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<document id="389C246038CE2663FFB6500EE3191AB7" ID-DOI="10.11646/zootaxa.5540.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14240496" ID-ZooBank="4DBD3A92-F14B-4C5B-95B3-2B430EC197DD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732832094216" checkinUser="plazi" docAuthor="Wu, Jingyu &amp; Liu, Xingyue" docDate="2024" docId="03B88790FFA8FFAA9EE7CD649D744A58" docLanguage="en" docName="zootaxa.5540.1.1.pdf" docOrigin="Zootaxa 5540 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5540.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Ankylopteryx delicatula Banks 1937" docType="treatment" docVersion="2" lastPageNumber="16" masterDocId="FF81FFE8FFA7FFA59E70CE029F264D33" masterDocTitle="Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China" masterLastPageNumber="169" masterPageNumber="1" pageNumber="16" updateTime="1732841452334" updateUser="ExternalLinkService">
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<mods:title id="18FAEBD9852C98696F64F8AF73159943">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:namePart id="BDD4A2FD04C19C6CCBC8C7EA0E2AECCE">Wu, Jingyu</mods:namePart>
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<mods:affiliation id="67EAE5BE94F99587963D400540EDE7C8">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:namePart id="2C3D4E9249FBC40039B3223393044E50">Liu, Xingyue</mods:namePart>
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<mods:affiliation id="37C630E429C8610EEB611B2895E355B6">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:date id="40EF43256F86FC418F0DE8B163CF1C60">2024</mods:date>
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<mods:number id="41D92F7FDB167EDE2452C183BE95D71E">5540</mods:number>
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<treatment id="03B88790FFA8FFAA9EE7CD649D744A58" LSID="urn:lsid:plazi:treatment:03B88790FFA8FFAA9EE7CD649D744A58" httpUri="http://treatment.plazi.org/id/03B88790FFA8FFAA9EE7CD649D744A58" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="C30B650DFFA8FFAA9EE7CD649DA64EB3" box="[151,640,870,897]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BAE3686FFA8FFAA9EE7CD649DA64EB3" blockId="15.[151,640,870,933]" box="[151,640,870,897]" pageId="15" pageNumber="16">
<emphasis id="B965EA94FFA8FFAA9EE7CD649DA64EB3" bold="true" box="[151,640,870,897]" pageId="15" pageNumber="16">
[2]
<taxonomicName id="4C114D05FFA8FFAA9EB1CD649DA64EB3" ID-CoL="5V4JT" authority="Banks, 1937" authorityName="Banks" authorityYear="1937" box="[193,640,870,897]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="delicatula">
<emphasis id="B965EA94FFA8FFAA9EB1CD649EC64EB2" bold="true" box="[193,480,870,897]" italics="true" pageId="15" pageNumber="16">Ankylopteryx delicatula</emphasis>
<bibRefCitation id="EF804B77FFA8FFAA9F98CD659DA64EB3" author="Banks, N." box="[488,640,870,897]" pageId="15" pageNumber="16" pagination="255 - 291" refId="ref88124" refString="Banks, N. (1937) Neuropteroid insects from Formosa. Philippine Journal of Science, 62, 255 - 291." type="journal article" year="1937">Banks, 1937</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
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<paragraph id="8BAE3686FFA8FFAA9EE7CD899E3F4E96" blockId="15.[151,640,870,933]" box="[151,281,907,933]" pageId="15" pageNumber="16">
(
<figureCitation id="132A2A03FFA8FFAA9EEFCD899FC14E96" box="[159,231,907,933]" captionStart="FIGURE 8" captionStartId="16.[151,250,1552,1576]" captionTargetBox="[192,1404,188,1518]" captionTargetId="figure-33@16.[183,1404,181,1528]" captionTargetPageId="16" captionText="FIGURE 8. Ankylopteryx delicatula Banks, 1937. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view. Scale bar: 5 mm (A), 1 mm (B–D)." figureDoi="http://doi.org/10.5281/zenodo.14240522" httpUri="https://zenodo.org/record/14240522/files/figure.png" pageId="15" pageNumber="16">Figs 8</figureCitation>
,
<figureCitation id="132A2A03FFA8FFAA9E84CD899E374E96" box="[244,273,907,933]" captionStart="FIGURE 79" captionStartId="142.[151,250,1894,1918]" captionTargetBox="[196,1392,1098,1870]" captionTargetId="figure-46@142.[196,1392,1098,1870]" captionTargetPageId="142" captionText="FIGURE 79. Distribution map of species of Ankylopteryx from China." figureDoi="http://doi.org/10.5281/zenodo.14240704" httpUri="https://zenodo.org/record/14240704/files/figure.png" pageId="15" pageNumber="16">79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFA8FFAA9EE7CDCD9B6C4ED4" box="[151,1098,975,1000]" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8BAE3686FFA8FFAA9EE7CDCD9B6C4ED4" blockId="15.[151,1098,975,1000]" box="[151,1098,975,1000]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB0111A8FFA8FFAA9EE7CDCD9B6C4ED4" box="[151,1098,975,1000]" pageId="15" pageNumber="16">
<treatmentCitation id="0AB01097FFA8FFAA9EE7CDCD9D624ED4" author="Banks, N." box="[151,580,975,999]" page="280" pageId="15" pageNumber="16" year="1937">
<taxonomicName id="4C114D05FFA8FFAA9EE7CDCD9D624ED4" ID-CoL="5V4JT" authority="Banks, 1937: 280" authorityName="Banks" authorityPageNumber="280" authorityYear="1937" box="[151,580,975,999]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="delicatula">
<emphasis id="B965EA94FFA8FFAA9EE7CDCD9EAE4ED4" box="[151,392,975,999]" italics="true" pageId="15" pageNumber="16">Ankylopteryx delicatula</emphasis>
<bibRefCitation id="EF804B77FFA8FFAA9FFFCDCD9D624ED4" author="Banks, N." box="[399,580,975,999]" pageId="15" pageNumber="16" pagination="255 - 291" refId="ref88124" refString="Banks, N. (1937) Neuropteroid insects from Formosa. Philippine Journal of Science, 62, 255 - 291." type="journal article" year="1937">Banks, 1937: 280</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type locality:
<collectingCountry id="F3067616FFA8FFAA9CAFCDD29C3F4ED4" box="[735,793,976,999]" name="Japan" pageId="15" pageNumber="16">Japan</collectingCountry>
(
<collectingRegion id="49D5F864FFA8FFAA9D56CDCD9CA14ED4" box="[806,903,975,999]" country="Japan" name="Okinawa" pageId="15" pageNumber="16">Okinawa</collectingRegion>
).
<materialsCitation id="3B793CDBFFA8FFAA9DE8CDCD9B634ED4" box="[920,1093,975,999]" collectionCode="MCZ" pageId="15" pageNumber="16" specimenCount="1" typeStatus="syntype">
<typeStatus id="54AA8824FFA8FFAA9DE8CDCD9CCD4ED4" box="[920,1003,975,999]" pageId="15" pageNumber="16" type="syntype">Syntype</typeStatus>
in
<collectionCode id="ED00AE43FFA8FFAA9A7BCDCD9B634ED4" box="[1035,1093,975,999]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="15" pageNumber="16" type="Museum">MCZ</collectionCode>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFA8FFAA9EE7CA179DAD49A8" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BAE3686FFA8FFAA9EE7CA179DAD49A8" blockId="15.[151,1437,1045,1899]" pageId="15" pageNumber="16">
<emphasis id="B965EA94FFA8FFAA9EE7CA179E34491C" bold="true" box="[151,274,1045,1071]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
This species is characterized by the presence of two black interantennal spots, two pale brown stripes below antennae, the gena marked with black spots, the pronotum with blackish-brown spots on the anterolateral angle, the mesonotum and metanotum with blackish-brown stripe, the quadrate cell
<emphasis id="B965EA94FFA8FFAA9A39CA5C9B434944" box="[1097,1125,1118,1143]" italics="true" pageId="15" pageNumber="16">im</emphasis>
of forewing, and the wings with brownish vittae along posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFA8FFAA9EB7CAA79C954BA0" pageId="15" pageNumber="16" type="description">
<paragraph id="8BAE3686FFA8FFAA9EB7CAA79A1649F3" blockId="15.[151,1437,1045,1899]" box="[199,1328,1189,1216]" pageId="15" pageNumber="16">
<emphasis id="B965EA94FFA8FFAA9EB7CAA79E7F498C" bold="true" box="[199,345,1189,1215]" pageId="15" pageNumber="16">Description.</emphasis>
Female. Body length
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, forewing length
<quantity id="4CE99B63FFA8FFAA9D0BCAA79CC849F3" box="[891,1006,1189,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="15" pageNumber="16" unit="mm" value="11.8">11.80 mm</quantity>
, hindwing length
<quantity id="4CE99B63FFA8FFAA9AC8CAA79A0A49F3" box="[1208,1324,1189,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="mm" value="11.3">11.30 mm</quantity>
.
</paragraph>
<paragraph id="8BAE3686FFA8FFAA9EB7CACB9B3E4818" blockId="15.[151,1437,1045,1899]" pageId="15" pageNumber="16">Head narrow, vertex raised. Compound eyes brown. Antennae yellow, scape and pedicel wider than flagellum; two interantennal spots small and black; two pale brown stripes below antennae. Clypeal margin with black stripes. Gena marked with black spots. Maxillary and labial palpi pale brownish yellow.</paragraph>
<paragraph id="8BAE3686FFA8FFAA9EB7CB349CD64840" blockId="15.[151,1437,1045,1899]" pageId="15" pageNumber="16">Pronotum posterior margin longer than anterior margin, with square shaped blackish-brown spots on anterolateral angle; mesonotum and metanotum with blackish-brown stripe along margin.</paragraph>
<paragraph id="8BAE3686FFA8FFAA9EB7CB7C9AB548A4" blockId="15.[151,1437,1045,1899]" box="[199,1427,1405,1432]" pageId="15" pageNumber="16">Legs with dense setae. Pro- and mesotibia with median black spot. Pretarsal claws brown, with basal dilation.</paragraph>
<paragraph id="8BAE3686FFA8FFAA9EB7CBA39BE84B14" blockId="15.[151,1437,1045,1899]" pageId="15" pageNumber="16">
Forewing broad, marked with brownish vittae along base of posterior margin. Pterostigma dark brown. Costal area broad basally, with 25 crossveins; 11 radial crossveins present. Base of Rs with pale brown markings.
<emphasis id="B965EA94FFA8FFAA9B0ACBC79ABB48EC" box="[1402,1437,1477,1503]" italics="true" pageId="15" pageNumber="16">m1</emphasis>
smaller than
<emphasis id="B965EA94FFA8FFAA9F57CBEB9E6E4B30" box="[295,328,1513,1539]" italics="true" pageId="15" pageNumber="16">m2</emphasis>
,
<emphasis id="B965EA94FFA8FFAA9F26CBE89E544B30" box="[342,370,1514,1539]" italics="true" pageId="15" pageNumber="16">im</emphasis>
quadrate; 8 crossveins present between Psm and Psc.
<emphasis id="B965EA94FFA8FFAA9DBBCBEB9CC04B30" box="[971,998,1513,1539]" italics="true" pageId="15" pageNumber="16">c1</emphasis>
smaller than
<emphasis id="B965EA94FFA8FFAA9A0BCBEB9BB04B30" box="[1147,1174,1513,1539]" italics="true" pageId="15" pageNumber="16">c2</emphasis>
,
<emphasis id="B965EA94FFA8FFAA9AD3CBEB9BEC4B30" box="[1187,1226,1513,1539]" italics="true" pageId="15" pageNumber="16">dcc</emphasis>
closed. A1 forked, A2 and A3 simple. Inner gradate series with 5 crossveins, outer gradate series with 6 crossveins.
</paragraph>
<paragraph id="8BAE3686FFA8FFAA9EB7C8339C954BA0" blockId="15.[151,1437,1045,1899]" pageId="15" pageNumber="16">Hindwing narrow, marked with brownish vittate along middle of posterior margin. Pterostigma brown. Costal area with 19 crossveins; 9 radial crossveins and 8 psm-psc crossveins present. A1 forked, A2 and A3 simple. Inner gradate series with 4 crossveins, outer gradate series with 5 crossveins.</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFA8FFAA9EB7C89F9B904BE8" pageId="15" pageNumber="16" type="materials_examined">
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<emphasis id="B965EA94FFA8FFAA9EB7C89F9E104B84" bold="true" box="[199,310,1693,1719]" pageId="15" pageNumber="16">Material.</emphasis>
<materialsCitation id="3B793CDBFFA8FFAA9F31C89F9A674B84" box="[321,1345,1693,1720]" collectingDate="1981-04-12" collectionCode="CAU" country="China" elevation="620" location="Sanchahe" municipality="Jinghong" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
1 ex,
<collectingCountry id="F3067616FFA8FFAA9FF5C89F9EED4B84" box="[389,459,1693,1719]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
,
<collectingRegion id="49D5F864FFA8FFAA9FA9C89C9D144B8B" box="[473,562,1694,1720]" country="China" name="Yunnan" pageId="15" pageNumber="16">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFA8FFAA9C30C89F9D8C4B84" box="[576,682,1693,1719]" pageId="15" pageNumber="16">Jinghong</collectingMunicipality>
,
<location id="8ECE605DFFA8FFAA9CC8C89F9C004B84" LSID="urn:lsid:plazi:treatment:03B88790FFA8FFAA9EE7CD649D744A58:8ECE605DFFA8FFAA9CC8C89F9C004B84" box="[696,806,1693,1719]" country="China" municipality="Jinghong" name="Sanchahe" pageId="15" pageNumber="16" stateProvince="Yunnan">Sanchahe</location>
,
<quantity id="4CE99B63FFA8FFAA9D44C89F9CA74B8B" box="[820,897,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="15" pageNumber="16" unit="m" value="620.0">
<elevation id="003CD1B5FFA8FFAA9D44C89F9CA74B8B" box="[820,897,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="15" pageNumber="16" unit="m" value="620.0">620 m</elevation>
</quantity>
,
<date id="FFAF1046FFA8FFAA9DE0C89F9B284B84" box="[912,1038,1693,1719]" pageId="15" pageNumber="16" value="1981-04-12">
<collectingDate id="EFEBE9AEFFA8FFAA9DE0C89F9B284B84" box="[912,1038,1693,1719]" pageId="15" pageNumber="16" value="1981-04-12">1981.IV.12</collectingDate>
</date>
, Chikun Yang (
<collectionCode id="ED00AE43FFA8FFAA9AB6C89F9A214B84" box="[1222,1287,1693,1719]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="15" pageNumber="16">CAU</collectionCode>
);
<specimenCount id="9D17FD0FFFA8FFAA9B6EC89F9A674B84" box="[1310,1345,1693,1719]" count="1" pageId="15" pageNumber="16" type="female">1♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B793CDBFFA8FFAA9B22C89F9B974BE8" collectingDate="2012-05-02" collectionCode="CAU" country="China" location="Tongbiguan Provincial Nature Reserve" municipality="Yingjiang" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Yunnan">
<collectingCountry id="F3067616FFA8FFAA9B22C89F9ABE4B84" box="[1362,1432,1693,1719]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
,
<collectingRegion id="49D5F864FFA8FFAA9EE7C8C09FD64BEF" box="[151,240,1730,1756]" country="China" name="Yunnan" pageId="15" pageNumber="16">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFA8FFAA9E8AC8C39E4D4BE8" box="[250,363,1729,1755]" pageId="15" pageNumber="16">Yingjiang</collectingMunicipality>
,
<location id="8ECE605DFFA8FFAA9F06C8C39C0A4BEF" LSID="urn:lsid:plazi:treatment:03B88790FFA8FFAA9EE7CD649D744A58:8ECE605DFFA8FFAA9F06C8C39C0A4BEF" box="[374,812,1729,1756]" country="China" municipality="Yingjiang" name="Tongbiguan Provincial Nature Reserve" pageId="15" pageNumber="16" stateProvince="Yunnan">Tongbiguan Provincial Nature Reserve</location>
,
<date id="FFAF1046FFA8FFAA9D47C8C39C8A4BE8" box="[823,940,1729,1756]" pageId="15" pageNumber="16" value="2012-05-02">
<collectingDate id="EFEBE9AEFFA8FFAA9D47C8C39C8A4BE8" box="[823,940,1729,1756]" pageId="15" pageNumber="16" value="2012-05-02">2012.V.02</collectingDate>
</date>
, Feiyang Liang (
<collectionCode id="ED00AE43FFA8FFAA9A19C8C39B804BE8" box="[1129,1190,1729,1755]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="15" pageNumber="16">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFA8FFAA9EB7C8E79CA64BCC" box="[199,896,1765,1792]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BAE3686FFA8FFAA9EB7C8E79CA64BCC" blockId="15.[151,1437,1045,1899]" box="[199,896,1765,1792]" pageId="15" pageNumber="16">
<emphasis id="B965EA94FFA8FFAA9EB7C8E79E474BCC" bold="true" box="[199,353,1765,1791]" pageId="15" pageNumber="16">Distribution.</emphasis>
<collectingCountry id="F3067616FFA8FFAA9F17C8E79E8C4BCC" box="[359,426,1765,1791]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
(
<collectingRegion id="49D5F864FFA8FFAA9FCBC8E49D324A33" box="[443,532,1766,1792]" country="China" name="Yunnan" pageId="15" pageNumber="16">Yunnan</collectingRegion>
,
<collectingCountry id="F3067616FFA8FFAA9C6EC8E49D804BCC" box="[542,678,1765,1792]" name="Hong Kong" pageId="15" pageNumber="16">Hong Kong</collectingCountry>
);
<collectingCountry id="F3067616FFA8FFAA9CCAC8E49DDC4A33" box="[698,762,1766,1792]" name="Japan" pageId="15" pageNumber="16">Japan</collectingCountry>
(
<collectingRegion id="49D5F864FFA8FFAA9D78C8E79C524BCC" box="[776,884,1765,1791]" country="Japan" name="Okinawa" pageId="15" pageNumber="16">Okinawa</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFA8FFAA9EB7C90B9D744A58" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BAE3686FFA8FFAA9EB7C90B9D744A58" blockId="15.[151,1437,1045,1899]" pageId="15" pageNumber="16">
<emphasis id="B965EA94FFA8FFAA9EB7C90B9E1D4A10" bold="true" box="[199,315,1801,1827]" pageId="15" pageNumber="16">Remarks.</emphasis>
<bibRefCitation id="EF804B77FFA8FFAA9F36C90B9EC74A10" author="Banks, N." box="[326,481,1801,1827]" pageId="15" pageNumber="16" pagination="255 - 291" refId="ref88124" refString="Banks, N. (1937) Neuropteroid insects from Formosa. Philippine Journal of Science, 62, 255 - 291." type="journal article" year="1937">Banks (1937)</bibRefCitation>
originally described this species without providing genital characters.
<bibRefCitation id="EF804B77FFA8FFAA9B6CC90B9FC44A7B" author="Tsukaguchi, S." pageId="15" pageNumber="16" refId="ref91372" refString="Tsukaguchi, S. (1995) Chrysopidae of Japan (Insecta, Neuroptera). Privately printed, Osaka, 224 pp." type="book" year="1995">Tsukaguchi (1995)</bibRefCitation>
redescribed this species and provided information about the male genitalia. The specimens we examined have lost their abdomens unfortunately.
</paragraph>
</subSubSection>
</treatment>
</document>
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