198 lines
28 KiB
XML
198 lines
28 KiB
XML
<document id="9FCE27025D1BC7275A574C0066571283" ID-DOI="10.11646/zootaxa.4105.2.1" ID-GBIF-Dataset="12e8319b-8b19-46db-a34a-764f91863bf8" ID-ISSN="1175-5326" ID-Zenodo-Dep="256481" ID-ZooBank="A4B3B85B-1E19-40C5-9AD5-C416FE0E24E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461134115519" checkinUser="plazi" docAuthor="Hara, Marcos Ryotaro" docDate="2016" docId="B409D267FFF4FFE2FF0FFE04FC0DFE77" docLanguage="en" docName="zootaxa.4105.2.1.pdf.imf" docOrigin="Zootaxa 4105 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nanophareus bicornutus Hara, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="114" masterDocId="4830AA1FFFFEFFEFFF98FF80FFFFFF82" masterDocTitle="Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae)" masterLastPageNumber="123" masterPageNumber="101" pageNumber="111" updateTime="1698598291331" updateUser="plazi">
|
||
<mods:mods id="BFC9AD5F6A1A593ACBE256A415EEA9DE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3AC94C599FFC91D1F72C5B5E20B5C7A1">
|
||
<mods:title id="3BEC9165FABBAA93648DD7DBBB8AA4C6">Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1EE49CAA09FD50A5F5DEC5A7581A9D7C" type="personal">
|
||
<mods:role id="F4B16EC0688F0ECB51C91DDCA78F15D6">
|
||
<mods:roleTerm id="C56744BC8725B1CBF7949682C22F40D2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8FBA06EAB41647421CC0422EFF4FFB01">Hara, Marcos Ryotaro</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="B22738E007076B6694F488A05A2B0262">text</mods:typeOfResource>
|
||
<mods:relatedItem id="3CFB4BC5FCCBF658CC0E455B7986B54D" type="host">
|
||
<mods:titleInfo id="A78BADED1CE4BE024955B77F91202A77">
|
||
<mods:title id="865B7531F76E0DB84A2654675D6EE461">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2B0A0BA0275EA8FEC1987CCA6B1A9AC8">
|
||
<mods:date id="8A47D876CA61AB3171EB8A62B53094E4">2016</mods:date>
|
||
<mods:detail id="5F1879630CC15A05E3C3AA5E6616D78F" type="volume">
|
||
<mods:number id="463FC2A3FD1C0D04B6827F1D309DFA99">4105</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="778B6673D8DDA084244491F647CCD0ED" type="issue">
|
||
<mods:number id="D651D6C867B186C49B4F340B75432F45">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4D70AE73C9786E24FDF18B772F47058F" unit="page">
|
||
<mods:start id="CE89B10F4F669FAD7130303ED8D6AF51">101</mods:start>
|
||
<mods:end id="D75A3B559650AD4CA1B6EAE3875CD793">123</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="135BCFC61D63C38A0DE5DC89C2E68D38">journal article</mods:classification>
|
||
<mods:identifier id="8B1D45B701B5F6A032FB96F1B6E82CB9" type="DOI">10.11646/zootaxa.4105.2.1</mods:identifier>
|
||
<mods:identifier id="D5F703425C752F1C72AF70CF32F294B5" type="GBIF-Dataset">12e8319b-8b19-46db-a34a-764f91863bf8</mods:identifier>
|
||
<mods:identifier id="EE069497FEB1EE9025B9B143ABEFB2CF" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="999AE5AEEAE200FD9721036927C9D6B5" type="Zenodo-Dep">256481</mods:identifier>
|
||
<mods:identifier id="D87E5D70F9CB53DE934D01499E876169" type="ZooBank">A4B3B85B-1E19-40C5-9AD5-C416FE0E24E9</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="B409D267FFF4FFE2FF0FFE04FC0DFE77" ID-DOI="http://doi.org/10.5281/zenodo.5662466" ID-GBIF-Taxon="119627572" ID-Zenodo-Dep="5662466" LSID="urn:lsid:plazi:treatment:B409D267FFF4FFE2FF0FFE04FC0DFE77" httpUri="http://treatment.plazi.org/id/B409D267FFF4FFE2FF0FFE04FC0DFE77" lastPageId="13" lastPageNumber="114" pageId="10" pageNumber="111">
|
||
<subSubSection id="74BA30FAFFF4FFE5FF0FFE04FE87FE42" pageId="10" pageNumber="111" type="nomenclature">
|
||
<paragraph id="3C1F6371FFF4FFE5FF0FFE04FDD5FE1D" blockId="10.[151,554,388,448]" box="[151,554,388,415]" pageId="10" pageNumber="111">
|
||
<heading id="6757D41DFFF4FFE5FF0FFE04FDD5FE1D" bold="true" box="[151,554,388,415]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
|
||
<emphasis id="0ED4BF63FFF4FFE5FF0FFE04FDD5FE1D" bold="true" box="[151,554,388,415]" pageId="10" pageNumber="111">
|
||
<taxonomicName id="FBA018F2FFF4FFE5FF0FFE04FE3AFE1C" ID-CoL="7XDTL" box="[151,453,388,414]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="bicornutus" status="sp. nov.">
|
||
<emphasis id="0ED4BF63FFF4FFE5FF0FFE04FE3AFE1C" bold="true" box="[151,453,388,414]" italics="true" pageId="10" pageNumber="111">Nanophareus bicornutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="15E70218FFF4FFE5FE54FE05FDD5FE1D" box="[460,554,389,415]" pageId="10" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF4FFE5FF0FFE27FE87FE42" blockId="10.[151,554,388,448]" box="[151,376,423,448]" pageId="10" pageNumber="111">
|
||
(
|
||
<figureCitation id="A49B7FF4FFF4FFE5FF07FE27FF18FE42" box="[159,231,423,448]" captionStart="FIGURE 2" captionStartId="11.[151,250,1856,1878]" captionTargetBox="[206,1381,198,1835]" captionTargetId="figure@11.[200,1386,193,1843]" captionTargetPageId="11" captionText="FIGURE 2. Nanophareus bicornutus sp. nov. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting tubercles of regular size); C, ocularium, anterior view; D, habitus, right lateral view; E, right tarsus IV, prolateral view; F, right pedipalp, prolateral view; G, ditto, retrolateral view. Abbreviations: A 1 – 4, scutal areas I – IV; Ca, carapace; Fh, frontal hump; Ft 1 – 3, free tergites I – III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum. A, D at same scale; F, G at same scale. Scale bars of A, B, D, F, G: 1 mm. Scale bars of C, E: 0.5 mm." httpUri="https://zenodo.org/record/256483/files/figure.png" pageId="10" pageNumber="111">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="A49B7FF4FFF4FFE5FF6BFE28FEFCFE42" box="[243,259,424,448]" captionStart="FIGURE 3" captionStartId="12.[151,250,1867,1889]" captionTargetBox="[159,1428,208,1829]" captionTargetId="figure@12.[156,1430,193,1845]" captionTargetPageId="12" captionText="FIGURE 3. Nanophareus bicornutus sp. nov. Male (holotype) right trochanter-metatarsus IV: A, dorsal view; B, prolateral view; C, ventral view; D, retrolateral view. Abbreviations: Pro, prolateral; Ret, retrolateral. A – D at same scale. Scale bars: 1 mm." httpUri="https://zenodo.org/record/256484/files/figure.png" pageId="10" pageNumber="111">3</figureCitation>
|
||
,
|
||
<figureCitation id="A49B7FF4FFF4FFE5FE97FE28FEE3FE42" box="[271,284,424,448]" captionStart="FIGURE 8" captionStartId="19.[151,250,1858,1880]" captionTargetBox="[218,1257,229,1784]" captionTargetId="figure@19.[169,1335,177,1817]" captionTargetPageId="19" captionText="FIGURE 8. Distal part of Nanophareus spp. penis: A, N. bicornutus sp. nov., dorsal view, B, ditto, right lateral view; C, N. maipu sp. nov., in dorsal view, D, ditto, right lateral view, E, ditto, ventral view; F, N. polyhastatus sp. nov., dorsal view, G, ditto, right lateral view. Abbreviations: bms, basal group of macrosetae (MS); Gl, glans; MS C, MS of group C; MS D, MS of group D; MS E, MS of group E; St, stylus of glans; Vpl, ventral plate; Vpr, ventral process of glans; Vt, ventral trichomes. A, B at same scale; C – E at same scale, F, G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256489/files/figure.png" pageId="10" pageNumber="111">8</figureCitation>
|
||
A–B, 9)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74BA30FAFFF4FFE5FF0FFE6FFCA5FDAF" pageId="10" pageNumber="111" type="materials_examined">
|
||
<paragraph id="3C1F6371FFF4FFE5FF0FFE6FFCA5FDAF" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
<emphasis id="0ED4BF63FFF4FFE5FF0FFE6FFEBBFD8A" bold="true" box="[151,324,495,520]" pageId="10" pageNumber="111">
|
||
<typeStatus id="E31BDDD3FFF4FFE5FF0FFE6FFF2DFD8A" box="[151,210,495,520]" pageId="10" pageNumber="111">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<collectingCountry id="44B723E1FFF4FFE5FED3FE70FE5EFD8A" box="[331,417,496,520]" name="Chile" pageId="10" pageNumber="111">CHILE</collectingCountry>
|
||
. V Región de Valparaíso: Zapallar (valle de Aconcagua),
|
||
<date id="481E45B1FFF4FFE5FBBBFE6FFB62FD8A" box="[1059,1181,495,520]" pageId="10" pageNumber="111" value="1950-11-27">27.xi.1950</date>
|
||
, Ross & Michelbacher
|
||
<emphasis id="0ED4BF63FFF4FFE5FF0FFD94FF46FDAE" box="[151,185,532,556]" italics="true" pageId="10" pageNumber="111">leg</emphasis>
|
||
., ma
|
||
<typeStatus id="E31BDDD3FFF4FFE5FF6FFD94FEA6FDAF" box="[247,345,532,557]" pageId="10" pageNumber="111" type="holotype">holotype</typeStatus>
|
||
(CASENT, originally in CASENT 9055060).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74BA30FAFFF4FFE5FF5FFDB7FC35FCCF" pageId="10" pageNumber="111" type="diagnosis">
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FFDB7FC35FCCF" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
<emphasis id="0ED4BF63FFF4FFE5FF5FFDB7FE48FDD2" bold="true" box="[199,439,567,592]" pageId="10" pageNumber="111">Diagnosis for males.</emphasis>
|
||
<taxonomicName id="FBA018F2FFF4FFE5FE25FDB8FD2EFDD2" box="[445,721,568,592]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="bicornutus" status="sp. nov.">
|
||
<emphasis id="0ED4BF63FFF4FFE5FE25FDB8FD2EFDD2" box="[445,721,568,592]" italics="true" pageId="10" pageNumber="111">Nanophareus bicornutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="15E70218FFF4FFE5FD41FDB7FCCCFDD2" box="[729,819,567,592]" pageId="10" pageNumber="111" rank="species">
|
||
<emphasis id="0ED4BF63FFF4FFE5FD41FDB7FCCCFDD2" bold="true" box="[729,819,567,592]" pageId="10" pageNumber="111">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
resembles
|
||
<taxonomicName id="FBA018F2FFF4FFE5FC28FDB9FB92FDD2" box="[944,1133,568,592]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="bosqenublado">
|
||
<emphasis id="0ED4BF63FFF4FFE5FC28FDB9FC3CFDD2" box="[944,963,569,592]" italics="true" pageId="10" pageNumber="111">N</emphasis>
|
||
.
|
||
<emphasis id="0ED4BF63FFF4FFE5FC48FDB8FB92FDD2" box="[976,1133,568,592]" italics="true" pageId="10" pageNumber="111">bosqenublado</emphasis>
|
||
</taxonomicName>
|
||
by the long coxa IV (close to posterior margin of dorsal scutum in dorsal view), coxa IV with prolateral apical apophysis with subbasal region bifid, trochanter IV with prodorsal apical large apophysis and femur IV with a retromedian spine. Can be distinguished from
|
||
<taxonomicName id="FBA018F2FFF4FFE5FEE1FD25FDC4FD3E" box="[377,571,676,700]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="bosqenublado">
|
||
<emphasis id="0ED4BF63FFF4FFE5FEE1FD25FE73FD3E" box="[377,396,677,700]" italics="true" pageId="10" pageNumber="111">N</emphasis>
|
||
.
|
||
<emphasis id="0ED4BF63FFF4FFE5FE06FD24FDC4FD3E" box="[414,571,676,700]" italics="true" pageId="10" pageNumber="111">bosqenublado</emphasis>
|
||
</taxonomicName>
|
||
by: unarmed frontal hump on anterior margin of dorsal scutum, low and narrow ocularium, and a retroapical conical long apophysis on trochanter IV. Can be distinguished from the other species of the genus by: low and narrow (less than ⅓ carapace width) ocularium, scutal area III with a pair of enlarged tubercles, long cheliceral segment I, pedipalpal coxa enlarged, long (length close to carapace length), and basally inflated, and trochanter IV with two dorsoapical large apophyses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74BA30FAFFF4FFE5FF5FFCD7FDC2FC16" pageId="10" pageNumber="111" type="etymology">
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FFCD7FDC2FC16" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
<emphasis id="0ED4BF63FFF4FFE5FF5FFCD7FEB1FCF2" bold="true" box="[199,334,855,880]" pageId="10" pageNumber="111">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="0ED4BF63FFF4FFE5FD90FCD8FDE1FCF2" box="[520,542,856,880]" italics="true" pageId="10" pageNumber="111">bi</emphasis>
|
||
, meaning two, and
|
||
<emphasis id="0ED4BF63FFF4FFE5FD67FCD9FCB4FCF2" box="[767,843,857,880]" italics="true" pageId="10" pageNumber="111">cornus</emphasis>
|
||
, meaning horns. In reference to the two dorsoapical horn-like apophyses on trochanter IV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74BA30FAFFF4FFE2FF5FFC1FFE60FE2F" lastPageId="13" lastPageNumber="114" pageId="10" pageNumber="111" type="description">
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FFC1FFBB0FB12" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
<emphasis id="0ED4BF63FFF4FFE5FF5FFC1FFEA6FC3A" bold="true" box="[199,345,927,952]" pageId="10" pageNumber="111">Description.</emphasis>
|
||
<emphasis id="0ED4BF63FFF4FFE5FEF8FC20FE66FC3A" box="[352,409,928,952]" italics="true" pageId="10" pageNumber="111">Male</emphasis>
|
||
(
|
||
<typeStatus id="E31BDDD3FFF4FFE5FE32FC1FFDECFC3A" box="[426,531,927,952]" pageId="10" pageNumber="111" type="holotype">holotype</typeStatus>
|
||
): Dorsum (
|
||
<figureCitation id="A49B7FF4FFF4FFE5FD0AFC1FFD2AFC3A" box="[658,725,927,952]" captionStart="FIGURE 2" captionStartId="11.[151,250,1856,1878]" captionTargetBox="[206,1381,198,1835]" captionTargetId="figure@11.[200,1386,193,1843]" captionTargetPageId="11" captionText="FIGURE 2. Nanophareus bicornutus sp. nov. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting tubercles of regular size); C, ocularium, anterior view; D, habitus, right lateral view; E, right tarsus IV, prolateral view; F, right pedipalp, prolateral view; G, ditto, retrolateral view. Abbreviations: A 1 – 4, scutal areas I – IV; Ca, carapace; Fh, frontal hump; Ft 1 – 3, free tergites I – III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum. A, D at same scale; F, G at same scale. Scale bars of A, B, D, F, G: 1 mm. Scale bars of C, E: 0.5 mm." httpUri="https://zenodo.org/record/256483/files/figure.png" pageId="10" pageNumber="111">Fig. 2</figureCitation>
|
||
A, C, D): Measurements: CL 1.90; CW 2.60; DSL 5.05; DSW 4.80; LI 10.15; LII 19.55; LIII 14.60; LIV 20.20. Anterior margin of carapace with unarmed, granulate frontal hump. Ocularium low, narrowed (less than ⅓ carapace width), smooth and unarmed. Carapace with sparse tubercles. Scutal areas I–IV with scattered tubercles; scutal area III with 1 pair of enlarged tubercles; scutal area IV undivided. Body shape
|
||
<typeStatus id="E31BDDD3FFF4FFE5FE30FBB1FE27FBCB" box="[424,472,1073,1097]" pageId="10" pageNumber="111">type</typeStatus>
|
||
gamma. Lateral margin of dorsal scutum with a row of enlarged tubercles inserted amongst small ones, more densely distributed between scutal grooves II and III. Posterior margin of dorsal scutum and free tergites I–III, each with a row of tubercles plus few tubercles on the corners.
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FFB1DFC94FB5A" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
Venter (
|
||
<figureCitation id="A49B7FF4FFF4FFE5FEB9FB1CFE97FB36" box="[289,360,1180,1205]" captionStart="FIGURE 2" captionStartId="11.[151,250,1856,1878]" captionTargetBox="[206,1381,198,1835]" captionTargetId="figure@11.[200,1386,193,1843]" captionTargetPageId="11" captionText="FIGURE 2. Nanophareus bicornutus sp. nov. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting tubercles of regular size); C, ocularium, anterior view; D, habitus, right lateral view; E, right tarsus IV, prolateral view; F, right pedipalp, prolateral view; G, ditto, retrolateral view. Abbreviations: A 1 – 4, scutal areas I – IV; Ca, carapace; Fh, frontal hump; Ft 1 – 3, free tergites I – III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum. A, D at same scale; F, G at same scale. Scale bars of A, B, D, F, G: 1 mm. Scale bars of C, E: 0.5 mm." httpUri="https://zenodo.org/record/256483/files/figure.png" pageId="10" pageNumber="111">Fig. 2</figureCitation>
|
||
B): Coxae I–IV and stigmatic area tuberculate; coxa I with enlarged tubercles in median rows and on apical region. Free sternites each with 1 row of tubercles.
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FFB64FEF9FAA2" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
Chelicerae: Segment I elongated, with
|
||
<emphasis id="0ED4BF63FFF4FFE5FD1EFB64FD3FFB7E" box="[646,704,1252,1276]" italics="true" pageId="10" pageNumber="111">bulla</emphasis>
|
||
slightly marked; fixed finger with 5 teeth; movable finger with 5–7 teeth.
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FFAACFCF3FA7A" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
Pedipalps (
|
||
<figureCitation id="A49B7FF4FFF4FFE5FEDBFAACFE79FAC6" box="[323,390,1324,1349]" captionStart="FIGURE 2" captionStartId="11.[151,250,1856,1878]" captionTargetBox="[206,1381,198,1835]" captionTargetId="figure@11.[200,1386,193,1843]" captionTargetPageId="11" captionText="FIGURE 2. Nanophareus bicornutus sp. nov. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting tubercles of regular size); C, ocularium, anterior view; D, habitus, right lateral view; E, right tarsus IV, prolateral view; F, right pedipalp, prolateral view; G, ditto, retrolateral view. Abbreviations: A 1 – 4, scutal areas I – IV; Ca, carapace; Fh, frontal hump; Ft 1 – 3, free tergites I – III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum. A, D at same scale; F, G at same scale. Scale bars of A, B, D, F, G: 1 mm. Scale bars of C, E: 0.5 mm." httpUri="https://zenodo.org/record/256483/files/figure.png" pageId="10" pageNumber="111">Fig. 2</figureCitation>
|
||
F, G): Coxa elongated (close to carapace length) and robust, basally inflated, with retroapical cluster of tubercles. Trochanter inflated and smooth, with 2–3 ventral setiferous tubercles (prolateral largest). Femur with sparse dorsal tubercles, without prolateral subapical seta, ventral side with 3–4 tubercles and 1 enlarged basal one. Patella obliquely inserted in tibia (see in lateral view). Tibia and tarsus with few scattered setiferous tubercles; tibia baso-ventrally oblique (seen in lateral view). Tibial setation: prolateral IIi, retrolateral I[Ii]i. Tarsal setation: prolateral iIii, iIiii, retrolateral iIii.
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF4FFE5FF5FF985FAC3F873" blockId="10.[151,1437,495,2033]" pageId="10" pageNumber="111">
|
||
Legs (
|
||
<figureCitation id="A49B7FF4FFF4FFE5FE88F984FEA4F99E" box="[272,347,1540,1565]" captionStart="FIGURE 2" captionStartId="11.[151,250,1856,1878]" captionTargetBox="[206,1381,198,1835]" captionTargetId="figure@11.[200,1386,193,1843]" captionTargetPageId="11" captionText="FIGURE 2. Nanophareus bicornutus sp. nov. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting tubercles of regular size); C, ocularium, anterior view; D, habitus, right lateral view; E, right tarsus IV, prolateral view; F, right pedipalp, prolateral view; G, ditto, retrolateral view. Abbreviations: A 1 – 4, scutal areas I – IV; Ca, carapace; Fh, frontal hump; Ft 1 – 3, free tergites I – III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum. A, D at same scale; F, G at same scale. Scale bars of A, B, D, F, G: 1 mm. Scale bars of C, E: 0.5 mm." httpUri="https://zenodo.org/record/256483/files/figure.png" pageId="10" pageNumber="111">Figs 2</figureCitation>
|
||
E, 3): Coxa II with 1 prolateral apophysis; coxa IV with 1 prolateral apical apophysis, dorsoventrally bifid (dorsal branch longer than ventral one) from subbasal region, directed backwards and 1 retrolateral, conical, short apical apophysis. Trochanter IV longer than wide, prolateral side with 1 conical, short, blunt submedian apophysis, its apex slightly curved dorsally, and 1 long, robust dorsoapical apophysis, this one bifid (prolateral branch longest), projected dorsally, its apex curved anteriorly; retrolateral side with a cluster of enlarged basal tubercles, 1 long, robust dorsoapical apophysis, this one curved dorsally, and 1 conical, short apical apophysis. Femora I–II unarmed; femur III with 2 ventral rows of tubercles increasing in size apically. Femur IV straight, with enlarged tubercles in the median regions of prolateral and retrodorsal rows of tubercles; retrolateral side with 1 basal, small, pointed spine, 1 median large spine, ventral side with an irregular retrolateral row of enlarged tubercles, and a prolateral row with 3–4 high, pointed tubercles; ventroapical side with 1 prolateral spine and 1 enlarged retrolateral tubercle. Patellae I–IV tuberculate, patellae I–III unarmed; patella IV with ventral enlarged, pointed tubercles. Tibiae I–II unarmed; tibia III with 2 ventral rows of tubercles slightly increasing in size apically. Tibia IV with 2 ventral rows of tubercles increasing in size subapically, retrolateral one with enlarged tubercles becoming high and pointed; ventroapical side unarmed. Tarsal segmentation: 6(3), 9–10(3), 6, 6.
|
||
</paragraph>
|
||
<caption id="68DF33F9FFF5FFE4FF0FF8C0FEBEF86D" httpUri="https://zenodo.org/record/256483/files/figure.png" pageId="11" pageNumber="112" targetBox="[206,1381,198,1835]" targetPageId="11">
|
||
<paragraph id="3C1F6371FFF5FFE4FF0FF8C0FEBEF86D" blockId="11.[151,1436,1856,2031]" pageId="11" pageNumber="112">
|
||
<emphasis id="0ED4BF63FFF5FFE4FF0FF8C0FEE9F8D4" bold="true" box="[151,278,1856,1878]" pageId="11" pageNumber="112">FIGURE 2.</emphasis>
|
||
<taxonomicName id="FBA018F2FFF5FFE4FEB8F8C0FDE2F8D4" box="[288,541,1856,1878]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="bicornutus" status="sp. nov.">
|
||
<emphasis id="0ED4BF63FFF5FFE4FEB8F8C0FDE2F8D4" box="[288,541,1856,1878]" italics="true" pageId="11" pageNumber="112">Nanophareus bicornutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="15E70218FFF5FFE4FDBFF8C0FD84F8D4" box="[551,635,1856,1878]" pageId="11" pageNumber="112" rank="species">
|
||
<emphasis id="0ED4BF63FFF5FFE4FDBFF8C0FD84F8D4" bold="true" box="[551,635,1856,1878]" pageId="11" pageNumber="112">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting tubercles of regular size); C, ocularium, anterior view; D, habitus, right lateral view; E, right tarsus IV, prolateral view; F, right pedipalp, prolateral view; G, ditto, retrolateral view. Abbreviations: A1–4, scutal areas I–IV; Ca, carapace; Fh, frontal hump; Ft1–3, free tergites I–III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum. A, D at same scale; F, G at same scale. Scale bars of A, B, D, F, G: 1 mm. Scale bars of C, E: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="68DF33F9FFF2FFE3FF0FF8CBFF39F81C" httpUri="https://zenodo.org/record/256484/files/figure.png" pageId="12" pageNumber="113" targetBox="[159,1428,208,1829]" targetPageId="12">
|
||
<paragraph id="3C1F6371FFF2FFE3FF0FF8CBFF39F81C" blockId="12.[151,1436,1867,1950]" pageId="12" pageNumber="113">
|
||
<emphasis id="0ED4BF63FFF2FFE3FF0FF8CBFEEAF8E3" bold="true" box="[151,277,1867,1889]" pageId="12" pageNumber="113">FIGURE 3.</emphasis>
|
||
<taxonomicName id="FBA018F2FFF2FFE3FE85F8CBFDE4F8E3" box="[285,539,1867,1889]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="bicornutus" status="sp. nov.">
|
||
<emphasis id="0ED4BF63FFF2FFE3FE85F8CBFDE4F8E3" box="[285,539,1867,1889]" italics="true" pageId="12" pageNumber="113">Nanophareus bicornutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="15E70218FFF2FFE3FDBBF8CBFD89F8E3" box="[547,630,1867,1889]" pageId="12" pageNumber="113" rank="species">
|
||
<emphasis id="0ED4BF63FFF2FFE3FDBBF8CBFD89F8E3" bold="true" box="[547,630,1867,1889]" pageId="12" pageNumber="113">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Male (holotype) right trochanter-metatarsus IV: A, dorsal view; B, prolateral view; C, ventral view; D, retrolateral view. Abbreviations: Pro, prolateral; Ret, retrolateral. A–D at same scale. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3C1F6371FFF3FFE2FF5FFF17FCE4FE9E" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="114">
|
||
Penis (
|
||
<figureCitation id="A49B7FF4FFF3FFE2FE8DFF17FEA7FF32" box="[277,344,151,176]" captionStart="FIGURE 8" captionStartId="19.[151,250,1858,1880]" captionTargetBox="[218,1257,229,1784]" captionTargetId="figure@19.[169,1335,177,1817]" captionTargetPageId="19" captionText="FIGURE 8. Distal part of Nanophareus spp. penis: A, N. bicornutus sp. nov., dorsal view, B, ditto, right lateral view; C, N. maipu sp. nov., in dorsal view, D, ditto, right lateral view, E, ditto, ventral view; F, N. polyhastatus sp. nov., dorsal view, G, ditto, right lateral view. Abbreviations: bms, basal group of macrosetae (MS); Gl, glans; MS C, MS of group C; MS D, MS of group D; MS E, MS of group E; St, stylus of glans; Vpl, ventral plate; Vpr, ventral process of glans; Vt, ventral trichomes. A, B at same scale; C – E at same scale, F, G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256489/files/figure.png" pageId="13" pageNumber="114">Fig. 8</figureCitation>
|
||
A, B): Glans with wide sac, its distal part projected dorsally; stylus elongated and curved, with ventral trichomes on median third; ventral process apex with nail head shape (apex with lateral projections), curved towards stylus. Ventral plate subrectagular, basally inflated, with 3 pairs of MS C (distal pair smallest), 1 pair of short MS D, 2 pairs of small MS E, 4 pairs of basal setae.
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF3FFE2FF5FFEA7FC18FE0A" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="114">Coloration: Body, pedipalps, patellae I–IV, tibia IV and apical regions of femora I–IV and tibiae I–III brown; coxa IV, femur IV and apophyses of coxa–trochanter IV dark brown; the remaining parts (chelicerae, trochanters I–IV, most part of femora and tibiae I–III, metatarsi–tarsi I–IV) pale brown.</paragraph>
|
||
<paragraph id="3C1F6371FFF3FFE2FF5FFE14FE60FE2F" blockId="13.[151,1436,151,501]" box="[199,415,404,429]" pageId="13" pageNumber="114">
|
||
<emphasis id="0ED4BF63FFF3FFE2FF5FFE14FEE4FE2E" box="[199,283,404,428]" italics="true" pageId="13" pageNumber="114">Female</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74BA30FAFFF3FFE2FF5FFE37FC0DFE77" pageId="13" pageNumber="114" type="materials_examined">
|
||
<paragraph id="3C1F6371FFF3FFE2FF5FFE37FCCCFE52" blockId="13.[151,1436,151,501]" box="[199,819,439,464]" pageId="13" pageNumber="114">
|
||
<emphasis id="0ED4BF63FFF3FFE2FF5FFE37FE9AFE52" bold="true" box="[199,357,439,464]" pageId="13" pageNumber="114">
|
||
<typeStatus id="E31BDDD3FFF3FFE2FF5FFE37FEFFFE52" box="[199,256,439,464]" pageId="13" pageNumber="114">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="44B723E1FFF3FFE2FEF4FE37FE53FE52" box="[364,428,439,464]" name="Chile" pageId="13" pageNumber="114">Chile</collectingCountry>
|
||
, V Región de Valparaíso, Zapallar.
|
||
</paragraph>
|
||
<paragraph id="3C1F6371FFF3FFE2FF5FFE5CFC0DFE77" blockId="13.[151,1436,151,501]" box="[199,1010,476,501]" pageId="13" pageNumber="114">
|
||
<emphasis id="0ED4BF63FFF3FFE2FF5FFE5CFD9FFE77" bold="true" box="[199,608,476,501]" pageId="13" pageNumber="114">
|
||
Geographical distribution (
|
||
<figureCitation id="A49B7FF4FFF3FFE2FD94FE5CFDAEFE77" box="[524,593,476,501]" captionStart="FIGURE 9" captionStartId="20.[151,250,1322,1344]" captionTargetBox="[155,1435,198,1296]" captionTargetId="figure@20.[151,1436,193,1301]" captionTargetPageId="20" captionText="FIGURE 9. Distribution of Nanophareus spp. The precise locality of N. palpalis is unknown." httpUri="https://zenodo.org/record/256490/files/figure.png" pageId="13" pageNumber="114">Fig. 9</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Known only from the
|
||
<typeStatus id="E31BDDD3FFF3FFE2FCF8FE5DFC6FFE77" box="[864,912,477,501]" pageId="13" pageNumber="114">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |