectotype</typeStatus>
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<collectionCode id="ED5A14BE4B1CFFB63A26FCDDED9D1D16" box="[365,433,846,872]" country="Sweden" httpUri="http://grbio.org/cool/32ji-dbbk" name="Paleontological Museum of Uppsala" pageId="5" pageNumber="142">PMU</collectionCode>
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.24105
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</emphasis>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351DFF04B1CFFB63BF6FCE1EDDF196B" pageId="5" pageNumber="142" type="description">
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<paragraph id="8BF48C7B4B1CFFB63BF6FCE1EFEB1A42" blockId="5.[189,1401,846,1084]" pageId="5" pageNumber="142">
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Understanding of the
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<typeStatus id="54F032D94B1CFFB63A82FCE1EE191DF2" box="[457,565,882,908]" pageId="5" pageNumber="142" type="lectotype">lectotype</typeStatus>
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was so far based on the hand-drawing of
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<bibRefCitation id="EFDAF18A4B1CFFB63F08FCE1E8DA1DF2" author="Anthula" box="[1091,1270,882,908]" firstAuthor="Anthula" pageId="5" pageNumber="142" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900</bibRefCitation>
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: pl. X(IX) fig. 2a–b) (see
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<figureCitation id="137090FE4B1CFFB63A3AFC06EDD41DD1" box="[369,504,917,943]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="5" pageNumber="142">Fig. 2L–M</figureCitation>
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). It is here properly illustrated (
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<figureCitation id="137090FE4B1CFFB6383EFC06EFD51DD1" box="[885,1017,917,943]" captionStart="Fig" captionStartId="3.[189,232,1857,1883]" captionTargetBox="[188,1399,225,1818]" captionTargetId="figure-12@3.[635,1349,809,1345]" captionTargetPageId="3" captionText="Fig. 2. A–K. Parahoplites abichi Anthula, 1900. A–B. Reproduction of the original hand-drawing. C–G. Lectotype (IPUW 1900-3a). G. Focus on its inner whorls. H–K. Syntype (IPUW 1900-3b), which is here excluded from the species. – L–R. Parahoplites aschiltaensis Anthula, 1900.L–M. Reproduction of the original hand-drawing. N–R. Lectotype (PMU.24105). R. Focus on its inner whorls. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269544" httpUri="https://zenodo.org/record/8269544/files/figure.png" pageId="5" pageNumber="142">Fig. 2N–R</figureCitation>
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). The
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<typeStatus id="54F032D94B1CFFB63F0EFC06E89D1DD1" box="[1093,1201,917,943]" pageId="5" pageNumber="142" type="lectotype">lectotype</typeStatus>
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shows the same state of preservation as that of
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<taxonomicName id="4C4BF7F84B1CFFB6397EFC2BEF751DAC" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[565,857,952,978]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="5" pageNumber="142" phylum="Mollusca" rank="species" species="abichi">
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<emphasis id="B93F50694B1CFFB6397EFC2BEF751DAC" box="[565,857,952,978]" italics="true" pageId="5" pageNumber="142">Protacanthoplites abichi</emphasis>
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</taxonomicName>
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described above. Peristome is not preserved. The body chamber occupies half of the outer whorl. Measurements are given in
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<tableCitation id="C6C9B9C04B1CFFB63F26FC48E8EA1D8B" box="[1133,1222,987,1013]" captionStart="Table 3" captionStartId="8.[191,261,265,291]" captionTargetPageId="8" captionText="Table 3. Dimensions for specimens of Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105) and paralectotypes (PMU.24107 and PMU.24108). *measurements taken between ventral ribs. Abbreviations: see Material and methods." httpUri="http://table.plazi.org/id/DF34DCF34B11FFBB3BF4FE9AEECC1F17" pageId="5" pageNumber="142" tableUuid="DF34DCF34B11FFBB3BF4FE9AEECC1F17">Table 3</tableCitation>
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. Its shell shape is characterised by a moderate size, discoidal (Ww/D ~ 0.35), weakly compressed (Ww/Wh ~ 0.82), subevolute (U/Wh ~ 0.76) and subvirgacone coiling (U/D ~ 0.33).
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</paragraph>
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<paragraph id="8BF48C7B4B1CFFB63BF6FBF0EE69187E" blockId="5.[187,1399,1123,1536]" pageId="5" pageNumber="142">
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The ontogenetic sequence develops four stages following the worn embryonic whorls (
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<figureCitation id="137090FE4B1CFFB63F91FBF0E91B1A03" box="[1242,1335,1123,1149]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[223,1364,265,1748]" captionTargetId="figure-81@6.[215,929,275,1535]" captionTargetPageId="6" captionText="Fig. 3. Illustration of the ontogenetic stages observed after the ammonitella (not seen), including Royerianum (green), Abichi (yellow), Crassicostatum-like (orange), Tobleri (violet), and Aschiltaensis (blue). A. Protacanthoplites abichi (Anthula, 1900), lectotype (IPUW 1900-3a). B. Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105). C. A. aschiltaensis, paralectotype (PMU.24107). D–J. Schematic evolution of the whorl section of the plexus P. abichi – A. aschiltaensis. D–E. After Bogdanova & Mikhailova (2016: fig. 57), not to scale. F. After IPUW 1900-3a, not to scale. G. After paralectotype (PMU.24108), at scale. H–J. After paralectotype (PMU.24107), at scale." figureDoi="http://doi.org/10.5281/zenodo.8269546" httpUri="https://zenodo.org/record/8269546/files/figure.png" pageId="5" pageNumber="142">Fig. 3B</figureCitation>
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) that closely match those of
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<taxonomicName id="4C4BF7F84B1CFFB63A81FB15EEC71ADE" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[458,747,1158,1184]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="5" pageNumber="142" phylum="Mollusca" rank="species" species="abichi">
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<emphasis id="B93F50694B1CFFB63A81FB15EEC71ADE" box="[458,747,1158,1184]" italics="true" pageId="5" pageNumber="142">Protacanthoplites abichi</emphasis>
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</taxonomicName>
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described above. The first visible whorls bear the latest part of a typical Royerianum stage that is well visible on the left face. It changes to an Abichi stage almost identical to that of
|
||
<taxonomicName id="4C4BF7F84B1CFFB63ABAFB5EEF3E1A99" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[497,786,1229,1255]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="5" pageNumber="142" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1CFFB63ABAFB5EEF3E1A99" box="[497,786,1229,1255]" italics="true" pageId="5" pageNumber="142">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
, except its more regular pattern of secondary ribs at the beginning. The Abichi stage occupies one whorl and a half. As growth proceeds, the primary ribs lose their tubercles, while secondaries become irregular, and start decreasing. Ribbing changes progressively on the late phragmocone into an alternation of simple, primary ribs, and generally one secondary, irregular, both in length and strength on the flank. The secondaries can be indistinctly branched on the primary ribs in the lower third of the flank or at mid-flank. All ribs are flat-topped or cuneiform over the venter. This is almost similar to the Crassicostatum-like stage of
|
||
<taxonomicName id="4C4BF7F84B1CFFB63F02FA33E9431BC4" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[1097,1391,1440,1466]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="5" pageNumber="142" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1CFFB63F02FA33E9431BC4" box="[1097,1391,1440,1466]" italics="true" pageId="5" pageNumber="142">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
, but differences are seen in the progressive change of the whorl section becoming weakly compressed, subrectangular, higher than wide.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1CFFB63BF6F9BBEDDF196B" blockId="5.[188,1399,1576,1814]" pageId="5" pageNumber="142">
|
||
The Crassicostatum-like stage changes again in the late phragmocone into a regular alternation of strong, slightly flexuous, primary ribs, with a slight retrocurvature at the umbilical margin, and a single secondary starting high on the flank. The maximum whorl width is reached in the lower third of the flank. Twenty-five ventral ribs are observed on the last half whorl. They are distinctly flat-topped over the venter. These features conform to the adult ontogeny of
|
||
<taxonomicName id="4C4BF7F84B1CFFB63832F926E8F918B0" baseAuthorityName="d'Orbigny" baseAuthorityYear="1841" box="[889,1237,1717,1743]" class="Cephalopoda" family="Deshayesitidae" genus="Colombiceras" kingdom="Animalia" pageId="5" pageNumber="142" phylum="Mollusca" rank="species" species="crassicostatum">
|
||
<emphasis id="B93F50694B1CFFB63832F926E8F918B0" box="[889,1237,1717,1743]" italics="true" pageId="5" pageNumber="142">Colombiceras crassicostatum</emphasis>
|
||
</taxonomicName>
|
||
referred to as the Tobleri stage (
|
||
<bibRefCitation id="EFDAF18A4B1CFFB63AC0F94BEE64188C" author="Frau" box="[395,584,1752,1778]" etAl="et al." firstAuthor="Frau" pageId="5" pageNumber="142" pagination="1 - 16" refId="ref11154" refString="Frau C. 2020 a. New insight on the age and significance of the Aparein black shales in the Basque- Cantabrian Basin, northern Spain. Newsletters on Stratigraphy 54 (1): 1 - 16. https: // doi. org / 10.1127 / nos / 2020 / 0583" type="journal article" year="2020">
|
||
Frau
|
||
<emphasis id="B93F50694B1CFFB63A8CF94AEDD5188C" box="[455,505,1752,1778]" italics="true" pageId="5" pageNumber="142">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
). The Tobleri stage starts around D ~
|
||
<quantity id="4CB3219E4B1CFFB638BEF94BE86A188C" box="[1013,1094,1752,1778]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="142" unit="mm" value="40.0">40 mm</quantity>
|
||
(estimation) and occupies the rest of the outer whorl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B1CFFB63BA1F8AEEE1E1928" box="[234,562,1852,1879]" pageId="5" pageNumber="142" type="materials_examined">
|
||
<paragraph id="8BF48C7B4B1CFFB63BA1F8AEEE1E1928" blockId="5.[189,1398,1852,2019]" box="[234,562,1852,1879]" pageId="5" pageNumber="142">
|
||
<materialsCitation id="3B2386264B1CFFB63BA1F8AEEE1E1928" ID-GBIF-Occurrence="4398934304" box="[234,562,1852,1879]" pageId="5" pageNumber="142" typeStatus="paralectotype">
|
||
<emphasis id="B93F50694B1CFFB63BA1F8AEEE1E1928" bold="true" box="[234,562,1852,1879]" pageId="5" pageNumber="142">
|
||
<typeStatus id="54F032D94B1CFFB63BA1F8AEEDBB1929" box="[234,407,1853,1879]" pageId="5" pageNumber="142" type="paralectotype">Paralectotype</typeStatus>
|
||
PMU.24106
|
||
</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B1CFFB63BF6F8F3EE92199D" pageId="5" pageNumber="142" type="description">
|
||
<paragraph id="8BF48C7B4B1CFFB63BF6F8F3EE92199D" blockId="5.[189,1398,1852,2019]" pageId="5" pageNumber="142">
|
||
This
|
||
<typeStatus id="54F032D94B1CFFB63BB7F8F3EDB61904" box="[252,410,1888,1914]" pageId="5" pageNumber="142" type="paralectotype">paralectotype</typeStatus>
|
||
was not figured by
|
||
<bibRefCitation id="EFDAF18A4B1CFFB639DBF8F3EF671904" author="Anthula" box="[656,843,1888,1914]" firstAuthor="Anthula" pageId="5" pageNumber="142" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900)</bibRefCitation>
|
||
. It corresponds to a fragment of a large-sized phragmocone displaying the features of the Tobleri stage, including a rather regular alternation of strong, moderately flexuous primary ribs, with a slight retrocurvature at the umbilical margin, and a single secondary of variable height (
|
||
<figureCitation id="137090FE4B1CFFB63923F85AEE9C199D" box="[616,688,1993,2019]" captionStart="Fig" captionStartId="7.[189,232,1998,2024]" captionTargetBox="[189,1398,961,1952]" captionTargetId="figure-268@7.[-381,1239,803,2018]" captionTargetPageId="7" captionText="Fig. 4. Acanthohoplites aschiltaensis (Anthula, 1900), paralectotype (PMU.24106). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269548" httpUri="https://zenodo.org/record/8269548/files/figure.png" pageId="5" pageNumber="142">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF34DCF34B1FFFB53BF6F968E8941996" ID-DOI="http://doi.org/10.5281/zenodo.8269546" ID-Zenodo-Dep="8269546" httpUri="https://zenodo.org/record/8269546/files/figure.png" pageId="6" pageNumber="143" startId="6.[189,232,1787,1813]" targetBox="[223,1364,265,1748]" targetPageId="6">
|
||
<paragraph id="8BF48C7B4B1FFFB53BF6F968E8941996" blockId="6.[187,1399,1787,2025]" pageId="6" pageNumber="143">
|
||
<emphasis id="B93F50694B1FFFB53BF6F968ED3D196B" bold="true" box="[189,273,1787,1813]" pageId="6" pageNumber="143">Fig. 3.</emphasis>
|
||
Illustration of the ontogenetic stages observed after the ammonitella (not seen), including Royerianum (green), Abichi (yellow), Crassicostatum-like (orange), Tobleri (violet), and Aschiltaensis (blue).
|
||
<emphasis id="B93F50694B1FFFB53A5EF8D2ED061925" bold="true" box="[277,298,1857,1883]" pageId="6" pageNumber="143">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C4BF7F84B1FFFB53A76F8D2EF1C1922" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[317,816,1857,1884]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="143" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B1FFFB53A76F8D2EE4E1925" box="[317,610,1857,1883]" italics="true" pageId="6" pageNumber="143">Protacanthoplites abichi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B1FFFB5393DF8D2EF041922" author="Anthula" box="[630,808,1857,1884]" firstAuthor="Anthula" pageId="6" pageNumber="143" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype (IPUW 1900-3a).
|
||
<emphasis id="B93F50694B1FFFB53FD9F8D1E88A1922" bold="true" box="[1170,1190,1858,1884]" pageId="6" pageNumber="143">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C4BF7F84B1FFFB53FF3F8D2EE3B1901" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="143" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B1FFFB53FF3F8D2ED791901" italics="true" pageId="6" pageNumber="143">Acanthohoplites aschiltaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B1FFFB53A29F8F6EE221901" author="Anthula" box="[354,526,1893,1919]" firstAuthor="Anthula" pageId="6" pageNumber="143" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype (PMU.24105).
|
||
<emphasis id="B93F50694B1FFFB5380AF8F7EF7A1900" bold="true" box="[833,854,1892,1918]" pageId="6" pageNumber="143">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C4BF7F84B1FFFB5382FF8F6E8371901" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[868,1051,1893,1919]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="143" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B1FFFB5382FF8F6E8371901" box="[868,1051,1893,1919]" italics="true" pageId="6" pageNumber="143">A. aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
, paralectotype (PMU.24107).
|
||
<emphasis id="B93F50694B1FFFB53BF0F81BECC219DC" bold="true" box="[187,238,1928,1954]" pageId="6" pageNumber="143">D–J</emphasis>
|
||
. Schematic evolution of the whorl section of the plexus
|
||
<emphasis id="B93F50694B1FFFB538D4F81AE8CD19DC" box="[927,1249,1928,1954]" italics="true" pageId="6" pageNumber="143">
|
||
<taxonomicName id="4C4BF7F84B1FFFB538D4F81AE82B19DC" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[927,1031,1928,1954]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="143" phylum="Mollusca" rank="species" species="abichi">P. abichi</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C4BF7F84B1FFFB53F62F81BE8CD19DC" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[1065,1249,1928,1954]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="143" phylum="Mollusca" rank="species" species="aschiltaensis">A. aschiltaensis</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B93F50694B1FFFB53FB9F81BE90519DC" bold="true" box="[1266,1321,1928,1954]" pageId="6" pageNumber="143">D–E</emphasis>
|
||
. After
|
||
<bibRefCitation id="EFDAF18A4B1FFFB53BF5F838EE6819BB" author="Bogdanova & Mikhailova" box="[190,580,1963,1989]" firstAuthor="Bogdanova" pageId="6" pageNumber="143" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova (2016</bibRefCitation>
|
||
: fig. 57), not to scale.
|
||
<emphasis id="B93F50694B1FFFB5381BF838EF4E19BB" bold="true" box="[848,866,1963,1989]" pageId="6" pageNumber="143">F</emphasis>
|
||
. After IPUW 1900-3a, not to scale.
|
||
<emphasis id="B93F50694B1FFFB53E5EF838E90619BB" bold="true" box="[1301,1322,1963,1989]" pageId="6" pageNumber="143">G</emphasis>
|
||
. After paralectotype (PMU.24108), at scale.
|
||
<emphasis id="B93F50694B1FFFB53930F85DEE821996" bold="true" box="[635,686,1998,2024]" pageId="6" pageNumber="143">H–J</emphasis>
|
||
. After paralectotype (PMU.24107), at scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C351DFF04B1EFFB43BA1FE9AEE1E1F5C" box="[234,562,264,291]" pageId="7" pageNumber="144" type="materials_examined">
|
||
<paragraph id="8BF48C7B4B1EFFB43BA1FE9AEE1E1F5C" blockId="7.[189,1400,264,502]" box="[234,562,264,291]" pageId="7" pageNumber="144">
|
||
<materialsCitation id="3B2386264B1EFFB43BA1FE9AEE1E1F5C" ID-GBIF-Occurrence="4398934305" box="[234,562,264,291]" pageId="7" pageNumber="144" typeStatus="paralectotype">
|
||
<emphasis id="B93F50694B1EFFB43BA1FE9AEE1E1F5C" bold="true" box="[234,562,264,291]" pageId="7" pageNumber="144">
|
||
<typeStatus id="54F032D94B1EFFB43BA1FE9AEDBB1F5D" box="[234,407,265,291]" pageId="7" pageNumber="144" type="paralectotype">Paralectotype</typeStatus>
|
||
PMU.24107
|
||
</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B1EFFBB3BF6FEBFEE9B1D6C" lastPageId="8" lastPageNumber="145" pageId="7" pageNumber="144" type="description">
|
||
<paragraph id="8BF48C7B4B1EFFB43BF6FEBFE8731F88" blockId="7.[189,1400,264,502]" pageId="7" pageNumber="144">
|
||
<bibRefCitation id="EFDAF18A4B1EFFB43BF6FEBFED5C1F38" author="Anthula" box="[189,368,300,326]" firstAuthor="Anthula" pageId="7" pageNumber="144" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900</bibRefCitation>
|
||
: pl. XI(X) fig. 1) provided a hand-drawing of this
|
||
<typeStatus id="54F032D94B1EFFB438ACFEBFE8A51F38" box="[999,1161,300,326]" pageId="7" pageNumber="144" type="paralectotype">paralectotype</typeStatus>
|
||
. It is here properly illustrated for the first time (
|
||
<figureCitation id="137090FE4B1EFFB43959FEDCEEB41F14" box="[530,664,335,362]" captionStart="Fig" captionStartId="9.[189,232,1998,2024]" captionTargetBox="[192,1398,605,1956]" captionTargetId="figure-117@9.[-49,1050,555,2021]" captionTargetPageId="9" captionText="Fig. 5. Acanthohoplites aschiltaensis (Anthula, 1900), paralectotype (PMU.24107). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269552" httpUri="https://zenodo.org/record/8269552/files/figure.png" pageId="7" pageNumber="144">Figs 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="137090FE4B1EFFB439EDFEDCEEDD1F17" box="[678,753,335,361]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[189,1398,429,1917]" captionTargetId="figure-44@10.[-351,971,289,2052]" captionTargetPageId="10" captionText="Fig. 6. Acanthohoplites aschiltaensis (Anthula, 1900). A–C. Paralectotype (PMU.24107). D–H. Paralectotype (PMU.24108). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269554" httpUri="https://zenodo.org/record/8269554/files/figure.png" pageId="7" pageNumber="144">6A–C</figureCitation>
|
||
). It corresponds to a 3D-preserved internal mould but of larger size compared to the
|
||
<typeStatus id="54F032D94B1EFFB4397DFEE1EE891FF2" box="[566,677,370,396]" pageId="7" pageNumber="144" type="lectotype">lectotype</typeStatus>
|
||
. Measurements are given in
|
||
<tableCitation id="C6C9B9C04B1EFFB43F4EFEE1E84E1FF2" box="[1029,1122,370,396]" captionStart="Table 3" captionStartId="8.[191,261,265,291]" captionTargetPageId="8" captionText="Table 3. Dimensions for specimens of Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105) and paralectotypes (PMU.24107 and PMU.24108). *measurements taken between ventral ribs. Abbreviations: see Material and methods." httpUri="http://table.plazi.org/id/DF34DCF34B11FFBB3BF4FE9AEECC1F17" pageId="7" pageNumber="144" tableUuid="DF34DCF34B11FFBB3BF4FE9AEECC1F17">Table 3</tableCitation>
|
||
. Estimated diameter is about
|
||
<quantity id="4CB3219E4B1EFFB43A4CFE06ED401FCE" box="[263,364,405,432]" metricMagnitude="-1" metricUnit="m" metricValue="2.9699999999999998" pageId="7" pageNumber="144" unit="mm" value="297.0">297 mm</quantity>
|
||
. Its phragmocone is broken, but the general shell shape is virgacone (U/D ~ 0.39) with extremely discoidal (Ww/D ~ 0.25), very evolute (U/Wh ~ 1.19) and weakly compressed coiling (Ww/ Wh ~ 0.77). Its last preserved sutures are approximated suggesting its maturity.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B1EFFBB3BF6FD8EEE9B1D6C" blockId="7.[189,1401,541,884]" lastBlockId="8.[188,1398,689,786]" lastPageId="8" lastPageNumber="145" pageId="7" pageNumber="144">
|
||
The first preserved whorls bear a late Abichi stage, consisting in strong primaries, simple or bifurcate, bearing tubercle-like thickenings at the point of furcation, and a variable number of strong secondaries. It gives way to a short Crassicostatum-like stage (estimation: D ~
|
||
<quantity id="4CB3219E4B1EFFB4389DFDF0E8021C00" box="[982,1070,611,638]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="144" unit="mm" value="35.0">35 mm</quantity>
|
||
), made of strong primaries, simple or bifurcate, with decreasing thickenings on the flank, and generally one strong secondary. All ribs are flat-topped or cuneiform over the venter. Then, a Tobleri stage develops at D ~
|
||
<quantity id="4CB3219E4B1EFFB43E6AFD39E95B1CBA" box="[1313,1399,682,709]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="144" unit="mm" value="50.0">50 mm</quantity>
|
||
(estimation) bearing spaced, sharper primary ribs, being slightly flexuous with a slight retrocurvature at the umbilical margin. Secondary ribs are unique, high on the flank, so that they are almost covered by the succeeding whorl. Ribbing changes in the last two whorls into more spaced and enlarged, more flexuous, primaries with a maximum on the body chamber. Primary ribs start with a distinctive retrocurvature at the umbilical margin except approaching the aperture, and are simple, rarely bifurcate, over the flank. The secondaries are unique, and generally start in the middle, or upper flank. This last ontogenetic stage is here referred to as the Aschiltaensis stage (
|
||
<figureCitation id="137090FE4B11FFBB3982FD47EF091C91" box="[713,805,724,751]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[223,1364,265,1748]" captionTargetId="figure-81@6.[215,929,275,1535]" captionTargetPageId="6" captionText="Fig. 3. Illustration of the ontogenetic stages observed after the ammonitella (not seen), including Royerianum (green), Abichi (yellow), Crassicostatum-like (orange), Tobleri (violet), and Aschiltaensis (blue). A. Protacanthoplites abichi (Anthula, 1900), lectotype (IPUW 1900-3a). B. Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105). C. A. aschiltaensis, paralectotype (PMU.24107). D–J. Schematic evolution of the whorl section of the plexus P. abichi – A. aschiltaensis. D–E. After Bogdanova & Mikhailova (2016: fig. 57), not to scale. F. After IPUW 1900-3a, not to scale. G. After paralectotype (PMU.24108), at scale. H–J. After paralectotype (PMU.24107), at scale." figureDoi="http://doi.org/10.5281/zenodo.8269546" httpUri="https://zenodo.org/record/8269546/files/figure.png" pageId="8" pageNumber="145">Fig. 3C</figureCitation>
|
||
). Number of ventral ribs cannot be estimated but it seems denser than during the Tobleri stage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF34DCF34B1EFFB43BF6F85DE95B1997" ID-DOI="http://doi.org/10.5281/zenodo.8269548" ID-Zenodo-Dep="8269548" box="[189,1399,1998,2025]" httpUri="https://zenodo.org/record/8269548/files/figure.png" pageId="7" pageNumber="144" startId="7.[189,232,1998,2024]" targetBox="[189,1398,961,1952]" targetPageId="7">
|
||
<paragraph id="8BF48C7B4B1EFFB43BF6F85DE95B1997" blockId="7.[189,1399,1998,2025]" box="[189,1399,1998,2025]" pageId="7" pageNumber="144">
|
||
<emphasis id="B93F50694B1EFFB43BF6F85DED271996" bold="true" box="[189,267,1998,2024]" pageId="7" pageNumber="144">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C4BF7F84B1EFFB43A5AF85DEF1F1996" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[273,819,1998,2024]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="7" pageNumber="144" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B1EFFB43A5AF85DEE431996" box="[273,623,1998,2024]" italics="true" pageId="7" pageNumber="144">Acanthohoplites aschiltaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B1EFFB43936F85DEF061996" author="Anthula" box="[637,810,1998,2024]" firstAuthor="Anthula" pageId="7" pageNumber="144" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paralectotype (PMU.24106). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF34DCF34B11FFBB3BF4FE9AEECC1F17" ID-Table-UUID="DF34DCF34B11FFBB3BF4FE9AEECC1F17" httpUri="http://table.plazi.org/id/DF34DCF34B11FFBB3BF4FE9AEECC1F17" pageId="8" pageNumber="145" startId="8.[191,261,265,291]" targetBox="[449,1145,427,613]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8BF48C7B4B11FFBB3BF4FE9AEECC1F17" blockId="8.[187,1399,265,362]" pageId="8" pageNumber="145">
|
||
<emphasis id="B93F50694B11FFBB3BF4FE9AED071F5D" bold="true" box="[191,299,265,291]" pageId="8" pageNumber="145">Table 3.</emphasis>
|
||
Dimensions for specimens of
|
||
<taxonomicName id="4C4BF7F84B11FFBB39F1FE9AE8DB1F5D" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[698,1271,265,291]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B11FFBB39F1FE9AE80C1F5D" box="[698,1056,265,291]" italics="true" pageId="8" pageNumber="145">Acanthohoplites aschiltaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B11FFBB3F73FE9AE8C21F5D" author="Anthula" box="[1080,1262,265,291]" firstAuthor="Anthula" pageId="8" pageNumber="145" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype (PMU.24105) and paralectotypes (PMU.24107 and PMU.24108). *measurements taken between ventral ribs. Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF48C7B4B11FFBB3A86FE38E8541C1B" pageId="8" pageNumber="145">
|
||
<table id="F94B7EDB4B11004C3A8AFE38E8551C1B" box="[449,1145,427,613]" gridcols="5" gridrows="4" pageId="8" pageNumber="145">
|
||
<tr id="357B8E394B11004C3A8AFE38E8551FB8" box="[449,1145,427,454]" gridrow="0" pageId="8" pageNumber="145">
|
||
<th id="76AAE7454B11004C3A8AFE38EE7D1FB8" box="[449,593,427,454]" gridcol="0" gridrow="0" pageId="8" pageNumber="145">
|
||
<emphasis id="B93F50694B11FFBB3A86FE38EE6A1FBB" bold="true" box="[461,582,427,453]" pageId="8" pageNumber="145">Specimen</emphasis>
|
||
</th>
|
||
<th id="76AAE7454B11004C39E6FE38EECC1FB8" box="[685,736,427,454]" gridcol="1" gridrow="0" pageId="8" pageNumber="145">
|
||
<emphasis id="B93F50694B11FFBB39F7FE3FEEFD1FB8" bold="true" box="[700,721,428,454]" pageId="8" pageNumber="145">D</emphasis>
|
||
</th>
|
||
<th id="76AAE7454B11004C387EFE38EF441FB8" box="[821,872,427,454]" gridcol="2" gridrow="0" pageId="8" pageNumber="145">
|
||
<emphasis id="B93F50694B11FFBB380FFE3FEF751FB8" bold="true" box="[836,857,428,454]" pageId="8" pageNumber="145">U</emphasis>
|
||
</th>
|
||
<th id="76AAE7454B11004C38F2FE38EFDC1FB8" box="[953,1008,427,454]" gridcol="3" gridrow="0" pageId="8" pageNumber="145">
|
||
<emphasis id="B93F50694B11FFBB38F2FE3FEFC21FB8" bold="true" box="[953,1006,428,454]" pageId="8" pageNumber="145">Wh*</emphasis>
|
||
</th>
|
||
<th id="76AAE7454B11004C3F75FE38E8551FB8" box="[1086,1145,427,454]" gridcol="4" gridrow="0" pageId="8" pageNumber="145">
|
||
<emphasis id="B93F50694B11FFBB3F75FE3FE8551FB8" bold="true" box="[1086,1145,428,454]" pageId="8" pageNumber="145">Ww*</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="357B8E394B11004C3A8AFE73E8551F85" box="[449,1145,480,507]" gridrow="1" pageId="8" pageNumber="145">
|
||
<th id="76AAE7454B11004C3A8AFE73EE7D1F85" box="[449,593,480,507]" gridcol="0" gridrow="1" pageId="8" pageNumber="145">PMU 24105</th>
|
||
<td id="76AAE7454B11004C39E6FE73EECC1F85" box="[685,736,480,507]" gridcol="1" gridrow="1" pageId="8" pageNumber="145">68.6</td>
|
||
<td id="76AAE7454B11004C387EFE73EF441F85" box="[821,872,480,507]" gridcol="2" gridrow="1" pageId="8" pageNumber="145">22.6</td>
|
||
<td id="76AAE7454B11004C38F2FE73EFDC1F85" box="[953,1008,480,507]" gridcol="3" gridrow="1" pageId="8" pageNumber="145">29.6</td>
|
||
<td id="76AAE7454B11004C3F75FE73E8551F85" box="[1086,1145,480,507]" gridcol="4" gridrow="1" pageId="8" pageNumber="145">24.3</td>
|
||
</tr>
|
||
<tr id="357B8E394B11004C3A8AFD86E8551C4E" box="[449,1145,533,560]" gridrow="2" pageId="8" pageNumber="145">
|
||
<th id="76AAE7454B11004C3A8AFD86EE7D1C4E" box="[449,593,533,560]" gridcol="0" gridrow="2" pageId="8" pageNumber="145">PMU 24107</th>
|
||
<td id="76AAE7454B11004C39E6FD86EECC1C4E" box="[685,736,533,560]" gridcol="1" gridrow="2" pageId="8" pageNumber="145">297</td>
|
||
<td id="76AAE7454B11004C387EFD86EF441C4E" box="[821,872,533,560]" gridcol="2" gridrow="2" pageId="8" pageNumber="145">117</td>
|
||
<td id="76AAE7454B11004C38F2FD86EFDC1C4E" box="[953,1008,533,560]" gridcol="3" gridrow="2" pageId="8" pageNumber="145">98</td>
|
||
<td id="76AAE7454B11004C3F75FD86E8551C4E" box="[1086,1145,533,560]" gridcol="4" gridrow="2" pageId="8" pageNumber="145">75.6</td>
|
||
</tr>
|
||
<tr id="357B8E394B11004C3A8AFDD8E8551C1B" box="[449,1145,587,613]" gridrow="3" pageId="8" pageNumber="145">
|
||
<th id="76AAE7454B11004C3A8AFDD8EE7D1C1B" box="[449,593,587,613]" gridcol="0" gridrow="3" pageId="8" pageNumber="145">PMU 24108</th>
|
||
<td id="76AAE7454B11004C39E6FDD8EECC1C1B" box="[685,736,587,613]" gridcol="1" gridrow="3" pageId="8" pageNumber="145">93</td>
|
||
<td id="76AAE7454B11004C387EFDD8EF441C1B" box="[821,872,587,613]" gridcol="2" gridrow="3" pageId="8" pageNumber="145">39</td>
|
||
<td id="76AAE7454B11004C38F2FDD8EFDC1C1B" box="[953,1008,587,613]" gridcol="3" gridrow="3" pageId="8" pageNumber="145">32.8</td>
|
||
<td id="76AAE7454B11004C3F75FDD8E8551C1B" box="[1086,1145,587,613]" gridcol="4" gridrow="3" pageId="8" pageNumber="145">30.8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C351DFF04B11FFBB3BA1FCAAEE1E1D2D" box="[234,562,825,851]" pageId="8" pageNumber="145" type="materials_examined">
|
||
<paragraph id="8BF48C7B4B11FFBB3BA1FCAAEE1E1D2D" blockId="8.[188,1400,825,1168]" box="[234,562,825,851]" pageId="8" pageNumber="145">
|
||
<materialsCitation id="3B2386264B11FFBB3BA1FCAAEE1E1D2D" ID-GBIF-Occurrence="4398934303" box="[234,562,825,851]" pageId="8" pageNumber="145" typeStatus="paralectotype">
|
||
<emphasis id="B93F50694B11FFBB3BA1FCAAEE1E1D2D" bold="true" box="[234,562,825,851]" pageId="8" pageNumber="145">
|
||
<typeStatus id="54F032D94B11FFBB3BA1FCAAEDBB1D2D" box="[234,407,825,851]" pageId="8" pageNumber="145" type="paralectotype">Paralectotype</typeStatus>
|
||
PMU.24108
|
||
</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B11FFBA3BF6FCCFEE8B1C67" lastPageId="9" lastPageNumber="146" pageId="8" pageNumber="145" type="description">
|
||
<paragraph id="8BF48C7B4B11FFBB3BF6FCCFEE0D1AEE" blockId="8.[188,1400,825,1168]" pageId="8" pageNumber="145">
|
||
<bibRefCitation id="EFDAF18A4B11FFBB3BF6FCCFED5D1D08" author="Anthula" box="[189,369,860,886]" firstAuthor="Anthula" pageId="8" pageNumber="145" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900</bibRefCitation>
|
||
: pl. X(IX) fig. 1) only gave an idealised hand-drawing of the cross section of this
|
||
<typeStatus id="54F032D94B11FFBB3BF6FCECED731DE7" box="[189,351,895,921]" pageId="8" pageNumber="145" type="paralectotype">paralectotype</typeStatus>
|
||
. A proper illustration is given here (
|
||
<figureCitation id="137090FE4B11FFBB3842FCECEFA21DE7" box="[777,910,895,921]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[189,1398,429,1917]" captionTargetId="figure-44@10.[-351,971,289,2052]" captionTargetPageId="10" captionText="Fig. 6. Acanthohoplites aschiltaensis (Anthula, 1900). A–C. Paralectotype (PMU.24107). D–H. Paralectotype (PMU.24108). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8269554" httpUri="https://zenodo.org/record/8269554/files/figure.png" pageId="8" pageNumber="145">Fig. 6D–H</figureCitation>
|
||
). It corresponds to a half of a moderately large phragmocone. Measurements are given in
|
||
<tableCitation id="C6C9B9C04B11FFBB39B3FC30EF7E1DC3" box="[760,850,931,957]" captionStart="Table 3" captionStartId="8.[191,261,265,291]" captionTargetPageId="8" captionText="Table 3. Dimensions for specimens of Acanthohoplites aschiltaensis (Anthula, 1900), lectotype (PMU.24105) and paralectotypes (PMU.24107 and PMU.24108). *measurements taken between ventral ribs. Abbreviations: see Material and methods." httpUri="http://table.plazi.org/id/DF34DCF34B11FFBB3BF4FE9AEECC1F17" pageId="8" pageNumber="145" tableUuid="DF34DCF34B11FFBB3BF4FE9AEECC1F17">Table 3</tableCitation>
|
||
. The broken part allows to observe the Abichi stage extending up to D ~
|
||
<quantity id="4CB3219E4B11FFBB3AA0FC55EE111D9E" box="[491,573,966,992]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="145" unit="mm" value="18.0">18 mm</quantity>
|
||
(estimation), and then, a part of the Crassicostatum-like stage. Beyond, the ontogeny is rather similar to the
|
||
<typeStatus id="54F032D94B11FFBB3935FC7AEEC61A7D" box="[638,746,1001,1027]" pageId="8" pageNumber="145" type="lectotype">lectotype</typeStatus>
|
||
although the Tobleri stage is more robust and shows great affinity with specimen PMU.24107. Indeed, the shell shape differs from the
|
||
<typeStatus id="54F032D94B11FFBB3FC0FB9FE8DB1A58" box="[1163,1271,1036,1062]" pageId="8" pageNumber="145" type="lectotype">lectotype</typeStatus>
|
||
by a more discoidal (Ww/D ~ 0.33) and weakly compressed (Ww/Wh ~ 0.94) coiling, while the umbilicus is more open (U/Wh ~ 1.19). Specimen PMU.24108 further displays a lower number of ventral ribs (estimated at ~ 21) on the last half whorl.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B11FFBB3BF6FB24ED6B1AAF" blockId="8.[189,327,1207,1233]" box="[189,327,1207,1233]" pageId="8" pageNumber="145">
|
||
<heading id="D0BC3B174B11FFBB3BF6FB24ED6B1AAF" bold="true" box="[189,327,1207,1233]" fontSize="11" level="3" pageId="8" pageNumber="145" reason="3">
|
||
<emphasis id="B93F50694B11FFBB3BF6FB24ED6B1AAF" bold="true" box="[189,327,1207,1233]" pageId="8" pageNumber="145">Suture line</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B11FFBB3BF6FB75EFF718D9" blockId="8.[188,1399,1254,1703]" pageId="8" pageNumber="145">
|
||
A partial suture line of
|
||
<taxonomicName id="4C4BF7F84B11FFBB3A9CFB75EF1A1B7E" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[471,822,1254,1280]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B11FFBB3A9CFB75EF1A1B7E" box="[471,822,1254,1280]" italics="true" pageId="8" pageNumber="145">Acanthohoplites aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
was drawn by
|
||
<bibRefCitation id="EFDAF18A4B11FFBB38A0FB75E8B71B7E" author="Anthula" box="[1003,1179,1254,1280]" firstAuthor="Anthula" pageId="8" pageNumber="145" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula (1900</bibRefCitation>
|
||
: pl. X(IX) fig. 3b) based on the lost
|
||
<typeStatus id="54F032D94B11FFBB3ACEFA99EDC81B5A" box="[389,484,1290,1316]" pageId="8" pageNumber="145" type="syntype">syntype</typeStatus>
|
||
. It is of quinquelobate type and shows a deep inner lateral lobe with distinctive central and inner branches. This drawing was commented on by
|
||
<bibRefCitation id="EFDAF18A4B11FFBB3881FABFE8BC1B38" author="Stoyanow A." box="[970,1168,1324,1350]" pageId="8" pageNumber="145" pagination="1 - 169" refId="ref7275" refString="Stoyanow A. 1949. Lower Cretaceous stratigraphy in southeastern Arizona. Memoir Geological Society of America 38: 1 - 169. https: // doi. org / 10.1130 / MEM 38 - p 1" type="journal article" year="1949">Stoyanow (1949</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDAF18A4B11FFBB3FD4FABFE8F11B39" author="Stoyanow A." box="[1183,1245,1324,1351]" pageId="8" pageNumber="145" pagination="607 - 610" refId="ref7312" refString="Stoyanow A. 1958. Suture of Acanthohoplites aschiltaensis (Anthula). Geological Society of America Bulletin 69 (5): 607 - 610. https: // doi. org / 10.1130 / 0016 - 7606 (1958) 69 [607: SOAAA] 2.0. CO; 2" type="journal article" year="1958">1958</bibRefCitation>
|
||
), and his re-examination of the
|
||
<typeStatus id="54F032D94B11FFBB3AFBFADCEE301B17" box="[432,540,1359,1385]" pageId="8" pageNumber="145" type="lectotype">lectotype</typeStatus>
|
||
PMU.24105 highlighted a distinctive symmetrically trifid first lateral lobe, with some asymmetry in its inner branch, and bifid saddles. The
|
||
<typeStatus id="54F032D94B11FFBB3F5DFAE0E8AE1BF3" box="[1046,1154,1395,1421]" pageId="8" pageNumber="145" type="lectotype">lectotype</typeStatus>
|
||
seems to be mature as it shows suture approximation at the end of the phragmocone. The suture morphogenesis of that species has then been examined by
|
||
<bibRefCitation id="EFDAF18A4B11FFBB393EFA2AEF7C1BAA" author="Drushchits" box="[629,848,1465,1492]" firstAuthor="Drushchits" pageId="8" pageNumber="145" pagination="105 - 113" refId="ref10891" refString="Drushchits V. V. 1956. On systematics of the family Parahoplitidae. Scientific reports of the Moscow State University, Geology 176: 105 - 113." type="journal article" year="1956">Drushchits (1956)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFDAF18A4B11FFBB38DEFA2AE9041BAD" author="Bogdanova & Mikhailova" box="[917,1320,1465,1491]" firstAuthor="Bogdanova" pageId="8" pageNumber="145" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova (2016)</bibRefCitation>
|
||
based on individuals from
|
||
<collectingRegion id="498F42994B11FFBB3AFBFA4EEE0D1B89" box="[432,545,1501,1527]" country="Russia" name="Dagestan" pageId="8" pageNumber="145">Dagestan</collectingRegion>
|
||
. The suture line consists of a bifid ventral lobe with a low bifid median saddle; rather symmetrical umbilical lobe with distinctive lateral branches; symmetrical, short and trifid first umbilical lobe; deep inner lateral lobe with distinctive central and inner branches; bifid dorsal lobe with variable digits; broad and high, asymmetrically bifid external and umbilical saddles. The first umbilical saddle (U1/I) is indistinctly asymmetrically trifid. There is no difference between the suture lines of
|
||
<taxonomicName id="4C4BF7F84B11FFBB3A50F91FEE1118D8" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[283,573,1676,1702]" class="Cephalopoda" family="Parahoplitidae" genus="Protacanthoplites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="abichi">
|
||
<emphasis id="B93F50694B11FFBB3A50F91FEE1118D8" box="[283,573,1676,1702]" italics="true" pageId="8" pageNumber="145">Protacanthoplites abichi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4BF7F84B11FFBB393DF91FEFF818D8" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[630,980,1676,1702]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B11FFBB393DF91FEFF818D8" box="[630,980,1676,1702]" italics="true" pageId="8" pageNumber="145">Acanthohoplites aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B11FFBB3BF6F95EECC21899" blockId="8.[189,238,1741,1767]" box="[189,238,1741,1767]" pageId="8" pageNumber="145">
|
||
<heading id="D0BC3B174B11FFBB3BF6F95EECC21899" bold="true" box="[189,238,1741,1767]" fontSize="11" level="3" pageId="8" pageNumber="145" reason="3">
|
||
<emphasis id="B93F50694B11FFBB3BF6F95EECC21899" bold="true" box="[189,238,1741,1767]" pageId="8" pageNumber="145">Age</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF48C7B4B11FFBA3BF6F96FEE8B1C67" blockId="8.[189,1401,1788,1991]" lastBlockId="9.[189,1399,265,537]" lastPageId="9" lastPageNumber="146" pageId="8" pageNumber="145">
|
||
There is no comprehensive description of the Ashil’ta locality that yielded the
|
||
<typeStatus id="54F032D94B11FFBB3FE5F96EE8CD1969" box="[1198,1249,1789,1815]" pageId="8" pageNumber="145">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C4BF7F84B11FFBB3BF6F88CEE081947" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[189,548,1823,1849]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B11FFBB3BF6F88CEE081947" box="[189,548,1823,1849]" italics="true" pageId="8" pageNumber="145">Acanthohoplites aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
. Nevertheless, the
|
||
<taxonomicName id="4C4BF7F84B11FFBB3855F88CEFF11947" authorityName="Tovbina" authorityYear="1970" box="[798,989,1823,1849]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="8" pageNumber="145" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93F50694B11FFBB3855F88CEFF11947" box="[798,989,1823,1849]" italics="true" pageId="8" pageNumber="145">Acanthohoplites</emphasis>
|
||
</taxonomicName>
|
||
-bearing beds of Akusha consist of a ~ 25 m-thick interval assigned to the ‘
|
||
<emphasis id="B93F50694B11FFBB3994F8D0E9091923" box="[735,1317,1859,1885]" italics="true" pageId="8" pageNumber="145">
|
||
<taxonomicName id="4C4BF7F84B11FFBB3994F8D0E8091923" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[735,1061,1859,1885]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="aschiltaensis">Acanthoplites aschiltaensis</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C4BF7F84B11FFBB3F7FF8D0E9091923" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[1076,1317,1859,1885]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="uhligi">Acanthoplites uhligi</taxonomicName>
|
||
</emphasis>
|
||
Zone’ (
|
||
<bibRefCitation id="EFDAF18A4B11FFBB3B8EF8F5EDA219FE" author="Mordvilko T. A." box="[197,398,1894,1920]" pageId="8" pageNumber="145" refId="ref6896" refString="Mordvilko T. A. 1962. Lower Cretaceous Deposits of the Southeast Northern Caucasus and Ciscaucasia), part 2. Akademii Nauk SSSR, Moscow - Leningrad." type="book" year="1962">Mordvilko 1962</bibRefCitation>
|
||
). This is the case in the reference Dargi River section (
|
||
<bibRefCitation id="EFDAF18A4B11FFBB3F71F8F5ECD019DD" author="Bogdanova & Mikhailova" firstAuthor="Bogdanova" pageId="8" pageNumber="145" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova 2016</bibRefCitation>
|
||
: figs 10–11). Most of the taxa from the ‘
|
||
<emphasis id="B93F50694B11FFBB39A2F81AE90519DD" box="[745,1321,1929,1956]" italics="true" pageId="8" pageNumber="145">
|
||
<taxonomicName id="4C4BF7F84B11FFBB39A2F81AE80019DD" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[745,1068,1929,1955]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="aschiltaensis">Acanthoplites aschiltaensis</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C4BF7F84B11FFBB3F71F81AE90519DD" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[1082,1321,1929,1955]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="8" pageNumber="145" phylum="Mollusca" rank="species" species="uhligi">Acanthoplites uhligi</taxonomicName>
|
||
</emphasis>
|
||
Zone’ of the Dargi River section derive from a siltstone horizon including fossiliferous phosphatic nodules.
|
||
<taxonomicName id="4C4BF7F84B10FFBA3BF6FE9AEE321F5D" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[189,542,265,291]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B10FFBA3BF6FE9AEE321F5D" box="[189,542,265,291]" italics="true" pageId="9" pageNumber="146">Acanthohoplites aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
co-occurs here with
|
||
<taxonomicName id="4C4BF7F84B10FFBA386AFE9AE8F71F5D" authority="Anthula, 1900" authorityName="Anthula" authorityYear="1900" box="[801,1243,265,291]" class="Cephalopoda" family="Deshayesitidae" genus="Parahoplites" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="sjogreni">
|
||
<emphasis id="B93F50694B10FFBA386AFE9AE80E1F5D" box="[801,1058,265,291]" italics="true" pageId="9" pageNumber="146">Parahoplites sjogreni</emphasis>
|
||
<bibRefCitation id="EFDAF18A4B10FFBA3F66FE9AE8F71F5D" author="Anthula" box="[1069,1243,265,291]" firstAuthor="Anthula" pageId="9" pageNumber="146" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BF7F84B10FFBA3FA1FE9AEEE31F38" authority="Michelin, 1834" authorityName="Michelin" authorityYear="1834" class="Cephalopoda" family="Phylloceratidae" genus="Phylloceras" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="velledae" subGenus="Hypophylloceras">
|
||
<emphasis id="B93F50694B10FFBA3FA1FE9AE95A1F5D" box="[1258,1398,265,291]" italics="true" pageId="9" pageNumber="146">Phylloceras</emphasis>
|
||
(
|
||
<emphasis id="B93F50694B10FFBA3B8CFEBFEDA31F38" box="[199,399,300,326]" italics="true" pageId="9" pageNumber="146">Hypophylloceras</emphasis>
|
||
)
|
||
<emphasis id="B93F50694B10FFBA3AEDFEBFEE241F38" box="[422,520,300,326]" italics="true" pageId="9" pageNumber="146">velledae</emphasis>
|
||
<bibRefCitation id="EFDAF18A4B10FFBA395FFEBFEEE31F38" author="Michelin H." box="[532,719,300,326]" pageId="9" pageNumber="146" pagination="35" refId="ref6813" refString="Michelin H. 1834. Coquilles fossiles de Gerodot (Aube). Magasine de Zoologie, Paris, n ° 3, Classe V: pl. 35. Available from https: // www. biodiversitylibrary. org / page / 50547302 [accessed 28 Jul. 2023]." type="journal article" year="1834">Michelin, 1834</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BF7F84B10FFBA39AAFEBEEFB51F39" box="[737,921,301,327]" class="Cephalopoda" family="Oppeliidae" genus="Aconeceras" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B93F50694B10FFBA39AAFEBEEF461F39" box="[737,874,301,327]" italics="true" pageId="9" pageNumber="146">Aconeceras</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, and the problematic
|
||
<taxonomicName id="4C4BF7F84B10FFBA3FE7FEBFE95A1F39" box="[1196,1398,300,327]" class="Cephalopoda" family="Douvilleiceratidae" genus="Cheloniceras" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B93F50694B10FFBA3FE7FEBFE9641F38" box="[1196,1352,300,326]" italics="true" pageId="9" pageNumber="146">Cheloniceras</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B10FFBA3B8EFEDCEE6E1F17" author="Bogdanova & Mikhailova" box="[197,578,335,362]" firstAuthor="Bogdanova" pageId="9" pageNumber="146" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova 2016</bibRefCitation>
|
||
). This horizon is of regional significance since it also crops out in the Rubas-chai section, in the vicinity of Khuchni (
|
||
<bibRefCitation id="EFDAF18A4B10FFBA39B5FEE1E8AE1FF2" author="Bogdanova & Mikhailova" box="[766,1154,370,397]" firstAuthor="Bogdanova" pageId="9" pageNumber="146" pagination="725 - 933" refId="ref10623" refString="Bogdanova T. N. & Mikhailova I. A. 2016. Middle Aptian biostratigraphy and ammonoids of the Northern Caucasus and Transcaspia. Paleontological Journal 50 (8): 725 - 933. https: // doi. org / 10.1134 / S 0031030115100019" type="journal article" year="2016">Bogdanova & Mikhailova 2016</bibRefCitation>
|
||
: fig. 12). A younger
|
||
<taxonomicName id="4C4BF7F84B10FFBA3BF6FE05ED501FCE" authorityName="Tovbina" authorityYear="1970" box="[189,380,406,432]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="9" pageNumber="146" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93F50694B10FFBA3BF6FE05ED501FCE" box="[189,380,406,432]" italics="true" pageId="9" pageNumber="146">Acanthohoplites</emphasis>
|
||
</taxonomicName>
|
||
-bearing horizon is documented in the upper part of the ‘
|
||
<emphasis id="B93F50694B10FFBA3F6DFE05ED821FAD" italics="true" pageId="9" pageNumber="146">
|
||
<taxonomicName id="4C4BF7F84B10FFBA3F6DFE05E9441FCE" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[1062,1384,406,432]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="aschiltaensis">Acanthoplites aschiltaensis</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C4BF7F84B10FFBA3BF6FE2AED821FAD" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[189,430,441,467]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="uhligi">Acanthoplites uhligi</taxonomicName>
|
||
</emphasis>
|
||
Zone’ in the Dargi River and Rubas-chai sections. Representatives assigned to
|
||
<taxonomicName id="4C4BF7F84B10FFBA3BF6FE4FED501F88" authorityName="Tovbina" authorityYear="1970" box="[189,380,476,502]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="9" pageNumber="146" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93F50694B10FFBA3BF6FE4FED501F88" box="[189,380,476,502]" italics="true" pageId="9" pageNumber="146">Acanthohoplites</emphasis>
|
||
</taxonomicName>
|
||
ex gr.
|
||
<taxonomicName id="4C4BF7F84B10FFBA3A99FE4FEE461F88" baseAuthorityName="Anthula" baseAuthorityYear="1899" box="[466,618,476,502]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthoplites" kingdom="Animalia" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B10FFBA3A99FE4FEE461F88" box="[466,618,476,502]" italics="true" pageId="9" pageNumber="146">aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
would be still present in the corresponding horizon but the fauna lacks illustration for further confirmation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF34DCF34B10FFBA3BF6F85DE95B1997" ID-DOI="http://doi.org/10.5281/zenodo.8269552" ID-Zenodo-Dep="8269552" box="[189,1399,1998,2025]" httpUri="https://zenodo.org/record/8269552/files/figure.png" pageId="9" pageNumber="146" startId="9.[189,232,1998,2024]" targetBox="[192,1398,605,1956]" targetPageId="9">
|
||
<paragraph id="8BF48C7B4B10FFBA3BF6F85DE95B1997" blockId="9.[189,1399,1998,2025]" box="[189,1399,1998,2025]" pageId="9" pageNumber="146">
|
||
<emphasis id="B93F50694B10FFBA3BF6F85DED271996" bold="true" box="[189,267,1998,2024]" pageId="9" pageNumber="146">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C4BF7F84B10FFBA3A5AF85DEF1F1996" authority="(Anthula, 1900)" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[273,819,1998,2024]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="9" pageNumber="146" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B10FFBA3A5AF85DEE431996" box="[273,623,1998,2024]" italics="true" pageId="9" pageNumber="146">Acanthohoplites aschiltaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDAF18A4B10FFBA3936F85DEF061996" author="Anthula" box="[637,810,1998,2024]" firstAuthor="Anthula" pageId="9" pageNumber="146" pagination="55 - 160" refId="ref10343" refString="Anthula D. J. 1900. Uber die Kreidefossilien des Kaukasus mit einem allgemeinen Uberblick uber die Entwicklung der Sedimentarbildungen des Kaukasus. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients 12 (for 1899): 55 - 160 (1 - 106)." type="journal article" year="1900">Anthula, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paralectotype (PMU.24107). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C351DFF04B13FFB93BF6FE9AEEFF1F5D" box="[189,723,264,291]" pageId="10" pageNumber="147" type="multiple">
|
||
<paragraph id="8BF48C7B4B13FFB93BF6FE9AEEFF1F5D" blockId="10.[189,723,264,291]" box="[189,723,264,291]" pageId="10" pageNumber="147">
|
||
<heading id="D0BC3B174B13FFB93BF6FE9AEEFF1F5D" bold="true" box="[189,723,264,291]" fontSize="11" level="3" pageId="10" pageNumber="147" reason="3">
|
||
<emphasis id="B93F50694B13FFB93BF6FE9AEEFF1F5D" bold="true" box="[189,723,264,291]" pageId="10" pageNumber="147">
|
||
Subspecies of
|
||
<taxonomicName id="4C4BF7F84B13FFB93A20FE9BEEFF1F5D" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[363,723,264,291]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="10" pageNumber="147" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B13FFB93A20FE9BEEFF1F5D" bold="true" box="[363,723,264,291]" italics="true" pageId="10" pageNumber="147">Acanthohoplites aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351DFF04B13FFB93BF6FEA4EEE01F0B" pageId="10" pageNumber="147" type="taxon_list">
|
||
<paragraph id="8BF48C7B4B13FFB93BF6FEA4EEE01F0B" blockId="10.[189,1399,311,373]" pageId="10" pageNumber="147">
|
||
<bibRefCitation id="EFDAF18A4B13FFB93BF6FEA4EE2B1F2C" author="Klein J. & Bogdanova T." box="[189,519,311,338]" pageId="10" pageNumber="147" refId="ref6504" refString="Klein J. & Bogdanova T. 2013. Lower Cretaceous Ammonites VI. Douvilleiceratoidea & Deshayesitoidea. Fossilium Catalogus I: Animalia - Pars 151. Backhuys Publishers, Leiden." type="book" year="2013">Klein & Bogdanova (2013)</bibRefCitation>
|
||
listed three subspecies of
|
||
<taxonomicName id="4C4BF7F84B13FFB93801FEA4E8861F2F" baseAuthorityName="Anthula" baseAuthorityYear="1900" box="[842,1194,311,337]" class="Cephalopoda" family="Deshayesitidae" genus="Acanthohoplites" kingdom="Animalia" order="Ammonitida" pageId="10" pageNumber="147" phylum="Mollusca" rank="species" species="aschiltaensis">
|
||
<emphasis id="B93F50694B13FFB93801FEA4E8861F2F" box="[842,1194,311,337]" italics="true" pageId="10" pageNumber="147">Acanthohoplites aschiltaensis</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<emphasis id="B93F50694B13FFB93E48FEA4ED1A1F0B" italics="true" pageId="10" pageNumber="147">Fossilium Catalogus</emphasis>
|
||
. Their validity is discussed below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |