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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="A748A1F3BC804CB9971EA5EF97702922" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Calosoma luxatum Say 1823" docType="treatment" docVersion="10" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="186" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="184" updateTime="1668154792637" updateUser="ExternalLinkService">
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<mods:title>Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bousquet, Yves</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
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<mods:date>2012</mods:date>
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<mods:number>2012-11-28</mods:number>
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<mods:number>245</mods:number>
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<treatment ID-GBIF-Taxon="127869052" LSID="urn:lsid:plazi:treatment:A748A1F3BC804CB9971EA5EF97702922" httpUri="http://treatment.plazi.org/id/A748A1F3BC804CB9971EA5EF97702922" lastPageId="185" lastPageNumber="186" pageId="183" pageNumber="184">
<subSubSection pageId="183" pageNumber="184" type="nomenclature">
<paragraph pageId="183" pageNumber="184">
<taxonomicName LSID="A748A1F3-BC80-4CB9-971E-A5EF97702922" authority="Say, 1823" authorityName="Say" authorityYear="1823" class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Calosoma luxatum" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="luxatum">Calosoma luxatum Say, 1823</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="185" lastPageNumber="186" pageId="183" pageNumber="184" type="reference_group">
<paragraph pageId="183" pageNumber="184">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="luxata">Calosoma luxata</taxonomicName>
Say, 1823b: 149. Type locality: &quot;Douglas Spring, Routt Co[unty], Colo[rado]&quot; (neotype label). Neotype (♂), designated by Lindroth and Freitag (1969: 331), in MCZ [# 33087]. Note. &quot;Arkansa [probably the Arkansas River since Say added &quot;found near the Rocky mountains
<normalizedToken originalValue="”]»">&quot;]&quot;</normalizedToken>
was the area originally cited by Say (1823b: 150).
</paragraph>
<paragraph pageId="183" pageNumber="184">
<taxonomicName authorityName="LeConte" authorityYear="1847" class="Hexapoda" family="Carabidae" genus="Carabus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="zimmermani">Carabus zimmermani</taxonomicName>
LeConte, 1847: 445. Type locality: &quot;Rocky Mountains&quot; (original citation). Holotype [by monotypy] in MCZ [# 638]. Synonymy established by Jeannel (1940: 175). Etymology. The species name honors Christian Zimmermann [1800-1867], an accomplished entomologist. Born in Germany, Zimmermann immigrated to the United States at the age of 31 and eventually settled in South Carolina where he divided his time between collecting trips along the East Coast and working on his collection at home. His collection was bought by Dr. Lewis of Philadelphia and from him by George Robert Crotch who sold it to the MCZ. According to Hagen (1889: 57), &quot;a great part [of
<normalizedToken originalValue="Zimmermanns">Zimmermann's</normalizedToken>
specimens] is in
<normalizedToken originalValue="Lecontes">Leconte's</normalizedToken>
collection, and can be recognized at once by the number on the pins in
<normalizedToken originalValue="Zimmermanns">Zimmermann's</normalizedToken>
hand-writing.&quot;
</paragraph>
<paragraph pageId="183" pageNumber="184">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="striatulum">Calosoma striatulum</taxonomicName>
LeConte, 1859c: 4 [primary homonym of
<taxonomicName class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="striatulum">
<emphasis italics="true" pageId="183" pageNumber="184">Calosoma striatulum</emphasis>
</taxonomicName>
Chevrolat, 1835]. Type locality: &quot;Milk river [probably in Montana]; Utah&quot; (original citation). Syntype(s) in MCZ [# 639]. Synonymy established by Burgess and Collins (1917: 120), confirmed by Lindroth (1961a: 54).
</paragraph>
<paragraph lastPageId="184" lastPageNumber="185" pageId="183" pageNumber="184">
<taxonomicName authorityName="F. Walker" authorityYear="1866" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="pimelioides">Callisthenes pimelioides</taxonomicName>
Walker, 1866: 312. Type locality: British Columbia (inferred from title of the book), restricted to
<normalizedToken originalValue="«Oliver»">&quot;Oliver&quot;</normalizedToken>
by Lindroth (1961a: 54). At least
<pageBreakToken pageId="184" pageNumber="185" start="start">one</pageBreakToken>
syntype in BMNH (Lindroth 1961a: 55). Synonymy established, under the name
<taxonomicName baseAuthorityName="LeConte" baseAuthorityYear="1847" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="zimmermani">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes zimmermani</emphasis>
</taxonomicName>
LeConte, by LeConte (1870: 399) and Horn (1870a: 70), confirmed by Lindroth (1961a: 54).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="luxatus">Callisthenes luxatus</taxonomicName>
var.
<emphasis italics="true" pageId="184" pageNumber="185">opacus</emphasis>
<normalizedToken originalValue="Géhin">Gehin</normalizedToken>
, 1885: 70. Type locality:
<normalizedToken originalValue="«Orégon»">&quot;Oregon&quot;</normalizedToken>
(original citation). Syntype(s) in MHNP (Deuve 1978: 252). Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 83).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1913" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="exaratus">Callisthenes exaratus</taxonomicName>
Casey, 1913: 72. Type locality: &quot;Placer Co[unty], California&quot; (original citation). Three syntypes [3 originally cited] in USNM [# 37132]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1913" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="tegulatus">Callisthenes tegulatus</taxonomicName>
Casey, 1913: 72. Type locality:
<normalizedToken originalValue="«California»">&quot;California&quot;</normalizedToken>
(original citation). One syntype in USNM [# 37128]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 83).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="tegulatus" subSpecies="viator">Callisthenes tegulatus viator</taxonomicName>
Casey, 1913: 72. Type locality:
<normalizedToken originalValue="«California»">&quot;California&quot;</normalizedToken>
(original citation). One syntype in USNM [# 37129]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 83).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1913" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="pustulosus">Callisthenes pustulosus</taxonomicName>
Casey, 1913: 73. Type locality: &quot;Yreka [Siskiyou County], California&quot; (original citation). One syntype in USNM [# 37127]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1913" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="diffractus">Callisthenes diffractus</taxonomicName>
Casey, 1913: 75. Type locality: &quot;Coolidge [McKinley County], New Mexico&quot; (original citation). One syntype in USNM [# 37126]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="reflexus">Callisthenes reflexus</taxonomicName>
Casey, 1920: 164. Type locality: &quot;northern Rocky Mountain region&quot; (original citation). One syntype in USNM [# 37130]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="utensis">Callisthenes utensis</taxonomicName>
Casey, 1920: 165. Type locality: &quot;Stockton [Tooele County], Utah&quot; (original citation). One syntype in USNM [# 37131]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="semotus">Callisthenes semotus</taxonomicName>
Casey, 1920: 166. Type locality: &quot;Stockton [Tooele County], Utah&quot; (original citation). One syntype in USNM [# 37133]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="debilis">Callisthenes debilis</taxonomicName>
Casey, 1920: 167. Type locality:
<normalizedToken originalValue="«Oregon»">&quot;Oregon&quot;</normalizedToken>
(original citation). One syntype in USNM [# 37134] and one in SIM (Hennessey 1990: 466). Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="parowanus">Callisthenes parowanus</taxonomicName>
Casey, 1920: 167. Type locality: &quot;Parowan [Iron County], Utah&quot; (original citation). One syntype in USNM [# 37136]. Synonymy established, under the name
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="subSpecies" species="luxatum" subSpecies="zimmermanni">
<emphasis italics="true" pageId="184" pageNumber="185">Callisthenes luxatum zimmermanni</emphasis>
</taxonomicName>
LeConte, by Breuning (1928b: 84).
</paragraph>
<paragraph pageId="184" pageNumber="185">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="striata">Calosoma striata</taxonomicName>
Breuning, 1928b: 86. Replacement name for
<taxonomicName class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="striatulum">
<emphasis italics="true" pageId="184" pageNumber="185">Calosoma striatulum</emphasis>
</taxonomicName>
LeConte, 1859.
</paragraph>
<paragraph pageId="185" pageNumber="186">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="subSpecies" species="zimmermanni" subSpecies="tahoensis">
<pageBreakToken pageId="185" pageNumber="186" start="start">Calosoma</pageBreakToken>
zimmermanni tahoensis
</taxonomicName>
Van Dyke, 1943: 18. Type locality: &quot;slopes of M[oun]t Tallac, near Lake Tahoe [Placer County], California&quot; (original citation). Holotype (♂) in CAS [# 5296]. Synonymy established, under the name
<taxonomicName authorityName="LeConte" authorityYear="1848" class="Hexapoda" family="Carabidae" genus="Calosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis italics="true" pageId="185" pageNumber="186">Calosoma zimmermanni</emphasis>
</taxonomicName>
LeConte, by Gidaspow (1959: 318).
</paragraph>
</subSubSection>
<subSubSection pageId="185" pageNumber="186" type="distribution">
<paragraph pageId="185" pageNumber="186">Distribution.</paragraph>
<paragraph pageId="185" pageNumber="186">
This species ranges from southern Saskatchewan to the Okanagan Valley in south-central British Columbia (Lindroth 1961a: 56), south to southeastern California, northwestern New Mexico (Casey, 1913: 75, as
<taxonomicName authorityName="Casey" authorityYear="1913" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="diffractus">
<emphasis italics="true" pageId="185" pageNumber="186">Callisthenes diffractus</emphasis>
</taxonomicName>
), and northern Kansas [see Gidaspow 1959: Fig. 11, as
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="luxatus">
<emphasis italics="true" pageId="185" pageNumber="186">Callisthenes luxatus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="F. Walker" authorityYear="1866" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="pimelioides">
<emphasis italics="true" pageId="185" pageNumber="186">Callisthenes pimelioides</emphasis>
</taxonomicName>
, and
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis italics="true" pageId="185" pageNumber="186">Callisthenes zimmermanni</emphasis>
</taxonomicName>
]. At least one specimen simply labeled from New Mexico is known (Gidaspow 1959: 311). The records from
<normalizedToken originalValue="“Oklahoma”">&quot;Oklahoma&quot;</normalizedToken>
and
<normalizedToken originalValue="“Texas”">&quot;Texas&quot;</normalizedToken>
(Burgess and Collins 1917: 121, as
<taxonomicName baseAuthorityName="LeConte" baseAuthorityYear="1847" class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="zimmermani">
<emphasis italics="true" pageId="185" pageNumber="186">Callisthenes zimmermani</emphasis>
</taxonomicName>
) need confirmation.
</paragraph>
</subSubSection>
<subSubSection pageId="185" pageNumber="186" type="records">
<paragraph pageId="185" pageNumber="186">Records.</paragraph>
<paragraph pageId="185" pageNumber="186">
<emphasis bold="true" pageId="185" pageNumber="186">CAN</emphasis>
: AB, BC, SK
<emphasis bold="true" pageId="185" pageNumber="186">USA</emphasis>
: AZ, CA, CO, ID, KS, MT, NE, NM, NV, OR, SD, UT, WA, WY [OK, TX]
</paragraph>
</subSubSection>
<subSubSection pageId="185" pageNumber="186" type="note">
<paragraph pageId="185" pageNumber="186">Note.</paragraph>
<paragraph pageId="185" pageNumber="186">
Gidaspow (1959) considered
<taxonomicName class="Insecta" family="Carabidae" genus="Callisthenes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="pimeloides">
<emphasis italics="true" pageId="185" pageNumber="186">Callisthenes pimeloides</emphasis>
</taxonomicName>
Walker [synonyms:
<emphasis italics="true" pageId="185" pageNumber="186">tegulatus</emphasis>
Casey,
<emphasis italics="true" pageId="185" pageNumber="186">viator</emphasis>
Casey,
<emphasis italics="true" pageId="185" pageNumber="186">pustulosus</emphasis>
Casey,
<emphasis italics="true" pageId="185" pageNumber="186">reflexus</emphasis>
Casey, and
<emphasis italics="true" pageId="185" pageNumber="186">parowanus</emphasis>
Casey] and
<taxonomicName class="Hexapoda" family="Carabidae" genus="Carabus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis italics="true" pageId="185" pageNumber="186">Carabus zimmermanni</emphasis>
</taxonomicName>
LeConte [synonyms:
<emphasis italics="true" pageId="185" pageNumber="186">opacus</emphasis>
<normalizedToken originalValue="Géhin">Gehin</normalizedToken>
,
<emphasis italics="true" pageId="185" pageNumber="186">exaratus</emphasis>
Casey,
<emphasis italics="true" pageId="185" pageNumber="186">debilis</emphasis>
Casey, and
<emphasis italics="true" pageId="185" pageNumber="186">tahoensis</emphasis>
Van Dyke] as distinct species but Lindroth (1961a: 55) argued that the structural differences noted between these forms are no more than intraspecific variation.
</paragraph>
</subSubSection>
</treatment>
</document>