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<document id="1656EB4CA5C5F51923361D6E9972CDB7" ID-CLB-Dataset="46261" ID-DOI="10.11646/zootaxa.3774.5.3" ID-GBIF-Dataset="a73d4668-01cd-4e4c-8173-e130939a89c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="226487" ID-ZooBank="091256F3-B410-42FD-AFA2-3415BA48DDBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460394050722" checkinUser="plazi" docAuthor="Santos, Bianca Tamires Silva Dos, Nascimento, Agata Tyanne Silva &amp; Fernandes, José Antônio Marin" docDate="2014" docId="5D0E87ABFFD8702BFF4D5A136FE4FEBF" docLanguage="en" docName="zt03774p459.pdf" docOrigin="Zootaxa 3774 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessa pudibunda" docType="key" docVersion="9" lastPageNumber="444" masterDocId="A137FFD3FFDA7028FFDA5F736A78FF9D" masterDocTitle="Proposition of a new species group in Edessa Fabricius, 1803 (Hemiptera: Heteroptera: Pentatomidae: Edessinae)" masterLastPageNumber="459" masterPageNumber="441" pageNumber="443" updateTime="1698298553916" updateUser="ExternalLinkService">
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<mods:title id="0E053DEC61248C104CD6918342E59A09">Proposition of a new species group in Edessa Fabricius, 1803 (Hemiptera: Heteroptera: Pentatomidae: Edessinae)</mods:title>
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<mods:namePart id="DFA3B884DE4AD8DE79B47530FF34B541">Santos, Bianca Tamires Silva Dos</mods:namePart>
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<mods:namePart id="3189BAAEF92BF6A6F7343DE3AF31F836">Nascimento, Agata Tyanne Silva</mods:namePart>
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<mods:namePart id="FED84F324745A6029FB5CEC433363909">Fernandes, José Antônio Marin</mods:namePart>
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<treatment id="5D0E87ABFFD8702BFF4D5A136FE4FEBF" ID-DOI="http://doi.org/10.5281/zenodo.6133717" ID-GBIF-Taxon="119417650" ID-Zenodo-Dep="6133717" LSID="urn:lsid:plazi:treatment:5D0E87ABFFD8702BFF4D5A136FE4FEBF" httpUri="http://treatment.plazi.org/id/5D0E87ABFFD8702BFF4D5A136FE4FEBF" lastPageId="3" lastPageNumber="444" pageId="2" pageNumber="443">
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<paragraph id="D51836BDFFD8702AFF4D5A13682FFAE7" blockId="2.[151,599,1376,1402]" box="[151,599,1376,1402]" pageId="2" pageNumber="443">
<heading id="8E5081D1FFD8702AFF4D5A13682FFAE7" bold="true" box="[151,599,1376,1402]" fontSize="11" level="1" pageId="2" pageNumber="443" reason="1">
<emphasis id="E7D3EAAFFFD8702AFF4D5A13682FFAE7" bold="true" box="[151,599,1376,1402]" pageId="2" pageNumber="443">
Key to the species group
<taxonomicName id="12A74D3EFFD8702AFE145A13682FFAE7" box="[462,599,1376,1402]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="pudibunda">pudibunda</taxonomicName>
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<keyStep id="6E532E18FFD8702AFF4D5AD56FE3FA6B" pageId="2" pageNumber="443">
<paragraph id="D51836BDFFD8702AFF4D5AD56FE4FA26" blockId="2.[151,1437,1446,2025]" box="[151,1436,1446,1467]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D5AD56FE4FA26" box="[151,1436,1446,1467]" pageId="2" pageNumber="443">
1. Body coarsely punctured. Metasternal process densely spotted. Connexivum without large yellow spots (
<figureCitation id="4D9C2A38FFD8702AFB145AD56F6CFA26" box="[1230,1300,1446,1467]" captionStart="FIGURES 42 48" captionStartId="16.[151,264,1877,1899]" captionTargetBox="[253,1358,215,1847]" captionTargetId="figure@16.[214,1372,193,1856]" captionTargetPageId="16" captionText="FIGURES 42 48. Photo of the species, dorsal view. 42 - Edessa godmani Distant, 1881; 43 — Edessa punctata sp. nov.; 44 — Edessa picticornis Stål, 1872; 45 — Edessa luteomaculata sp. nov.; 46 — Edessa pudibunda Stål, 1862; 47 — Edessa nigriventris sp. nov.; 48 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226495/files/figure.png" pageId="2" pageNumber="443">Fig. 42</figureCitation>
)......... 2
</keyLead>
</paragraph>
<paragraph id="D51836BDFFD8702AFF4D5AB76FE3FA6B" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D5AB76FE3FA6B" pageId="2" pageNumber="443">
- Body thinly punctured. Metasternal process not spotted or with a few tiny spots. Connexivum with large yellow spots (
<figureCitation id="4D9C2A38FFD8702AFA955AB76FEDFA44" box="[1359,1429,1476,1497]" captionStart="FIGURES 42 48" captionStartId="16.[151,264,1877,1899]" captionTargetBox="[253,1358,215,1847]" captionTargetId="figure@16.[214,1372,193,1856]" captionTargetPageId="16" captionText="FIGURES 42 48. Photo of the species, dorsal view. 42 - Edessa godmani Distant, 1881; 43 — Edessa punctata sp. nov.; 44 — Edessa picticornis Stål, 1872; 45 — Edessa luteomaculata sp. nov.; 46 — Edessa pudibunda Stål, 1862; 47 — Edessa nigriventris sp. nov.; 48 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226495/files/figure.png" pageId="2" pageNumber="443">Fig. 48</figureCitation>
).................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E532E18FFD8702AFF4D5A8D6FE2F9F6" pageId="2" pageNumber="443">
<paragraph id="D51836BDFFD8702AFF4D5A8D6FE3F9AC" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D5A8D6FE3F9AC" pageId="2" pageNumber="443">
2. Paramere with distal part elliptical, smaller than superior process of genital cup (
<figureCitation id="4D9C2A38FFD8702AFBDE5A8D6E39F98E" box="[1028,1089,1534,1555]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 6. Edessa godmani. 1 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="2" pageNumber="443">Fig. 2</figureCitation>
). Medial margins of gonocoxites 8 forming a U-shaped wide excavation (
<figureCitation id="4D9C2A38FFD8702AFD8A596F68F7F9AC" box="[592,655,1564,1585]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 6. Edessa godmani. 1 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="2" pageNumber="443">Fig. 5</figureCitation>
).........................................................
<taxonomicName id="12A74D3EFFD8702AFA9D59686FE3F9AC" box="[1351,1435,1563,1585]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="godmani">
<emphasis id="E7D3EAAFFFD8702AFA9D59686FE3F9AC" box="[1351,1435,1563,1585]" italics="true" pageId="2" pageNumber="443">godmani</emphasis>
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</paragraph>
<paragraph id="D51836BDFFD8702AFF4D594A6FE2F9F6" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D594A6FE2F9F6" pageId="2" pageNumber="443">
- Paramere with distal part rounded, larger than superior process of the genital cup (
<figureCitation id="4D9C2A38FFD8702AFBD3594A6E3DF9D3" box="[1033,1093,1593,1614]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="2" pageNumber="443">Fig. 8</figureCitation>
). Medial margins of gonocoxites 8 forming a narrow V-shaped excavation (
<figureCitation id="4D9C2A38FFD8702AFDB9592568AAF9F6" box="[611,722,1622,1643]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1804,1826]" captionTargetBox="[218,1400,221,1771]" captionTargetId="figure@6.[178,1409,193,1783]" captionTargetPageId="6" captionText="FIGURES 7 12. Edessa punctata sp. nov. 7 9, Male, pygophore; 7 — dorsal; 8 — posterior; 9 — ventral. 10 — Metasternal process. 11 12, Female, genital plates, ventral and posterior respectively. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226489/files/figure.png" pageId="2" pageNumber="443">Fig. 11, 12</figureCitation>
)...................................................
<taxonomicName id="12A74D3EFFD8702AFA9C59256FE2F9F6" box="[1350,1434,1622,1643]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="E7D3EAAFFFD8702AFA9C59256FE2F9F6" box="[1350,1434,1622,1643]" italics="true" pageId="2" pageNumber="443">punctata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E532E18FFD8702AFF4D59076FE3F97C" pageId="2" pageNumber="443">
<paragraph id="D51836BDFFD8702AFF4D59076FE3F93B" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D59076FE3F93B" pageId="2" pageNumber="443">
3. Apex of scutellum concolorous or slightly faded but never distinctly yellow (e.g.
<figureCitation id="4D9C2A38FFD8702AFBDB59076E33F914" box="[1025,1099,1652,1673]" captionStart="FIGURES 42 48" captionStartId="16.[151,264,1877,1899]" captionTargetBox="[253,1358,215,1847]" captionTargetId="figure@16.[214,1372,193,1856]" captionTargetPageId="16" captionText="FIGURES 42 48. Photo of the species, dorsal view. 42 - Edessa godmani Distant, 1881; 43 — Edessa punctata sp. nov.; 44 — Edessa picticornis Stål, 1872; 45 — Edessa luteomaculata sp. nov.; 46 — Edessa pudibunda Stål, 1862; 47 — Edessa nigriventris sp. nov.; 48 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226495/files/figure.png" pageId="2" pageNumber="443">Fig. 44</figureCitation>
). Ventral surface of abdomen dark yellow (e.g.
<figureCitation id="4D9C2A38FFD8702AFE8359E26BDCF93B" box="[345,420,1681,1702]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Fig. 51</figureCitation>
), sometimes with small lateral suffused areas but never black spots............................. 4
</keyLead>
</paragraph>
<paragraph id="D51836BDFFD8702AFF4D59DD6FE3F97C" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D59DD6FE3F97C" pageId="2" pageNumber="443">
- Apex of scutellum distinctly yellow (e.g.
<figureCitation id="4D9C2A38FFD8702AFDAB59DD68C0F95E" box="[625,696,1710,1731]" captionStart="FIGURES 42 48" captionStartId="16.[151,264,1877,1899]" captionTargetBox="[253,1358,215,1847]" captionTargetId="figure@16.[214,1372,193,1856]" captionTargetPageId="16" captionText="FIGURES 42 48. Photo of the species, dorsal view. 42 - Edessa godmani Distant, 1881; 43 — Edessa punctata sp. nov.; 44 — Edessa picticornis Stål, 1872; 45 — Edessa luteomaculata sp. nov.; 46 — Edessa pudibunda Stål, 1862; 47 — Edessa nigriventris sp. nov.; 48 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226495/files/figure.png" pageId="2" pageNumber="443">Fig. 43</figureCitation>
). Ventral surface of abdomen mostly black or with large black spots and suffused areas (
<figureCitation id="4D9C2A38FFD8702AFE8359BF6BB4F97C" box="[345,460,1740,1761]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Figs. 5355</figureCitation>
)............................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E532E18FFD8702AFF4D599A6FE4F80C" pageId="2" pageNumber="443">
<paragraph id="D51836BDFFD8702AFF4D599A6FE3F8A4" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D599A6FE3F8A4" pageId="2" pageNumber="443">
4. Metasternal process with tiny dark punctures and spots (
<figureCitation id="4D9C2A38FFD8702AFCD4599A6918F963" box="[782,864,1769,1790]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Fig. 49</figureCitation>
). Ventral surface of abdomen with dense, coarse and dark punctures (
<figureCitation id="4D9C2A38FFD8702AFE9158756BEAF886" box="[331,402,1798,1819]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Fig. 49</figureCitation>
). Parameres tall, reaching or slightly surpassing dorsal rim (
<figureCitation id="4D9C2A38FFD8702AFC1558756E6EF886" box="[975,1046,1798,1819]" captionStart="FIGURES 13 18" captionStartId="7.[151,264,1800,1822]" captionTargetBox="[172,1436,225,1775]" captionTargetId="figure@7.[151,1436,193,1778]" captionTargetPageId="7" captionText="FIGURES 13 18. Edessa picticornis. 13 15, Male, pygophore; 13 — dorsal; 14 — posterior; 15 — ventral. 16 17, Female, genital plates, ventral and posterior respectively. 18 — Metasternal process. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226490/files/figure.png" pageId="2" pageNumber="443">Fig. 14</figureCitation>
). Outline of the gonocoxites 8 W-shaped (
<figureCitation id="4D9C2A38FFD8702AFF3258576B2CF8A4" box="[232,340,1828,1849]" captionStart="FIGURES 13 18" captionStartId="7.[151,264,1800,1822]" captionTargetBox="[172,1436,225,1775]" captionTargetId="figure@7.[151,1436,193,1778]" captionTargetPageId="7" captionText="FIGURES 13 18. Edessa picticornis. 13 15, Male, pygophore; 13 — dorsal; 14 — posterior; 15 — ventral. 16 17, Female, genital plates, ventral and posterior respectively. 18 — Metasternal process. (Scale = 1 mm)." httpUri="https://zenodo.org/record/226490/files/figure.png" pageId="2" pageNumber="443">Figs.16, 17</figureCitation>
).................................................................................
<taxonomicName id="12A74D3EFFD8702AFAEF58576FE3F8A4" box="[1333,1435,1828,1849]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="picticornis">
<emphasis id="E7D3EAAFFFD8702AFAEF58576FE3F8A4" box="[1333,1435,1828,1849]" italics="true" pageId="2" pageNumber="443">picticornis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D51836BDFFD8702AFF4D58326FE4F80C" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D58326FE4F80C" pageId="2" pageNumber="443">
- Metasternal process not punctured or spotted (
<figureCitation id="4D9C2A38FFD8702AFD7F58326897F8CB" box="[677,751,1857,1878]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Fig. 52</figureCitation>
). Ventral surface of abdomen with fine, more sparse and brown punctures (
<figureCitation id="4D9C2A38FFD8702AFEC7582D6B1EF8EE" box="[285,358,1886,1907]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Fig. 52</figureCitation>
). Parameres short, reaching at most the base of proctiger (
<figureCitation id="4D9C2A38FFD8702AFC40582D699BF8EE" box="[922,995,1886,1907]" captionStart="FIGURES 19 25" captionStartId="9.[151,264,1770,1792]" captionTargetBox="[164,1416,220,1748]" captionTargetId="figure@9.[151,1436,193,1748]" captionTargetPageId="9" captionText="FIGURES 19 25. Edessa luteomaculata sp. nov. 19 21, Male, pygophore; 19 — dorsal; 20 — posterior; 21 — ventral. 22 23, Female, genital plates, ventral and posterior respectively; 24 — lateral view. 25 — Metasternal process; (Scale = 1 mm)." httpUri="https://zenodo.org/record/226491/files/figure.png" pageId="2" pageNumber="443">Fig. 20</figureCitation>
). Outline of the gonocoxites 8 rounded (
<figureCitation id="4D9C2A38FFD8702AFAB1582D6B66F80C" captionStart="FIGURES 19 25" captionStartId="9.[151,264,1770,1792]" captionTargetBox="[164,1416,220,1748]" captionTargetId="figure@9.[151,1436,193,1748]" captionTargetPageId="9" captionText="FIGURES 19 25. Edessa luteomaculata sp. nov. 19 21, Male, pygophore; 19 — dorsal; 20 — posterior; 21 — ventral. 22 23, Female, genital plates, ventral and posterior respectively; 24 — lateral view. 25 — Metasternal process; (Scale = 1 mm)." httpUri="https://zenodo.org/record/226491/files/figure.png" pageId="2" pageNumber="443">Figs. 22, 23</figureCitation>
)...................................................................................
<taxonomicName id="12A74D3EFFD8702AFAC958086FE4F80C" box="[1299,1436,1915,1937]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="luteomaculata">
<emphasis id="E7D3EAAFFFD8702AFAC958086FE4F80C" box="[1299,1436,1915,1937]" italics="true" pageId="2" pageNumber="443">luteomaculata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E532E18FFD8702AFF4D58EA6FE3F874" pageId="2" pageNumber="443">
<paragraph id="D51836BDFFD8702AFF4D58EA6FE4F833" blockId="2.[151,1437,1446,2025]" box="[151,1436,1945,1966]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D58EA6FE4F833" box="[151,1436,1945,1966]" pageId="2" pageNumber="443">
5. First three antennal segments suffused and not spotted. Body ventrally mostly black (
<figureCitation id="4D9C2A38FFD8702AFBC858EA6E21F833" box="[1042,1113,1945,1966]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="2" pageNumber="443">Fig. 55</figureCitation>
). Legs without dark spots.... 6
</keyLead>
</paragraph>
<paragraph id="D51836BDFFD8702AFF4D58C56FE3F874" blockId="2.[151,1437,1446,2025]" pageId="2" pageNumber="443">
<keyLead id="6E569588FFD8702AFF4D58C56FE3F874" pageId="2" pageNumber="443">
- First three antennal segments light with dense dark spots. Body ventrally dark yellow to brown with large black spots and suffused areas (fig. 53). Legs with small black spots....................................................
<taxonomicName id="12A74D3EFFD8702AFAEF58A06FE3F874" box="[1333,1435,2003,2025]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="E7D3EAAFFFD8702AFAEF58A06FE3F874" box="[1333,1435,2003,2025]" italics="true" pageId="2" pageNumber="443">pudibunda</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6E532E18FFD9702BFF4D5FE46FE4FEBF" pageId="3" pageNumber="444">
<paragraph id="D51836BDFFD9702BFF4D5FE46FE1FF57" blockId="3.[151,1436,151,290]" pageId="3" pageNumber="444">
<keyLead id="6E569588FFD9702BFF4D5FE46FE1FF57" pageId="3" pageNumber="444">
6. Body dorsally green to greenish-brown with corium brown. Thorax ventrally black with some yellow spots (
<figureCitation id="4D9C2A38FFD9702BFB365FE46F4BFF31" box="[1260,1331,151,172]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="3" pageNumber="444">Fig. 54</figureCitation>
). Evaporatorium completely black. Paramere broad (
<figureCitation id="4D9C2A38FFD9702BFDA15FC668BBFF57" box="[635,707,181,202]" captionStart="FIGURES 32 36" captionStartId="13.[151,264,1730,1752]" captionTargetBox="[258,1371,221,1708]" captionTargetId="figure@13.[206,1380,193,1709]" captionTargetPageId="13" captionText="FIGURES 32 36. Edessa nigriventris sp. nov. 32 34, Male, pygophore; 32 — dorsal; 33 — posterior; 34 — ventral. 35 — Female, genital plates. 36 — Metasternal process; (Scale = 1 mm)." httpUri="https://zenodo.org/record/226493/files/figure.png" pageId="3" pageNumber="444">Fig. 33</figureCitation>
). Outline of gonocoxites 8 rounded (
<figureCitation id="4D9C2A38FFD9702BFBF95FC66E11FF57" box="[1059,1129,181,202]" captionStart="FIGURES 32 36" captionStartId="13.[151,264,1730,1752]" captionTargetBox="[258,1371,221,1708]" captionTargetId="figure@13.[206,1380,193,1709]" captionTargetPageId="13" captionText="FIGURES 32 36. Edessa nigriventris sp. nov. 32 34, Male, pygophore; 32 — dorsal; 33 — posterior; 34 — ventral. 35 — Female, genital plates. 36 — Metasternal process; (Scale = 1 mm)." httpUri="https://zenodo.org/record/226493/files/figure.png" pageId="3" pageNumber="444">Fig. 35</figureCitation>
)...............
<taxonomicName id="12A74D3EFFD9702BFAF35FC66FE1FF57" box="[1321,1433,181,202]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="E7D3EAAFFFD9702BFAF35FC66FE1FF57" box="[1321,1433,181,202]" italics="true" pageId="3" pageNumber="444">nigriventris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D51836BDFFD9702BFF4D5FA16FE4FEBF" blockId="3.[151,1436,151,290]" pageId="3" pageNumber="444">
<keyLead id="6E569588FFD9702BFF4D5FA16FE4FEBF" pageId="3" pageNumber="444">
- Body dorsally greenish-brown to dark brown with parts of pronotum and scutellum almost black. Thorax ventrally variegated black, brown and yellow (
<figureCitation id="4D9C2A38FFD9702BFE005F9C685AFE99" box="[474,546,239,260]" captionStart="FIGURES 49 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="3" pageNumber="444">Fig. 55</figureCitation>
). Evaporatorium black with yellow spots. Paramere narrow (
<figureCitation id="4D9C2A38FFD9702BFBB25F9C6EC8FE99" box="[1128,1200,239,260]" captionStart="FIGURES 37 41" captionStartId="15.[151,264,1834,1856]" captionTargetBox="[235,1341,207,1809]" captionTargetId="figure@15.[195,1391,193,1813]" captionTargetPageId="15" captionText="FIGURES 37 41. Edessa nigridorsata sp. nov. 37 39, Male, pygophore; 37 — dorsal; 38 — posterior; 39 — ventral. 40 — Female, genital plates. 41 — Metasternal process; (Scale = 1 mm)." httpUri="https://zenodo.org/record/226494/files/figure.png" pageId="3" pageNumber="444">Fig. 38</figureCitation>
). Outline of gonocoxites 8 W-shaped (
<figureCitation id="4D9C2A38FFD9702BFEBA5E7E6BDDFEBF" box="[352,421,269,290]" captionStart="FIGURES 37 41" captionStartId="15.[151,264,1834,1856]" captionTargetBox="[235,1341,207,1809]" captionTargetId="figure@15.[195,1391,193,1813]" captionTargetPageId="15" captionText="FIGURES 37 41. Edessa nigridorsata sp. nov. 37 39, Male, pygophore; 37 — dorsal; 38 — posterior; 39 — ventral. 40 — Female, genital plates. 41 — Metasternal process; (Scale = 1 mm)." httpUri="https://zenodo.org/record/226494/files/figure.png" pageId="3" pageNumber="444">Fig. 40</figureCitation>
)..........................................................................
<taxonomicName id="12A74D3EFFD9702BFAFE5E7F6FE4FEBF" box="[1316,1436,268,290]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="nigridorsata">
<emphasis id="E7D3EAAFFFD9702BFAFE5E7F6FE4FEBF" box="[1316,1436,268,290]" italics="true" pageId="3" pageNumber="444">nigridorsata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>