364 lines
59 KiB
XML
364 lines
59 KiB
XML
<document id="A5AAA9443834D6B543DC6AAA7674F35F" ID-CLB-Dataset="7965" ID-DOI="10.11646/zootaxa.4938.3.3" ID-GBIF-Dataset="fd22468d-37ba-4056-806f-2d12af30bdf0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4569679" ID-ZooBank="1619AE6B-60C3-46D2-85DF-18C03FCEA722" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614597623634" checkinUser="plazi" docAuthor="Trivedi, Jigneshkumar N., Mitra, Santanu & Ng, Peter K. L." docDate="2021" docId="03B8878DFFA0FF82FF33FE362AE33F2E" docLanguage="en" docName="zootaxa.4938.3.3.pdf" docOrigin="Zootaxa 4938 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tritodynamia bengalensis Trivedi & Mitra & Ng 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="329" masterDocId="FF81FFF5FFA1FF86FFA4FF81282D3E6C" masterDocTitle="Tritodynamia bengalensis n. sp., a new species of brachyuran crab from West Bengal state, India (Decapoda: Brachyura: Macrophthalmidae)" masterLastPageNumber="330" masterPageNumber="325" pageNumber="326" updateTime="1698931644514" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="40F4674454C852E9F8BAB91D1EE1694A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B72B6E765FF61202FF28F0129DCDCE2F">
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<mods:title id="FBF922E4E4327E2EE420ACBA064FC9BB">Tritodynamia bengalensis n. sp., a new species of brachyuran crab from West Bengal state, India (Decapoda: Brachyura: Macrophthalmidae)</mods:title>
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<mods:name id="3AC8914E4AE75A8E4E43D66A98FBDE61" type="personal">
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<mods:roleTerm id="6588C0E71D1467BF5BEA50F686AF69E9">Author</mods:roleTerm>
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<mods:namePart id="D6E379035BCE84B5DCF47D6DFB4F5937">Trivedi, Jigneshkumar N.</mods:namePart>
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<mods:affiliation id="1CCC44C46CC465CAF1EC145853A25D8E">Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India.</mods:affiliation>
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<mods:name id="2A42C8E16A76FD0E7CFAC5CDADB7E9B3" type="personal">
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<mods:roleTerm id="C9E01326E3B38F6E55BF8E04DAC319F5">Author</mods:roleTerm>
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<mods:namePart id="E7279A211DDE454974B95D74FE963899">Mitra, Santanu</mods:namePart>
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<mods:affiliation id="08091A79C472F4C99375EA9A2313E7C1">Zoological Survey of India, F. P. S. Building, 27 J. L. Nehru Road, Kolkata- 700016, West Bengal, India.</mods:affiliation>
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</mods:name>
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<mods:name id="6857BC57D4C7D0C31D5FF49440D6FAF8" type="personal">
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<mods:roleTerm id="B5D9726CEEC635CB346621DCC4C2DABB">Author</mods:roleTerm>
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<mods:namePart id="E750F9C6EB7BEB59BB713AB3B94CE0AD">Ng, Peter K. L.</mods:namePart>
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<mods:nameIdentifier id="471B0F3F517B03B007FAA0F0EFB41A9B" type="ORCID">0000-0001-5946-0608</mods:nameIdentifier>
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<mods:affiliation id="49720C08A6F03AD3CCD33EEBEF805D59">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, 117377 Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:nameIdentifier id="0DF1E2D4B980F6BE7ABE778CCA06A676" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:title id="7E0870300D8C68CA280AFD8C01C345A7">Zootaxa</mods:title>
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<mods:part id="E042542BC80B57CA959CCD5A85069135">
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<mods:date id="26B2FD894000F5E9A5D51CD84885B5E6">2021</mods:date>
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<mods:number id="1DB2FA38A173A9AC64D7AC878DF66904">2021-03-01</mods:number>
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<mods:number id="15DE9BCA22F83A5ABB519BF0AC5B8388">4938</mods:number>
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<treatment id="03B8878DFFA0FF82FF33FE362AE33F2E" ID-DOI="http://doi.org/10.5281/zenodo.4569683" ID-GBIF-Taxon="179981683" ID-Zenodo-Dep="4569683" LSID="urn:lsid:plazi:treatment:03B8878DFFA0FF82FF33FE362AE33F2E" httpUri="http://treatment.plazi.org/id/03B8878DFFA0FF82FF33FE362AE33F2E" lastPageId="4" lastPageNumber="329" pageId="1" pageNumber="326">
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<subSubSection id="C30B6510FFA0FF87FF33FE362A373FBD" box="[151,538,439,466]" pageId="1" pageNumber="326" type="nomenclature">
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<paragraph id="8BAE369BFFA0FF87FF33FE362A373FBD" blockId="1.[151,538,439,501]" box="[151,538,439,466]" pageId="1" pageNumber="326">
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<heading id="D0E681F7FFA0FF87FF33FE362A373FBD" bold="true" box="[151,538,439,466]" fontSize="11" level="1" pageId="1" pageNumber="326" reason="1">
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<emphasis id="B965EA89FFA0FF87FF33FE362A373FBD" bold="true" box="[151,538,439,466]" pageId="1" pageNumber="326">
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<taxonomicName id="4C114D18FFA0FF87FF33FE3629FD3FBE" ID-CoL="86YBY" authority="Trivedi & Mitra & Ng, 2021" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" box="[151,464,439,466]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="326" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
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<emphasis id="B965EA89FFA0FF87FF33FE3629FD3FBE" bold="true" box="[151,464,439,466]" italics="true" pageId="1" pageNumber="326">Tritodynamia bengalensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25657F2FFA0FF87FE73FE362A373FBD" box="[471,538,439,465]" pageId="1" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B6510FFA0FF87FF33FE5A293F3F99" box="[151,274,475,501]" pageId="1" pageNumber="326" type="description">
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<paragraph id="8BAE369BFFA0FF87FF33FE5A293F3F99" blockId="1.[151,538,439,501]" box="[151,274,475,501]" pageId="1" pageNumber="326">
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(
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<figureCitation id="132A2A1EFFA0FF87FF3BFE5A29243F99" box="[159,265,475,501]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 3" captionStartId-0="2.[151,250,1964,1988]" captionStartId-1="4.[151,250,1631,1655]" captionTargetBox-0="[159,1429,594,1938]" captionTargetBox-1="[165,1424,348,1598]" captionTargetId-0="figure-189@2.[159,1429,594,1938]" captionTargetId-1="figure-107@4.[151,1436,345,1608]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." captionText-1="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi-0="http://doi.org/10.5281/zenodo.4569681" figureDoi-1="http://doi.org/10.5281/zenodo.4569687" httpUri-0="https://zenodo.org/record/4569681/files/figure.png" httpUri-1="https://zenodo.org/record/4569687/files/figure.png" pageId="1" pageNumber="326">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B6510FFA0FF87FF33FDA22CB43C0D" pageId="1" pageNumber="326" type="materials_examined">
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<paragraph id="8BAE369BFFA0FF87FF33FDA22CB43C0D" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="326">
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<emphasis id="B965EA89FFA0FF87FF33FDA229AC3C51" bold="true" box="[151,385,547,573]" pageId="1" pageNumber="326">Material examined.</emphasis>
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<materialsCitation id="3B793CC6FFA0FF87FE2DFDA22CB43C0D" ID-GBIF-Occurrence="3043833301" collectingDate="2018-03-23" collectionCode="ZSIC" collectorName="S. Mitra." country="India" elevation="7" location="Shankarpur" pageId="1" pageNumber="326" specimenCode="ZSIC 8619" specimenCount="1" stateProvince="West Bengal" typeStatus="holotype">
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<typeStatus id="54AA8839FFA0FF87FE2DFDA229D93C51" box="[393,500,547,573]" pageId="1" pageNumber="326" type="holotype">Holotype</typeStatus>
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, male (CL 5.0 mm, CW
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<quantity id="4CE99B7EFFA0FF87FCAEFDA22B4F3C52" box="[778,866,547,574]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="326" unit="mm" value="9.5">9.5 mm</quantity>
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) (
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<specimenCode id="DBB79EE0FFA0FF87FCDDFDA22BD73C52" box="[889,1018,547,574]" collectionCode="ZSIC" country="India" httpUri="http://grbio.org/cool/5icv-29es" name="Zoological Survey of India" pageId="1" pageNumber="326">ZSIC 8619</specimenCode>
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/2),
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<elevation id="003CD1A8FFA0FF87FB81FDA22C903C51" box="[1061,1213,547,574]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="326" unit="m" value="6.5" valueMax="7.0" valueMin="6.0">
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<quantity id="4CE99B7EFFA0FF87FB81FDA22C593C52" box="[1061,1140,547,574]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="326" unit="m" value="6.5" valueMax="7.0" valueMin="6.0">6 –7 m</quantity>
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depth
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</elevation>
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, muddy bottom, off
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<location id="8ECE6040FFA0FF87FF33FDC629353C0D" LSID="urn:lsid:plazi:treatment:03B8878DFFA0FF82FF33FE362AE33F2E:8ECE6040FFA0FF87FF33FDC629353C0D" box="[151,280,583,609]" country="India" name="Shankarpur" pageId="1" pageNumber="326" stateProvince="West Bengal">Shankarpur</location>
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fishing harbor,
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<collectingRegion id="49D5F879FFA0FF87FE6DFDC92A773C0D" box="[457,602,583,610]" country="India" name="West Bengal" pageId="1" pageNumber="326">West Bengal</collectingRegion>
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,
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<collectingCountry id="F306760BFFA0FF87FDC1FDC62A8C3C0D" box="[613,673,583,609]" name="India" pageId="1" pageNumber="326">India</collectingCountry>
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,
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<date id="FFAF105BFFA0FF87FD09FDC62B763C0D" box="[685,859,583,609]" pageId="1" pageNumber="326" value="2018-03-23">
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<collectingDate id="EFEBE9B3FFA0FF87FD09FDC62B763C0D" box="[685,859,583,609]" pageId="1" pageNumber="326" value="2018-03-23">23 March 2018</collectingDate>
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</date>
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, from dredge, coll.
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<collectorName id="26E4534DFFA0FF87FB91FDC62CB43C0D" box="[1077,1177,583,609]" pageId="1" pageNumber="326">S. Mitra.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B6510FFA0FF84FF63FDEA28E73C51" lastPageId="2" lastPageNumber="327" pageId="1" pageNumber="326" type="description">
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<paragraph id="8BAE369BFFA0FF87FF63FDEA2B8D3DA5" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="326">
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<emphasis id="B965EA89FFA0FF87FF63FDEA29743CE9" bold="true" box="[199,345,619,645]" pageId="1" pageNumber="326">Description.</emphasis>
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Carapace elliptical, transversely elongate (
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<figureCitation id="132A2A1EFFA0FF87FC91FDEA2BB53CE9" box="[821,920,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Figs. 1A</figureCitation>
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, 2A), 1.9 times wider than long; dorsal surface slightly convex longitudinally, glabrous, sparsely punctate. Front slightly directed downwards (
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<figureCitation id="132A2A1EFFA0FF87FB7BFD0E2D153CC5" box="[1247,1336,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Fig. 1C</figureCitation>
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), frontal margin cristate, almost straight, 0.4 times of width between external orbital angles; supraorbital margin cristate with slight notch on inner angle; infraorbital margin present only on outer third, composed of row of small granules; suborbital crista lined with small granules; external orbital angle approximately right-angled, directed anteriorly, ventral surface gently swollen (
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<figureCitation id="132A2A1EFFA0FF87FE5FFC9E2A733D55" box="[507,606,799,825]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Figs. 1C</figureCitation>
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). Anterolateral margin cristate, lined with small granules, margin slightly convex behind external orbital angle; posterolateral margin cristate, lined with small granules, converging posteriorly; posterolateral region clearly demarcated from dorsal surface by sinuous line of granules; forming facet-like structure; posterior carapace margin wide, slightly convex (
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<figureCitation id="132A2A1EFFA0FF87FC94FC0A2BBE3DC9" box="[816,915,907,933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Figs. 1A</figureCitation>
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, 2A). Posterior margin of epistome with broad median triangular structure, lateral margins gently concave (Fig. 2B).
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</paragraph>
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<paragraph id="8BAE369BFFA0FF87FF63FC552AE63A85" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="326">
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Eye well developed. Antennules typical for the genus. Antennae short with flagellum as long as width of orbit (Fig. 2A). Third maxillipeds narrow, leaving large median opening when closed; ischium slightly shorter than merus, medially concave, inner margin finely toothed, with numerous long setae; outer margin also toothed medially; merus widening distally, with shallow longitudinal grove along inner margin, inner margin finely toothed, with numerous long setae; carpus widening distally, setose; propodus with foliaceous distoextensor lobe, distally rounded with long setae reaching setae of distal margin of dactylus; dactylus attached with proximoflexer angle of propodus, slightly longer than propodus, elongate-elliptical, reaching far beyond foliaceous lobe of propodus, fringed with long setae on margins (
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<figureCitation id="132A2A1EFFA0FF87FD88FB4E2AA03A85" box="[556,653,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Figs. 1B</figureCitation>
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, 2C).
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</paragraph>
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<paragraph id="8BAE369BFFA0FF87FF63FB7229A53865" blockId="1.[151,1437,547,2014]" pageId="1" pageNumber="326">
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Chelipeds symmetrical, equal in size (
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<figureCitation id="132A2A1EFFA0FF87FDDCFB722AD23B62" box="[632,767,1267,1294]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Figs. 1A, B</figureCitation>
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); merus with distodorsal margin lined with small granules; carpus rhomboidal, with distodorsal projection, strong external dorsolateral angle having some subconical granules (Fig. 2G); palm with convex outer surface, dorsal surface with row of small granules; 2 grooves on lower margin, both continuing to pollex, both bordered by cristae lined with granules (Fig. 2E), lower most cristae on ventral surface ends in deep concavity on base of pollex (Fig. 2F), upper groove from proximal end of palm, lined with granules throughout; fingers leaving gape when closed; dactylus with 2 median teeth on occlusal margin, distal half with low teeth, pollex with obtuse basal tooth, rest of the margin with low teeth, finger tips corneous, slender, pointed (
|
||
<figureCitation id="132A2A1EFFA0FF87FF5FFA6E29603865" box="[251,333,1519,1545]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Fig.1A</figureCitation>
|
||
, 2E).
|
||
</paragraph>
|
||
<paragraph id="8BAE369BFFA0FF84FF63F9922A0D3F2D" blockId="1.[151,1437,547,2014]" lastBlockId="2.[151,1437,151,573]" lastPageId="2" lastPageNumber="327" pageId="1" pageNumber="326">
|
||
Ambulatory legs (P2–P5) relatively stout (
|
||
<figureCitation id="132A2A1EFFA0FF87FD02F9922B013842" box="[678,812,1555,1582]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="1" pageNumber="326">Figs. 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A1EFFA0FF87FC9DF9922B6A3841" box="[825,839,1555,1581]" captionStart="FIGURE" captionStartId="3.[151,250,1073,1097]" captionTargetBox="[156,1419,194,1038]" captionTargetId="figure-17@3.[151,1436,181,1049]" captionTargetPageId="3" captionText="FIGURE 2 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, carapace, eyestalks, antennules and antennae, dorsal view; B, epistome, ventral view; C, third maxilliped, external view; D, pleon; E, right chela, external view; F, right chela, ventral view; G, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4569685" httpUri="https://zenodo.org/record/4569685/files/figure.png" pageId="1" pageNumber="326">2</figureCitation>
|
||
A–D), setose; P3 longest, P5 shortest. P2 with merus about 3.2 times as long as wide, with row of granules expanding distally on anterodorsal surface, rows of strong granules posterodorsal and posteroventral margins; dactylus tapering distally, covered with long setae (
|
||
<figureCitation id="132A2A1EFFA0FF87FA97F9DA2DA23819" box="[1331,1423,1627,1653]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="1" pageNumber="326">Fig. 3A</figureCitation>
|
||
). P3 combined length of merus to dactylus about 1.7 times CW; ischium with granules on dorsal surface, posterior margin with strong projecting tooth; merus stout, about 2.5 times as long as wide, anterior and posterior surfaces with numerous rounded, conical granules, posterodorsal and posteroventral margins lined with numerous conical granules; carpus expanding distally, anterior margin smooth, posterodorsal and posteroventral margin with granules, dorsal surface with 3 rows of granules; propodus 3.6 times as long as wide; anterior margin with line of granules; dorsal surface with 2 rows of granules, first row running parallel to anterior margin, second row of granules running parallel to posterior margin, row of long setae present on mid line, posterodorsal margin with row of 18 uneven strong dentiform granules, strong teeth present in proximal half; dactylus tapering distally, covered with long setae (Fig. 2B). P4 fringed with numerous long plumose setae; ischium without projecting teeth on posterior margin; merus stout; 2.5 times as long as wide, anterior, posterior surfaces with numerous rounded or subconical granules, mainly on posterodorsal, posteroventral margins, carpus anterior, posterior margin, dorsal, ventral surface smooth; propodus with convex anterior and posterior margins, 2 times as long as wide, without granules; dactylus slender, tapering distally (
|
||
<figureCitation id="132A2A1EFFA3FF84FEFAFF5E299A3E95" box="[350,439,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="2" pageNumber="327">Fig. 3C</figureCitation>
|
||
). P5 shortest, most slender, fringed with numerous long plumose setae; merus 3.1 times as long as wide, smooth, carpus smooth, longer than propodus, propodus smooth, shorter than dactylus, dactylus slender, tapering distally (
|
||
<figureCitation id="132A2A1EFFA3FF84FE1DFEA62A393F2D" box="[441,532,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="2" pageNumber="327">Fig. 3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAE369BFFA3FF84FF63FECA28E73C51" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="327">
|
||
Thoracic sternum broad (
|
||
<figureCitation id="132A2A1EFFA3FF84FE7BFECA2A183F09" box="[479,565,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="2" pageNumber="327">Fig. 1B</figureCitation>
|
||
); sternites 1 and 2 completely fused, forming subtriangular structure, medially depressed, appearing sunken; separated from sternite 3 by prominent deeply concave suture. Sternopleonal cavity deep, reaching to just before suture between sternites 2 and 3 (Fig. 2B). Male pleon obtusely triangular with all somites and telson free; somite 6 broadly trapezoidal with lateral margins slightly convex; telson rounded, inserted into distal margin of somite 6 (
|
||
<figureCitation id="132A2A1EFFA3FF84FE4DFE5A2A6C3F99" box="[489,577,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="2" pageNumber="327">Fig. 1D</figureCitation>
|
||
). G1 relatively stout, inner margin with long plumose setae, distal part curved with long tapering chitinous process, mostly obscured by dense setae (
|
||
<figureCitation id="132A2A1EFFA3FF84FC66FE7E2C163C75" box="[962,1083,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="2" pageNumber="327">Fig. 3E, F</figureCitation>
|
||
). G2 small with blunt tip (
|
||
<figureCitation id="132A2A1EFFA3FF84FACBFE7E28903C51" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="2" pageNumber="327">Fig. 3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6E6613FFA3FF84FF33F82D2A4C3984" ID-DOI="http://doi.org/10.5281/zenodo.4569681" ID-Zenodo-Dep="4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="2" pageNumber="327" startId="2.[151,250,1964,1988]" targetBox="[159,1429,594,1938]" targetPageId="2">
|
||
<paragraph id="8BAE369BFFA3FF84FF33F82D2A4C3984" blockId="2.[151,1436,1964,2024]" pageId="2" pageNumber="327">
|
||
<emphasis id="B965EA89FFA3FF84FF33F82D292039A8" bold="true" box="[151,269,1964,1988]" pageId="2" pageNumber="327">FIGURE 1</emphasis>
|
||
<taxonomicName id="4C114D18FFA3FF84FEB0F82D2A3639A8" authority="Trivedi & Mitra & Ng, 2021" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" box="[276,539,1964,1988]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="327" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
|
||
<emphasis id="B965EA89FFA3FF84FEB0F82D2A3639A8" box="[276,539,1964,1988]" italics="true" pageId="2" pageNumber="327">Tritodynamia bengalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965EA89FFA3FF84FD85F82D2A7639A8" bold="true" box="[545,603,1964,1988]" pageId="2" pageNumber="327">
|
||
<taxonomicNameLabel id="A25657F2FFA3FF84FD85F82D2A7639A8" box="[545,603,1964,1988]" pageId="2" pageNumber="327" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E6613FFA2FF85FF33FBB02C803AFD" ID-DOI="http://doi.org/10.5281/zenodo.4569685" ID-Zenodo-Dep="4569685" httpUri="https://zenodo.org/record/4569685/files/figure.png" pageId="3" pageNumber="328" startId="3.[151,250,1073,1097]" targetBox="[156,1419,194,1038]" targetPageId="3">
|
||
<paragraph id="8BAE369BFFA2FF85FF33FBB02C803AFD" blockId="3.[151,1436,1073,1169]" pageId="3" pageNumber="328">
|
||
<emphasis id="B965EA89FFA2FF85FF33FBB028D73A25" bold="true" box="[151,250,1073,1097]" pageId="3" pageNumber="328">FIGURE</emphasis>
|
||
2
|
||
<taxonomicName id="4C114D18FFA2FF85FEB0FBB02A363A25" authority="Trivedi & Mitra & Ng, 2021" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" box="[276,539,1073,1097]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
|
||
<emphasis id="B965EA89FFA2FF85FEB0FBB02A363A25" box="[276,539,1073,1097]" italics="true" pageId="3" pageNumber="328">Tritodynamia bengalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965EA89FFA2FF85FD85FBB02A763A25" bold="true" box="[545,603,1073,1097]" pageId="3" pageNumber="328">
|
||
<taxonomicNameLabel id="A25657F2FFA2FF85FD85FBB02A763A25" box="[545,603,1073,1097]" pageId="3" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, carapace, eyestalks, antennules and antennae, dorsal view; B, epistome, ventral view; C, third maxilliped, external view; D, pleon; E, right chela, external view; F, right chela, ventral view; G, carpus of left cheliped, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30B6510FFA2FF85FF63FB3E2D2E3AB5" box="[199,1283,1215,1242]" pageId="3" pageNumber="328" type="distribution">
|
||
<paragraph id="8BAE369BFFA2FF85FF63FB3E2D2E3AB5" blockId="3.[151,1437,1215,2034]" box="[199,1283,1215,1242]" pageId="3" pageNumber="328">
|
||
<emphasis id="B965EA89FFA2FF85FF63FB3E294C3AB5" bold="true" box="[199,353,1215,1241]" pageId="3" pageNumber="328">Distribution.</emphasis>
|
||
So far known only from the
|
||
<typeStatus id="54AA8839FFA2FF85FD01FB412AF83AB6" box="[677,725,1216,1242]" pageId="3" pageNumber="328">type</typeStatus>
|
||
locality off Shankarpur,
|
||
<collectingRegion id="49D5F879FFA2FF85FC4FFB412C9B3AB6" box="[1003,1206,1215,1242]" country="India" name="West Bengal" pageId="3" pageNumber="328">West Bengal state</collectingRegion>
|
||
,
|
||
<collectingCountry id="F306760BFFA2FF85FB65FB3E2CD33AB5" box="[1217,1278,1215,1241]" name="India" pageId="3" pageNumber="328">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B6510FFA2FF85FF63FB622D7E3A91" box="[199,1363,1251,1278]" pageId="3" pageNumber="328" type="etymology">
|
||
<paragraph id="8BAE369BFFA2FF85FF63FB622D7E3A91" blockId="3.[151,1437,1215,2034]" box="[199,1363,1251,1278]" pageId="3" pageNumber="328">
|
||
<emphasis id="B965EA89FFA2FF85FF63FB6229643A91" bold="true" box="[199,329,1251,1277]" pageId="3" pageNumber="328">Etymology.</emphasis>
|
||
The new species is named after the state of
|
||
<collectingRegion id="49D5F879FFA2FF85FCB8FB652B893A91" box="[796,932,1251,1278]" country="India" name="West Bengal" pageId="3" pageNumber="328">West Bengal</collectingRegion>
|
||
in
|
||
<collectingCountry id="F306760BFFA2FF85FC61FB622BD33A91" box="[965,1022,1251,1277]" name="India" pageId="3" pageNumber="328">India</collectingCountry>
|
||
, the
|
||
<typeStatus id="54AA8839FFA2FF85FB94FB652C733A92" box="[1072,1118,1252,1278]" pageId="3" pageNumber="328">type</typeStatus>
|
||
locality of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B6510FFA2FF85FF63FA862D573B29" pageId="3" pageNumber="328" type="biology_ecology">
|
||
<paragraph id="8BAE369BFFA2FF85FF63FA862D573B29" blockId="3.[151,1437,1215,2034]" pageId="3" pageNumber="328">
|
||
<emphasis id="B965EA89FFA2FF85FF63FA8629073B4D" bold="true" box="[199,298,1287,1313]" pageId="3" pageNumber="328">Ecology.</emphasis>
|
||
<materialsCitation id="3B793CC6FFA2FF85FE8BFA862D583B29" ID-GBIF-Occurrence="3043833302" country="India" county="The" elevation="7" location="Shankarpur" municipality="Midnapore district" pageId="3" pageNumber="328" specimenCount="1" stateProvince="West Bengal" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="54AA8839FFA2FF85FEC5FA8629EC3B4D" box="[353,449,1287,1313]" pageId="3" pageNumber="328" type="holotype">holotype</typeStatus>
|
||
was collected by fishing vessels using dredge from a depth of
|
||
<quantity id="4CE99B7EFFA2FF85FBC8FA862C993B4E" box="[1132,1204,1287,1314]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="328" unit="m" value="6.5" valueMax="7.0" valueMin="6.0">
|
||
<elevation id="003CD1A8FFA2FF85FBC8FA862C993B4E" box="[1132,1204,1287,1314]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="328" unit="m" value="6.5" valueMax="7.0" valueMin="6.0">6–7 m</elevation>
|
||
</quantity>
|
||
, off
|
||
<location id="8ECE6040FFA2FF85FB41FA862D4E3B4D" LSID="urn:lsid:plazi:treatment:03B8878DFFA0FF82FF33FE362AE33F2E:8ECE6040FFA2FF85FB41FA862D4E3B4D" box="[1253,1379,1287,1313]" country="India" county="The" municipality="Midnapore district" name="Shankarpur" pageId="3" pageNumber="328" stateProvince="West Bengal">Shankarpur</location>
|
||
fishing harbor located in
|
||
<collectingMunicipality id="6BCAACE1FFA2FF85FE25FAAA2A613B29" box="[385,588,1323,1349]" pageId="3" pageNumber="328">Midnapore district</collectingMunicipality>
|
||
of
|
||
<collectingRegion id="49D5F879FFA2FF85FDCBFAAD2B193B2A" box="[623,820,1323,1350]" country="India" name="West Bengal" pageId="3" pageNumber="328">West Bengal state</collectingRegion>
|
||
,
|
||
<collectingCountry id="F306760BFFA2FF85FC9BFAAA2B563B29" box="[831,891,1323,1349]" name="India" pageId="3" pageNumber="328">India</collectingCountry>
|
||
.
|
||
<collectingCounty id="62CF4E17FFA2FF85FC21FAAA2B9D3B29" box="[901,944,1323,1349]" pageId="3" pageNumber="328">The</collectingCounty>
|
||
bottom substrate was recorded as muddy
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B6510FFA2FF82FF63FACE2AE33F2E" lastPageId="4" lastPageNumber="329" pageId="3" pageNumber="328" type="discussion">
|
||
<paragraph id="8BAE369BFFA2FF85FF63FACE2DA4399E" blockId="3.[151,1437,1215,2034]" pageId="3" pageNumber="328">
|
||
<emphasis id="B965EA89FFA2FF85FF63FACE29163B05" bold="true" box="[199,315,1359,1385]" pageId="3" pageNumber="328">Remarks.</emphasis>
|
||
<taxonomicName id="4C114D18FFA2FF85FEE5FACE2A4F3B05" authority="Trivedi & Mitra & Ng, 2021" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" box="[321,610,1359,1385]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
|
||
<emphasis id="B965EA89FFA2FF85FEE5FACE2A4F3B05" box="[321,610,1359,1385]" italics="true" pageId="3" pageNumber="328">Tritodynamia bengalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965EA89FFA2FF85FDCCFACE2A883B05" bold="true" box="[616,677,1359,1385]" pageId="3" pageNumber="328">
|
||
<taxonomicNameLabel id="A25657F2FFA2FF85FDCCFACE2A883B05" box="[616,677,1359,1385]" pageId="3" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="4C114D18FFA2FF85FCCBFAD12CE23B06" authority="Yang & Tang, 2005" authorityName="Yang & Tang" authorityYear="2005" box="[879,1231,1359,1386]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="B965EA89FFA2FF85FCCBFAD12BD93B05" box="[879,1012,1359,1385]" italics="true" pageId="3" pageNumber="328">T. bidentata</emphasis>
|
||
<bibRefCitation id="EF804B6AFFA2FF85FC5CFAD12CE23B06" author="Yang, S. & Tang, B." box="[1016,1231,1359,1386]" pageId="3" pageNumber="328" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
(from Bohai,
|
||
<collectingCountry id="F306760BFFA2FF85FACEFACE28983BE2" name="China" pageId="3" pageNumber="328">China</collectingCountry>
|
||
), and
|
||
<taxonomicName id="4C114D18FFA2FF85FF52FAF52A7F3BE1" authority="Anker & Ng, 2014" authorityName="Anker & Ng" authorityYear="2014" box="[246,594,1395,1422]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="serratipes">
|
||
<emphasis id="B965EA89FFA2FF85FF52FAF529AD3BE1" box="[246,384,1396,1421]" italics="true" pageId="3" pageNumber="328">T. serratipes</emphasis>
|
||
<bibRefCitation id="EF804B6AFFA2FF85FE20FAF22A7F3BE1" author="Anker, A. & Ng, P. K. L." box="[388,594,1395,1422]" pageId="3" pageNumber="328" pagination="369 - 376" refId="ref2866" refString="Anker, A. & Ng, P. K. L. (2014) Tritodynamia serratipes sp. nov., a new marine crab from Singapore (Crustacea: Decapoda: Brachyura: Macrophthalmidae). Zootaxa, 3826 (2), 369 - 376. https: // doi. org / 10.11646 / zootaxa. 3826.2.6" type="journal article" year="2014">Anker & Ng, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(from
|
||
<collectingCountry id="F306760BFFA2FF85FD39FAF22B3D3BE2" box="[669,784,1395,1422]" name="Singapore" pageId="3" pageNumber="328">Singapore</collectingCountry>
|
||
) in its relatively wide carapace which is about two times as wide as along, possessing a relatively broad P3 merus, and with the P3 length much less than two times the width of the carapace (cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FEF2FA3D2A003BBA" author="Yang, S. & Tang, B." box="[342,557,1467,1494]" pageId="3" pageNumber="328" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF804B6AFFA2FF85FD93FA3A2B2A3BB9" author="Anker, A. & Ng, P. K. L." box="[567,775,1467,1494]" pageId="3" pageNumber="328" pagination="369 - 376" refId="ref2866" refString="Anker, A. & Ng, P. K. L. (2014) Tritodynamia serratipes sp. nov., a new marine crab from Singapore (Crustacea: Decapoda: Brachyura: Macrophthalmidae). Zootaxa, 3826 (2), 369 - 376. https: // doi. org / 10.11646 / zootaxa. 3826.2.6" type="journal article" year="2014">Anker & Ng 2014</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C114D18FFA2FF85FCBEFA3A2C113BB9" authority="Trivedi & Mitra & Ng, 2021" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" box="[794,1084,1467,1493]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
|
||
<emphasis id="B965EA89FFA2FF85FCBEFA3A2C113BB9" box="[794,1084,1467,1493]" italics="true" pageId="3" pageNumber="328">Tritodynamia bengalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965EA89FFA2FF85FBE7FA3A2CAF3BB9" bold="true" box="[1091,1154,1467,1493]" pageId="3" pageNumber="328">
|
||
<taxonomicNameLabel id="A25657F2FFA2FF85FBE7FA3A2CAF3BB9" box="[1091,1154,1467,1493]" pageId="3" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C114D18FFA2FF85FAB3FA3D2DB13BB9" authorityName="Yang & Tang" authorityYear="2005" box="[1303,1436,1467,1493]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="B965EA89FFA2FF85FAB3FA3D2DB13BB9" box="[1303,1436,1467,1493]" italics="true" pageId="3" pageNumber="328">T. bidentata</emphasis>
|
||
</taxonomicName>
|
||
markedly in having the carapace elliptical in shape (
|
||
<figureCitation id="132A2A1EFFA2FF85FD4DFA5E2B633B95" box="[745,846,1503,1529]" captionStart="FIGURE 1" captionStartId="2.[151,250,1964,1988]" captionTargetBox="[159,1429,594,1938]" captionTargetId="figure-189@2.[159,1429,594,1938]" captionTargetPageId="2" captionText="FIGURE 1 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, dorsal view; B, ventral view; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.4569681" httpUri="https://zenodo.org/record/4569681/files/figure.png" pageId="3" pageNumber="328">Figs. 1A</figureCitation>
|
||
, 2A) (versus broadly trapezoidal in
|
||
<taxonomicName id="4C114D18FFA2FF85FB4DFA612D5D3B95" authorityName="Yang & Tang" authorityYear="2005" box="[1257,1392,1503,1529]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="B965EA89FFA2FF85FB4DFA612D5D3B95" box="[1257,1392,1503,1529]" italics="true" pageId="3" pageNumber="328">T. bidentata</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FF33F98529433872" author="Yang, S. & Tang, B." box="[151,366,1539,1566]" pageId="3" pageNumber="328" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang 2005</bibRefCitation>
|
||
: fig. 1A); the dactylus of the third maxilliped is only slightly longer than the propodus (Fig. 2C) (versus the dactylus of the third maxilliped is prominently longer than the propodus in
|
||
<taxonomicName id="4C114D18FFA2FF85FBF2F9A92CF7382D" authorityName="Yang & Tang" authorityYear="2005" box="[1110,1242,1575,1601]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="B965EA89FFA2FF85FBF2F9A92CF7382D" box="[1110,1242,1575,1601]" italics="true" pageId="3" pageNumber="328">T. bidentata</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FAADF9A928FE380A" author="Yang, S. & Tang, B." pageId="3" pageNumber="328" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang 2005</bibRefCitation>
|
||
: fig. 1B); the ventral margin of the P3 merus is distinctly serrated with sharp granules (
|
||
<figureCitation id="132A2A1EFFA2FF85FB30F9CA2CC43809" box="[1172,1257,1611,1637]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="3" pageNumber="328">Fig. 3B</figureCitation>
|
||
) (versus ventral margin of the P3 merus more weakly granulated and not appearing serrate; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FBA0F9F12CF138E6" author="Yang, S. & Tang, B." box="[1028,1244,1647,1674]" pageId="3" pageNumber="328" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang 2005</bibRefCitation>
|
||
: fig. 1E); and the G1 distal part possesses a long tapering chitinous process (
|
||
<figureCitation id="132A2A1EFFA2FF85FC8AF9122B8C38C2" box="[814,929,1683,1710]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="3" pageNumber="328">Fig. 3E, F</figureCitation>
|
||
) (versus distal part divided into two rounded lobes without any elongate chitinous process in
|
||
<taxonomicName id="4C114D18FFA2FF85FD18F9392B6E38BD" authorityName="Yang & Tang" authorityYear="2005" box="[700,835,1719,1745]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="B965EA89FFA2FF85FD18F9392B6E38BD" box="[700,835,1719,1745]" italics="true" pageId="3" pageNumber="328">T. bidentata</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FCDCF9392C7838BE" author="Yang, S. & Tang, B." box="[888,1109,1719,1746]" pageId="3" pageNumber="328" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang 2005</bibRefCitation>
|
||
: fig. 2A).
|
||
<taxonomicName id="4C114D18FFA2FF85FB6FF93628DE3899" authority="Trivedi & Mitra & Ng, 2021" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
|
||
<emphasis id="B965EA89FFA2FF85FB6FF93628DE3899" italics="true" pageId="3" pageNumber="328">Tritodynamia bengalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B965EA89FFA2FF85FF5EF95A29143899" bold="true" box="[250,313,1755,1781]" pageId="3" pageNumber="328">
|
||
<taxonomicNameLabel id="A25657F2FFA2FF85FF5EF95A29143899" box="[250,313,1755,1781]" pageId="3" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears to be closest to
|
||
<taxonomicName id="4C114D18FFA2FF85FDE9F95D2AF53899" authorityName="Anker & Ng" authorityYear="2014" box="[589,728,1756,1781]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="serratipes">
|
||
<emphasis id="B965EA89FFA2FF85FDE9F95D2AF53899" box="[589,728,1756,1781]" italics="true" pageId="3" pageNumber="328">T. serratipes</emphasis>
|
||
</taxonomicName>
|
||
in having a P3 merus which is also prominently serrated along its ventral margin; it can nevertheless be separated by the dactylus of the third maxilliped being relatively shorter, only slightly longer than the propodus (Fig. 2C) (versus the dactylus of the third maxilliped is much longer than the propodus in
|
||
<taxonomicName id="4C114D18FFA2FF85FE80F8C9299D390D" authorityName="Anker & Ng" authorityYear="2014" box="[292,432,1864,1889]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="serratipes">
|
||
<emphasis id="B965EA89FFA2FF85FE80F8C9299D390D" box="[292,432,1864,1889]" italics="true" pageId="3" pageNumber="328">T. serratipes</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FE46F8C62A9E390D" author="Anker, A. & Ng, P. K. L." box="[482,691,1863,1890]" pageId="3" pageNumber="328" pagination="369 - 376" refId="ref2866" refString="Anker, A. & Ng, P. K. L. (2014) Tritodynamia serratipes sp. nov., a new marine crab from Singapore (Crustacea: Decapoda: Brachyura: Macrophthalmidae). Zootaxa, 3826 (2), 369 - 376. https: // doi. org / 10.11646 / zootaxa. 3826.2.6" type="journal article" year="2014">Anker & Ng 2014</bibRefCitation>
|
||
: fig. 1E); there is only one strong and one low tooth at the base of pollex (Fig. 2E) (versus with two strong teeth present in proximal half of pollex in
|
||
<taxonomicName id="4C114D18FFA2FF85FBE9F8ED2CF739E9" authorityName="Anker & Ng" authorityYear="2014" box="[1101,1242,1900,1925]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="serratipes">
|
||
<emphasis id="B965EA89FFA2FF85FBE9F8ED2CF739E9" box="[1101,1242,1900,1925]" italics="true" pageId="3" pageNumber="328">T. serratipes</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FAA9F8EA28FE39C5" author="Anker, A. & Ng, P. K. L." pageId="3" pageNumber="328" pagination="369 - 376" refId="ref2866" refString="Anker, A. & Ng, P. K. L. (2014) Tritodynamia serratipes sp. nov., a new marine crab from Singapore (Crustacea: Decapoda: Brachyura: Macrophthalmidae). Zootaxa, 3826 (2), 369 - 376. https: // doi. org / 10.11646 / zootaxa. 3826.2.6" type="journal article" year="2014">Anker & Ng 2014</bibRefCitation>
|
||
: figs. 1F, 3C); the P3 propodus is proportionately longer and is relatively less strongly serrated with small 18 teeth on the ventral margin (
|
||
<figureCitation id="132A2A1EFFA2FF85FE77F8322A0439A1" box="[467,553,1971,1997]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="3" pageNumber="328">Fig. 3B</figureCitation>
|
||
) (versus proportionately shorter and strongly serrated along the ventral margin with 15 large teeth in
|
||
<taxonomicName id="4C114D18FFA2FF85FE28F8592A3A399D" authorityName="Anker & Ng" authorityYear="2014" box="[396,535,2008,2033]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="serratipes">
|
||
<emphasis id="B965EA89FFA2FF85FE28F8592A3A399D" box="[396,535,2008,2033]" italics="true" pageId="3" pageNumber="328">T. serratipes</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF804B6AFFA2FF85FDECF8562B3A399D" author="Anker, A. & Ng, P. K. L." box="[584,791,2007,2034]" pageId="3" pageNumber="328" pagination="369 - 376" refId="ref2866" refString="Anker, A. & Ng, P. K. L. (2014) Tritodynamia serratipes sp. nov., a new marine crab from Singapore (Crustacea: Decapoda: Brachyura: Macrophthalmidae). Zootaxa, 3826 (2), 369 - 376. https: // doi. org / 10.11646 / zootaxa. 3826.2.6" type="journal article" year="2014">Anker & Ng 2014</bibRefCitation>
|
||
: figs. 1J, K, 3A). Males are not known for
|
||
<taxonomicName id="4C114D18FFA2FF85FB53F8592DAF399D" authorityName="Anker & Ng" authorityYear="2014" box="[1271,1410,2008,2033]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="serratipes">
|
||
<emphasis id="B965EA89FFA2FF85FB53F8592DAF399D" box="[1271,1410,2008,2033]" italics="true" pageId="3" pageNumber="328">T. serratipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAE369BFFA5FF82FF63FF162AE33F2E" blockId="4.[151,1436,151,322]" pageId="4" pageNumber="329">
|
||
The G1 structure of
|
||
<taxonomicName id="4C114D18FFA5FF82FE0CFF162A6B3EDD" authorityName="Trivedi & Mitra & Ng" authorityYear="2021" box="[424,582,151,177]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="bengalensis">
|
||
<emphasis id="B965EA89FFA5FF82FE0CFF162A6B3EDD" box="[424,582,151,177]" italics="true" pageId="4" pageNumber="329">T. bengalensis</emphasis>
|
||
</taxonomicName>
|
||
is unusual in that it is relatively slender, is not bilobed distally and possesses an elongate distal process which is sharply bent (
|
||
<figureCitation id="132A2A1EFFA5FF82FD1EFF3A2B033EB9" box="[698,814,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1631,1655]" captionTargetBox="[165,1424,348,1598]" captionTargetId="figure-107@4.[151,1436,345,1608]" captionTargetPageId="4" captionText="FIGURE 3 Tritodynamia bengalensis n. sp., holotype, male (CL 5.0 mm, CW 9.5 mm) (ZSIC 8619/2) West Bengal, India. A, right P2, dorsal view; B, right P3, dorsal view; C, right P4, dorsal view; D, right P5, dorsal view; E, left G1(ventral view); F, left G1, distal part(denuded); G, left G2 (denuded)." figureDoi="http://doi.org/10.5281/zenodo.4569687" httpUri="https://zenodo.org/record/4569687/files/figure.png" pageId="4" pageNumber="329">Fig. 3E, F</figureCitation>
|
||
). All other species of
|
||
<taxonomicName id="4C114D18FFA5FF82FB85FF3A2C953EB9" authorityName="Ortmann" authorityYear="1894" box="[1057,1208,187,213]" class="Malacostraca" family="Macrophthalmidae" genus="Tritodynamia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="329" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B965EA89FFA5FF82FB85FF3A2C953EB9" box="[1057,1208,187,213]" italics="true" pageId="4" pageNumber="329">Tritodynamia</emphasis>
|
||
</taxonomicName>
|
||
have G1s which are distinctly stouter, and the distal part is distinctly bilobed and/or rounded, without any elongate processes (cf.
|
||
<bibRefCitation id="EF804B6AFFA5FF82FAC0FF5E28FE3F71" author="Shen, C." pageId="4" pageNumber="329" pagination="1 - 10" refId="ref3567" refString="Shen, C. (1932) The Brachyuran Crustacea of North China. In: Zoologica Sinica. Series A (Invertebrates of China). 9 (1). The Fan Institute of Biology, Peking [Beijing], pp. i-x + 1 - 321, pls. 1 - 10, 1 map." type="book chapter" year="1932">Shen 1932</bibRefCitation>
|
||
: text-fig. 70b, 72c;
|
||
<bibRefCitation id="EF804B6AFFA5FF82FE0EFE822A0A3F71" author="Chen, H." box="[426,551,259,285]" pageId="4" pageNumber="329" pagination="334 - 337" refId="ref2972" refString="Chen, H. (1979) A new species of Tritodynamia (Crustacea: Brachyura) from Fujian Province, China. Oceanologia et Limnologia Sinica, 10 (4), 334 - 337. [in Chinese with English abstract]" type="journal article" year="1979">Chen 1979</bibRefCitation>
|
||
: fig.1-10, 1-11;
|
||
<bibRefCitation id="EF804B6AFFA5FF82FD7CFE822BAD3F71" author="Dai, A. - Y. & Feng, Z. - Q. & Song, Y. - Z. & Chen, G. - X." box="[728,896,259,285]" pageId="4" pageNumber="329" pagination="129 - 143" refId="ref3016" refString="Dai, A. - Y., Feng, Z. - Q., Song, Y. - Z. & Chen, G. - X. (1980) New species and new records of family Pinnotheridae from Hainan Island. Acta Zootaxonomica Sinica, 5 (2), 129 - 143. [in Chinese with English summary]" type="journal article" year="1980">
|
||
Dai
|
||
<emphasis id="B965EA89FFA5FF82FCA2FE822B133F71" box="[774,830,259,285]" italics="true" pageId="4" pageNumber="329">et al.</emphasis>
|
||
1980
|
||
</bibRefCitation>
|
||
: figs. 10E, 11E;
|
||
<bibRefCitation id="EF804B6AFFA5FF82FB90FE822CD03F71" author="Yang, S. & Sun, X." box="[1076,1277,259,285]" pageId="4" pageNumber="329" pagination="1 - 3" refId="ref3730" refString="Yang, S. & Sun, X. (1996) A new species of Tritodynamia from China (Crustacea: Decapoda: Pinnotheridae). Memoirs of Beijing Natural History Museum, 55, 1 - 3." type="journal article" year="1996">Yang & Sun 1996</bibRefCitation>
|
||
: fig. 1K;
|
||
<bibRefCitation id="EF804B6AFFA5FF82FAC6FE8229033F2D" author="Yang, S. & Tang, B." pageId="4" pageNumber="329" pagination="781 - 786" refId="ref3770" refString="Yang, S. & Tang, B. (2005) A new species of Tritodynamia (Decapoda, Brachyura, Pinnotheridae) from the Bohai Sea, China, with a key to the genus Tritodynami a Ortmann, 1894. Crustaceana, 78, 781 - 786. https: // doi. org / 10.1163 / 156854005774445528" type="journal article" year="2005">Yang & Tang 2005</bibRefCitation>
|
||
: fig. 2A;
|
||
<bibRefCitation id="EF804B6AFFA5FF82FE33FEA62A423F2D" author="Naruse, T. & Ng, P. K. L." box="[407,623,295,321]" pageId="4" pageNumber="329" pagination="291 - 298" refId="ref3267" refString="Naruse, T. & Ng, P. K. L. (2010) A new species of Tritodynamia Ortmann, 1894 (Decapoda, Brachyura, Macrophthalmidae) from Singapore. Crustaceana, 83, 291 - 298. https: // doi. org / 10.1163 / 001121610 X 12627655658249" type="journal article" year="2010">Naruse & Ng 2010</bibRefCitation>
|
||
: fig. 1f).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |