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<document id="A00164AC820F7DD17D76E568F7602A58" ID-CLB-Dataset="46261" ID-DOI="10.11646/zootaxa.3774.5.3" ID-GBIF-Dataset="a73d4668-01cd-4e4c-8173-e130939a89c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="226487" ID-ZooBank="091256F3-B410-42FD-AFA2-3415BA48DDBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460394050722" checkinUser="plazi" docAuthor="Santos, Bianca Tamires Silva Dos, Nascimento, Agata Tyanne Silva &amp; Fernandes, José Antônio Marin" docDate="2014" docId="5D0E87ABFFDF702DFF4D5EE76E79F8F2" docLanguage="en" docName="zt03774p459.pdf" docOrigin="Zootaxa 3774 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessa punctata Santos &amp; Fernandes, sp. nov." docType="treatment" docVersion="9" lastPageNumber="446" masterDocId="A137FFD3FFDA7028FFDA5F736A78FF9D" masterDocTitle="Proposition of a new species group in Edessa Fabricius, 1803 (Hemiptera: Heteroptera: Pentatomidae: Edessinae)" masterLastPageNumber="459" masterPageNumber="441" pageNumber="446" updateTime="1698298553916" updateUser="ExternalLinkService">
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<mods:title id="9C5E7D648BF0C8C361B762B5B7C598B9">Proposition of a new species group in Edessa Fabricius, 1803 (Hemiptera: Heteroptera: Pentatomidae: Edessinae)</mods:title>
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<mods:namePart id="CB14C9B7501640F10345E9D9D361083D">Santos, Bianca Tamires Silva Dos</mods:namePart>
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<mods:namePart id="74F0880E41C5573243DB51D9CF6593C7">Nascimento, Agata Tyanne Silva</mods:namePart>
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<mods:namePart id="3F606624FCD73377661E903C3B9077A2">Fernandes, José Antônio Marin</mods:namePart>
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<treatment id="5D0E87ABFFDF702DFF4D5EE76E79F8F2" ID-DOI="http://doi.org/10.5281/zenodo.6133721" ID-GBIF-Taxon="119417647" ID-Zenodo-Dep="6133721" LSID="urn:lsid:plazi:treatment:5D0E87ABFFDF702DFF4D5EE76E79F8F2" httpUri="http://treatment.plazi.org/id/5D0E87ABFFDF702DFF4D5EE76E79F8F2" lastPageNumber="446" pageId="5" pageNumber="446">
<subSubSection id="9DBD6536FFDF702DFF4D5EE76BE1FE52" pageId="5" pageNumber="446" type="nomenclature">
<paragraph id="D51836BDFFDF702DFF4D5EE768BEFE33" blockId="5.[151,710,404,463]" box="[151,710,404,430]" pageId="5" pageNumber="446">
<heading id="8E5081D1FFDF702DFF4D5EE768BEFE33" bold="true" box="[151,710,404,430]" fontSize="11" level="1" pageId="5" pageNumber="446" reason="1">
<emphasis id="E7D3EAAFFFDF702DFF4D5EE768BEFE33" bold="true" box="[151,710,404,430]" pageId="5" pageNumber="446">
<taxonomicName id="12A74D3EFFDF702DFF4D5EE76819FE33" authority="Santos &amp; Fernandes" authorityName="Santos &amp; Fernandes" box="[151,609,404,430]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="E7D3EAAFFFDF702DFF4D5EE76B24FE33" bold="true" box="[151,348,404,430]" italics="true" pageId="5" pageNumber="446">Edessa punctata</emphasis>
Santos &amp; Fernandes
</taxonomicName>
<taxonomicNameLabel id="FCE057D4FFDF702DFDB25EE768BEFE33" box="[616,710,404,430]" pageId="5" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D51836BDFFDF702DFF4D5EC56BE1FE52" blockId="5.[151,710,404,463]" box="[151,409,438,463]" pageId="5" pageNumber="446">
(
<figureCitation id="4D9C2A38FFDF702DFF455EC56B6AFE52" box="[159,274,438,463]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Figs.712</figureCitation>
; 43; 50; 56)
</paragraph>
</subSubSection>
<subSubSection id="9DBD6536FFDF702DFF4D5E8D6B79FCC6" pageId="5" pageNumber="446" type="etymology">
<paragraph id="D51836BDFFDF702DFF4D5E8D69E6FD8A" blockId="5.[151,1437,510,1903]" box="[151,926,510,535]" pageId="5" pageNumber="446">
<emphasis id="E7D3EAAFFFDF702DFF4D5E8D6B66FD8A" bold="true" box="[151,286,510,535]" pageId="5" pageNumber="446">Etymology.</emphasis>
Name refers to dense and dark punctation of the species.
</paragraph>
<paragraph id="D51836BDFFDF702DFF1D5D5168DCFDC2" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
<typeStatus id="0A1C881FFFDF702DFF1D5D516B57FDA6" box="[199,303,546,571]" pageId="5" pageNumber="446" type="holotype">Holotype</typeStatus>
♂.
<collectingCountry id="ADB0762DFFDF702DFEBF5D50686AFDA6" box="[357,530,547,571]" name="Costa Rica" pageId="5" pageNumber="446">COSTA RICA</collectingCountry>
,
<emphasis id="E7D3EAAFFFDF702DFDFE5D5168CDFDA6" bold="true" box="[548,693,546,571]" pageId="5" pageNumber="446">Puntarenas:</emphasis>
Finca Cafrosa, Embalce,
<quantity id="125F9B58FFDF702DFC355D506E47FDA7" box="[1007,1087,547,571]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="446" unit="m" value="800.0">800 m</quantity>
, NO de Tigra,
<quantity id="125F9B58FFDF702DFB245D506F24FDA7" box="[1278,1372,547,571]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="5" pageNumber="446" unit="m" value="1280.0">1280 m</quantity>
. L_S 317800_596200#7893.
<date id="A119107DFFDF702DFE7B5D356BC4FDC2" box="[417,444,582,607]" pageId="5" pageNumber="446" value="1996-07-12">12</date>
-VII-1996. (INBIO).
</paragraph>
<paragraph id="D51836BDFFDF702DFF1D5D186B79FCC6" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
<typeStatus id="0A1C881FFFDF702DFF1D5D186B40FD1E" box="[199,312,619,643]" pageId="5" pageNumber="446" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="ADB0762DFFDF702DFE985D186B92FD1E" box="[322,490,619,643]" name="Costa Rica" pageId="5" pageNumber="446">COSTA RICA</collectingCountry>
,
<emphasis id="E7D3EAAFFFDF702DFE2F5D1968FEFD1E" bold="true" box="[501,646,618,643]" pageId="5" pageNumber="446">Puntarenas:</emphasis>
♀ same data as
<typeStatus id="0A1C881FFFDF702DFC985D1969DCFD1E" box="[834,932,618,643]" pageId="5" pageNumber="446" type="holotype">holotype</typeStatus>
(INBIO); ♀ Mellizas Sabalito,
<quantity id="125F9B58FFDF702DFAD05D186F2AFD1F" box="[1290,1362,619,643]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="446" unit="m" value="700.0">700 m</quantity>
SE de Mellizas, Sector Las Tablas,
<quantity id="125F9B58FFDF702DFE3A5DFD6841FD3B" box="[480,569,654,679]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="446" unit="m" value="1500.0">1500 m</quantity>
,
<date id="A119107DFFDF702DFD9C5DFC697FFD3A" box="[582,775,654,679]" pageId="5" pageNumber="446" value="1995-09-21" valueMax="1995-09-24" valueMin="1995-09-21">21 a 24-IX-1995</date>
, E. Navarro, L_S315500_598500#6143 (INBIO); ♀
<quantity id="125F9B58FFDF702DFAB85DFC6FE4FD3A" box="[1378,1436,654,679]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="446" unit="km" value="2.0">2 km</quantity>
WSW Monteverde Cloud Forest Reserve,
<date id="A119107DFFDF702DFD5B5DC1693AFD56" box="[641,834,690,715]" pageId="5" pageNumber="446" value="1995-08-15" valueMax="1995-08-16" valueMin="1995-08-15">1516-VIII-1995</date>
(JEE); ♂3♀ Monteverde Cloud For. Res.
<date id="A119107DFFDF702DFAE85DC06AAAFD72" pageId="5" pageNumber="446" value="1984-05-27" valueMax="1984-05-31" valueMin="1984-05-27">2731-V- 1984</date>
, Riley, Rider and LeDoux (DAR).
<emphasis id="E7D3EAAFFFDF702DFD8F5DA568B8FD72" bold="true" box="[597,704,726,751]" pageId="5" pageNumber="446">Cartago:</emphasis>
♂ Muneco,
<date id="A119107DFFDF702DFC965DA469BEFD72" box="[844,966,727,751]" pageId="5" pageNumber="446" value="1982-05-31">31-V-1982</date>
, L.J. Barkley (DAR). PANAM,
<emphasis id="E7D3EAAFFFDF702DFAF65DA56FE4FD72" bold="true" box="[1324,1436,726,751]" pageId="5" pageNumber="446">Chiriqui:</emphasis>
♂ Volcan Area, 5500,
<date id="A119107DFFDF702DFE435D886866FC8E" box="[409,542,763,787]" pageId="5" pageNumber="446" value="1973-04-10">10-IV-1973</date>
, D. Engleman (UNSM);
<specimenCount id="C3A1FD34FFDF702DFCEE5D896921FC8E" box="[820,857,762,787]" pageId="5" pageNumber="446" type="generic">1♂</specimenCount>
6♀
<quantity id="125F9B58FFDF702DFC5B5D8869CBFC8E" box="[897,947,762,787]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="5" pageNumber="446" unit="mi" value="8.0">8 mi</quantity>
NE El Vocan, 5900,
<date id="A119107DFFDF702DFB7D5D886F52FC8E" box="[1191,1322,763,787]" pageId="5" pageNumber="446" value="1974-07-08">8-VII-1974</date>
, C.W. and L. OBrien and
<collectingCountry id="ADB0762DFFDF702DFE8A5C6D6BCCFCAA" box="[336,436,798,823]" name="Marshall Islands" pageId="5" pageNumber="446">Marshall</collectingCountry>
(DAR); ♂♀ same data (UFRG); ♀ Bambito, El Volcan,
<date id="A119107DFFDF702DFB905C6C6E9BFCAA" box="[1098,1251,799,823]" pageId="5" pageNumber="446" value="1999-08-31">31-VIII-1999</date>
, J. C. Schaffner (TAMU).
</paragraph>
</subSubSection>
<subSubSection id="9DBD6536FFDF702DFF1D5C156838F90A" pageId="5" pageNumber="446" type="description">
<paragraph id="D51836BDFFDF702DFF1D5C1568D8FC3E" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
<emphasis id="E7D3EAAFFFDF702DFF1D5C156B05FCE2" bold="true" box="[199,381,870,895]" pageId="5" pageNumber="446">Measurements.</emphasis>
Head length (2.1); head width (3.23.7); pronotal length (2.32.8); pronotal width (6.57.5); total length (1113); abdominal width (6.57).
</paragraph>
<paragraph id="D51836BDFFDF702DFF1D5CDD6BF5FAB3" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
Body greenish-brown dorsally (
<figureCitation id="4D9C2A38FFDF702DFDED5CDD68F5FC5A" box="[567,653,942,967]" captionStart="FIGURES 42 48" captionStartId="16.[151,264,1877,1899]" captionTargetBox="[253,1358,215,1847]" captionTargetId="figure@16.[214,1372,193,1856]" captionTargetPageId="16" captionText="FIGURES 42 48. Photo of the species, dorsal view. 42 - Edessa godmani Distant, 1881; 43 — Edessa punctata sp. nov.; 44 — Edessa picticornis Stål, 1872; 45 — Edessa luteomaculata sp. nov.; 46 — Edessa pudibunda Stål, 1862; 47 — Edessa nigriventris sp. nov.; 48 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226495/files/figure.png" pageId="5" pageNumber="446">Fig. 43</figureCitation>
) and yellow to dark brown ventrally (
<figureCitation id="4D9C2A38FFDF702DFB895CDD6ED4FC5A" box="[1107,1196,942,967]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="5" pageNumber="446">Fig. 50</figureCitation>
); densely punctured, punctures black, sometimes surrounded by suffused spots on scutellum, corium and abdomen; some parts with dark spots instead of punctures. Head uniformly and densely punctured. Mandibular plates gently rugose and level with clypeus and cephalic disc. Ventral surface of head dark yellow and punctured, dark spotted below antenniferous tubercles. First three antennal segments light brown and spotted; fourth and fifth segments brown and not spotted. Pronotum brown with dense dark punctures, anterior half lighter than posterior; anterolateral margin reddish. Apex of scutellum with a large yellow spot. Corium with a dark faded oblique spot. Ventral punctures of thorax black. Metasternal process and evaporatorium with dense dark spots. Metasternal process light brown, arms of bifurcation strong and divergent. Legs densely dark spotted. Connexivum greenish-brown with dark punctures and a medial small calloused area yellow. Ventral surface of abdomen densely punctured. Female without a rounded pair of dark spots on segment VII.
</paragraph>
<paragraph id="D51836BDFFDF702DFF1D5A4969ACFA7E" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
Male: Pygophore (
<figureCitation id="4D9C2A38FFDF702DFE7F5A49686EFACE" box="[421,534,1338,1363]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Figs. 79</figureCitation>
) brown. Dorsal rim with a dark spot restricted to middle region. Superior processes of the genital cup slightly concave and elliptical (
<figureCitation id="4D9C2A38FFDF702DFCE75A2D69D3FAEA" box="[829,939,1374,1399]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Figs. 78</figureCitation>
); margin slightly tumid and reaching dorsal rim (
<figureCitation id="4D9C2A38FFDF702DFF155AF16B6DFA06" box="[207,277,1410,1435]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 8</figureCitation>
). Paramere pale with margins brown; dorsal part elliptical, ventral process long and laterally directed (
<figureCitation id="4D9C2A38FFDF702DFF455AD56A9DFA22" box="[159,229,1446,1471]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 8</figureCitation>
). Proctiger with two almost inconspicuous keels; posterior face ogive-like (
<figureCitation id="4D9C2A38FFDF702DFB935AD56EF7FA22" box="[1097,1167,1446,1471]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 8</figureCitation>
). Ventral rim yellow to brown with tiny dark punctures; expansions rounded and swollen (
<figureCitation id="4D9C2A38FFDF702DFC5B5AB969BDFA7E" box="[897,965,1482,1507]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="D51836BDFFDF702DFF1D5A9D6838F90A" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
Female: Gonocoxites 8 strongly convex with dense dark punctures (
<figureCitation id="4D9C2A38FFDF702DFC005A9D6E48F99A" box="[986,1072,1518,1543]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 12</figureCitation>
), excavation of medial margins deep, V-shaped (
<figureCitation id="4D9C2A38FFDF702DFE8559616BC0F9B6" box="[351,440,1554,1579]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 11</figureCitation>
), narrower than gonocoxite 9, rounded distally and leaving visible medially tumid gonapophyses 8. Gonocoxite 9 medially carinate; constricted just below punctate part, basal part turned ventrad forming a small conical projection. Laterotergites 9 punctured and not grooved, acuminate apices surpassing the band uniting laterotergites 8 (
<figureCitation id="4D9C2A38FFDF702DFE3B590D684BF90A" box="[481,563,1662,1687]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="5" pageNumber="446">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9DBD6536FFDF702DFF1D59D16850F8D6" pageId="5" pageNumber="446" type="discussion">
<paragraph id="D51836BDFFDF702DFF1D59D16850F8D6" blockId="5.[151,1437,510,1903]" pageId="5" pageNumber="446">
<emphasis id="E7D3EAAFFFDF702DFF1D59D16B37F926" bold="true" box="[199,335,1698,1723]" pageId="5" pageNumber="446">Comments.</emphasis>
The species
<taxonomicName id="12A74D3EFFDF702DFE3859D0681CF927" box="[482,612,1699,1722]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="E7D3EAAFFFDF702DFE3859D0681CF927" box="[482,612,1699,1722]" italics="true" pageId="5" pageNumber="446">E. punctata</emphasis>
</taxonomicName>
and
<taxonomicName id="12A74D3EFFDF702DFD4159D06965F926" box="[667,797,1699,1723]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="godmani">
<emphasis id="E7D3EAAFFFDF702DFD4159D06965F926" box="[667,797,1699,1723]" italics="true" pageId="5" pageNumber="446">E. godmani</emphasis>
</taxonomicName>
are found at altitudes ranging from 700 to 1,000 meters. They share several characteristics (see comments of
<taxonomicName id="12A74D3EFFDF702DFD2359B46906F943" box="[761,894,1734,1758]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="godmani">
<emphasis id="E7D3EAAFFFDF702DFD2359B46906F943" box="[761,894,1734,1758]" italics="true" pageId="5" pageNumber="446">E. godmani</emphasis>
</taxonomicName>
), but they can be separate by the shape of the excavation of medial margins of gonocoxites 8 and shape of the paramere.
<taxonomicName id="12A74D3EFFDF702DFC0059986EEAF89F" box="[986,1170,1771,1795]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="E7D3EAAFFFDF702DFC0059986EEAF89F" box="[986,1170,1771,1795]" italics="true" pageId="5" pageNumber="446">Edessa punctata</emphasis>
</taxonomicName>
is more rounded, larger and darker than
<taxonomicName id="12A74D3EFFDF702DFE8A587C6BAAF8BB" box="[336,466,1806,1830]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="godmani">
<emphasis id="E7D3EAAFFFDF702DFE8A587C6BAAF8BB" box="[336,466,1806,1830]" italics="true" pageId="5" pageNumber="446">E. godmani</emphasis>
</taxonomicName>
. Both species can be separate from remaining species of the group by the denser and more coarse punctation of the body.
</paragraph>
</subSubSection>
<subSubSection id="9DBD6536FFDF702DFF1D58256E79F8F2" box="[199,1025,1878,1903]" pageId="5" pageNumber="446" type="distribution">
<paragraph id="D51836BDFFDF702DFF1D58256E79F8F2" blockId="5.[151,1437,510,1903]" box="[199,1025,1878,1903]" pageId="5" pageNumber="446">
<emphasis id="E7D3EAAFFFDF702DFF1D58256B21F8F2" bold="true" box="[199,345,1878,1903]" pageId="5" pageNumber="446">Distribution</emphasis>
(
<figureCitation id="4D9C2A38FFDF702DFEB258256BC2F8F2" box="[360,442,1878,1903]" captionStart="FIGURE 56" captionStartId="18.[151,250,1159,1181]" captionTargetBox="[168,1417,206,1121]" captionTargetId="figure@18.[151,1436,171,1138]" captionTargetPageId="18" captionText="FIGURE 56. Distribution of the species of the pudibunda group." httpUri="https://zenodo.org/record/226497/files/figure.png" pageId="5" pageNumber="446">Fig. 56</figureCitation>
).
<collectingCountry id="ADB0762DFFDF702DFE0A58246837F8F2" box="[464,591,1878,1903]" name="Costa Rica" pageId="5" pageNumber="446">Costa Rica</collectingCountry>
: Puntarenas, Cartago. Panam: Chiriqu.
</paragraph>
</subSubSection>
</treatment>
</document>