402 lines
53 KiB
XML
402 lines
53 KiB
XML
<document id="6133998617D0B61A8BA7299E5F278729" ID-CLB-Dataset="28054" ID-DOI="10.11646/zootaxa.4508.2.1" ID-GBIF-Dataset="da79b654-bd50-4715-8ed3-80fa7a00d469" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606934" ID-ZooBank="5E2BC894-1919-4F63-8EF5-BAAC91913388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585631772" checkinUser="plazi" docAuthor="Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel" docDate="2018" docId="F95B87D7FFA4950CFF49FAF6D120587E" docLanguage="en" docName="zootaxa.4508.2.1.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila (Vovidesa) chiapensis Tang & Skelley & Pérez-Farrera 2018, new species" docType="treatment" docVersion="7" lastPageNumber="171" masterDocId="0562FFAFFFB59518FFDEFF8BD26C5B01" masterDocTitle="Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)" masterLastPageNumber="178" masterPageNumber="151" pageNumber="168" updateTime="1698712984586" updateUser="ExternalLinkService">
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<mods:title id="E7CFA56EAE018C26010417D93BCF6BD3">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="436C359AF39D799007EB15F08AFD27F7">Tang, William</mods:namePart>
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<mods:namePart id="3411BDC5B5AC844F7F3D9B041C6FF6FB">Skelley, Paul</mods:namePart>
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<mods:namePart id="DCA0249F5089D27335F7CEB8E2004B1A">Pérez-Farrera, Miguel Angel</mods:namePart>
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<mods:typeOfResource id="B2C813BC33FFFC5CF860C3C7C307B938">text</mods:typeOfResource>
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<mods:title id="FAF8E1DE3AE0B35C78EC62D95AA76F0E">Zootaxa</mods:title>
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<mods:part id="1B8E983EB9A91FDBB9472BADD4DC81CD">
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<mods:date id="0270366DBF14303832B6E3A807FDB5DD">2018</mods:date>
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<mods:number id="1A66B15DD7296CAEC1CF16A2C18DF020">2018-10-31</mods:number>
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<mods:number id="0342392E63153A181FEB01E84970F878">4508</mods:number>
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<mods:identifier id="B7566808BB39FCA8D5F94E5C8835387E" type="DOI">10.11646/zootaxa.4508.2.1</mods:identifier>
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<treatment id="F95B87D7FFA4950CFF49FAF6D120587E" ID-DOI="http://doi.org/10.5281/zenodo.5958040" ID-GBIF-Taxon="154495720" ID-Zenodo-Dep="5958040" LSID="urn:lsid:plazi:treatment:F95B87D7FFA4950CFF49FAF6D120587E" httpUri="http://treatment.plazi.org/id/F95B87D7FFA4950CFF49FAF6D120587E" lastPageId="20" lastPageNumber="171" pageId="17" pageNumber="168">
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<subSubSection id="39E8654AFFA49509FF49FAF6D63A5E96" box="[151,1110,1405,1431]" pageId="17" pageNumber="168" type="nomenclature">
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<paragraph id="714D36C1FFA49509FF49FAF6D63A5E96" blockId="17.[151,1110,1405,1464]" box="[151,1110,1405,1431]" pageId="17" pageNumber="168">
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<heading id="2A0581ADFFA49509FF49FAF6D63A5E96" bold="true" box="[151,1110,1405,1431]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
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<emphasis id="4386EAD3FFA49509FF49FAF6D63A5E96" bold="true" box="[151,1110,1405,1431]" pageId="17" pageNumber="168">
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<taxonomicName id="B6F24D42FFA49509FF49FAF6D1D65E96" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,954,1405,1431]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Vovidesa">
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<emphasis id="4386EAD3FFA49509FF49FAF6D34B5E96" bold="true" box="[151,295,1405,1431]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFA49509FEE9FAF6D3F25E96" bold="true" box="[311,414,1405,1431]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
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)
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<emphasis id="4386EAD3FFA49509FE6CFAF6D05D5E96" bold="true" box="[434,561,1405,1431]" italics="true" pageId="17" pageNumber="168">chiapensis</emphasis>
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Tang, Skelley & Pérez-Farrera
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</taxonomicName>
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,
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<taxonomicNameLabel id="58B557A8FFA49509FC19FAF6D63A5E96" box="[967,1110,1405,1431]" pageId="17" pageNumber="168" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFA49509FF49FA14D3235EB9" box="[151,335,1439,1464]" pageId="17" pageNumber="168" type="description">
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<paragraph id="714D36C1FFA49509FF49FA14D3235EB9" blockId="17.[151,1110,1405,1464]" box="[151,335,1439,1464]" pageId="17" pageNumber="168">
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(
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<figureCitation id="E9C92A44FFA49509FF41FA14D2935EB9" box="[159,255,1439,1464]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="17" pageNumber="168">Figs. 1E</figureCitation>
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,
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<figureCitation id="E9C92A44FFA49509FED2FA14D3755EB9" box="[268,281,1439,1464]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="17" pageNumber="168">6</figureCitation>
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A–I)
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFA49509FF49FA6CD0325D8E" pageId="17" pageNumber="168" type="diagnosis">
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<paragraph id="714D36C1FFA49509FF49FA6CD0325D8E" blockId="17.[151,1437,1511,2005]" pageId="17" pageNumber="168">
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<emphasis id="4386EAD3FFA49509FF49FA6CD30D5D01" bold="true" box="[151,353,1511,1536]" pageId="17" pageNumber="168">Adult diagnosis.</emphasis>
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Distinguished from other
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<taxonomicName id="B6F24D42FFA49509FD74FA63D1DA5D01" authority="(Vovidesa)" baseAuthorityName="Vovidesa" box="[682,950,1512,1536]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="155" phylum="Tracheophyta" rank="genus">
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<emphasis id="4386EAD3FFA49509FD74FA63D15D5D01" box="[682,817,1512,1536]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFA49509FC94FA63D1C75D01" box="[842,939,1512,1536]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
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)
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</taxonomicName>
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spp. by male lacking subapical medial emarginations on tibiae; submentum of both sexes with sparse punctures and setae; female elytra dulled medially and laterally; female abdomen strongly dulled, obscuring punctures; male genitalia with median lobe laterally flattened and curved; and with a known distribution in
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<collectingCountry id="09E57651FFA49509FCC3F9DFD1195D6C" box="[797,885,1620,1645]" name="Mexico" pageId="17" pageNumber="168">Mexico</collectingCountry>
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, state of
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<collectingRegion id="B336F823FFA49509FC3DF9DFD62C5D6C" box="[995,1088,1620,1645]" country="Mexico" name="Chiapas" pageId="17" pageNumber="168">Chiapas</collectingRegion>
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, on
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<taxonomicName id="B6F24D42FFA49509FBABF9DFD71C5D6D" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[1141,1392,1620,1644]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="alvarezii">
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<emphasis id="4386EAD3FFA49509FBABF9DFD71C5D6D" box="[1141,1392,1620,1644]" italics="true" pageId="17" pageNumber="168">Ceratozamia alvarezii</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="B6F24D42FFA49509FA5CF9DFD3695D91" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="mirandae">
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<emphasis id="4386EAD3FFA49509FA5CF9DFD7F95D6A" box="[1410,1429,1620,1643]" italics="true" pageId="17" pageNumber="168">C</emphasis>
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.
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<emphasis id="4386EAD3FFA49509FF49F9F3D3695D91" box="[151,261,1656,1680]" italics="true" pageId="17" pageNumber="168">mirandae</emphasis>
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</taxonomicName>
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<emphasis id="4386EAD3FFA49509FEDBF9F3D3FA5D8E" box="[261,406,1656,1680]" italics="true" pageId="17" pageNumber="168">
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,
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<taxonomicName id="B6F24D42FFA49509FECEF9F3D3FA5D8E" authorityName="D. W. Stevenson" authorityYear="1982" box="[272,406,1656,1679]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="norstogii">C. norstogii</taxonomicName>
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</emphasis>
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, and
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<taxonomicName id="B6F24D42FFA49509FE0DF9F3D0345D91" box="[467,600,1656,1680]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="vovidesii">
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<emphasis id="4386EAD3FFA49509FE0DF9F3D38A5D8E" box="[467,486,1656,1679]" italics="true" pageId="17" pageNumber="168">C</emphasis>
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.
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<emphasis id="4386EAD3FFA49509FE2AF9F3D0345D91" box="[500,600,1656,1680]" italics="true" pageId="17" pageNumber="168">vovidesii</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="39E8654AFFA4950AFF19F917D28F5CDD" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168" type="description">
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<paragraph id="714D36C1FFA49509FF19F917D6855C45" blockId="17.[151,1437,1511,2005]" pageId="17" pageNumber="168">
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<emphasis id="4386EAD3FFA49509FF19F917D38C5DB4" bold="true" box="[199,480,1692,1717]" pageId="17" pageNumber="168">Adult male description.</emphasis>
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Length
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<quantity id="B60A9B24FFA49509FDE0F917D08F5DB5" box="[574,739,1692,1717]" metricMagnitude="-3" metricUnit="m" metricValue="4.015" metricValueMax="4.49" metricValueMin="3.54" pageId="17" pageNumber="168" unit="mm" value="4.015" valueMax="4.49" valueMin="3.54">3.54–4.49 mm</quantity>
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, width
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<quantity id="B60A9B24FFA49509FCE8F917D1B65DB5" box="[822,986,1692,1717]" metricMagnitude="-3" metricUnit="m" metricValue="1.645" metricValueMax="1.79" metricValueMin="1.5" pageId="17" pageNumber="168" unit="mm" value="1.645" valueMax="1.79" valueMin="1.5">1.50–1.79 mm</quantity>
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(n = 20). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
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||
<figureCitation id="E9C92A44FFA49509FAD4F934D7345DD9" box="[1290,1368,1727,1752]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="17" pageNumber="168">Figs. 6</figureCitation>
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A–E), head and pronotum concolorous and on some slightly darker than the yellowish-brown elytra; dorsal surface punctate, shining but weakly alutaceous, short procumbent hairs associated with punctation, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
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</paragraph>
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<paragraph id="714D36C1FFA4950AFF19F8C4D3DB5AAC" blockId="17.[151,1437,1511,2005]" lastBlockId="18.[151,1437,151,2012]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
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<emphasis id="4386EAD3FFA49509FF19F8C4D36B5C69" bold="true" box="[199,263,1871,1896]" pageId="17" pageNumber="168">Head</emphasis>
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in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
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<figureCitation id="E9C92A44FFA49509FEC9F8FFD3CC5C8D" box="[279,416,1908,1933]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="17" pageNumber="168">Figs. 6A, D</figureCitation>
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), average distance between closest punctures 2× width of puncture; width
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||
<quantity id="B60A9B24FFA49509FB2FF8FFD7F45C8D" box="[1265,1432,1908,1933]" metricMagnitude="-4" metricUnit="m" metricValue="8.3" metricValueMax="9.1" metricValueMin="7.5" pageId="17" pageNumber="168" unit="mm" value="0.83" valueMax="0.91" valueMin="0.75">0.75–0.91 mm</quantity>
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; dorsal interocular distance
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<quantity id="B60A9B24FFA49509FE3DF813D0FE5CB1" box="[483,658,1944,1969]" metricMagnitude="-4" metricUnit="m" metricValue="5.449999999999999" metricValueMax="6.0" metricValueMin="4.9" pageId="17" pageNumber="168" unit="mm" value="0.5449999999999999" valueMax="0.6" valueMin="0.49">0.49–0.60 mm</quantity>
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, head width/dorsal interocular distance ratio 1.45–1.56, ventral interocular distance
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<quantity id="B60A9B24FFA49509FEA3F837D0485CD5" box="[381,548,1980,2005]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.5" metricValueMin="3.5999999999999996" pageId="17" pageNumber="168" unit="mm" value="0.405" valueMax="0.45" valueMin="0.36">0.36–0.45 mm</quantity>
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, head width/ventral interocular distance ratio 1.96–2.22. Eye with large black facets. Antennal length approximately equal to pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate, antennomere II subequal to III; IV–VIII small, width equals length; club fairly large, IX and X similar in length, XI slightly longer, globular with acuminate apex. Clypeus truncate anteriorly, moderately punctate, with narrow margin. Mentum and submentum with moderate punctation, distance between nearest punctures ~ 2× own width, each puncture with a long seta ~ 2–3× width of eye facet and reclining anteriad (often abraded). Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
|
||
<figureCitation id="E9C92A44FFA7950AFE8DFE1FD3C65AAC" box="[339,426,404,429]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Fig. 6E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFA7950AFF19FE3CD6B15FE9" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="169">
|
||
<emphasis id="4386EAD3FFA7950AFF19FE3CD34C5AD1" bold="true" box="[199,288,439,464]" pageId="18" pageNumber="169">Thorax</emphasis>
|
||
with pronotum weakly transverse in dorsal view with marginal beads basally and laterally, anterior margin with weak bead, reduced to absent medially; length/width ratio (PL/PW) = 0.76–0.83, rectangular; anterior and posterior angles present, anterior angles projecting; lateral carinae expanding in basal 1/10 to parallel sides, sometimes slightly emarginated, parallel for 50% of length, then gradually converging to anterior angles; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/5 length of posterior margin from posterior corners and touching and just anterior to marginal bead, pores mark base of longitudinal furrows onto disc. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae ~ 1/3 length of eye. Hypomeron laterally apparently lacking punctures, some with minute setae visible; medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.67–1.74, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size, scutellary striole extending ¼ elytral length, with 9–16 punctures; intervals of striae with shallow punctures similar to those of striae, obscuring striae. All punctures of elytral bearing a single fine, procumbent seta; seta often only visible in profile, extending posteriorly to, over or beyond next puncture (often abraded), interval puncture setae longer than strial puncture setae. Punctation on meso- and metaventrites moderate to dense, distance between nearest punctures 1–2× width of punctures, puncture depth shallow. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over 3/4 metaventrite length. Legs with procoxa oval; mesocoxa globular; metacoxa transversely elongateoval; trochanters obliquely truncate apically; in males all femora swollen, stout, dorso-ventral width greater than eye length; all tibiae lacking shallow medial subapical emargination, obliquely truncate at apex, with complete fringe of fine spinules on anterior and posterior margins; protibia swollen, not flattened in cross section notably larger than other tibiae, triangularly dilated to apex; mesotibia weakly swollen, moderately triangularly dilated to apex; metatibia narrowed, weakly dilating to midlength then nearly parallel to apex; pro- and mesotarsi greatly enlarged, corresponding tarsomere I width ~ 3× own length, metatarsomere I width ~ own length.
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFA7950AFF19FB7FD2A65C04" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="169">
|
||
<emphasis id="4386EAD3FFA7950AFF19FB7FD32F5E0C" bold="true" box="[199,323,1268,1293]" pageId="18" pageNumber="169">Abdomen.</emphasis>
|
||
Ventrite I with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites strongly alutaceous across entire surface, bearing indistinct punctures, distance to nearest puncture ~ 1× width of puncture, punctures bearing mostly reclining setae; ventrites I–V with setae length ~ 2–3× width of puncture; I–V each with 2 or more median pairs of longer, semi-erect sensory hairs (obscured with other setation), V with additional 4–5 pairs of long semi-erect hairs located postero-laterally, but not at submargin. Male genitalia with genital capsule fringed with fine setae; tegmen sclerotized, triangular, laterally compressed; anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly (
|
||
<figureCitation id="E9C92A44FFA7950AFBAEF9BCD6D15D51" box="[1136,1213,1591,1616]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Figs. 6</figureCitation>
|
||
G–H); apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately half its own width, in dorso-ventral view length/width ratio = 2.14–2.67; apically with setae, length of longest setae> width of paramere in dorso-ventral view (
|
||
<figureCitation id="E9C92A44FFA7950AFDA9F92FD0A45DBC" box="[631,712,1700,1725]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">
|
||
Fig.
|
||
<date id="054C1001FFA7950AFD70F92FD0A45DBC" box="[686,712,1700,1725]" pageId="18" pageNumber="169">6I</date>
|
||
</figureCitation>
|
||
); median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to a point. Ratio of penile strut length to median lobe length 1.93–2.19 (n = 2) (
|
||
<figureCitation id="E9C92A44FFA7950AFAB1F94CD2D15C04" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Fig. 6H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFA7950AFF19F884D28F5CDD" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="169">
|
||
<emphasis id="4386EAD3FFA7950AFF19F884D3725C29" bold="true" box="[199,286,1807,1832]" pageId="18" pageNumber="169">Female</emphasis>
|
||
generally similar to male, except for sexually dimorphisms. Female pronotum slightly more transverse than male, PL/PW = 0.75–0.81; elytra more strongly dulled and alutaceous laterally and over most of disc; legs unmodified, fore and middle legs not swollen, femora dorso-ventral width about equal to eye, all tibia gradually dilating to narrowly triangular apex, all legs with tarsomere I width ~ own length (
|
||
<figureCitation id="E9C92A44FFA7950AFBB3F8F7D6D35C94" box="[1133,1215,1916,1941]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Figs. 6</figureCitation>
|
||
A–C); metaventrite flattened medially, not impressed; abdomen strongly alutaceous, obscuring punctures, less densely punctate and setose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="258D6649FFA6950BFF49F808D2A25CF4" httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="19" pageNumber="170" startId="19.[151,250,1923,1945]" targetBox="[203,1389,193,1908]" targetPageId="19">
|
||
<paragraph id="714D36C1FFA6950BFF49F808D2A25CF4" blockId="19.[151,1437,1923,2037]" pageId="19" pageNumber="170">
|
||
<emphasis id="4386EAD3FFA6950BFF49F808D3785C98" bold="true" box="[151,276,1923,1945]" pageId="19" pageNumber="170">FIGURE 6.</emphasis>
|
||
Photographs of
|
||
<taxonomicName id="B6F24D42FFA6950BFE63F808D1705C98" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[445,796,1923,1945]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="19" pageNumber="170" phylum="Tracheophyta" rank="subGenus" subGenus="Uovidesa">
|
||
<emphasis id="4386EAD3FFA6950BFE63F808D0545C98" box="[445,568,1923,1945]" italics="true" pageId="19" pageNumber="170">Ceratophila</emphasis>
|
||
(
|
||
<emphasis id="4386EAD3FFA6950BFD96F808D0CE5C98" box="[584,674,1923,1945]" italics="true" pageId="19" pageNumber="170">Uovidesa</emphasis>
|
||
)
|
||
<emphasis id="4386EAD3FFA6950BFD6FF808D1705C98" box="[689,796,1923,1945]" italics="true" pageId="19" pageNumber="170">chiapensis</emphasis>
|
||
</taxonomicName>
|
||
collected from
|
||
<taxonomicName id="B6F24D42FFA6950BFC61F80FD6CF5C98" box="[959,1187,1923,1945]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="19" pageNumber="170" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||
<emphasis id="4386EAD3FFA6950BFC61F80FD6CF5C98" box="[959,1187,1923,1945]" italics="true" pageId="19" pageNumber="170">Ceratozamia vovidesii</emphasis>
|
||
</taxonomicName>
|
||
(A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="39E8654AFFA1950CFF19FF1CD7E959CD" pageId="20" pageNumber="171" type="materials_examined">
|
||
<paragraph id="714D36C1FFA1950CFF19FF1CD66F5BB1" blockId="20.[151,1437,151,896]" box="[199,1027,151,176]" pageId="20" pageNumber="171">
|
||
<materialsCitation id="C19A3C9CFFA1950CFF19FF1CD66F5BB1" ID-GBIF-Occurrence="2028138746" box="[199,1027,151,176]" country="Mexico" elevation="957" location="Jaltenango" pageId="20" pageNumber="171" specimenCount="957" stateProvince="Chiapas" typeStatus="holotype">
|
||
<emphasis id="4386EAD3FFA1950CFF19FF1CD3095BB1" bold="true" box="[199,357,151,176]" pageId="20" pageNumber="171">
|
||
<typeStatus id="AE498863FFA1950CFF19FF1CD36E5BB1" box="[199,258,151,176]" pageId="20" pageNumber="171">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="09E57651FFA1950CFEB2FF1CD3A85BB1" box="[364,452,151,176]" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingCountry>
|
||
,
|
||
<collectingRegion id="B336F823FFA1950CFE11FF1CD0405BB1" box="[463,556,151,176]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||
,
|
||
<location id="742D601AFFA1950CFDE6FF1CD0DF5BB1" LSID="urn:lsid:plazi:treatment:F95B87D7FFA4950CFF49FAF6D120587E:742D601AFFA1950CFDE6FF1CD0DF5BB1" box="[568,691,151,176]" country="Mexico" name="Jaltenango" pageId="20" pageNumber="171" stateProvince="Chiapas">Jaltenango</location>
|
||
, [GPS coord. omitted],
|
||
<specimenCount id="67F4FD48FFA1950CFC62FF1CD66F5BB1" box="[956,1027,151,176]" pageId="20" pageNumber="171" type="generic">
|
||
<elevation id="FADFD1F2FFA1950CFC62FF1CD1935BB1" box="[956,1023,151,176]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">
|
||
<quantity id="B60A9B24FFA1950CFC62FF1CD1935BB1" box="[956,1023,151,176]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">957m</quantity>
|
||
</elevation>
|
||
.
|
||
</specimenCount>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFA1950CFF19FF37D76A5BD5" blockId="20.[151,1437,151,896]" box="[199,1286,188,213]" pageId="20" pageNumber="171">
|
||
<emphasis id="4386EAD3FFA1950CFF19FF37D3765BD4" bold="true" box="[199,282,188,213]" pageId="20" pageNumber="171">Range.</emphasis>
|
||
Sierra Madre de
|
||
<collectingRegion id="B336F823FFA1950CFE03FF37D0565BD4" box="[477,570,188,213]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCountry id="09E57651FFA1950CFD9BFF37D0F15BD4" box="[581,669,188,213]" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingCountry>
|
||
, inhabiting male cones of four species of
|
||
<taxonomicName id="B6F24D42FFA1950CFBB1FF37D76C5BD2" box="[1135,1280,188,211]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4386EAD3FFA1950CFBB1FF37D76C5BD2" box="[1135,1280,188,211]" italics="true" pageId="20" pageNumber="171">Ceratozamia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="714D36C1FFA1950CFF19FF54D7E959CD" blockId="20.[151,1437,151,896]" pageId="20" pageNumber="171">
|
||
<emphasis id="4386EAD3FFA1950CFF19FF54D05B5BF9" bold="true" box="[199,567,223,248]" pageId="20" pageNumber="171">
|
||
Material examined.
|
||
<typeStatus id="AE498863FFA1950CFE14FF54D05B5BF9" box="[458,567,223,248]" pageId="20" pageNumber="171" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(by designation) male of
|
||
<taxonomicName id="B6F24D42FFA1950CFC59FF6BD6445BF9" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[903,1064,224,248]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="chiapensis">
|
||
<emphasis id="4386EAD3FFA1950CFC59FF6BD6445BF9" box="[903,1064,224,248]" italics="true" pageId="20" pageNumber="171">C. chiapensis</emphasis>
|
||
</taxonomicName>
|
||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||
<collectingCountry id="09E57651FFA1950CFD57FE8FD0915A1D" box="[649,765,260,284]" name="Mexico" pageId="20" pageNumber="171">MEXICO</collectingCountry>
|
||
,
|
||
<collectingRegion id="B336F823FFA1950CFCD3FE8FD1065A1C" box="[781,874,260,285]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||
, Jaltenango, [GPS coord. omitted],
|
||
<quantity id="B60A9B24FFA1950CFAD1FE8FD73D5A1C" box="[1295,1361,260,285]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">957m</quantity>
|
||
, ex
|
||
<specimenCount id="67F4FD48FFA1950CFA59FE8ED7F05A1D" box="[1415,1436,261,284]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
<taxonomicName id="B6F24D42FFA1950CFF49FEA3D3F85A41" box="[151,404,296,320]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||
<emphasis id="4386EAD3FFA1950CFF49FEA3D3F85A41" box="[151,404,296,320]" italics="true" pageId="20" pageNumber="171">Ceratozamia vovidesii</emphasis>
|
||
</taxonomicName>
|
||
cone,
|
||
<date id="054C1001FFA1950CFE01FEA3D0375A41" box="[479,603,296,320]" pageId="20" pageNumber="171" value="2017-05-21">
|
||
21-
|
||
<bibRefCitation id="15634B30FFA1950CFDDCFEA3D0375A41" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[514,603,296,320]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||
</date>
|
||
, H. Gomez Dominguez & W.Tang”; 2) [rectangular; red; printed in black ink]
|
||
<typeStatus id="AE498863FFA1950CFF0CFEC7D30A5A65" box="[210,358,332,356]" pageId="20" pageNumber="171" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="67F4FD48FFA1950CFEABFEC6D3E65A65" box="[373,394,333,356]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
<taxonomicName id="B6F24D42FFA1950CFE46FEC7D6045A65" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[408,1128,332,357]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4386EAD3FFA1950CFE46FEC7D0C85A65" box="[408,676,332,356]" italics="true" pageId="20" pageNumber="171">Ceratophila chiapensis</emphasis>
|
||
<bibRefCitation id="15634B30FFA1950CFD6CFEC6D6045A65" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[690,1128,332,357]" pageId="20" pageNumber="171" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
. Deposited in the FSCA.
|
||
<typeStatus id="AE498863FFA1950CFF49FEE4D2905A89" box="[151,252,367,392]" pageId="20" pageNumber="171" type="allotype">
|
||
<emphasis id="4386EAD3FFA1950CFF49FEE4D2905A89" bold="true" box="[151,252,367,392]" pageId="20" pageNumber="171">Allotype</emphasis>
|
||
</typeStatus>
|
||
(FSCA) same data as
|
||
<typeStatus id="AE498863FFA1950CFE2AFEE4D03A5A89" box="[500,598,367,392]" pageId="20" pageNumber="171" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="67F4FD48FFA1950CFD55FEE4D0985A89" box="[651,756,367,392]" pageId="20" pageNumber="171" type="adult">175 adult</specimenCount>
|
||
<typeStatus id="AE498863FFA1950CFD24FEE4D11C5A89" box="[762,880,367,392]" pageId="20" pageNumber="171" type="paratype">
|
||
<emphasis id="4386EAD3FFA1950CFD24FEE4D11C5A89" bold="true" box="[762,880,367,392]" pageId="20" pageNumber="171">paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<emphasis id="4386EAD3FFA1950CFCA0FEE4D60A5A89" bold="true" box="[894,1126,367,392]" pageId="20" pageNumber="171">
|
||
<collectingCountry id="09E57651FFA1950CFCA0FEE4D1945A89" box="[894,1016,367,392]" name="Mexico" pageId="20" pageNumber="171">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="B336F823FFA1950CFBDAFEE4D60A5A89" box="[1028,1126,367,392]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||
</emphasis>
|
||
, Ejido La Sombra, Mun. de Villa Flores,
|
||
<taxonomicName id="B6F24D42FFA1950CFEF3FE1FD05E5AAD" box="[301,562,404,428]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||
<emphasis id="4386EAD3FFA1950CFEF3FE1FD05E5AAD" box="[301,562,404,428]" italics="true" pageId="20" pageNumber="171">Ceratozamia mirandae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="054C1001FFA1950CFD9DFE1FD0C35AAD" box="[579,687,404,428]" pageId="20" pageNumber="171" value="1994-05-01">1-V-1994</date>
|
||
, M. A. Perez-Farrera (3); Mpio. Villa Flores,
|
||
<quantity id="B60A9B24FFA1950CFB1CFE1FD76D5AAC" box="[1218,1281,404,429]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="171" unit="km" value="2.0">2 km</quantity>
|
||
. Oeste Ejido: “La Sombra”,
|
||
<quantity id="B60A9B24FFA1950CFEE0FE3CD3ED5AD1" box="[318,385,439,464]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="171" unit="m" value="850.0">850m</quantity>
|
||
, oak forest on
|
||
<taxonomicName id="B6F24D42FFA1950CFDEEFE33D1435ACE" authorityName="D. W. Stevenson" authorityYear="1982" box="[560,815,440,463]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="norstogii">
|
||
<emphasis id="4386EAD3FFA1950CFDEEFE33D1435ACE" box="[560,815,440,463]" italics="true" pageId="20" pageNumber="171">Ceratozamia norstogii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="054C1001FFA1950CFC9EFE3CD1C15AD1" box="[832,941,439,464]" pageId="20" pageNumber="171" value="1995-05-01">1-V-1995</date>
|
||
, M. A. Perez, (4); Mpio. Cintalpa, Rancho Cafetal,
|
||
<taxonomicName id="B6F24D42FFA1950CFF22FE57D3975AF5" box="[252,507,476,500]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="alvaresii">
|
||
<emphasis id="4386EAD3FFA1950CFF22FE57D3975AF5" box="[252,507,476,500]" italics="true" pageId="20" pageNumber="171">Ceratozamia alvaresii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="054C1001FFA1950CFDD1FE57D0EA5AF4" box="[527,646,476,501]" pageId="20" pageNumber="171" value="1996-03-04">4-III-1996</date>
|
||
, A. Vovides, 1A, (1); Mpo. Cintalapa, Ejido Flor de
|
||
<collectingRegion id="B336F823FFA1950CFACDFE57D71C5AF4" box="[1299,1392,476,501]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||
, ex
|
||
<taxonomicName id="B6F24D42FFA1950CFF49FD8BD3F05916" authorityName="D. W. Stevenson" authorityYear="1982" box="[151,412,512,535]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="norstogii">
|
||
<emphasis id="4386EAD3FFA1950CFF49FD8BD3F05916" box="[151,412,512,535]" italics="true" pageId="20" pageNumber="171">Ceratozamia norstogii</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="67F4FD48FFA1950CFE72FD8BD3AD5916" box="[428,449,512,535]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
cone;
|
||
<date id="054C1001FFA1950CFDC3FE74D0FD5919" box="[541,657,511,536]" pageId="20" pageNumber="171" value="2010-02">Feb 2010</date>
|
||
, M.A. Perez Farrera (1); Mpo. Villa Flores, Ejido La Sombra,
|
||
<taxonomicName id="B6F24D42FFA1950CFF49FDAFD3C9593D" box="[151,421,548,572]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||
<emphasis id="4386EAD3FFA1950CFF49FDAFD3C9593D" box="[151,421,548,572]" italics="true" pageId="20" pageNumber="171">Ceratozamia mirandae</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="67F4FD48FFA1950CFE69FDAED3A0593D" box="[439,460,549,572]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
cone,
|
||
<date id="054C1001FFA1950CFDF2FDAFD0E1593D" box="[556,653,548,572]" pageId="20" pageNumber="171" value="2010-11">XI-2010</date>
|
||
, M.A. Perez Farrera (25); Champerico, [GPS coord. omitted],
|
||
<taxonomicName id="B6F24D42FFA1950CFF49FDC3D3F35961" box="[151,415,584,608]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||
<emphasis id="4386EAD3FFA1950CFF49FDC3D3F35961" box="[151,415,584,608]" italics="true" pageId="20" pageNumber="171">Ceratozamia mirandae</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="67F4FD48FFA1950CFE72FDC3D3AD595E" box="[428,449,584,607]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
cone,
|
||
<date id="054C1001FFA1950CFDC8FDC3D0FE5961" box="[534,658,584,608]" pageId="20" pageNumber="171" value="2017-05-20">
|
||
20-
|
||
<bibRefCitation id="15634B30FFA1950CFDE4FDC3D0FE5961" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[570,658,584,608]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||
</date>
|
||
, H. Gomez Dominguez & W.Tang (46); Jaltenango, [GPS coord. omitted],
|
||
<quantity id="B60A9B24FFA1950CFED5FDE7D3225984" box="[267,334,620,645]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">957m</quantity>
|
||
, ex
|
||
<specimenCount id="67F4FD48FFA1950CFE57FDE6D3F25985" box="[393,414,621,644]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
<taxonomicName id="B6F24D42FFA1950CFE73FDE7D0DC5985" box="[429,688,620,644]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||
<emphasis id="4386EAD3FFA1950CFE73FDE7D0DC5985" box="[429,688,620,644]" italics="true" pageId="20" pageNumber="171">Ceratozamia vovidesii</emphasis>
|
||
</taxonomicName>
|
||
cone,
|
||
<date id="054C1001FFA1950CFCD6FDE7D1EF5985" box="[776,899,620,644]" pageId="20" pageNumber="171" value="2017-05-21">
|
||
21-
|
||
<bibRefCitation id="15634B30FFA1950CFCF4FDE7D1EF5985" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[810,899,620,644]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||
</date>
|
||
, H. Gomez Dominguez & W.Tang (71); La Concordia, Rancho Los Pinos, ex
|
||
<specimenCount id="67F4FD48FFA1950CFDF8FD1BD05759A6" box="[550,571,656,679]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||
<taxonomicName id="B6F24D42FFA1950CFD9BFD1BD12859A9" box="[581,836,656,680]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||
<emphasis id="4386EAD3FFA1950CFD9BFD1BD12859A9" box="[581,836,656,680]" italics="true" pageId="20" pageNumber="171">Ceratozamia vovidesii</emphasis>
|
||
</taxonomicName>
|
||
cone,
|
||
<date id="054C1001FFA1950CFC4AFD1BD66159A9" box="[916,1037,656,680]" pageId="20" pageNumber="171" value="2017-06-01">
|
||
1-
|
||
<bibRefCitation id="15634B30FFA1950CFC7BFD1BD66159A9" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[933,1037,656,680]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">VI-2017</bibRefCitation>
|
||
</date>
|
||
, M.A. Pérez Farrera & H. Gómez Domínguez (22).
|
||
<typeStatus id="AE498863FFA1950CFE83FD3ED3A659CC" box="[349,458,693,717]" pageId="20" pageNumber="171" type="paratype">Paratypes</typeStatus>
|
||
deposited at ANIC, BMNH, FSCA, IEXA, INBio, MNHN, NZAC, UNAM, USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39E8654AFFA1950CFF19FD5CD63659F1" box="[199,1114,727,752]" pageId="20" pageNumber="171" type="etymology">
|
||
<paragraph id="714D36C1FFA1950CFF19FD5CD63659F1" blockId="20.[151,1437,151,896]" box="[199,1114,727,752]" pageId="20" pageNumber="171">
|
||
<emphasis id="4386EAD3FFA1950CFF19FD5CD32259F1" bold="true" box="[199,334,727,752]" pageId="20" pageNumber="171">Etymology.</emphasis>
|
||
This species is named for the Mexican state where it was discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39E8654AFFA1950CFF19FD77D120587E" pageId="20" pageNumber="171" type="discussion">
|
||
<paragraph id="714D36C1FFA1950CFF19FD77D120587E" blockId="20.[151,1437,151,896]" pageId="20" pageNumber="171">
|
||
<emphasis id="4386EAD3FFA1950CFF19FD77D3575814" bold="true" box="[199,315,764,789]" pageId="20" pageNumber="171">Remarks.</emphasis>
|
||
This species coinhabits male cones of at least four species of
|
||
<taxonomicName id="B6F24D42FFA1950CFBE0FD77D6A35812" box="[1086,1231,764,787]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4386EAD3FFA1950CFBE0FD77D6A35812" box="[1086,1231,764,787]" italics="true" pageId="20" pageNumber="171">Ceratozamia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="B6F24D42FFA1950CFB3BFD77D7025815" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[1253,1390,764,788]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||
<emphasis id="4386EAD3FFA1950CFB3BFD77D6945812" box="[1253,1272,764,787]" italics="true" pageId="20" pageNumber="171">C</emphasis>
|
||
.
|
||
<emphasis id="4386EAD3FFA1950CFAD3FD77D7025815" box="[1293,1390,764,788]" italics="true" pageId="20" pageNumber="171">alvarezii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B6F24D42FFA1950CFA5CFD77D3695839" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||
<emphasis id="4386EAD3FFA1950CFA5CFD77D7F95812" box="[1410,1429,764,787]" italics="true" pageId="20" pageNumber="171">C</emphasis>
|
||
.
|
||
<emphasis id="4386EAD3FFA1950CFF49FCABD3695839" box="[151,261,800,824]" italics="true" pageId="20" pageNumber="171">mirandae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4386EAD3FFA1950CFEDBFCABD3D95836" box="[261,437,800,824]" italics="true" pageId="20" pageNumber="171">
|
||
,
|
||
<taxonomicName id="B6F24D42FFA1950CFEC3FCABD3DE5836" authorityName="D. W. Stevenson" authorityYear="1982" box="[285,434,800,823]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="norstogii">C. norstogii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="B6F24D42FFA1950CFE17FCABD0355839" box="[457,601,800,824]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||
<emphasis id="4386EAD3FFA1950CFE17FCABD3B05836" box="[457,476,800,823]" italics="true" pageId="20" pageNumber="171">C</emphasis>
|
||
.
|
||
<emphasis id="4386EAD3FFA1950CFE28FCABD0355839" box="[502,601,800,824]" italics="true" pageId="20" pageNumber="171">vovidesii</emphasis>
|
||
</taxonomicName>
|
||
) with
|
||
<taxonomicName id="B6F24D42FFA1950CFD65FCABD6655836" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[699,1033,800,824]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4386EAD3FFA1950CFD65FCABD12F5839" box="[699,835,800,824]" italics="true" pageId="20" pageNumber="171">Ceratophila</emphasis>
|
||
(
|
||
<emphasis id="4386EAD3FFA1950CFC81FCABD1155836" box="[863,889,800,823]" italics="true" pageId="20" pageNumber="171">C.</emphasis>
|
||
)
|
||
<emphasis id="4386EAD3FFA1950CFC48FCABD6655836" box="[918,1033,800,823]" italics="true" pageId="20" pageNumber="171">picipennis</emphasis>
|
||
</taxonomicName>
|
||
and an undescribed species of
|
||
<taxonomicName id="B6F24D42FFA1950CFF49FCCFD35B585D" authorityName="Reitter" authorityYear="1875" box="[151,311,836,860]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4386EAD3FFA1950CFF49FCCFD35B585D" box="[151,311,836,860]" italics="true" pageId="20" pageNumber="171">Pharaxonotha</emphasis>
|
||
</taxonomicName>
|
||
. In nine collections from four host species listed above,
|
||
<taxonomicName id="B6F24D42FFA1950CFBD6FCCFD7F0585D" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1032,1436,836,860]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||
<emphasis id="4386EAD3FFA1950CFBD6FCCFD6FC585D" box="[1032,1168,836,860]" italics="true" pageId="20" pageNumber="171">Ceratophila</emphasis>
|
||
(
|
||
<emphasis id="4386EAD3FFA1950CFB77FCCFD766585D" box="[1193,1290,836,860]" italics="true" pageId="20" pageNumber="171">Vovidesa</emphasis>
|
||
)
|
||
<emphasis id="4386EAD3FFA1950CFAF8FCCFD7F0585D" box="[1318,1436,836,860]" italics="true" pageId="20" pageNumber="171">chiapensis</emphasis>
|
||
</taxonomicName>
|
||
accounted for 6% of the adult pharaxonothine beetles present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |