159 lines
15 KiB
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159 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.699.12031" ID-GBIF-Dataset="dbf6bb9f-7ea4-4fa3-83ee-f58263427e39" ID-PMC="PMC5673864" ID-Pensoft-Pub="1313-2970-699-1" ID-PubMed="29200927" ID-ZBK="0F92401F3F7C4896AD9D72BC84348C7D" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-699-1" ModsDocOrigin="ZooKeys 699" ModsDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" checkinTime="1505836577638" checkinUser="pensoft" docAuthor="Cherman, Mariana Alejandra, Mise, Kleber Makoto, Moron †, Miguel Angel, Vaz-de-Mello, Fernando Z. & Almeida, Lucia Massutti de" docDate="2017" docId="382FE4C71CFF95BB7F5E24C0DE8347C9" docLanguage="en" docName="ZooKeys 699: 1-120" docOrigin="ZooKeys 699" docSource="http://dx.doi.org/10.3897/zookeys.699.12031" docTitle="Liogenys laminiceps Moser 1919" docType="treatment" docVersion="4" lastPageNumber="41" masterDocId="FFD9636FB945C41DFF8AB133FFFA9D76" masterDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="38" updateTime="1668164813467" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cherman, Mariana Alejandra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mise, Kleber Makoto</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moron †, Miguel Angel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vaz-de-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Almeida, Lucia Massutti de</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>699</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>120</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.699.12031</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.699.12031</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-699-1</mods:identifier>
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<mods:identifier type="ZBK">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
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<mods:identifier type="ZooBank">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134311193" LSID="urn:lsid:plazi:treatment:382FE4C71CFF95BB7F5E24C0DE8347C9" httpUri="http://treatment.plazi.org/id/382FE4C71CFF95BB7F5E24C0DE8347C9" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38">
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<subSubSection pageId="37" pageNumber="38" type="nomenclature">
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<paragraph pageId="37" pageNumber="38">
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<taxonomicName authority="Moser, 1919" authorityName="Moser" authorityYear="1919" class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys laminiceps" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="laminiceps">Liogenys laminiceps Moser, 1919</taxonomicName>
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Figs 65, 91
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</paragraph>
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</subSubSection>
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<subSubSection pageId="38" pageNumber="39" type="reference_group">
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<paragraph pageId="38" pageNumber="39">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys laminiceps" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="laminiceps">
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<pageBreakToken pageId="38" pageNumber="39" start="start">Liogenys</pageBreakToken>
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laminiceps
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</taxonomicName>
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Moser, 1919: 16 (orig. desc.);
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<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 227 (check.);
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<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
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: 38 (key);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" pagination="1 - 458" title="A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)." url="https://doi.org/10.11646/zootaxa.211.1.1" volume="211" year="2003">Evans 2003</bibRefCitation>
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: 210 (check.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://digitalcommons.unl.edu/entomologypapers/2" year="2005">Evans and Smith 2005</bibRefCitation>
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: 174 (check.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://museum.unl.edu/research/entomology/SSSA/nwmelos.htm" year="2009">Evans and Smith 2009</bibRefCitation>
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: 178 (check.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="38" pageNumber="39" type="type material">
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<paragraph pageId="38" pageNumber="39">Type material.</paragraph>
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<paragraph pageId="38" pageNumber="39">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys laminiceps" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="laminiceps">Liogenys laminiceps</taxonomicName>
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male holotype (ZMHB): [white printed] "Brasilia/ [handwritten] Sao Paulo", [white handwritten] "
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="laminiceps" pageId="38" pageNumber="39" rank="species" species="laminiceps">laminiceps</taxonomicName>
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/Mos/Typen", [light red printed]
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<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
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, [white printed] "
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="laminiceps" pageId="38" pageNumber="39" rank="species" species="laminiceps">laminiceps</taxonomicName>
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/Mos.", [red printed] "HOLOTYPUS/
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="laminiceps" pageId="38" pageNumber="39" rank="species" species="laminiceps">laminiceps</taxonomicName>
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Moser, 1919/labelled by MNHUB 2013". Genitalia mounted.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="38" pageNumber="39" type="diagnosis">
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<paragraph pageId="38" pageNumber="39">Diagnosis.</paragraph>
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<paragraph pageId="38" pageNumber="39">Body and elytra reddish brown; elongate, widest at posterior third; pronotum darker; clypeal emargination shallow, rounded and wide; outer sides of anterior teeth parallel; clypeal lateral margin convex, with a rounded projection, obtuse angle between outer side of anterior teeth and clypeal lateral projection; antenna 9-articulated; pronotal disc punctures very sparse; pronotal posterior corners sub-angled, obtuse; mesotibia quadrate in cross section; metafemur with thick and erect bristles on posterior margin; pygidium convex; in males, parameres widened from the midline towards the apex, maximum width subapically; apex rounded; inner margins convergent (Fig. 65F).</paragraph>
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<caption pageId="38" pageNumber="39">
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<paragraph pageId="38" pageNumber="39">
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Figure 65.
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys laminiceps" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="laminiceps">Liogenys laminiceps</taxonomicName>
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Moser. A Dorsal view B Lateral view C Frontal view D Clypeus and pronotum E Pygidium F Parameres, dorsal view G Parameres, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="redescription">
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<paragraph pageId="38" pageNumber="39">Redescription.</paragraph>
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<paragraph lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
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Length: 9.8 mm; width: 5.5 mm. Reddish brown. Head: distance between eyes nearly twice the width of one eye; frons shorter than clypeus; clypeus bristled anteriorly; clypeal emargination rounded, shallow and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; clypeal lateral margin convex, with a rounded projection; distance between this projection and anterior tooth shorter than basal width of anterior tooth, distance between clypeal lateral projection and anterior margin of eye shorter than one eye; obtuse angle between outer side of anterior teeth and clypeal lateral projection; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width slightly wider than the apex; fovea deep and elongate, extending to or past the transverse midline of the palpomere; labium transversely carinated, as wide as it is long; antenna 9-articulated, lamellae lighter in color than flagellum and equal in length. Thorax: anterior margin of pronotum straight, flanged throughout; maximum length of pronotum exceeding the length of tarsomeres I, II and III together; disc glabrous, punctures very sparse and coarse; pronotal posterior corners sub-angled, obtuse; proepisternum with long bristles; mesepisternum scaly; sides of metasternum scaly and bristled, few long bristles on the anterior margin; distance between meso- and metacoxae up
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<pageBreakToken pageId="39" pageNumber="40" start="start">to</pageBreakToken>
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twice longer than the metacoxa; scutellum rounded, coarsely punctured. Elytra: shiny, glabrous, uniform reddish brown; elytra more than three times longer than the pronotum; elytral suture and elytron unicolored, distinctly elevated; all four elytral ridges barely noticeable. Legs: procoxa scaly on infra-carinal and outer surface, punctures visible at 12
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; three protibial teeth, middle and apical equal in size, the three teeth equally spaced; protibial inner apical spur present; mesofemural disc setose, with a row of long bristles on anterior and posterior margins; mesotibia quadrate in cross section, disc finely sculptured; two mesotibial transverse carinae, the apical one incomplete; basal apophysis of metacoxa produced beyond the outer margin of trochanter; metafemur with thick and erect bristles on posterior margin; inner margin of metatibia carinated towards apex; inner surface setose; disc finely sculptured; metatibial transverse carina present posteriorly and posterior discontinuous longitudinal carina; metatibial apical spurs of different lengths, the longest equal in length to the diameter of the tibial apex; basal metatarsomere wider than tarsomere II and equal in length; protarsomere II short and wide; pro- and mesotarsomeres I to IV enlarged, protarsomeres slightly wider than the mesotarsomeres and more than twice as wide as metatarsi; claw bifid, symmetrical, superior tooth longer and as wide as the inferior; distance between teeth as long as the inferior tooth. Abdomen: band of scales visible at the lowest magnification beneath the outer margin of elytra; ventrites bristled on disc and sides; propygidium visible, bristled and scaly; pygidium convex, sub-trapezoidal, wide, pygidial width not exceeding distance between spiracles of pro
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<pageBreakToken pageId="40" pageNumber="41" start="start">pygidium</pageBreakToken>
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, pygidial disc bristled throughout; pygidial apex rounded. Parameres: basal region with parameral split at 2/3; parameres widened from the midline towards the apex, maximum width subapically (Fig. 65F); apex rounded; inner margins convergent. In lateral view parameres concave (Fig. 65G).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="type-locality">
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<paragraph pageId="40" pageNumber="41">Type-locality.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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BRAZIL.
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Paulo.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="distribution">
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<paragraph pageId="40" pageNumber="41">Geographical distribution.</paragraph>
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<paragraph pageId="40" pageNumber="41">BRAZIL (SP).</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="remarks">
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<paragraph pageId="40" pageNumber="41">Remarks.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys laminiceps" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="laminiceps">Liogenys laminiceps</taxonomicName>
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is one of the few species with antenna 9-articulated, together with
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<taxonomicName lsidName="L. sinuaticeps" pageId="40" pageNumber="41" rank="species" species="sinuaticeps">L. sinuaticeps</taxonomicName>
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(Fig. 72) and
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<taxonomicName lsidName="L. flavida" pageId="40" pageNumber="41" rank="species" species="flavida">L. flavida</taxonomicName>
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, a southern Argentinian species. The last two species are easily differentiated from
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<taxonomicName lsidName="L. laminiceps" pageId="40" pageNumber="41" rank="species" species="laminiceps">L. laminiceps</taxonomicName>
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because they are yellowish and with sides of body being almost parallel. The female of
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<taxonomicName lsidName="L. laminiceps" pageId="40" pageNumber="41" rank="species" species="laminiceps">L. laminiceps</taxonomicName>
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remains unknown.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |