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<document id="7CC6F1D3E235A3329C8AA10FE11787C1" ID-CLB-Dataset="27992" ID-DOI="10.11646/zootaxa.4570.1.1" ID-GBIF-Dataset="66601a1e-ba16-41b2-b182-0fd80e75ccba" ID-ISSN="1175-5326" ID-Zenodo-Dep="2608527" ID-ZooBank="EF369E98-EBD9-4647-B081-65AD1794A27C" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553591750157" checkinUser="plazi" docAuthor="Soto, Joan J. &amp; Peña, Álvaro L." docDate="2019" docId="03B887B7A460FFFCE983FD87FE739C38" docLanguage="en" docName="zootaxa.4570.1.1.pdf" docOrigin="Zootaxa 4570 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thuiaria longstaffi Hickson &amp; Gravely 1907" docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FF81FFCFA46EFFF2E914FFEFFFBB9B7E" masterDocTitle="Benthic hydroids (Cnidaria, Hydrozoa) from the Weddell Sea (Antarctica)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="15" updateTime="1698713421926" updateUser="ExternalLinkService">
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<mods:title id="EB8761BCEA1C36CC3BDF274C9B5BB2D0">Benthic hydroids (Cnidaria, Hydrozoa) from the Weddell Sea (Antarctica)</mods:title>
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<mods:namePart id="34E6165927619878FA9353B303822A7B">Soto, Joan J.</mods:namePart>
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<treatment id="03B887B7A460FFFCE983FD87FE739C38" ID-DOI="http://doi.org/10.5281/zenodo.5934899" ID-GBIF-Taxon="154497176" ID-Zenodo-Dep="5934899" LSID="urn:lsid:plazi:treatment:03B887B7A460FFFCE983FD87FE739C38" httpUri="http://treatment.plazi.org/id/03B887B7A460FFFCE983FD87FE739C38" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="C30B652AA460FFFCE983FD87FD7099FC" box="[151,715,616,642]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BAE36A1A460FFFCE983FD87FD7099FC" blockId="14.[151,715,616,675]" box="[151,715,616,642]" pageId="14" pageNumber="15">
<heading id="D0E681CDA460FFFCE983FD87FD7099FC" bold="true" box="[151,715,616,642]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B965EAB3A460FFFCE983FD87FD7099FC" bold="true" box="[151,715,616,642]" pageId="14" pageNumber="15">
<emphasis id="B965EAB3A460FFFCE983FD87FEAB99FC" bold="true" box="[151,272,616,642]" italics="true" pageId="14" pageNumber="15">Tubularia</emphasis>
<taxonomicName id="4C114D22A460FFFCE803FD87FD7099FC" authority="Hickson &amp; Gravely, 1907" authorityName="Hickson &amp; Gravely" authorityYear="1907" box="[279,715,616,642]" class="Hydrozoa" family="Sertulariidae" genus="Thuiaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="longstaffi">
<emphasis id="B965EAB3A460FFFCE803FD87FE3299FC" bold="true" box="[279,393,616,642]" italics="true" pageId="14" pageNumber="15">longstaffi</emphasis>
<bibRefCitation id="EF804B50A460FFFCE885FD87FD7099FC" author="Hickson, S. J. &amp; Gravely, F. H." box="[401,715,616,642]" pageId="14" pageNumber="15" pagination="1 - 34" refId="ref42832" refString="Hickson, S. J. &amp; Gravely, F. H. (1907) Coelenterata. II. Hydroid zoophytes. National Antarctic Expedition (S. S. Discovery) 1901 - 1904, Natural History, 3, 1 - 34." type="journal article" year="1907">Hickson &amp; Gravely, 1907</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30B652AA460FFFCE983FD65FEB699DD" box="[151,269,650,675]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BAE36A1A460FFFCE983FD65FEB699DD" blockId="14.[151,715,616,675]" box="[151,269,650,675]" pageId="14" pageNumber="15">
(
<figureCitation id="132A2A24A460FFFCE98BFD65FF5999DD" box="[159,226,650,675]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
ai)
</paragraph>
</subSubSection>
<subSubSection id="C30B652AA460FFFCE983FD3DFAAC9871" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BAE36A1A460FFFCE983FD3DFAAC9871" blockId="14.[151,1436,722,1863]" pageId="14" pageNumber="15">
<emphasis id="B965EAB3A460FFFCE983FD3DFE3B9995" bold="true" box="[151,384,722,747]" pageId="14" pageNumber="15">Material examined.</emphasis>
<emphasis id="B965EAB3A460FFFCE893FD3CFDB09995" box="[391,523,723,747]" italics="true" pageId="14" pageNumber="15">ANT XVII/3</emphasis>
:
<emphasis id="B965EAB3A460FFFCEB0EFD3DFDEC9995" bold="true" box="[538,599,722,747]" pageId="14" pageNumber="15">111-5</emphasis>
, few stems, up to
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high, on
<taxonomicName id="4C114D22A460FFFCEAC8FD3CFBC29995" authorityName="Pena Cantero &amp; Vervoort" authorityYear="1996" box="[988,1145,723,747]" class="Hydrozoa" family="Kirchenpaueriidae" genus="Oswaldella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="terranovae">
<emphasis id="B965EAB3A460FFFCEAC8FD3CFBC29995" box="[988,1145,723,747]" italics="true" pageId="14" pageNumber="15">O. terranovae</emphasis>
</taxonomicName>
;
<emphasis id="B965EAB3A460FFFCED93FD3DFB789995" bold="true" box="[1159,1219,722,747]" pageId="14" pageNumber="15">111-7</emphasis>
, some stems, up to
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high, with developing gonophores;
<emphasis id="B965EAB3A460FFFCEB6EFD19FD0D9871" bold="true" box="[634,694,758,783]" pageId="14" pageNumber="15">111-9</emphasis>
, some stems, up to
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, with mature gonophores.
</paragraph>
</subSubSection>
<subSubSection id="C30B652AA460FFFCE9D3FCF5FE299DED" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BAE36A1A460FFFCE9D3FCF5FE4498BD" blockId="14.[151,1436,722,1863]" pageId="14" pageNumber="15">
<emphasis id="B965EAB3A460FFFCE9D3FCF5FE80984D" bold="true" box="[199,315,794,819]" pageId="14" pageNumber="15">Remarks.</emphasis>
The specimens examined here concurs with all the diagnostic features mentioned in the redescription of the
<typeStatus id="54AA8803A460FFFCE873FCD0FE2C9829" box="[359,407,831,855]" pageId="14" pageNumber="15">type</typeStatus>
material of
<taxonomicName id="4C114D22A460FFFCEB30FCD0FD109829" authorityName="Hickson &amp; Gravely" authorityYear="1907" box="[548,683,830,855]" class="Hydrozoa" family="Sertulariidae" genus="Thuiaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="longstaffi">
<emphasis id="B965EAB3A460FFFCEB30FCD0FD109829" box="[548,683,830,855]" italics="true" pageId="14" pageNumber="15">T. longstaffi</emphasis>
</taxonomicName>
by Peña Cantero (2017): yellow stems, in entwined clusters, tube diameter increasing distally (
<figureCitation id="132A2A24A460FFFCE8CAFC8DFD8B9805" box="[478,560,866,891]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4a</figureCitation>
), with transversal ridges; polyps large, up to
<date id="FFAF1061A460FFFCED39FC8DFBEA9804" box="[1069,1105,866,891]" pageId="14" pageNumber="15">5 x</date>
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, with an oral crown of 2937 tentacles and an aboral one of 1219 tentacles;
<typeStatus id="54AA8803A460FFFCEA55FC68FCCA98E1" box="[833,881,903,927]" pageId="14" pageNumber="15">type</typeStatus>
and size of nematocysts, and gonophores as cryptomedusoids (
<figureCitation id="132A2A24A460FFFCE872FC45FE0398BD" box="[358,440,938,963]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4d</figureCitation>
, gh).
</paragraph>
<paragraph id="8BAE36A1A460FFFCE9D3FC21FCA39EA1" blockId="14.[151,1436,722,1863]" pageId="14" pageNumber="15">
Additional knowledge regarding sexual dimorphism and early developmental stages is drawn from the present study. Male blastostyles elongate, c.
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long and with up to 17 stalked gonophores as cryptomedusoids (
<figureCitation id="132A2A24A460FFFCEC2AFC1DFA349F75" box="[1342,1423,1010,1035]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4c</figureCitation>
). Gonophore maturity increasing distally: recently formed gonophores located proximally, spherical, c. 300 µm in diameter, and completely filled with spermatogenic tissue (reddish in preserved specimens); mid-term gonophores, c. 600 µm in diameter, filled for 2/3 to 3/4 with spermatogenic tissue, with 45 rudimentary tentacular bulbs and slightly protruded aperture (
<figureCitation id="132A2A24A460FFFCE8C5FB6DFD9F9FE5" box="[465,548,1154,1179]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4c</figureCitation>
); fully grown male gonophores located distally, spherical, c.
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in diameter, almost completely filled with whitish sperm cells between aperture and spermatogenic tissue, rudimentary tentacular bulbs difficult to appreciate at this stage (
<figureCitation id="132A2A24A460FFFCEBF3FB25FC879F9D" box="[743,828,1226,1251]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4d</figureCitation>
). Female blastostyles elongate, up to
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long, and with up to 11 stalked gonophores as cryptomedusoids, fully matured ones scattered along the blastostyle (
<figureCitation id="132A2A24A460FFFCEC54FB01FA2B9E79" box="[1344,1424,1262,1287]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4f</figureCitation>
). Developing gonophores sphaerical, c. 250600 µm; fully grown female gonophores oblong in lateral view, c.
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wide
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x
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mm high, with a digitiform process running on proximal section, with 45 rudimentary tentacular bulbs, and with up to four actinulae, of which 23 developing and 12 mature (
<figureCitation id="132A2A24A460FFFCED09FAB5FBDA9E0D" box="[1053,1121,1370,1395]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
gh). Actinula resembling a
<emphasis id="B965EAB3A460FFFCE983FA91FEBD9EE9" box="[151,262,1406,1431]" italics="true" pageId="14" pageNumber="15">Tubularia</emphasis>
polyp, with 10 elongate aboral tentacles and four short oral ones. Basal section dense, with two constrictions, one at the base of aboral tentacles, the other at the basal fourth, delimiting a disc. Basal disc with radial grooves, probably with adhesive function (
<figureCitation id="132A2A24A460FFFCEBA9FA29FCB09EA1" box="[701,779,1478,1503]" captionStart="FIGURE 4" captionStartId="15.[151,250,1875,1897]" captionTargetBox="[166,1416,272,1843]" captionTargetId="figure@15.[151,1436,267,1853]" captionTargetPageId="15" captionText="FIGURE 4. ai Tubularia longstaffi: a polyp-carrying tube; b male polyp; c blastostyle with developing male gonophores; d blastostyle with fully-grown male gonophores; e female polyp; f blastostyle with fully-grown female gonophores; gh detail of female gonophores carrying actinulae; i actinulae in different stages of development. jo Zanclea hicksoni: j early-developing polyp, k expanded young polyps, l expanded polyps with hydrorhizal filaments, m contracted polyp, no detail of polyps showing tentacle arrangement in linear clusters (arrows)." httpUri="https://zenodo.org/record/2608535/files/figure.png" pageId="14" pageNumber="15">
Fig.
<date id="FFAF1061A460FFFCEBE5FA29FCB09EA1" box="[753,779,1478,1503]" pageId="14" pageNumber="15">4i</date>
</figureCitation>
).
</paragraph>
<paragraph id="8BAE36A1A460FFFCE9D3FA05FE299DED" blockId="14.[151,1436,722,1863]" pageId="14" pageNumber="15">Cnidome composed by small stenoteles [range 9.011.0 x 7.08.5 µm, mean 10.1±0.7 x 7.8±0.6 µm (n=18)], large stenoteles [range 14.015.5 x 10.012.5 µm, mean 14.4±0.4 x 10.7±0.7 µm (n=17)], tear-shaped haplonemes [range 13.516.5 x 4.05.5 µm, mean 14.9±1.1 x 4.9±0.6 µm (n=16)], rounded haplonemes [range 15.516.5 x 10.011.5 µm, mean 16.0±0.5 x 10.9±0.5 µm (n=15)] and desmonemes [range 5.57.0 x 4.06.0 µm, mean 6.4±0.6 x 4.8±0.6 µm (n=16)].</paragraph>
</subSubSection>
<subSubSection id="C30B652AA460FFFCE9D3F971FE739C38" pageId="14" pageNumber="15" type="biology_ecology">
<paragraph id="8BAE36A1A460FFFCE9D3F971FE739C38" blockId="14.[151,1436,722,1863]" pageId="14" pageNumber="15">
<emphasis id="B965EAB3A460FFFCE9D3F971FE4E9DC9" bold="true" box="[199,501,1694,1719]" pageId="14" pageNumber="15">Ecology and distribution.</emphasis>
<emphasis id="B965EAB3A460FFFCE8E8F971FD609DC9" box="[508,731,1694,1719]" italics="true" pageId="14" pageNumber="15">
Tubularia
<taxonomicName id="4C114D22A460FFFCEB65F971FD609DC9" authorityName="Hickson &amp; Gravely" authorityYear="1907" box="[625,731,1694,1719]" class="Hydrozoa" family="Sertulariidae" genus="Thuiaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="longstaffi">longstaffi</taxonomicName>
</emphasis>
was only known from the Ross Sea (
<bibRefCitation id="EF804B50A460FFFCED6DF971FA2C9DC9" author="Hickson, S. J. &amp; Gravely, F. H." box="[1145,1431,1694,1719]" pageId="14" pageNumber="15" pagination="1 - 34" refId="ref42832" refString="Hickson, S. J. &amp; Gravely, F. H. (1907) Coelenterata. II. Hydroid zoophytes. National Antarctic Expedition (S. S. Discovery) 1901 - 1904, Natural History, 3, 1 - 34." type="journal article" year="1907">Hickson &amp; Gravely 1907</bibRefCitation>
; Peña Cantero 2017), reported at depths between 222 and
<quantity id="4CE99B44A460FFFCEA5EF92DFC2C9DA4" box="[842,919,1730,1755]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="14" pageNumber="15" unit="m" value="630.0">630 m</quantity>
(Peña Cantero 2017); present material was collected at
<quantity id="4CE99B44A460FFFCE834F909FE3E9D80" box="[288,389,1766,1791]" metricMagnitude="1" metricUnit="m" metricValue="6.45" metricValueMax="6.7" metricValueMin="6.2" pageId="14" pageNumber="15" unit="m" value="64.5" valueMax="67.0" valueMin="62.0">6267 m</quantity>
depth, in the vicinity of Cape
<collectingCountry id="F3067631A460FFFCEBF5F909FCF69D81" box="[737,845,1766,1791]" name="Norway" pageId="14" pageNumber="15">Norvegia</collectingCountry>
. This constitutes the third record of the species, and the first evidence from both the Weddell Sea and West
<collectingCountry id="F3067631A460FFFCEA22F8E5FC159C5D" box="[822,942,1802,1827]" name="Antarctica" pageId="14" pageNumber="15">Antarctica</collectingCountry>
. We tentatively assign a circum-Antarctic distribution to
<taxonomicName id="4C114D22A460FFFCE828F8C0FE7A9C39" authorityName="Hickson &amp; Gravely" authorityYear="1907" box="[316,449,1838,1863]" class="Hydrozoa" family="Sertulariidae" genus="Thuiaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="longstaffi">
<emphasis id="B965EAB3A460FFFCE828F8C0FE7A9C39" box="[316,449,1838,1863]" italics="true" pageId="14" pageNumber="15">T. longstaffi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>