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<document id="53E637E6369D5D9605659867D3828E3A" ID-CLB-Dataset="4090" ID-DOI="10.5852/ejt.2021.773.1511" ID-GBIF-Dataset="38bb3ec2-51c6-4ffe-bf28-6f28266c2eaa" ID-ISSN="2118-9773" ID-Zenodo-Dep="5536387" ID-ZooBank="FD4A74C5-9746-4DB0-A252-82711FEB13E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632919232762" checkinUser="felipe" docAuthor="Martínez, Leonel, Sánchez-Ruiz, Alexander &amp; Bonaldo, Alexandre B." docDate="2021" docId="03B887D5FF92FFBF55DBF98AFC49D14F" docLanguage="en" docName="ejt-773_61-79.pdf" docOrigin="European Journal of Taxonomy 773" docSource="http://dx.doi.org/10.5852/ejt.2021.773.1511" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Medionops blades Sanchez-Ruiz &amp; Brescovit 2017" docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="FF81FFADFF91FFBB543CFFB1FFDED713" masterDocTitle="The spider genus Medionops Sánchez-Ruiz &amp; Brescovit (Araneae: Caponiidae) in Colombia, with the description of four new species" masterLastPageNumber="79" masterPageNumber="61" pageNumber="64" updateTime="1699458186880" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="85AB1A104A999B71227C60903ABFE984">The spider genus Medionops Sánchez-Ruiz &amp; Brescovit (Araneae: Caponiidae) in Colombia, with the description of four new species</mods:title>
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<mods:namePart id="551E287A4EB6A578A22A0A30A2A3C0F5">Martínez, Leonel</mods:namePart>
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<mods:affiliation id="BE4BA9F0D771A2C4AD7CB732F1F895EA">Grupo de Investigación Biodiversidad del Caribe Colombiano, Semillero de Investigación Sistemática de Artrópodos Neotropicales, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia.</mods:affiliation>
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<mods:namePart id="8F66D12494569CF5D4A63229A4F26274">Sánchez-Ruiz, Alexander</mods:namePart>
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<mods:affiliation id="582399FDF5C43DE6077D14DA1023651C">Museu Paraense Emílio Goeldi, Coordenação de Zoologia, Laboratório de Aracnologia, Av. Perimetral, 1901, Terra Firme, CEP 66077 - 830, Belém, Pará, Brazil.</mods:affiliation>
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<mods:affiliation id="38DD7AD98AB122B31DBB960E097246E6">Museu Paraense Emílio Goeldi, Coordenação de Zoologia, Laboratório de Aracnologia, Av. Perimetral, 1901, Terra Firme, CEP 66077 - 830, Belém, Pará, Brazil.</mods:affiliation>
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Sánchez-Ruiz &amp; Brescovit, 2017
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<paragraph id="8BAE36C3FF92FFB8569FF9EEFC4FD16A" blockId="3.[487,1101,1595,1657]" box="[675,913,1631,1657]" pageId="3" pageNumber="64">
<figureCitation id="132A2A46FF92FFB8569FF9EEFD32D16A" box="[675,748,1631,1657]" captionStart="Fig" captionStartId="5.[189,232,1786,1812]" captionTargetBox="[226,1366,279,1736]" captionTargetId="figure-12@5.[210,1378,265,1748]" captionTargetPageId="5" captionText="Fig. 1. Medionops blades Sánchez-Ruiz &amp; Brescovit, 2017. A–B, E–F. Male, Boyacá population (IAvH-I 3381). C–D, G–H. Female, Boyacá population (IAvH-I 3382). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; ess = external sclerotization around spiracles; go = genital opening; re = receptaculum; ue = uterus externo." figureDoi="http://doi.org/10.5281/zenodo.5536389" httpUri="https://zenodo.org/record/5536389/files/figure.png" pageId="3" pageNumber="64">Figs 1</figureCitation>
, 
<figureCitation id="132A2A46FF92FFB856C6F9EEFCD5D16A" box="[762,779,1631,1657]" captionStart="Fig" captionStartId="14.[187,230,1998,2024]" captionTargetBox="[230,1361,1150,1933]" captionTargetId="figure-321@14.[189,1398,1136,1959]" captionTargetPageId="14" captionText="Fig. 6. Distribution map of Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017." figureDoi="http://doi.org/10.5281/zenodo.5536399" httpUri="https://zenodo.org/record/5536399/files/figure.png" pageId="3" pageNumber="64">6</figureCitation>
, 
<figureCitation id="132A2A46FF92FFB85724F9EEFCE0D16A" box="[792,830,1631,1657]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[308,1294,450,1852]" captionTargetId="figure-57@15.[264,1323,427,1857]" captionTargetPageId="15" captionText="Fig. 7. Drawings of the male copulatory bulbs (retrolateral view) in Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017. A. M. blades Sánchez-Ruiz &amp; Brescovit, 2017. B. M. carolinae sp. nov. C. M. cauca sp. nov. D. M. luiscarlosi sp. nov. E. M. santarosa sp. nov. Scale bars = 0.5 mm. Abbreviations: e = embolus; t = tegulum." figureDoi="http://doi.org/10.5281/zenodo.5536401" httpUri="https://zenodo.org/record/5536401/files/figure.png" pageId="3" pageNumber="64">7A</figureCitation>
, 
<figureCitation id="132A2A46FF92FFB85776F9EEFC4FD16A" box="[842,913,1631,1657]" captionStart="Fig" captionStartId="16.[189,232,1857,1883]" captionTargetBox="[199,1388,463,1818]" captionTargetId="figure-79@16.[189,1398,456,1821]" captionTargetPageId="16" captionText="Fig. 8. Habitats of Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017. A–B. Type locality of M. blades Sánchez-Ruiz &amp; Brescovit, 2017, Páramo de Monserrate, Cundimarca department, Colombia. C–D. Type locality of M. carolinae sp. nov., Sutamarchán, Boyacá department, Colombia. E–F. Habitat of female paratype (IAvH-I 3771) of M. carolinae sp. nov., Sector Carrizal, Santuario de Flora y Fauna Iguaque, Boyacá department, Colombia." figureDoi="http://doi.org/10.5281/zenodo.5536403" httpUri="https://zenodo.org/record/5536403/files/figure.png" pageId="3" pageNumber="64">8A–B</figureCitation>
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<emphasis id="B965EAD1FF92FFB85481F911FE4BD1A9" box="[189,405,1696,1722]" italics="true" pageId="3" pageNumber="64">Medionops blades</emphasis>
Sánchez-Ruiz &amp; Brescovit 2017: 122
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, figs 1e, 2a–l, 3a–l, 4a–l, 5a–l, 6a–e (
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).
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<emphasis id="B965EAD1FF92FFB85481F950FE4BD1E8" box="[189,405,1761,1787]" italics="true" pageId="3" pageNumber="64">Medionops blades</emphasis>
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– 
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<bibRefCitation id="EF804B32FF92FFB8558FF950FC00D1EF" author="Sanchez-Ruiz A. &amp; Brescovit A. D. &amp; Bonaldo A. B." box="[435,990,1761,1788]" firstAuthor="Sanchez-Ruiz" journalOrPublisher="Zootaxa" page="485" pageId="3" pageNumber="64" pagination="461 - 486" part="4751" refId="ref7792" refString="Sanchez-Ruiz A., Brescovit A. D. &amp; Bonaldo A. B. 2020. Revision of the spider genus Nyetnops Platnick &amp; Lise (Araneae: Caponiidae) with proposition of the new genus Nopsma, from Central and South America. Zootaxa 4751 (3): 461 - 486. https: // doi. org / 10.11646 / zootaxa. 4751.3.3" title="Revision of the spider genus Nyetnops Platnick &amp; Lise (Araneae: Caponiidae) with proposition of the new genus Nopsma, from Central and South America" type="journal article" year="2020">Sánchez-Ruiz, Brescovit &amp; Bonaldo 2020: 485</bibRefCitation>
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, fig. 19e (
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).
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<emphasis id="B965EAD1FF92FFB85481F892FEE9D02E" bold="true" box="[189,311,1827,1853]" pageId="3" pageNumber="64">Diagnosis</emphasis>
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<paragraph id="8BAE36C3FF92FFBF5481F8E0FCA5D655" blockId="3.[187,1399,1873,1970]" lastBlockId="4.[188,1399,265,327]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
Males resemble those of 
<taxonomicName id="4C114D40FF92FFB855CDF8E3FD53D078" authority="Martínez &amp; Sánchez-Ruiz &amp; Bonaldo, 2021" authorityName="Martínez &amp; Sánchez-Ruiz &amp; Bonaldo" authorityYear="2021" box="[497,653,1873,1899]" class="Arachnida" family="Caponiidae" genus="Medionops" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B965EAD1FF92FFB855CDF8E3FD53D078" box="[497,653,1873,1899]" italics="true" pageId="3" pageNumber="64">M. carolinae</emphasis>
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in having similar copulatory bulb shape, but can be distinguished by the straighter embolus tip and by the size of the tegulum, only ½ as long as the cymbium length (
<figureCitation id="132A2A46FF92FFB85528F829FE51D0A1" box="[276,399,1944,1970]" captionStart="Fig" captionStartId="5.[189,232,1786,1812]" captionTargetBox="[226,1366,279,1736]" captionTargetId="figure-12@5.[210,1378,265,1748]" captionTargetPageId="5" captionText="Fig. 1. Medionops blades Sánchez-Ruiz &amp; Brescovit, 2017. A–B, E–F. Male, Boyacá population (IAvH-I 3381). C–D, G–H. Female, Boyacá population (IAvH-I 3382). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; ess = external sclerotization around spiracles; go = genital opening; re = receptaculum; ue = uterus externo." figureDoi="http://doi.org/10.5281/zenodo.5536389" httpUri="https://zenodo.org/record/5536389/files/figure.png" pageId="3" pageNumber="64">Figs 1E–F</figureCitation>
, 
<figureCitation id="132A2A46FF92FFB855A7F829FE1ED0A1" box="[411,448,1944,1970]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[308,1294,450,1852]" captionTargetId="figure-57@15.[264,1323,427,1857]" captionTargetPageId="15" captionText="Fig. 7. Drawings of the male copulatory bulbs (retrolateral view) in Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017. A. M. blades Sánchez-Ruiz &amp; Brescovit, 2017. B. M. carolinae sp. nov. C. M. cauca sp. nov. D. M. luiscarlosi sp. nov. E. M. santarosa sp. nov. Scale bars = 0.5 mm. Abbreviations: e = embolus; t = tegulum." figureDoi="http://doi.org/10.5281/zenodo.5536401" httpUri="https://zenodo.org/record/5536401/files/figure.png" pageId="3" pageNumber="64">7A</figureCitation>
) (in 
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<emphasis id="B965EAD1FF92FFB855C4F828FD4ED0A1" box="[504,656,1944,1970]" italics="true" pageId="3" pageNumber="64">M. carolinae</emphasis>
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<taxonomicNameLabel id="A25657AAFF92FFB856ABF829FD2AD0A1" box="[663,756,1944,1970]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
, only ⅓ as long, 
<figureCitation id="132A2A46FF92FFB85786F829FBEAD0A1" box="[954,1076,1944,1970]" captionStart="Fig" captionStartId="7.[189,232,1820,1846]" captionTargetBox="[206,1372,302,1771]" captionTargetId="figure-12@7.[189,1398,286,1783]" captionTargetPageId="7" captionText="Fig. 2. Medionops carolinae sp. nov. A–B, E–F. Holotype, ♂ (IAvH-I 3782). C–D, G–H. Paratype, ♀ (IAvH-I 3771). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; b = area below genital opening; ess = external sclerotization around spiracles; go = genital opening." figureDoi="http://doi.org/10.5281/zenodo.5536391" httpUri="https://zenodo.org/record/5536391/files/figure.png" pageId="3" pageNumber="64">Figs 2E–F</figureCitation>
, 
<figureCitation id="132A2A46FF92FFB85003F829FBBBD0A1" box="[1087,1125,1944,1970]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[308,1294,450,1852]" captionTargetId="figure-57@15.[264,1323,427,1857]" captionTargetPageId="15" captionText="Fig. 7. Drawings of the male copulatory bulbs (retrolateral view) in Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017. A. M. blades Sánchez-Ruiz &amp; Brescovit, 2017. B. M. carolinae sp. nov. C. M. cauca sp. nov. D. M. luiscarlosi sp. nov. E. M. santarosa sp. nov. Scale bars = 0.5 mm. Abbreviations: e = embolus; t = tegulum." figureDoi="http://doi.org/10.5281/zenodo.5536401" httpUri="https://zenodo.org/record/5536401/files/figure.png" pageId="3" pageNumber="64">7B</figureCitation>
). Females of 
<taxonomicName id="4C114D40FF92FFB8513EF828FAA8D0A1" authorityName="Sanchez-Ruiz &amp; Brescovit" authorityYear="2017" box="[1282,1398,1944,1970]" class="Arachnida" family="Caponiidae" genus="Medionops" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="blades">
<emphasis id="B965EAD1FF92FFB8513EF828FAA8D0A1" box="[1282,1398,1944,1970]" italics="true" pageId="3" pageNumber="64">M. blades</emphasis>
</taxonomicName>
differ from those of 
<taxonomicName id="4C114D40FF95FFBF559AFEBBFDE3D630" authority="Martínez &amp; Sánchez-Ruiz &amp; Bonaldo, 2021" authorityName="Martínez &amp; Sánchez-Ruiz &amp; Bonaldo" authorityYear="2021" box="[422,573,265,291]" class="Arachnida" family="Caponiidae" genus="Medionops" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B965EAD1FF95FFBF559AFEBBFDE3D630" box="[422,573,265,291]" italics="true" pageId="4" pageNumber="65">M. carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657AAFF95FFBF5678FEBBFD40D637" box="[580,670,266,292]" pageId="4" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
by the wide, concave area below the genital opening (
<figureCitation id="132A2A46FF95FFBF512FFEB8FAB2D630" box="[1299,1388,265,291]" captionStart="Fig" captionStartId="5.[189,232,1786,1812]" captionTargetBox="[226,1366,279,1736]" captionTargetId="figure-12@5.[210,1378,265,1748]" captionTargetPageId="5" captionText="Fig. 1. Medionops blades Sánchez-Ruiz &amp; Brescovit, 2017. A–B, E–F. Male, Boyacá population (IAvH-I 3381). C–D, G–H. Female, Boyacá population (IAvH-I 3382). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; ess = external sclerotization around spiracles; go = genital opening; re = receptaculum; ue = uterus externo." figureDoi="http://doi.org/10.5281/zenodo.5536389" httpUri="https://zenodo.org/record/5536389/files/figure.png" pageId="4" pageNumber="65">Fig. 1G</figureCitation>
) (in 
<taxonomicName id="4C114D40FF95FFBF54D9FE9CFEA3D655" authority="Martínez &amp; Sánchez-Ruiz &amp; Bonaldo, 2021" authorityName="Martínez &amp; Sánchez-Ruiz &amp; Bonaldo" authorityYear="2021" box="[229,381,300,326]" class="Arachnida" family="Caponiidae" genus="Medionops" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B965EAD1FF95FFBF54D9FE9CFEA3D655" box="[229,381,300,326]" italics="true" pageId="4" pageNumber="65">M. carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657AAFF95FFBF55B9FE9CFE3CD654" box="[389,482,301,327]" pageId="4" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
, narrower and triangular, 
<figureCitation id="132A2A46FF95FFBF572CFE9DFCB0D655" box="[784,878,300,326]" captionStart="Fig" captionStartId="7.[189,232,1820,1846]" captionTargetBox="[206,1372,302,1771]" captionTargetId="figure-12@7.[189,1398,286,1783]" captionTargetPageId="7" captionText="Fig. 2. Medionops carolinae sp. nov. A–B, E–F. Holotype, ♂ (IAvH-I 3782). C–D, G–H. Paratype, ♀ (IAvH-I 3771). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; b = area below genital opening; ess = external sclerotization around spiracles; go = genital opening." figureDoi="http://doi.org/10.5281/zenodo.5536391" httpUri="https://zenodo.org/record/5536391/files/figure.png" pageId="4" pageNumber="65">Fig. 2G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6548FF95FFBF5481FEDCFAA9D4F8" pageId="4" pageNumber="65" type="materials_examined">
<paragraph id="8BAE36C3FF95FFBF5481FEDCFE73D694" blockId="4.[189,429,365,391]" box="[189,429,365,391]" pageId="4" pageNumber="65">
<heading id="D0E681AFFF95FFBF5481FEDCFE73D694" bold="true" box="[189,429,365,391]" fontSize="11" level="3" pageId="4" pageNumber="65" reason="3">
<emphasis id="B965EAD1FF95FFBF5481FEDCFE73D694" bold="true" box="[189,429,365,391]" pageId="4" pageNumber="65">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF54D6FE2DFCB3D6EE" blockId="4.[189,1399,412,509]" pageId="4" pageNumber="65">
<materialsCitation id="3B793C9EFF95FFBF54D6FE2DFCB3D6EE" ID-GBIF-Occurrence="3382549301" collectingDate="1968-07-23" collectionCode="AMNH" collectorName="H. Sturm" country="Colombia" elevation="2920" latitude="4.6" location="Paramo de Monserrate" longLatPrecision="1306" longitude="-74.05" municipality="Cundimarca" pageId="4" pageNumber="65" specimenCount="1" specimenCount-male="1" stateProvince="Bogota" typeStatus="holotype">
<emphasis id="B965EAD1FF95FFBF54D6FE2DFE83D6A5" bold="true" box="[234,349,412,438]" pageId="4" pageNumber="65">
<typeStatus id="54AA8861FF95FFBF54D6FE2DFE83D6A5" box="[234,349,412,438]" pageId="4" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3067653FF95FFBF54F4FE0EFEBDD6CA" box="[200,355,447,473]" name="Colombia" pageId="4" pageNumber="65">COLOMBIA</collectingCountry>
• 
<specimenCount id="9D17FD4AFF95FFBF55BEFE71FE45D6C9" box="[386,411,448,474]" pageId="4" pageNumber="65" type="male">♂</specimenCount>
; 
<collectingMunicipality id="6BCAACB9FF95FFBF5597FE0EFDE4D6CA" box="[427,570,447,473]" pageId="4" pageNumber="65">Cundimarca</collectingMunicipality>
department, 
<location id="8ECE6018FF95FFBF56E7FE0EFC30D6C9" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF56E7FE0EFC30D6C9" box="[731,1006,447,474]" country="Colombia" latitude="4.6" longLatPrecision="1306" longitude="-74.05" municipality="Cundimarca" name="Paramo de Monserrate" pageId="4" pageNumber="65" stateProvince="Bogota">Páramo de Monserrate</location>
, 
<locationDeviation id="9A1C2FE1FF95FFBF57C1FE0EFADCD6CA" box="[1021,1282,447,474]" pageId="4" pageNumber="65">
<quantity id="4CE99B26FF95FFBF57C1FE0EFB94D6CA" box="[1021,1098,447,473]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="65" unit="km" value="40.0">40 km</quantity>
NE of Bogotá
</locationDeviation>
; 
<geoCoordinate id="EE255004FF95FFBF512DFE0FFAACD6C9" box="[1297,1394,446,474]" degrees="4" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="65" precision="925" value="4.6">4°36′ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF5481FE50FEFED6EE" box="[189,288,481,509]" degrees="74" direction="west" minutes="3" orientation="longitude" pageId="4" pageNumber="65" precision="925" value="-74.05">74°3′ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF5564FE52FE6AD6EE" box="[344,436,483,509]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="4" pageNumber="65" unit="m" value="2920.0">
<elevation id="003CD1F0FF95FFBF5564FE52FE6AD6EE" box="[344,436,483,509]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="4" pageNumber="65" unit="m" value="2920.0">2920 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF55FCFE52FD8AD6EF" box="[448,596,482,509]" pageId="4" pageNumber="65" value="1968-07-23">
<collectingDate id="EFEBE9EBFF95FFBF55FCFE52FD8AD6EF" box="[448,596,482,509]" pageId="4" pageNumber="65" value="1968-07-23">23 Jul. 1968</collectingDate>
</date>
; 
<collectorName id="26E45315FF95FFBF565CFE52FD12D6EE" box="[608,716,483,509]" pageId="4" pageNumber="65">H. Sturm</collectorName>
leg.; 
<collectionCode id="ED00AE06FF95FFBF5730FE52FCB4D6EE" box="[780,874,483,509]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="65">AMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF54D6FD95FC1CD572" blockId="4.[189,962,548,610]" pageId="4" pageNumber="65">
<materialsCitation id="3B793C9EFF95FFBF54D6FD95FC61D572" ID-GBIF-Occurrence="3382549314" collectingDate="1968-07-23" collectionCode="AMNH" collectorName="de Monserrate &amp; H. Sturm" country="Colombia" elevation="2920" latitude="4.6" location="COLOMBIA" longLatPrecision="1306" longitude="-74.05" municipality="Cundimarca" pageId="4" pageNumber="65" specimenCount="1" specimenCount-female="1" stateProvince="Bogota" typeStatus="paratype">
<emphasis id="B965EAD1FF95FFBF54D6FD95FE83D52D" bold="true" box="[234,349,548,574]" pageId="4" pageNumber="65">
<typeStatus id="54AA8861FF95FFBF54D6FD95FE83D52D" box="[234,349,548,574]" pageId="4" pageNumber="65" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F3067653FF95FFBF5481FDF6FE86D572" box="[189,344,583,609]" name="Colombia" pageId="4" pageNumber="65">COLOMBIA</collectingCountry>
• 
<specimenCount id="9D17FD4AFF95FFBF554DFDF6FE40D571" box="[369,414,583,610]" pageId="4" pageNumber="65" type="female">1 ♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED00AE06FF95FFBF575DFDF6FC61D572" box="[865,959,583,609]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="65">AMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FD39FE69D5B1" blockId="4.[189,439,648,674]" box="[189,439,648,674]" pageId="4" pageNumber="65">
<heading id="D0E681AFFF95FFBF5481FD39FE69D5B1" bold="true" box="[189,439,648,674]" fontSize="11" level="3" pageId="4" pageNumber="65" reason="3">
<emphasis id="B965EAD1FF95FFBF5481FD39FE69D5B1" bold="true" box="[189,439,648,674]" pageId="4" pageNumber="65">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FD06FAA9D4F8" blockId="4.[189,1399,694,1004]" pageId="4" pageNumber="65">
<materialsCitation id="3B793C9EFF95FFBF5481FD06FBB4D5E7" ID-GBIF-Occurrence="3382549310" collectingDate="2015-06-08" collectingDateMax="2015-06-10" collectingDateMin="2015-06-08" collectorName="Y. Cifuentes &amp; J. Moreno" country="Colombia" elevation="3502" latitude="5.809111" location="Vereda Cuaguani" longLatPrecision="1" longitude="-73.25303" municipality="Sotaquira" pageId="4" pageNumber="65" specimenCode="IAvH-I 3381" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca department">
<collectingCountry id="F3067653FF95FFBF5481FD06FE86D5C2" box="[189,344,695,721]" name="Colombia" pageId="4" pageNumber="65">COLOMBIA</collectingCountry>
– 
<emphasis id="B965EAD1FF95FFBF5549FD07FDB2D5C2" bold="true" box="[373,620,694,721]" pageId="4" pageNumber="65">
<collectingRegion id="49D5F821FF95FFBF5549FD07FDB2D5C2" box="[373,620,694,721]" country="Colombia" name="Boyaca" pageId="4" pageNumber="65">Boyacá department</collectingRegion>
</emphasis>
• 
<specimenCount id="9D17FD4AFF95FFBF56BAFD06FD6AD5C1" box="[646,692,695,722]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BCAACB9FF95FFBF56FDFD06FCEAD5C2" box="[705,820,695,721]" pageId="4" pageNumber="65">Sotaquirá</collectingMunicipality>
, 
<location id="8ECE6018FF95FFBF5703FD06FBD2D5C2" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF5703FD06FBD2D5C2" box="[831,1036,695,721]" country="Colombia" latitude="5.809111" longLatPrecision="1" longitude="-73.25303" municipality="Sotaquira" name="Vereda Cuaguani" pageId="4" pageNumber="65" stateProvince="Boyaca department">Vereda Cuaguaní</location>
; 
<geoCoordinate id="EE255004FF95FFBF5024FD07FB6BD5C1" box="[1048,1205,694,722]" degrees="5" direction="north" minutes="48" orientation="latitude" pageId="4" pageNumber="65" precision="1" seconds="32.8" value="5.809111">5°48′32.8″ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF50FDFD07FAACD5C1" box="[1217,1394,694,722]" degrees="73" direction="west" minutes="15" orientation="longitude" pageId="4" pageNumber="65" precision="1" seconds="10.9" value="-73.25303">73°15′10.9″ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF54D2FD6BFE90D5E6" box="[238,334,730,757]" metricMagnitude="3" metricUnit="m" metricValue="3.502" pageId="4" pageNumber="65" unit="m" value="3502.0">
<elevation id="003CD1F0FF95FFBF54D2FD6BFE90D5E6" box="[238,334,730,757]" metricMagnitude="3" metricUnit="m" metricValue="3.502" pageId="4" pageNumber="65" unit="m" value="3502.0">3502 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF5563FD6BFDFED5E6" box="[351,544,730,757]" pageId="4" pageNumber="65" value="2015-06-08" valueMax="2015-06-10" valueMin="2015-06-08">
<collectingDate id="EFEBE9EBFF95FFBF5563FD6BFDFED5E6" box="[351,544,730,757]" pageId="4" pageNumber="65" value="2015-06-08" valueMax="2015-06-10" valueMin="2015-06-08">8–10 Jun. 2015</collectingDate>
</date>
; 
<collectorName id="26E45315FF95FFBF560CFD6AFD1BD5E7" box="[560,709,730,757]" pageId="4" pageNumber="65">Y. Cifuentes</collectorName>
and 
<collectorName id="26E45315FF95FFBF573BFD6AFC5DD5E6" box="[775,899,731,757]" pageId="4" pageNumber="65">J. Moreno</collectorName>
leg.; 
<specimenCode id="DBB79EB8FF95FFBF57F2FD6AFBB4D5E7" box="[974,1130,730,757]" pageId="4" pageNumber="65">IAvH-I 3381</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B793C9EFF95FFBF50B0FD6BFD5DD428" ID-GBIF-Occurrence="3382549311" collectingDate="2014-12-21" collectorName="J. Barrera &amp; M. Castro &amp; S. Mondragon" country="Colombia" elevation="3554" latitude="5.85225" location="Parque Municipal Rancheria" longLatPrecision="1" longitude="-73.13975" municipality="Paipa" pageId="4" pageNumber="65" specimenCode="IAvH-I 3391" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="9D17FD4AFF95FFBF50B0FD6BFB61D5E6" box="[1164,1215,730,757]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BCAACB9FF95FFBF50ECFD6BFACAD5E7" box="[1232,1300,730,756]" pageId="4" pageNumber="65">Paipa</collectingMunicipality>
, 
<location id="8ECE6018FF95FFBF5119FD6AFE66D404" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF5119FD6AFE66D404" country="Colombia" latitude="5.85225" longLatPrecision="1" longitude="-73.13975" municipality="Paipa" name="Parque Municipal Rancheria" pageId="4" pageNumber="65" stateProvince="Boyaca">Parque Municipal Ranchería</location>
; 
<geoCoordinate id="EE255004FF95FFBF55F8FD4DFDAED40B" box="[452,624,764,792]" degrees="05" direction="north" minutes="51" orientation="latitude" pageId="4" pageNumber="65" precision="1" seconds="08.1" value="5.85225">05°51′08.1″ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF5641FD4DFCF0D40B" box="[637,814,764,792]" degrees="73" direction="west" minutes="08" orientation="longitude" pageId="4" pageNumber="65" precision="1" seconds="23.1" value="-73.13975">73°08′23.1″ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF5754FD4CFC1AD40B" box="[872,964,765,792]" metricMagnitude="3" metricUnit="m" metricValue="3.554" pageId="4" pageNumber="65" unit="m" value="3554.0">
<elevation id="003CD1F0FF95FFBF5754FD4CFC1AD40B" box="[872,964,765,792]" metricMagnitude="3" metricUnit="m" metricValue="3.554" pageId="4" pageNumber="65" unit="m" value="3554.0">3554 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF57EDFD4FFBACD40B" box="[977,1138,766,792]" pageId="4" pageNumber="65" value="2014-12-21">
<collectingDate id="EFEBE9EBFF95FFBF57EDFD4FFBACD40B" box="[977,1138,766,792]" pageId="4" pageNumber="65" value="2014-12-21">21 Dec. 2014</collectingDate>
</date>
; 
<collectorName id="26E45315FF95FFBF5043FD4FFB2BD40B" box="[1151,1269,766,792]" pageId="4" pageNumber="65">J. Barrera</collectorName>
, 
<collectorName id="26E45315FF95FFBF513DFD4FFAA9D40B" box="[1281,1399,766,792]" pageId="4" pageNumber="65">M. Castro</collectorName>
and 
<collectorName id="26E45315FF95FFBF54CFFC90FE41D428" box="[243,415,801,827]" pageId="4" pageNumber="65">S. Mondragón</collectorName>
leg.; 
<specimenCode id="DBB79EB8FF95FFBF55D4FC90FD5DD428" box="[488,643,801,827]" pageId="4" pageNumber="65">IAvH-I 3391</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B793C9EFF95FFBF5698FC90FCACD44D" ID-GBIF-Occurrence="3382549309" collectingDate="2014-12-15" collectingDateMax="2014-12-21" collectingDateMin="2014-12-15" collectorName="J. Barrera &amp; M. Castro &amp; S. Mondragon" country="Colombia" elevation="3390" latitude="5.849056" location="Parque Municipal Rancheria" longLatPrecision="1" longitude="-73.12967" municipality="Paipa" pageId="4" pageNumber="65" specimenCode="IAvH-I 3401" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="9D17FD4AFF95FFBF5698FC90FD09D428" box="[676,727,801,827]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
; same collection data as for preceding; 
<geoCoordinate id="EE255004FF95FFBF50FEFCAEFAACD428" box="[1218,1394,799,827]" degrees="05" direction="north" minutes="50" orientation="latitude" pageId="4" pageNumber="65" precision="1" seconds="56.6" value="5.849056">05°50′56.6″ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF5481FCF3FEB3D44C" box="[189,365,834,863]" degrees="73" direction="west" minutes="07" orientation="longitude" pageId="4" pageNumber="65" precision="1" seconds="46.8" value="-73.12967">73°07′46.8″ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF559EFCF5FE23D44C" box="[418,509,836,863]" metricMagnitude="3" metricUnit="m" metricValue="3.3899999999999997" pageId="4" pageNumber="65" unit="m" value="3390.0">
<elevation id="003CD1F0FF95FFBF559EFCF5FE23D44C" box="[418,509,836,863]" metricMagnitude="3" metricUnit="m" metricValue="3.3899999999999997" pageId="4" pageNumber="65" unit="m" value="3390.0">3390 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF5634FCF5FD0CD44D" box="[520,722,836,863]" pageId="4" pageNumber="65" value="2014-12-15" valueMax="2014-12-21" valueMin="2014-12-15">
<collectingDate id="EFEBE9EBFF95FFBF5634FCF5FD0CD44D" box="[520,722,836,863]" pageId="4" pageNumber="65" value="2014-12-15" valueMax="2014-12-21" valueMin="2014-12-15">15–21 Dec. 2014</collectingDate>
</date>
; 
<specimenCode id="DBB79EB8FF95FFBF56E1FCF5FCACD44D" box="[733,882,836,862]" pageId="4" pageNumber="65">IAvH-I 3401</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B793C9EFF95FFBF57B4FCF5FBECD492" ID-GBIF-Occurrence="3382549305" collectingDate="2014-12-15" collectingDateMax="2014-12-21" collectingDateMin="2014-12-15" collectorName="J. Barrera &amp; M. Castro &amp; S. Mondragon" country="Colombia" elevation="3392" latitude="5.8492503" location="Parque Municipal Rancheria" longLatPrecision="1" longitude="-73.12967" municipality="Paipa" pageId="4" pageNumber="65" specimenCode="IAvH-I 3411" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="9D17FD4AFF95FFBF57B4FCF5FC68D44C" box="[904,950,836,863]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
; same collection data as for preceding; 
<geoCoordinate id="EE255004FF95FFBF5481FCD7FEB6D491" box="[189,360,870,898]" degrees="05" direction="north" minutes="50" orientation="latitude" pageId="4" pageNumber="65" precision="1" seconds="57.3" value="5.8492503">05°50′57.3″ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF5548FCD7FDFBD491" box="[372,549,870,898]" degrees="73" direction="west" minutes="07" orientation="longitude" pageId="4" pageNumber="65" precision="1" seconds="46.8" value="-73.12967">73°07′46.8″ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF5661FCD6FD67D491" box="[605,697,871,898]" metricMagnitude="3" metricUnit="m" metricValue="3.392" pageId="4" pageNumber="65" unit="m" value="3392.0">
<elevation id="003CD1F0FF95FFBF5661FCD6FD67D491" box="[605,697,871,898]" metricMagnitude="3" metricUnit="m" metricValue="3.392" pageId="4" pageNumber="65" unit="m" value="3392.0">3392 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF56F9FCD6FC4FD492" box="[709,913,871,898]" pageId="4" pageNumber="65" value="2014-12-15" valueMax="2014-12-21" valueMin="2014-12-15">
<collectingDate id="EFEBE9EBFF95FFBF56F9FCD6FC4FD492" box="[709,913,871,898]" pageId="4" pageNumber="65" value="2014-12-15" valueMax="2014-12-21" valueMin="2014-12-15">15–21 Dec. 2014</collectingDate>
</date>
; 
<specimenCode id="DBB79EB8FF95FFBF57A1FCD6FBECD492" box="[925,1074,871,897]" pageId="4" pageNumber="65">IAvH-I 3411</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B793C9EFF95FFBF5076FCD6FD85D4B7" ID-GBIF-Occurrence="3382549316" collectingDate="2014-12-15" collectingDateMax="2014-12-21" collectingDateMin="2014-12-15" collectorName="J. Barrera &amp; M. Castro &amp; S. Mondragon" country="Colombia" elevation="3392" latitude="5.8492503" location="Parque Municipal Rancheria" longLatPrecision="1" longitude="-73.12967" municipality="Paipa" pageId="4" pageNumber="65" specimenCode="IAvH-I 3382" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="9D17FD4AFF95FFBF5076FCD6FBA6D491" box="[1098,1144,871,898]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D17FD4AFF95FFBF50B8FCD6FB6FD491" box="[1156,1201,871,898]" pageId="4" pageNumber="65" type="female">1 ♀</specimenCount>
; same collection data as for preceding; 
<specimenCode id="DBB79EB8FF95FFBF55F8FC3AFD85D4B7" box="[452,603,906,933]" pageId="4" pageNumber="65">IAvH-I 3382</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B793C9EFF95FFBF5648FC3BFB08D4DB" ID-GBIF-Occurrence="3382549302" collectingDate="2001-07-11" collectorName="J. Barrera &amp; B. Robles" country="Colombia" county="Chaina" elevation="2730" latitude="5.4166665" location="Santuario de Flora" longLatPrecision="1305" longitude="-73.45" municipality="Santuario de Flora" pageId="4" pageNumber="65" specimenCode="IAvH-I 3773" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="9D17FD4AFF95FFBF5648FC3BFD7AD4B6" box="[628,676,906,933]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
; 
<collectingCounty id="62CF4E4FFF95FFBF568DFC3BFCD9D4B7" box="[689,775,906,932]" pageId="4" pageNumber="65">Chaina</collectingCounty>
, 
<location id="8ECE6018FF95FFBF572FFC3BFC33D4B7" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF572FFC3BFC33D4B7" box="[787,1005,906,932]" country="Colombia" county="Chaina" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Santuario de Flora" name="Santuario de Flora" pageId="4" pageNumber="65" stateProvince="Boyaca">Santuario de Flora</location>
y 
<location id="8ECE6018FF95FFBF5037FC3AFB62D4B6" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF5037FC3AFB62D4B6" box="[1035,1212,907,933]" country="Colombia" county="Chaina" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Santuario de Flora" name="Fauna Iguaque" pageId="4" pageNumber="65" stateProvince="Boyaca">Fauna Iguaque</location>
, 
<location id="8ECE6018FF95FFBF50F4FC3BFAACD4B7" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF50F4FC3BFAACD4B7" box="[1224,1394,906,932]" country="Colombia" county="Chaina" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" municipality="Santuario de Flora" name="Sector Chaina" pageId="4" pageNumber="65" stateProvince="Boyaca">Sector Chaina</location>
; 
<geoCoordinate id="EE255004FF95FFBF5481FC1DFEC5D4DB" box="[189,283,940,968]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="4" pageNumber="65" precision="925" value="5.4166665">5°25′ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF551BFC1DFE47D4DB" box="[295,409,940,968]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="4" pageNumber="65" precision="925" value="-73.45">73°27′ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF55EEFC1FFDF1D4DB" box="[466,559,942,968]" metricMagnitude="3" metricUnit="m" metricValue="2.73" pageId="4" pageNumber="65" unit="m" value="2730.0">
<elevation id="003CD1F0FF95FFBF55EEFC1FFDF1D4DB" box="[466,559,942,968]" metricMagnitude="3" metricUnit="m" metricValue="2.73" pageId="4" pageNumber="65" unit="m" value="2730.0">2730 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF5607FC1FFD11D4DB" box="[571,719,941,968]" pageId="4" pageNumber="65" value="2001-07-11">
<collectingDate id="EFEBE9EBFF95FFBF5607FC1FFD11D4DB" box="[571,719,941,968]" pageId="4" pageNumber="65" value="2001-07-11">11 Jul. 2001</collectingDate>
</date>
; 
<collectorName id="26E45315FF95FFBF56E7FC1FFC90D4DB" box="[731,846,942,968]" pageId="4" pageNumber="65">J. Barrera</collectorName>
and 
<collectorName id="26E45315FF95FFBF57B4FC1FFC25D4D4" box="[904,1019,941,968]" pageId="4" pageNumber="65">B. Robles</collectorName>
leg.; 
<specimenCode id="DBB79EB8FF95FFBF5003FC1FFB08D4DB" box="[1087,1238,942,968]" pageId="4" pageNumber="65">IAvH-I 3773</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B793C9EFF95FFBF50D3FC1FFAACD4F8" ID-GBIF-Occurrence="3382549304" collectingDate="2002-10-05" collectingDateMax="2002-10-09" collectingDateMin="2002-10-05" collectorName="P. Reina" country="Colombia" elevation="2840" latitude="5.7333336" location="Quebrada Los Mudos" longLatPrecision="1305" longitude="-73.433334" municipality="Sector Chaina" pageId="4" pageNumber="65" specimenCode="IAvH-I 3774" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="9D17FD4AFF95FFBF50D3FC1FFAC1D4DB" box="[1263,1311,942,968]" pageId="4" pageNumber="65" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BCAACB9FF95FFBF5117FC1FFECDD4F8" pageId="4" pageNumber="65">Sector Chaina</collectingMunicipality>
, 
<location id="8ECE6018FF95FFBF5521FC60FDC4D4F8" LSID="urn:lsid:plazi:treatment:03B887D5FF92FFBF55DBF98AFC49D14F:8ECE6018FF95FFBF5521FC60FDC4D4F8" box="[285,538,977,1003]" country="Colombia" latitude="5.7333336" longLatPrecision="1305" longitude="-73.433334" municipality="Sector Chaina" name="Quebrada Los Mudos" pageId="4" pageNumber="65" stateProvince="Boyaca">Quebrada Los Mudos</location>
; 
<geoCoordinate id="EE255004FF95FFBF5618FC7EFDA1D4F8" box="[548,639,975,1003]" degrees="5" direction="north" minutes="44" orientation="latitude" pageId="4" pageNumber="65" precision="925" value="5.7333336">5°44′ N</geoCoordinate>
, 
<geoCoordinate id="EE255004FF95FFBF56B5FC7EFD26D4F8" box="[649,760,975,1003]" degrees="73" direction="west" minutes="26" orientation="longitude" pageId="4" pageNumber="65" precision="925" value="-73.433334">73°26′ W</geoCoordinate>
; alt. 
<quantity id="4CE99B26FF95FFBF5710FC60FC5BD4F8" box="[812,901,977,1003]" metricMagnitude="3" metricUnit="m" metricValue="2.84" pageId="4" pageNumber="65" unit="m" value="2840.0">
<elevation id="003CD1F0FF95FFBF5710FC60FC5BD4F8" box="[812,901,977,1003]" metricMagnitude="3" metricUnit="m" metricValue="2.84" pageId="4" pageNumber="65" unit="m" value="2840.0">2840 m</elevation>
</quantity>
; 
<date id="FFAF1003FF95FFBF57ACFC60FBE8D4F8" box="[912,1078,977,1004]" pageId="4" pageNumber="65" value="2002-10-05" valueMax="2002-10-09" valueMin="2002-10-05">
<collectingDate id="EFEBE9EBFF95FFBF57ACFC60FBE8D4F8" box="[912,1078,977,1004]" pageId="4" pageNumber="65" value="2002-10-05" valueMax="2002-10-09" valueMin="2002-10-05">5–9 Oct. 2002</collectingDate>
</date>
; 
<collectorName id="26E45315FF95FFBF507CFC60FB43D4F8" box="[1088,1181,977,1003]" pageId="4" pageNumber="65">P. Reina</collectorName>
leg.; 
<specimenCode id="DBB79EB8FF95FFBF50E7FC60FAACD4F8" box="[1243,1394,977,1003]" pageId="4" pageNumber="65">IAvH-I 3774</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6548FF95FFBF5481FBA3FBCCD2F8" pageId="4" pageNumber="65" type="description">
<paragraph id="8BAE36C3FF95FFBF5481FBA3FE5ED33F" blockId="4.[189,384,1042,1068]" box="[189,384,1042,1068]" pageId="4" pageNumber="65">
<heading id="D0E681AFFF95FFBF5481FBA3FE5ED33F" bold="true" box="[189,384,1042,1068]" fontSize="11" level="3" pageId="4" pageNumber="65" reason="3">
<emphasis id="B965EAD1FF95FFBF5481FBA3FE5ED33F" bold="true" box="[189,384,1042,1068]" pageId="4" pageNumber="65">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FBF0FEBED3B1" blockId="4.[189,1398,1089,1186]" pageId="4" pageNumber="65">
The 
<typeStatus id="54AA8861FF95FFBF54CEFBF0FE87D348" box="[242,345,1089,1115]" pageId="4" pageNumber="65" type="holotype">holotype</typeStatus>
was found in the Andean moorland complex of the eastern hills of 
<collectingCountry id="F3067653FF95FFBF5050FBF0FB3ED348" box="[1132,1248,1089,1115]" name="Colombia" pageId="4" pageNumber="65">Colombia</collectingCountry>
(
<figureCitation id="132A2A46FF95FFBF50CCFBF0FAB0D348" box="[1264,1390,1089,1115]" captionStart="Fig" captionStartId="16.[189,232,1857,1883]" captionTargetBox="[199,1388,463,1818]" captionTargetId="figure-79@16.[189,1398,456,1821]" captionTargetPageId="16" captionText="Fig. 8. Habitats of Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017. A–B. Type locality of M. blades Sánchez-Ruiz &amp; Brescovit, 2017, Páramo de Monserrate, Cundimarca department, Colombia. C–D. Type locality of M. carolinae sp. nov., Sutamarchán, Boyacá department, Colombia. E–F. Habitat of female paratype (IAvH-I 3771) of M. carolinae sp. nov., Sector Carrizal, Santuario de Flora y Fauna Iguaque, Boyacá department, Colombia." figureDoi="http://doi.org/10.5281/zenodo.5536403" httpUri="https://zenodo.org/record/5536403/files/figure.png" pageId="4" pageNumber="65">Fig. 8A–B</figureCitation>
) which is predominantly populated by plant species such as 
<taxonomicName id="4C114D40FF95FFBF5745FBD5FA9BD36D" authority="Humb. &amp; Bonpl." authorityName="Humb. &amp; Bonpl." box="[889,1349,1124,1150]" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="grandiflora">
<emphasis id="B965EAD1FF95FFBF5745FBD5FBA9D36D" box="[889,1143,1124,1150]" italics="true" pageId="4" pageNumber="65">Espeletia grandiflora</emphasis>
Humb. &amp; Bonpl.
</taxonomicName>
and 
<taxonomicName id="4C114D40FF95FFBF5482FB39FEBED3B1" box="[190,352,1160,1186]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAD1FF95FFBF5482FB39FEE6D3B1" box="[190,312,1160,1186]" italics="true" pageId="4" pageNumber="65">Cupressus</emphasis>
sp.
</taxonomicName>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FB79FE91D3F1" blockId="4.[189,335,1224,1250]" box="[189,335,1224,1250]" pageId="4" pageNumber="65">
<heading id="D0E681AFFF95FFBF5481FB79FE91D3F1" bold="true" box="[189,335,1224,1250]" fontSize="11" level="3" pageId="4" pageNumber="65" reason="3">
<emphasis id="B965EAD1FF95FFBF5481FB79FE91D3F1" bold="true" box="[189,335,1224,1250]" pageId="4" pageNumber="65">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FB46FC62D202" blockId="4.[189,956,1271,1297]" box="[189,956,1271,1297]" pageId="4" pageNumber="65">Male and female described by Sánchez-Ruiz &amp; Brescovit (2017).</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FA89FEEAD241" blockId="4.[189,308,1336,1362]" box="[189,308,1336,1362]" pageId="4" pageNumber="65">
<heading id="D0E681AFFF95FFBF5481FA89FEEAD241" bold="true" box="[189,308,1336,1362]" fontSize="11" level="3" pageId="4" pageNumber="65" reason="3">
<emphasis id="B965EAD1FF95FFBF5481FA89FEEAD241" bold="true" box="[189,308,1336,1362]" pageId="4" pageNumber="65">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481FAD6FBCCD2F8" blockId="4.[188,1399,1383,1515]" pageId="4" pageNumber="65">
Males from the 
<collectingRegion id="49D5F821FF95FFBF554FFAD6FD8BD292" box="[371,597,1383,1409]" country="Colombia" name="Boyaca" pageId="4" pageNumber="65">Boyacá department</collectingRegion>
population (
<figureCitation id="132A2A46FF95FFBF56DBFAD6FCBAD292" box="[743,868,1383,1409]" captionStart="Fig" captionStartId="5.[189,232,1786,1812]" captionTargetBox="[226,1366,279,1736]" captionTargetId="figure-12@5.[210,1378,265,1748]" captionTargetPageId="5" captionText="Fig. 1. Medionops blades Sánchez-Ruiz &amp; Brescovit, 2017. A–B, E–F. Male, Boyacá population (IAvH-I 3381). C–D, G–H. Female, Boyacá population (IAvH-I 3382). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; ess = external sclerotization around spiracles; go = genital opening; re = receptaculum; ue = uterus externo." figureDoi="http://doi.org/10.5281/zenodo.5536389" httpUri="https://zenodo.org/record/5536389/files/figure.png" pageId="4" pageNumber="65">Fig. 1A–B</figureCitation>
) have the embolus tip somewhat more curved than those from 
<collectingRegion id="49D5F821FF95FFBF5540FA3BFD6DD2B7" box="[380,691,1418,1444]" country="Colombia" name="Cundinamarca" pageId="4" pageNumber="65">Cundinamarca department</collectingRegion>
(see images from 
<typeStatus id="54AA8861FF95FFBF57B1FA3AFC15D2B6" box="[909,971,1419,1445]" pageId="4" pageNumber="65">types</typeStatus>
in Sánchez-Ruiz &amp; Brescovit 2017: figs 2h, 6d). Females from the 
<collectingRegion id="49D5F821FF95FFBF5614FA1FFD5ED2DB" box="[552,640,1454,1480]" country="Colombia" name="Boyaca" pageId="4" pageNumber="65">Boyacá</collectingRegion>
population (
<figureCitation id="132A2A46FF95FFBF572AFA1CFCAFD2DB" box="[790,881,1453,1480]" captionStart="Fig" captionStartId="5.[189,232,1786,1812]" captionTargetBox="[226,1366,279,1736]" captionTargetId="figure-12@5.[210,1378,265,1748]" captionTargetPageId="5" captionText="Fig. 1. Medionops blades Sánchez-Ruiz &amp; Brescovit, 2017. A–B, E–F. Male, Boyacá population (IAvH-I 3381). C–D, G–H. Female, Boyacá population (IAvH-I 3382). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, dorsal view. D. Habitus, ventral view. E. Left palp, retrolateral view. F. Left palp, prolateral view. G. External genitalia, ventral view. H. Internal genitalia, dorsal view. Scale bars: A–D = 1.5 mm; E–F = 0.7 mm; G = 0.5 mm; H = 0.2 mm. Abbreviations: ap = anterior plate; ess = external sclerotization around spiracles; go = genital opening; re = receptaculum; ue = uterus externo." figureDoi="http://doi.org/10.5281/zenodo.5536389" httpUri="https://zenodo.org/record/5536389/files/figure.png" pageId="4" pageNumber="65">Fig. 1G</figureCitation>
) have a narrower and more sclerotized area under the genital opening than those from the 
<collectingRegion id="49D5F821FF95FFBF56E6FA60FC5BD2F8" box="[730,901,1489,1515]" country="Colombia" name="Cundinamarca" pageId="4" pageNumber="65">Cundinamarca</collectingRegion>
population.
</paragraph>
</subSubSection>
<subSubSection id="C30B6548FF95FFBF5481F9A3FC49D14F" box="[189,919,1554,1628]" pageId="4" pageNumber="65" type="distribution">
<paragraph id="8BAE36C3FF95FFBF5481F9A3FE86D13F" blockId="4.[189,344,1554,1580]" box="[189,344,1554,1580]" pageId="4" pageNumber="65">
<heading id="D0E681AFFF95FFBF5481F9A3FE86D13F" bold="true" box="[189,344,1554,1580]" fontSize="11" level="3" pageId="4" pageNumber="65" reason="3">
<emphasis id="B965EAD1FF95FFBF5481F9A3FE86D13F" bold="true" box="[189,344,1554,1580]" pageId="4" pageNumber="65">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36C3FF95FFBF5481F9F0FC49D14F" blockId="4.[189,919,1601,1628]" box="[189,919,1601,1628]" pageId="4" pageNumber="65">
Known from 
<collectingRegion id="49D5F821FF95FFBF5567F9F0FDD8D148" box="[347,518,1601,1627]" country="Colombia" name="Cundinamarca" pageId="4" pageNumber="65">Cundinamarca</collectingRegion>
and 
<collectingRegion id="49D5F821FF95FFBF5603F9F0FD49D148" box="[575,663,1601,1627]" country="Colombia" name="Boyaca" pageId="4" pageNumber="65">Boyacá</collectingRegion>
in 
<collectingCountry id="F3067653FF95FFBF5681F9F0FCEFD148" box="[701,817,1601,1627]" name="Colombia" pageId="4" pageNumber="65">Colombia</collectingCountry>
(
<figureCitation id="132A2A46FF95FFBF577CF9F0FC56D14F" box="[832,904,1601,1628]" captionStart="Fig" captionStartId="14.[187,230,1998,2024]" captionTargetBox="[230,1361,1150,1933]" captionTargetId="figure-321@14.[189,1398,1136,1959]" captionTargetPageId="14" captionText="Fig. 6. Distribution map of Colombian species of Medionops Sánchez-Ruiz &amp; Brescovit, 2017." figureDoi="http://doi.org/10.5281/zenodo.5536399" httpUri="https://zenodo.org/record/5536399/files/figure.png" pageId="4" pageNumber="65">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>