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27 KiB
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113 lines
27 KiB
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<document id="F4D83E3F0DF1CE9CF4D8FFF0453A2B79" ID-DOI="10.17109/AZH.64.4.277.2018" ID-ISSN="2064-2474" ID-Zenodo-Dep="12626311" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638277157792" checkinUser="felipe" docAuthor="Nabozhenko, Maxim & Ando, Kiyoshi" docDate="2018" docId="03BBAE26FFF5FFE5FE5FA5FE555D3980" docLanguage="en" docName="ActaZoolAcadSciH.64.4.277-327.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 64 (4)" docSource="http://dx.doi.org/10.17109/azh.64.4.277.2018" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.8:ActaZoolAcadSciH.2015-2023.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-2023.journal_article" docStyleVersion="8" docTitle="Cylindrinotina Espanol 1956" docType="treatment" docVersion="3" lastPageNumber="293" masterDocId="FF82D65EFFFAFFF5FFCCA16D56513801" masterDocTitle="Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region" masterLastPageNumber="327" masterPageNumber="277" pageNumber="292" updateTime="1719929912661" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="7C7D23F462A15AD3E7CB63AD1EADD230">Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region</mods:title>
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Subtribe
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<taxonomicName id="4C1264B3FFF5FFFAFDC4A5FE55D03CAF" authority="Espanol, 1956" authorityName="Espanol" authorityYear="1956" box="[520,897,1171,1198]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="292" phylum="Arthropoda" rank="subTribe" subTribe="Cylindrinotina">
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Cylindrinotina
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<bibRefCitation id="EF8362C1FFF5FFFAFD03A5FE55D03CAF" author="ESPANOL, F." box="[719,897,1171,1198]" pageId="15" pageNumber="292" pagination="83 - 123" refId="ref17772" refString="ESPANOL, F. (1956): Los Probaticus de Espana (Col. Tenebrionidae). - Eos, Revista Espanola de Entomologia 32: 83 - 123." type="journal article" year="1956">Español, 1956</bibRefCitation>
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(
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<figureCitation id="132903B5FFF5FFFAFD81A5DB54963CD0" box="[589,711,1206,1233]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig" captionStart-11="Fig" captionStart-12="Fig" captionStart-13="Fig" captionStart-14="Fig" captionStart-15="Fig" captionStart-16="Fig" captionStart-17="Fig" captionStart-18="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="17.[174,210,1457,1479]" captionStartId-1="20.[136,172,1483,1505]" captionStartId-10="36.[136,172,1459,1481]" captionStartId-11="37.[174,210,1545,1567]" captionStartId-12="38.[135,171,1552,1574]" captionStartId-13="39.[174,210,1493,1515]" captionStartId-14="40.[136,172,1462,1484]" captionStartId-15="41.[174,210,1458,1480]" captionStartId-16="42.[136,172,1458,1480]" captionStartId-17="44.[136,172,1525,1547]" captionStartId-18="45.[174,210,1526,1548]" captionStartId-2="21.[174,210,1525,1547]" captionStartId-3="23.[174,210,1462,1484]" captionStartId-4="24.[137,173,1473,1495]" captionStartId-5="26.[138,174,1501,1523]" captionStartId-6="29.[175,211,1499,1521]" captionStartId-7="30.[136,172,1528,1550]" captionStartId-8="31.[174,210,1550,1572]" captionStartId-9="32.[136,172,1496,1518]" captionTargetBox-0="[190,1108,185,1431]" captionTargetBox-1="[136,1088,741,1458]" captionTargetBox-10="[137,1088,192,1429]" captionTargetBox-11="[206,1099,869,1523]" captionTargetBox-12="[145,1079,734,1532]" captionTargetBox-13="[174,1126,338,1456]" captionTargetBox-14="[178,1047,190,1436]" captionTargetBox-15="[174,1126,742,1432]" captionTargetBox-16="[137,1088,202,1424]" captionTargetBox-17="[153,1078,189,1497]" captionTargetBox-18="[178,1122,408,1502]" captionTargetBox-2="[177,1126,798,1501]" captionTargetBox-3="[175,1124,193,1429]" captionTargetBox-4="[140,1081,642,1444]" captionTargetBox-5="[138,1088,854,1477]" captionTargetBox-6="[181,1121,518,1478]" captionTargetBox-7="[151,1074,797,1499]" captionTargetBox-8="[178,1120,1043,1522]" captionTargetBox-9="[136,1088,459,1472]" captionTargetId-0="figure-17@17.[188,1111,182,1432]" captionTargetId-1="figure-209@20.[136,1088,741,1458]" captionTargetId-10="figure-19@36.[137,1088,192,1432]" captionTargetId-11="figure-203@37.[201,1099,869,1523]" captionTargetId-12="figure-215@38.[145,1079,734,1532]" captionTargetId-13="figure-56@39.[174,1126,335,1466]" captionTargetId-14="figure-19@40.[178,1047,189,1439]" captionTargetId-15="figure-170@41.[174,1126,742,1432]" captionTargetId-16="figure-19@42.[136,1088,189,1424]" captionTargetId-17="figure-18@44.[147,1078,186,1497]" captionTargetId-18="figure-63@45.[178,1122,408,1502]" captionTargetId-2="figure-168@21.[174,1126,798,1504]" captionTargetId-3="figure-17@23.[174,1124,189,1429]" captionTargetId-4="figure-134@24.[136,1088,638,1444]" captionTargetId-5="figure-184@26.[136,1088,850,1480]" captionTargetId-6="figure-88@29.[180,1121,518,1478]" captionTargetId-7="figure-172@30.[151,1074,797,1499]" captionTargetId-8="figure-259@31.[174,1126,1037,1529]" captionTargetId-9="figure-84@32.[136,1088,459,1472]" captionTargetPageId-0="17" captionTargetPageId-1="20" captionTargetPageId-10="36" captionTargetPageId-11="37" captionTargetPageId-12="38" captionTargetPageId-13="39" captionTargetPageId-14="40" captionTargetPageId-15="41" captionTargetPageId-16="42" captionTargetPageId-17="44" captionTargetPageId-18="45" captionTargetPageId-2="21" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="26" captionTargetPageId-6="29" captionTargetPageId-7="30" captionTargetPageId-8="31" captionTargetPageId-9="32" captionText-0="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," captionText-1="Fig. 8. Nalassus (s. str.) brunneus brunneus, details of structure: A = head and prothorax, ventral view, B = head, ventral view (arrow shows weakly developed posterior grooves beginning at lower aspect of eyes), C = prosternum, D = prosternal process, procoxae and" captionText-10="Fig. 18. Asialassus cordicollis, details of structure: A = head, ventral view, B = male prosternum, C = male prosternal process, D = female prothorax and fore legs, ventral view, E = female prosternal process, ventral view, F = female prothorax, posterior transverse view (lateral arrows show elevate margin of procoxae, middle arrow shows depressed prosternal process between procoxae), G = male protibia, H = male mesotibia" captionText-11="Fig. 19. Asialassus amamiensis, holotype, female: A = habitus, B = protibia, C = prothorax, ventral view, D = labels of the holotype" captionText-12="Fig. 20. Asialassus amamiensis, male, details of structure: A = head, B = prosternum, C = prosternal process, ventral view, D = prosternal process, lateral view" captionText-13="Fig. 21. Asialassus kimurai, holotype and paratype: A = holotype, male, habitus, B = paratype, female, habitus, C = male protibia, dorsally, D = male protibia, ventrally, E = male mesotibia, F = male prothorax ventro-laterally, G = male prothorax, ventrally, H = aedea-" captionText-14="Fig. 22. Asialassus kimurai, holotype and paratype of the new junior synonym Tarpela tokunoshimana: A = holotype, male, habitus, B = paratype, female, habitus, C = male prothorax, ventrally, D = male prothorax, laterally, E = male protibia, dorsally, F = male protibia, ventrally, G = male mesotibia, dorsally, H = male mesotibia, ventrally, I = labels of the holo-" captionText-15="Fig. 23. Asialassus kimurai, details of structure, male: A = head, ventral view, B = prothorax, ventral view, C = prosternal process, D = protibia, ventrally" captionText-16="Fig. 24. Asialassus tsushimanus, male, holotype: A = habitus, B = protibia, laterally, C = protibia, ventrally, D = mesotibia, E = aedeagus, dorsal view, F = prothorax, ventral view, G =" captionText-17="Fig. 25. Asialassus todai, male, holotype: A = habitus, B = prothorax, ventral view, C = prothorax, lateral view, D = protibia, dorsal view, E = protibia, ventral view, F = mesotibia, G =" captionText-18="Fig. 26. Asialassus adachii, male, holotype: A = habitus, B = protibia dorsally, C = protibia ventrally, D = mesotibia, E = prothorax, ventral view, F = prosternal process, lateral view, G = aedeagus, H = labels of the holotype" captionText-2="Fig. 9. Nalassus brunneus konoi, holotype, female: A = habitus, B = labels, C = protibia, D = mesotibia" captionText-3="Fig. 10. Nalassus (s. str.) akitai, holotype and paratype: A = male, holotype, habitus, B = fe-" captionText-4="Fig. 11. Nalassus (s. str.) kawabatai, holotype and paratype. A = male, holotype, habitus, B = aedeagus, dorsal view, C = female paratype, habitus, D = male protibia, E = male mesotibia, F = male metatibia, G = male profemur, inner side, H = male prothorax, ventral view, I =" captionText-5="Fig. 12. Nalassus (s. str.) from Taiwan, habitus. A = N. formosanus, female, B = N. zoltani, male, C = N. pilushenmuus, male" captionText-6="Fig. 13. Nalassus (Nipponalassus) andoi, holotype and paratype. A = male, holotype, habitus, B = female paratype, habitus, C = male fore leg, dorsal margin, D = male profemur, inner side, E = aedeagus, dorsal view, F = prothorax, ventral view, G = abdominal ventrite 5, H = labels of the holotype" captionText-7="Fig. 14. Nalassus (Nipponalassus) andoi, details of structure. A = Head, ventral view, B = prothorax and base of fore legs, ventral side, C = male meso- and metaventrite, middle legs" captionText-8="Fig. 15. Nalassus (Nipponalassus) lewisi, holotype and paratype.A = male, holotype, habitus, B = female paratype, habitus, C = labels of the holotype, D = aedeagus, dorsal view" captionText-9="Fig. 16. Nalassus (Nipponalassus) lewisi (A, C–F) and N. (Nipponalassus) andoi (B), details of structure: A = head, ventral view, B = male abdominal ventrite 5, C = male meso- and metaventrite, middle legs basally, D = male abdominal ventrites, E = male profemur, inner side, F = male profemur dorsally" figureDoi-0="http://doi.org/10.5281/zenodo.12626333" figureDoi-1="http://doi.org/10.5281/zenodo.12626335" figureDoi-10="http://doi.org/10.5281/zenodo.12626355" figureDoi-11="http://doi.org/10.5281/zenodo.12626357" figureDoi-12="http://doi.org/10.5281/zenodo.12626359" figureDoi-13="http://doi.org/10.5281/zenodo.12626361" figureDoi-14="http://doi.org/10.5281/zenodo.12626363" figureDoi-15="http://doi.org/10.5281/zenodo.12626365" figureDoi-16="http://doi.org/10.5281/zenodo.12626367" figureDoi-17="http://doi.org/10.5281/zenodo.12626371" figureDoi-18="http://doi.org/10.5281/zenodo.12626375" figureDoi-2="http://doi.org/10.5281/zenodo.12626337" figureDoi-3="http://doi.org/10.5281/zenodo.12626339" figureDoi-4="http://doi.org/10.5281/zenodo.12626341" figureDoi-5="http://doi.org/10.5281/zenodo.12626343" figureDoi-6="http://doi.org/10.5281/zenodo.12626345" figureDoi-7="http://doi.org/10.5281/zenodo.12626347" figureDoi-8="http://doi.org/10.5281/zenodo.12626349" figureDoi-9="http://doi.org/10.5281/zenodo.12626353" httpUri-0="https://zenodo.org/record/12626333/files/figure.png" httpUri-1="https://zenodo.org/record/12626335/files/figure.png" httpUri-10="https://zenodo.org/record/12626355/files/figure.png" httpUri-11="https://zenodo.org/record/12626357/files/figure.png" httpUri-12="https://zenodo.org/record/12626359/files/figure.png" httpUri-13="https://zenodo.org/record/12626361/files/figure.png" httpUri-14="https://zenodo.org/record/12626363/files/figure.png" httpUri-15="https://zenodo.org/record/12626365/files/figure.png" httpUri-16="https://zenodo.org/record/12626367/files/figure.png" httpUri-17="https://zenodo.org/record/12626371/files/figure.png" httpUri-18="https://zenodo.org/record/12626375/files/figure.png" httpUri-2="https://zenodo.org/record/12626337/files/figure.png" httpUri-3="https://zenodo.org/record/12626339/files/figure.png" httpUri-4="https://zenodo.org/record/12626341/files/figure.png" httpUri-5="https://zenodo.org/record/12626343/files/figure.png" httpUri-6="https://zenodo.org/record/12626345/files/figure.png" httpUri-7="https://zenodo.org/record/12626347/files/figure.png" httpUri-8="https://zenodo.org/record/12626349/files/figure.png" httpUri-9="https://zenodo.org/record/12626353/files/figure.png" pageId="15" pageNumber="292">Figs 7–26</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="8BAD1F30FFF5FFFAFF26A58355403D76" blockId="15.[173,1127,1262,1601]" pageId="15" pageNumber="292">
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General differential morphology. Adult. Head with (
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<figureCitation id="132903B5FFF5FFFAFCD8A58355223D02" box="[788,883,1262,1284]" captionStart="Fig" captionStartId="30.[136,172,1528,1550]" captionTargetBox="[151,1074,797,1499]" captionTargetId="figure-172@30.[151,1074,797,1499]" captionTargetPageId="30" captionText="Fig. 14. Nalassus (Nipponalassus) andoi, details of structure. A = Head, ventral view, B = prothorax and base of fore legs, ventral side, C = male meso- and metaventrite, middle legs" figureDoi="http://doi.org/10.5281/zenodo.12626347" httpUri="https://zenodo.org/record/12626347/files/figure.png" pageId="15" pageNumber="292">Figs 14A</figureCitation>
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,
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<figureCitation id="132903B5FFF5FFFAFCB0A58355F73D05" box="[892,934,1262,1284]" captionStart="Fig" captionStartId="36.[136,172,1459,1481]" captionTargetBox="[137,1088,192,1429]" captionTargetId="figure-19@36.[137,1088,192,1432]" captionTargetPageId="36" captionText="Fig. 18. Asialassus cordicollis, details of structure: A = head, ventral view, B = male prosternum, C = male prosternal process, D = female prothorax and fore legs, ventral view, E = female prosternal process, ventral view, F = female prothorax, posterior transverse view (lateral arrows show elevate margin of procoxae, middle arrow shows depressed prosternal process between procoxae), G = male protibia, H = male mesotibia" figureDoi="http://doi.org/10.5281/zenodo.12626355" httpUri="https://zenodo.org/record/12626355/files/figure.png" pageId="15" pageNumber="292">18A</figureCitation>
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) or rarely without posterior grooves (
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<figureCitation id="132903B5FFF5FFFAFEBAA46657873D21" box="[374,470,1291,1313]" captionStart="Fig" captionStartId="38.[135,171,1552,1574]" captionTargetBox="[145,1079,734,1532]" captionTargetId="figure-215@38.[145,1079,734,1532]" captionTargetPageId="38" captionText="Fig. 20. Asialassus amamiensis, male, details of structure: A = head, B = prosternum, C = prosternal process, ventral view, D = prosternal process, lateral view" figureDoi="http://doi.org/10.5281/zenodo.12626359" httpUri="https://zenodo.org/record/12626359/files/figure.png" pageId="15" pageNumber="292">Figs 20A</figureCitation>
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,
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<figureCitation id="132903B5FFF5FFFAFE2DA466545A3D20" box="[481,523,1291,1313]" captionStart="Fig" captionStartId="41.[174,210,1458,1480]" captionTargetBox="[174,1126,742,1432]" captionTargetId="figure-170@41.[174,1126,742,1432]" captionTargetPageId="41" captionText="Fig. 23. Asialassus kimurai, details of structure, male: A = head, ventral view, B = prothorax, ventral view, C = prosternal process, D = protibia, ventrally" figureDoi="http://doi.org/10.5281/zenodo.12626365" httpUri="https://zenodo.org/record/12626365/files/figure.png" pageId="15" pageNumber="292">23A</figureCitation>
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) or with deep wrinkle (
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<figureCitation id="132903B5FFF5FFFAFCC5A46655083D21" box="[777,857,1291,1313]" captionStart="Fig" captionStartId="20.[136,172,1483,1505]" captionTargetBox="[136,1088,741,1458]" captionTargetId="figure-209@20.[136,1088,741,1458]" captionTargetPageId="20" captionText="Fig. 8. Nalassus (s. str.) brunneus brunneus, details of structure: A = head and prothorax, ventral view, B = head, ventral view (arrow shows weakly developed posterior grooves beginning at lower aspect of eyes), C = prosternum, D = prosternal process, procoxae and" figureDoi="http://doi.org/10.5281/zenodo.12626335" httpUri="https://zenodo.org/record/12626335/files/figure.png" pageId="15" pageNumber="292">Figs 8B</figureCitation>
|
||
,
|
||
<figureCitation id="132903B5FFF5FFFAFCA8A46655DE3D20" box="[868,911,1291,1313]" captionStart="Fig" captionStartId="32.[136,172,1496,1518]" captionTargetBox="[136,1088,459,1472]" captionTargetId="figure-84@32.[136,1088,459,1472]" captionTargetPageId="32" captionText="Fig. 16. Nalassus (Nipponalassus) lewisi (A, C–F) and N. (Nipponalassus) andoi (B), details of structure: A = head, ventral view, B = male abdominal ventrite 5, C = male meso- and metaventrite, middle legs basally, D = male abdominal ventrites, E = male profemur, inner side, F = male profemur dorsally" figureDoi="http://doi.org/10.5281/zenodo.12626353" httpUri="https://zenodo.org/record/12626353/files/figure.png" pageId="15" pageNumber="292">16A</figureCitation>
|
||
) beginning at lower aspect of the eyes. Labrofrontoclypeal membrane always exposed. Abdominal ventrite 5 not beaded along outer margin in most of all taxa excluding
|
||
<emphasis id="B966C322FFF5FFFAFCE6A42855EE3D5B" box="[810,959,1349,1370]" italics="true" pageId="15" pageNumber="292">Microdocnemis</emphasis>
|
||
Nabozhenko et Keskin, 2010 (but often with finely beaded posterior end).
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFF5FFE5FF26A413556A3946" blockId="15.[173,1127,1262,1601]" lastBlockId="16.[135,1090,190,385]" lastPageId="16" lastPageNumber="293" pageId="15" pageNumber="292">
|
||
Male genitalia. Apical piece flattened laterally at apex (
|
||
<figureCitation id="132903B5FFF5FFFAFC99A41355ED3D95" box="[853,956,1406,1428]" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="15" pageNumber="292">Figs 7B,F</figureCitation>
|
||
, 17B,G,L), with punctation, each puncture with very small seta directed toward the apex (‘nalassoid’
|
||
<typeStatus id="54A9A192FFF5FFFAFF62A4D5568C3DCC" box="[174,221,1464,1485]" pageId="15" pageNumber="292">type</typeStatus>
|
||
of genitalia) or apical piece flattened dorso-ventrally at apex (
|
||
<figureCitation id="132903B5FFF5FFFAFCBDA4D555943DCF" box="[881,965,1464,1486]" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="15" pageNumber="292">Figs 7G</figureCitation>
|
||
), neither punctured nor setiferous (‘cylindrinotoid’
|
||
<typeStatus id="54A9A192FFF5FFFAFD89A4B854253DEB" box="[581,628,1493,1514]" pageId="15" pageNumber="292">type</typeStatus>
|
||
of genitalia). Alae of apical piece moderately long, merged, concealed only half of median lobe, not reaching base of basal piece (
|
||
<figureCitation id="132903B5FFF5FFFAFBF5A49C575D3E22" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="15" pageNumber="292">Figs 7A,E,I,O</figureCitation>
|
||
, 17A,F,K). Basal piece separated from alae by connecting membrane only in apical half, with connecting membrane between margins in basal half. Median lobe acute apically, only with one apex, with weakly sclerotized baculi, which are not connected at base. Female genital tubes. Spermatheca composed of single long tube and short accessory gland (
|
||
<figureCitation id="132903B5FFEAFFE5FF19A195577E390C" box="[213,303,248,270]" captionStart="Fig" captionStartId="17.[174,210,1457,1479]" captionTargetBox="[190,1108,185,1431]" captionTargetId="figure-17@17.[188,1111,182,1432]" captionTargetPageId="17" captionText="Fig. 7. The genus Nalassus, genital structure: N. (s. str.) pilushenmuus (A–D), N. (s. str.) zoltani (E–H), N. (Nipponalassus) andoi (I–M), N. (Nipponalassus) lewisi (O–R): A, E, I, O = aedeagus, ventral view, B, F, J, P = aedeagus, lateral view, C, G, K, Q = male inner sternite VIII, D, H, L, R = gastral spicula, M = female genital tubes (ag – accessory gland of spermatheca," figureDoi="http://doi.org/10.5281/zenodo.12626333" httpUri="https://zenodo.org/record/12626333/files/figure.png" pageId="16" pageNumber="293">Figs 7M</figureCitation>
|
||
, 17P). Only the subgenus
|
||
<taxonomicName id="4C1264B3FFEAFFE5FD9EA19555FE390F" authority="Nabozhenko, 2006" authorityName="Nabozhenko" authorityYear="2006" box="[594,943,248,270]" class="Insecta" family="Tenebrionidae" genus="Caucasohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFEAFFE5FD9EA195548F390C" box="[594,734,248,269]" italics="true" pageId="16" pageNumber="293">Caucasohelops</emphasis>
|
||
Nabozhenko, 2006
|
||
</taxonomicName>
|
||
of the genus
|
||
<taxonomicName id="4C1264B3FFEAFFE5FF44A07857AF392A" authority="Nabozhenko, 2006" authorityName="Nabozhenko" authorityYear="2006" box="[136,510,277,299]" class="Insecta" family="Tenebrionidae" genus="Eustenomacidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966C322FFEAFFE5FF44A078577F392B" box="[136,302,277,298]" italics="true" pageId="16" pageNumber="293">Eustenomacidius</emphasis>
|
||
Nabozhenko, 2006
|
||
</taxonomicName>
|
||
has multibranched spermatheca (NABOZHENKO 2006). Valve between spermathecal and accessory gland ducts is absent.
|
||
</paragraph>
|
||
<paragraph id="8BAD1F30FFEAFFE5FF09A022555D3980" blockId="16.[135,1090,190,385]" pageId="16" pageNumber="293">Larva. Abdominal segment IX with small cylindrical or conical protuberances at base of urogomphi. Abdominal tergit VIII without protuberances.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |