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<document id="B7469BF2CB9D2192A462DFB316A43D44" ID-CLB-Dataset="27760" ID-DOI="10.11646/zootaxa.4532.1.4" ID-GBIF-Dataset="0f50b518-84b6-4931-be29-80fb5f7ccbc1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615065" ID-ZooBank="4CAC2D1A-7AFF-4570-A074-45223090C546" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787744637" checkinUser="plazi" docAuthor="Nakano, Takafumi, Tomikawa, Ko &amp; Grygier, Mark J." docDate="2018" docId="03D287D9FF8EFFAF338AF9AF3DA721B1" docLanguage="en" docName="zootaxa.4532.1.4.pdf" docOrigin="Zootaxa 4532 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procrangonyx japonicus" docType="treatment" docVersion="8" masterDocId="FFEBFFA1FF8DFFAC331DFD003834267E" masterDocTitle="Rediscovered syntypes of Procrangonyx japonicus, with nomenclatural consideration of some crangonyctoidean subterranean amphipods (Crustacea: Amphipoda: Allocrangonyctidae, Niphargidae, Pseudocrangonyctidae)" updateTime="1698714997393" updateUser="ExternalLinkService">
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<mods:title id="D28F25CBE7CDFF1F9ED555A47FE2351E">Rediscovered syntypes of Procrangonyx japonicus, with nomenclatural consideration of some crangonyctoidean subterranean amphipods (Crustacea: Amphipoda: Allocrangonyctidae, Niphargidae, Pseudocrangonyctidae)</mods:title>
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<mods:namePart id="02DC17E5840D999D54E3CE3E510CCF1D">Tomikawa, Ko</mods:namePart>
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<mods:namePart id="F31B25B9E3B240ED76C554B3FE8C1F99">Grygier, Mark J.</mods:namePart>
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<treatment id="03D287D9FF8EFFAF338AF9AF3DA721B1" ID-DOI="http://doi.org/10.5281/zenodo.5965991" ID-GBIF-Taxon="154521196" ID-Zenodo-Dep="5965991" LSID="urn:lsid:plazi:treatment:03D287D9FF8EFFAF338AF9AF3DA721B1" httpUri="http://treatment.plazi.org/id/03D287D9FF8EFFAF338AF9AF3DA721B1" pageId="3">
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<taxonomicName id="4C7B4D4CFF8EFFAF338AF9AF3A5022B4" ID-CoL="4MP9K" authority="(Ueno, 1930)" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[151,612,1199,1226]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8EFFAF338AF9AF3A5022B4" bold="true" box="[151,612,1199,1226]" pageId="3">
<emphasis id="B90FEADDFF8EFFAF338AF9AF39F422B7" bold="true" box="[151,448,1199,1225]" italics="true" pageId="3">Procrangonyx japonicus</emphasis>
(
<bibRefCitation id="EFEA4B3EFF8EFFAF32D2F9B03A6822B4" author="Ueno, M." box="[463,604,1200,1226]" pageId="3" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
)
</emphasis>
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</paragraph>
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<paragraph id="8BC436CFFF8EFFAF338AF9D2393F2295" blockId="3.[151,612,1199,1259]" box="[151,267,1234,1259]" pageId="3">
(
<figureCitation id="13402A4AFF8EFFAF3382F9D238D12295" box="[159,229,1234,1259]" captionStart="FIGURE 1" captionStartId="3.[151,250,1077,1099]" captionTargetBox="[150,1436,770,1055]" captionTargetId="figure@3.[150,1436,770,1055]" captionTargetPageId="3" captionText="FIGURE 1. Procrangonyx japonicus (Uéno), syntypes, KUZ Z1759. (A) Photograph showing two individuals. (B) Historical handwritten label. Scale bar, 5 mm." httpUri="https://zenodo.org/record/2615067/files/figure.png" pageId="3">Figs 1</figureCitation>
,
<figureCitation id="13402A4AFF8EFFAF33EEF9D339362295" box="[243,258,1235,1259]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1939]" captionTargetBox="[159,1430,199,1889]" captionTargetId="figure@4.[150,1436,195,1895]" captionTargetPageId="4" captionText="FIGURE 2. Procrangonyx japonicus (Uéno), syntype, KUZ Z1759. (A) Photograph of left uropods 13 of one of the syntypes, lateral view. (B) Photograph of the terminal part of left uropod 3, lateral view. (C) Line drawing of the terminal part of uropod 3, lateral view. Scale bars, 0.5 mm (A), 0.1 mm (B, C)." httpUri="https://zenodo.org/record/2615069/files/figure.png" pageId="3">2</figureCitation>
)
</paragraph>
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<paragraph id="8BC436CFFF8EFFAF338AF8163AAC2355" blockId="3.[151,917,1301,1354]" box="[151,664,1301,1323]" pageId="3">
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<taxonomicName id="4C7B4D4CFF8EFFAF338AF8163A1D2355" authority="Ueno, 1930: 21" authorityName="Ueno" authorityPageNumber="21" authorityYear="1930" box="[151,553,1301,1323]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8EFFAF338AF81639B32355" box="[151,391,1302,1323]" italics="true" pageId="3">Eucrangonyx japonicus</emphasis>
<treatmentCitation id="0ADA10DEFF8EFFAF3290F8153A1D2355" author="Ueno, M." box="[397,553,1301,1323]" page="21" pageId="3" year="1930">
<bibRefCitation id="EFEA4B3EFF8EFFAF3290F8153A322355" author="Ueno, M." box="[397,518,1301,1323]" pageId="3" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
: 21
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23, fig. 1.
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</paragraph>
<paragraph id="8BC436CFFF8EFFAF338AF8353BA12334" blockId="3.[151,917,1301,1354]" box="[151,917,1332,1354]" pageId="3">
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<taxonomicName id="4C7B4D4CFF8EFFAF338AF83539B22334" ID-CoL="6FJ88" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[151,390,1333,1354]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8EFFAF338AF83539B22334" box="[151,390,1333,1354]" italics="true" pageId="3">Eocrangonyx japonicus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0ADA10DEFF8EFFAF32BEF8343ACC2334" author="Tomikawa, K. &amp; Shinoda, S." box="[419,760,1332,1354]" page="585" pageId="3" year="2016">
<bibRefCitation id="EFEA4B3EFF8EFFAF32BEF8343AF32334" author="Tomikawa, K. &amp; Shinoda, S." box="[419,711,1332,1354]" pageId="3" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda, 2016</bibRefCitation>
: 585
</treatmentCitation>
593, figs. 26.
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</paragraph>
</subSubSection>
<subSubSection id="C3616544FF8EFFAF338AF8763C3223A9" pageId="3" type="materials_examined">
<paragraph id="8BC436CFFF8EFFAF338AF8763C3223A9" blockId="3.[151,1436,1398,2000]" pageId="3">
<emphasis id="B90FEADDFF8EFFAF338AF87639B523F1" bold="true" box="[151,385,1398,1423]" pageId="3">Material examined.</emphasis>
<materialsCitation id="3B133C92FF8EFFAF3297F8773C3623A9" ID-GBIF-Occurrence="2117757563" collectingDate="1930-01-26" collectionCode="KUZ" country="Japan" location="Article" municipality="Code" pageId="3" specimenCode="Z1759" specimenCount="2" stateProvince="Tokyo" typeStatus="syntype">
<typeStatus id="54C0886DFF8EFFAF3297F87739D823F0" box="[394,492,1399,1422]" pageId="3" type="syntype">
<emphasis id="B90FEADDFF8EFFAF3297F87739D823F0" box="[394,492,1399,1422]" italics="true" pageId="3">Syntypes</emphasis>
</typeStatus>
.
<collectionCode id="ED6AAE0AFF8EFFAF32E1F8783A0223F1" box="[508,566,1400,1423]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3">KUZ</collectionCode>
<specimenCode id="DBDD9EB4FF8EFFAF3123F8763ABD23F1" box="[574,649,1398,1423]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3">Z1759</specimenCode>
, two individuals, collected from underground water in
<collectingRegion id="49BFF82DFF8EFFAF37E2F8763D7D23F1" box="[1279,1353,1398,1423]" country="Japan" name="Tokyo" pageId="3">Tokyo</collectingRegion>
,
<collectingCountry id="F36C765FFF8EFFAF3648F8783DAC23F1" box="[1365,1432,1400,1423]" name="Japan" pageId="3">Japan</collectingCountry>
, on
<collectingDate id="EF81E9E7FF8EFFAF33A6F89B394223CD" box="[187,374,1435,1460]" pageId="3" value="1930-01-26">
<date id="FFC5100FFF8EFFAF33A6F89B394223CD" box="[187,374,1435,1460]" pageId="3" value="1930-01-26">26 January 1930</date>
</collectingDate>
(
<bibRefCitation id="EFEA4B3EFF8EFFAF329BF89B3A6123CD" author="Ueno, M." box="[390,597,1435,1460]" pageId="3" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">
Uéno 1930;
<figureCitation id="13402A4AFF8EFFAF3109F89B3A6123CD" box="[532,597,1435,1460]" captionStart="FIGURE 1" captionStartId="3.[151,250,1077,1099]" captionTargetBox="[150,1436,770,1055]" captionTargetId="figure@3.[150,1436,770,1055]" captionTargetPageId="3" captionText="FIGURE 1. Procrangonyx japonicus (Uéno), syntypes, KUZ Z1759. (A) Photograph showing two individuals. (B) Historical handwritten label. Scale bar, 5 mm." httpUri="https://zenodo.org/record/2615067/files/figure.png" pageId="3">Fig. 1</figureCitation>
</bibRefCitation>
); these two specimens, upon which the original description of the species was based, are automatically fixed as
<typeStatus id="54C0886DFF8EFFAF3122F8C03A9523A6" box="[575,673,1472,1496]" pageId="3" type="syntype">syntypes</typeStatus>
under
<location id="8EA46014FF8EFFAF31F3F8BE3B0823A9" LSID="urn:lsid:plazi:treatment:03D287D9FF8EFFAF338AF9AF3DA721B1:8EA46014FF8EFFAF31F3F8BE3B0823A9" box="[750,828,1470,1495]" country="Japan" municipality="Code" name="Article" pageId="3" stateProvince="Tokyo">Article</location>
73.2 of the
<collectingMunicipality id="6BA0ACB5FF8EFFAF30DEF8BE3C3623A9" box="[963,1026,1470,1495]" pageId="3">Code</collectingMunicipality>
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.
</paragraph>
</subSubSection>
<subSubSection id="C3616544FF8EFFAF33DAF8E33C592061" pageId="3" type="description">
<paragraph id="8BC436CFFF8EFFAF33DAF8E33C592061" blockId="3.[151,1436,1398,2000]" pageId="3">
<emphasis id="B90FEADDFF8EFFAF33DAF8E33A322382" bold="true" box="[199,518,1507,1532]" pageId="3">Redescription of uropod 3.</emphasis>
Left uropod 3 of one individual still discernable. Inner ramus absent; outer ramus uniarticulate, with two long, robust setae and seven short, simple, slender setae (
<figureCitation id="13402A4AFF8EFFAF3707FB063C542061" box="[1050,1120,1542,1567]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1939]" captionTargetBox="[159,1430,199,1889]" captionTargetId="figure@4.[150,1436,195,1895]" captionTargetPageId="4" captionText="FIGURE 2. Procrangonyx japonicus (Uéno), syntype, KUZ Z1759. (A) Photograph of left uropods 13 of one of the syntypes, lateral view. (B) Photograph of the terminal part of left uropod 3, lateral view. (C) Line drawing of the terminal part of uropod 3, lateral view. Scale bars, 0.5 mm (A), 0.1 mm (B, C)." httpUri="https://zenodo.org/record/2615069/files/figure.png" pageId="3">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616544FF8EFFAF33DAFB2B3DA721B1" pageId="3" type="discussion">
<paragraph id="8BC436CFFF8EFFAF33DAFB2B3DA721B1" blockId="3.[151,1436,1398,2000]" pageId="3">
<emphasis id="B90FEADDFF8EFFAF33DAFB2B390F203A" bold="true" box="[199,315,1579,1604]" pageId="3">Remarks.</emphasis>
It was hard to observe the detailed morphological characteristics of the
<typeStatus id="54C0886DFF8EFFAF376EFB2C3CE1203A" box="[1139,1237,1580,1604]" pageId="3" type="syntype">syntypes</typeStatus>
due to their poor condition. The preservative solution of the specimens appears to have dried up, whereupon the amphipods became flattened against the bottom of the glass tube and were damaged (
<figureCitation id="13402A4AFF8EFFAF3067FB733BEC20F5" box="[890,984,1651,1676]" captionStart="FIGURE 1" captionStartId="3.[151,250,1077,1099]" captionTargetBox="[150,1436,770,1055]" captionTargetId="figure@3.[150,1436,770,1055]" captionTargetPageId="3" captionText="FIGURE 1. Procrangonyx japonicus (Uéno), syntypes, KUZ Z1759. (A) Photograph showing two individuals. (B) Historical handwritten label. Scale bar, 5 mm." httpUri="https://zenodo.org/record/2615067/files/figure.png" pageId="3">Figs 1A</figureCitation>
,
<figureCitation id="13402A4AFF8EFFAF30F9FB733C3E20F5" box="[996,1034,1651,1675]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1939]" captionTargetBox="[159,1430,199,1889]" captionTargetId="figure@4.[150,1436,195,1895]" captionTargetPageId="4" captionText="FIGURE 2. Procrangonyx japonicus (Uéno), syntype, KUZ Z1759. (A) Photograph of left uropods 13 of one of the syntypes, lateral view. (B) Photograph of the terminal part of left uropod 3, lateral view. (C) Line drawing of the terminal part of uropod 3, lateral view. Scale bars, 0.5 mm (A), 0.1 mm (B, C)." httpUri="https://zenodo.org/record/2615069/files/figure.png" pageId="3">2A</figureCitation>
). However, the left uropod 3 of one of them was sufficiently well preserved to confirm that its outer ramus comprises only a single article. Therefore,
<bibRefCitation id="EFEA4B3EFF8EFFAF338AFBBB390C20AD" author="Ueno, M." box="[151,312,1723,1747]" pageId="3" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uénos (1930)</bibRefCitation>
original description of uropod 3 was correct. This means that
<taxonomicName id="4C7B4D4CFF8EFFAF30F2FBBC3C4C20AD" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[1007,1144,1724,1747]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8EFFAF30F2FBBC3C4C20AD" box="[1007,1144,1724,1747]" italics="true" pageId="3">P. japonicus</emphasis>
</taxonomicName>
is variable in this respect, the outer ramus being either uniarticulate (present
<typeStatus id="54C0886DFF8EFFAF31D6FBDF3B102089" box="[715,804,1759,1783]" pageId="3" type="syntype">syntype</typeStatus>
) or biarticulate (
<bibRefCitation id="EFEA4B3EFF8EFFAF30C1FBDE3D232089" author="Tomikawa, K. &amp; Shinoda, S." box="[988,1303,1758,1783]" pageId="3" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda 2016</bibRefCitation>
). Molecular phylogenetic analysis has shown that
<taxonomicName id="4C7B4D4CFF8EFFAF3121FA043C0F2165" authority="Stock &amp; Jo, 1990" authorityName="Stock &amp; Jo" authorityYear="1990" box="[572,1083,1795,1820]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="species" species="primoryensis">
<emphasis id="B90FEADDFF8EFFAF3121FA043B472165" box="[572,883,1796,1819]" italics="true" pageId="3">Procrangonyx primoryensis</emphasis>
<bibRefCitation id="EFEA4B3EFF8EFFAF3067FA033C0F2165" author="Stock, J. H. &amp; Jo, Y. W." box="[890,1083,1795,1820]" pageId="3" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo, 1990</bibRefCitation>
</taxonomicName>
is a member of a monophyletic lineage consisting otherwise only of
<taxonomicName id="4C7B4D4CFF8EFFAF317AFA263B042141" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[615,816,1830,1855]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF317AFA263B042141" box="[615,816,1830,1855]" italics="true" pageId="3">Pseudocrangonyx</emphasis>
</taxonomicName>
species (Tomikawa
<emphasis id="B90FEADDFF8EFFAF3729FA283C432141" box="[1076,1143,1830,1855]" italics="true" pageId="3">et al.</emphasis>
2016). It is clear that
<taxonomicName id="4C7B4D4CFF8EFFAF338AFA4C3902211D" authorityName=", Schellenberg" authorityYear="1934" box="[151,310,1868,1891]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF338AFA4C3902211D" box="[151,310,1868,1891]" italics="true" pageId="3">Procrangonyx</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D4CFF8EFFAF3296FA4B3A60211A" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[395,596,1867,1892]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF3296FA4B3A60211A" box="[395,596,1867,1892]" italics="true" pageId="3">Pseudocrangonyx</emphasis>
</taxonomicName>
are phylogenetically close, and
<taxonomicName id="4C7B4D4CFF8EFFAF3717FA4C3C9D211D" authorityName=", Schellenberg" authorityYear="1934" box="[1034,1193,1868,1891]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF3717FA4C3C9D211D" box="[1034,1193,1868,1891]" italics="true" pageId="3">Procrangonyx</emphasis>
</taxonomicName>
, despite its being nomenclaturally available as demonstrated above, could potentially be reduced to an invalid junior synonym of
<taxonomicName id="4C7B4D4CFF8EFFAF338AFA93395421D2" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[151,352,1939,1964]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF338AFA93395421D2" box="[151,352,1939,1964]" italics="true" pageId="3">Pseudocrangonyx</emphasis>
</taxonomicName>
, as previous studies have implied (
<bibRefCitation id="EFEA4B3EFF8EFFAF31EFFA933C1221D5" author="Sidorov, D. &amp; Holsinger, J. R." box="[754,1062,1939,1964]" pageId="3" pagination="417 - 430" refId="ref6467" refString="Sidorov, D. [A.] &amp; Holsinger, J. R. (2007) Procrangonyx stygoedincus, a new species of subterranean amphipod (Pseudocrangonyctidae) from the Far East of Russia, with remarks on biogeographic relationships. Crustaceana, 80, 417 - 430. https: // doi. org / 10.1163 / 156854007780440984" type="journal article" year="2007">Sidorov &amp; Holsinger 2007</bibRefCitation>
;
<bibRefCitation id="EFEA4B3EFF8EFFAF372EFA933D4521D2" author="Tomikawa, K. &amp; Shinoda, S." box="[1075,1393,1939,1964]" pageId="3" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda 2016</bibRefCitation>
). In this context, a molecular phylogenetic study of the relations of
<taxonomicName id="4C7B4D4CFF8EFFAF3047FAB83C5921B1" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[858,1133,1976,1999]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8EFFAF3047FAB83C5921B1" box="[858,1133,1976,1999]" italics="true" pageId="3">Procrangonyx japonicus</emphasis>
</taxonomicName>
is a necessary future step.
</paragraph>
</subSubSection>
</treatment>
</document>