262 lines
36 KiB
XML
262 lines
36 KiB
XML
<document id="ACFE9633A6BD7DE43FFBBD417C64FEDE" ID-CLB-Dataset="27714" ID-DOI="10.11646/zootaxa.4543.1.1" ID-GBIF-Dataset="22ef92b8-1d9c-4e9f-b01c-ce1f5d3faa7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617607" ID-ZooBank="31ABFA6E-6126-4603-B84F-4BEC7632D1E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927968043" checkinUser="plazi" docAuthor="Fresneda, Javier, Faille, Arnaud, Fery, Hans & Ribera, Ignacio" docDate="2019" docId="03FB8C32F800FFAEFF60F89FFBE948D1" docLanguage="en" docName="zootaxa.4543.1.1.pdf" docOrigin="Zootaxa 4543 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speonemadus Jeannel 1922" docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FFC2F44AF819FFB4FFF7FF98FFAD4B19" masterDocTitle="A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="26" updateTime="1698715325246" updateUser="ExternalLinkService">
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<mods:title id="E74CAC46696C30C7E145701A1C8268C3">A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)</mods:title>
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<mods:namePart id="3E5980C195B918C811BB71A1226330F1">Fresneda, Javier</mods:namePart>
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<mods:namePart id="BEF6229669956FA5C4672E49B78443BE">Faille, Arnaud</mods:namePart>
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<mods:namePart id="5A6DDC99D0DC8ECE7DE244811BA898FF">Fery, Hans</mods:namePart>
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<mods:namePart id="680AE0101F2644B61AA9FC2336A8330A">Ribera, Ignacio</mods:namePart>
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<mods:title id="6C043254C9BFA046DAB02C31B807EF4D">Zootaxa</mods:title>
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<mods:date id="E7055BA2CA7EBDC90E6EFCBADBD1A850">2019</mods:date>
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<mods:number id="AF3C5C44F450F12E033CF663456607A3">4543</mods:number>
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<treatment id="03FB8C32F800FFAEFF60F89FFBE948D1" ID-DOI="http://doi.org/10.5281/zenodo.5927156" ID-GBIF-Taxon="154529610" ID-Zenodo-Dep="5927156" LSID="urn:lsid:plazi:treatment:03FB8C32F800FFAEFF60F89FFBE948D1" httpUri="http://treatment.plazi.org/id/03FB8C32F800FFAEFF60F89FFBE948D1" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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<subSubSection id="C3486EAFF800FFADFF60F89FFE454C38" box="[151,488,1799,1825]" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8BED3D24F800FFADFF60F89FFE454C38" blockId="25.[151,488,1799,1825]" box="[151,488,1799,1825]" pageId="25" pageNumber="26">
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<heading id="D0A58A48F800FFADFF60F89FFE454C38" bold="true" box="[151,488,1799,1825]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="B926E136F800FFADFF60F89FFE454C38" bold="true" box="[151,488,1799,1825]" pageId="25" pageNumber="26">
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Phylogeny of
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<taxonomicName id="4C5246A7F800FFADFEB7F89FFE454C38" ID-CoL="8X7VV" authorityName="Jeannel" authorityYear="1922" box="[320,488,1799,1825]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B926E136F800FFADFEB7F89FFE454C38" bold="true" box="[320,488,1799,1825]" italics="true" pageId="25" pageNumber="26">Speonemadus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3486EAFF800FFAEFF60F8D5FBE948D1" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
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<paragraph id="8BED3D24F800FFAEFF60F8D5FE7A4AC9" blockId="25.[151,1436,1869,2002]" lastBlockId="26.[151,1436,151,968]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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After the description of the two new species and the reinstatement of
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<taxonomicName id="4C5246A7F800FFADFC24F8D6FBE54C7F" authorityName="Kraatz" authorityYear="1870" box="[979,1096,1869,1894]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="B926E136F800FFADFC24F8D6FBE54C7F" box="[979,1096,1869,1894]" italics="true" pageId="25" pageNumber="26">S. gracilis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5246A7F800FFADFB7FF8D6FAB64C7F" box="[1160,1307,1869,1894]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis id="B926E136F800FFADFB7FF8D6FAB64C7F" box="[1160,1307,1869,1894]" italics="true" pageId="25" pageNumber="26">S. pulchellus</emphasis>
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</taxonomicName>
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, the genus
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<taxonomicName id="4C5246A7F800FFADFF60F8E9FE9C4C93" authorityName="Jeannel" authorityYear="1922" box="[151,305,1905,1930]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B926E136F800FFADFF60F8E9FE9C4C93" box="[151,305,1905,1930]" italics="true" pageId="25" pageNumber="26">Speonemadus</emphasis>
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</taxonomicName>
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adds to 16 species, of which our molecular phylogeny includes ten (
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<tableCitation id="C6D0089FF800FFADFBAFF8E9FB164C93" box="[1112,1211,1905,1930]" captionStart="TABLE 1" captionStartId="1.[151,239,908,931]" captionText="TABLE 1. Species recognised in the genus Speonemadus, with the morphological species groups sensu Giachino & Vailati (1993) and Reboleira et al. (2017), and their phylogenetic placement (see Fig. 44). With question marks, species with no molecular data, which phylogenetic placement is hypothesised based on morphological similarities." pageId="25" pageNumber="26">Tables 1</tableCitation>
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,
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<tableCitation id="C6D0089FF800FFADFB3AF8EAFB704C93" box="[1229,1245,1906,1930]" captionStart="TABLE 2" captionStartId="3.[147,233,448,470]" captionText="TABLE 2. Material used in the molecular phylogeny of Speonemadus, with locality data and EMBL accession numbers (in bold, newly obtained sequences)." pageId="25" pageNumber="26">2</tableCitation>
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). Other than the newly described
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<taxonomicName id="4C5246A7F800FFADFEA4F80EFE104CB7" authority="Fresneda & Faille & Fery & Ribera, 2019" authorityName="Fresneda & Faille & Fery & Ribera" authorityYear="2019" box="[339,445,1942,1966]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
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<emphasis id="B926E136F800FFADFEA4F80EFE104CB7" box="[339,445,1942,1966]" italics="true" pageId="25" pageNumber="26">S. comasi</emphasis>
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</taxonomicName>
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<emphasis id="B926E136F800FFADFE33F80DFDAE4CB7" bold="true" box="[452,515,1941,1966]" pageId="25" pageNumber="26">
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<taxonomicNameLabel id="A2155C4DF800FFADFE33F80DFDAE4CB7" box="[452,515,1941,1966]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C5246A7F800FFADFDE7F80EFD584CB7" baseAuthorityName="Murray" baseAuthorityYear="1856" box="[528,757,1942,1966]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="transversostriatus">
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<emphasis id="B926E136F800FFADFDE7F80EFD584CB7" box="[528,757,1942,1966]" italics="true" pageId="25" pageNumber="26">S. transversostriatus</emphasis>
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</taxonomicName>
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is a missing species with an uncertain phylogenetic position, the only member of its species group sensu
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<bibRefCitation id="EFC340D5F800FFADFD49F821FC504CCB" author="Giachino, P. M. & Vailati, D." box="[702,1021,1977,2002]" pageId="25" pageNumber="26" refId="ref13309" refString="Giachino, P. M. & Vailati, D. (1993) Revisione degli Anemadinae. Monografie di " Natura Bresciana " No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino & Vailati (1993)</bibRefCitation>
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||
. These authors did not include
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<taxonomicName id="4C5246A7F800FFAEFA70F822FECC4BA9" baseAuthorityName="Murray" baseAuthorityYear="1856" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="26" lastPageNumber="27" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="transversostriatus">
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<emphasis id="B926E136F800FFAEFA70F822FECC4BA9" italics="true" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">S. transversostriatus</emphasis>
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</taxonomicName>
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in the
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<taxonomicName id="4C5246A7F803FFAEFE41FF0FFDB04BA9" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[438,541,151,176]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B926E136F803FFAEFE41FF0FFDB04BA9" box="[438,541,151,176]" italics="true" pageId="26" pageNumber="27">escalerai</emphasis>
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</taxonomicName>
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-group due to some differences in the sexual dimorphism of males: they have simple protibiae, without the characteristic keel of the species of the
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<taxonomicName id="4C5246A7F803FFAEFC3BFF24FB9E4BCC" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[972,1075,188,213]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B926E136F803FFAEFC3BFF24FB9E4BCC" box="[972,1075,188,213]" italics="true" pageId="26" pageNumber="27">escalerai</emphasis>
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</taxonomicName>
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-group (
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<bibRefCitation id="EFC340D5F803FFAEFB66FF24FA3C4BCD" author="Reboleira, A. S. & Fresneda, J. & Salgado, J. M." box="[1169,1425,188,213]" pageId="26" pageNumber="27" pagination="1 - 23" refId="ref14511" refString="Reboleira, A. S., Fresneda, J. & Salgado, J. M. (2017) A new species of Speonemadus from Portugal with the revision of the escalerai - group (Coleoptera, Leiodidae). European Journal of Taxonomy, 261, 1 - 23. https: // doi. org / 10.5852 / ejt. 2017.261" type="journal article" year="2017">
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Reboleira
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<emphasis id="B926E136F803FFAEFAFCFF25FAED4BCC" box="[1291,1344,188,213]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
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. 2017
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</bibRefCitation>
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). However, all other characters agree with the typical morphology of the
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<taxonomicName id="4C5246A7F803FFAEFC35FF47FB844BE1" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[962,1065,223,248]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B926E136F803FFAEFC35FF47FB844BE1" box="[962,1065,223,248]" italics="true" pageId="26" pageNumber="27">escalerai</emphasis>
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</taxonomicName>
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-group: elytral sculpture of "
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<typeStatus id="54E98386F803FFAEFA9EFF78FA314BE1" box="[1385,1436,224,248]" pageId="26" pageNumber="27">type</typeStatus>
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c", median lobe of the aedeagus bottle-shaped, parameres of the aedeagus only shortly longer than the median lobe (
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<bibRefCitation id="EFC340D5F803FFAEFF69FEBFFE194A59" author="Giachino, P. M. & Vailati, D." box="[158,436,295,320]" pageId="26" pageNumber="27" refId="ref13309" refString="Giachino, P. M. & Vailati, D. (1993) Revisione degli Anemadinae. Monografie di " Natura Bresciana " No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino & Vailati 1993</bibRefCitation>
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), and females with elytral apex more acute than males (strongly acute and with a cleavage in the females of the
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<taxonomicName id="4C5246A7F803FFAEFE7DFED4FE5C4A7C" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[394,497,332,357]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B926E136F803FFAEFE7DFED4FE5C4A7C" box="[394,497,332,357]" italics="true" pageId="26" pageNumber="27">escalerai</emphasis>
|
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</taxonomicName>
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-group,
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<bibRefCitation id="EFC340D5F803FFAEFDBDFED4FD6D4A7D" author="Blas, M." box="[586,704,332,357]" pageId="26" pageNumber="27" pagination="251 - 255" refId="ref12162" refString="Blas, M. (1984) El genero Hormosacus Jeannel, 1936 y su distribucion geografica. (Col. Catopidae). Memoires de Biospeologie, 11, 251 - 255." type="journal article" year="1984">Blas 1984</bibRefCitation>
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). It seems thus likely that
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<taxonomicName id="4C5246A7F803FFAEFC1AFED5FB784A7D" baseAuthorityName="Murray" baseAuthorityYear="1856" box="[1005,1237,333,356]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="transversostriatus">
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<emphasis id="B926E136F803FFAEFC1AFED5FB784A7D" box="[1005,1237,333,356]" italics="true" pageId="26" pageNumber="27">S. transversostriatus</emphasis>
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</taxonomicName>
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is either sister or within the clade formed by the species of the
|
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<taxonomicName id="4C5246A7F803FFAEFD53FEE8FC8A4A91" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[676,807,367,392]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B926E136F803FFAEFD53FEE8FC8A4A91" box="[676,807,367,392]" italics="true" pageId="26" pageNumber="27">S. escalerai</emphasis>
|
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</taxonomicName>
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-group. The other four missing species (
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<taxonomicName id="4C5246A7F803FFAEFB04FEE8FADA4A91" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1267,1399,367,392]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B926E136F803FFAEFB04FEE8FADA4A91" box="[1267,1399,367,392]" italics="true" pageId="26" pageNumber="27">S. escalerai</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C5246A7F803FFAEFA70FEE8FEB54AB4" authorityName="Reboleira, Fresneda & Salgado" authorityYear="2017" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="algarvensis">
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<emphasis id="B926E136F803FFAEFA70FEE8FEB54AB4" italics="true" pageId="26" pageNumber="27">S. algarvensis</emphasis>
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</taxonomicName>
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,
|
||
<taxonomicName id="4C5246A7F803FFAEFEDEFE0DFE794AB4" box="[297,468,404,429]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="orchesioides">
|
||
<emphasis id="B926E136F803FFAEFEDEFE0DFE9A4AB5" box="[297,311,405,428]" italics="true" pageId="26" pageNumber="27">S</emphasis>
|
||
.
|
||
<emphasis id="B926E136F803FFAEFEB0FE0CFE794AB4" box="[327,468,404,429]" italics="true" pageId="26" pageNumber="27">orchesioides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5246A7F803FFAEFDF8FE0DFD1C4AB4" baseAuthorityName="Reiche" baseAuthorityYear="1864" box="[527,689,404,429]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subcostatus">
|
||
<emphasis id="B926E136F803FFAEFDF8FE0DFD1C4AB4" box="[527,689,404,429]" italics="true" pageId="26" pageNumber="27">S. subcostatus</emphasis>
|
||
</taxonomicName>
|
||
) are morphologically very close to some species included in the phylogeny, as already noted.
|
||
</paragraph>
|
||
<paragraph id="8BED3D24F803FFAEFF30FE44FB30480D" blockId="26.[151,1436,151,968]" pageId="26" pageNumber="27">
|
||
Of the species groups defined in
|
||
<bibRefCitation id="EFC340D5F803FFAEFDCEFE44FCCD4AED" author="Giachino, P. M. & Vailati, D." box="[569,864,476,501]" pageId="26" pageNumber="27" refId="ref13309" refString="Giachino, P. M. & Vailati, D. (1993) Revisione degli Anemadinae. Monografie di " Natura Bresciana " No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino & Vailati (1993)</bibRefCitation>
|
||
, we recovered as monophyletic the studied species of the
|
||
<emphasis id="B926E136F803FFAEFF2AFE67FEE04901" box="[221,333,511,536]" italics="true" pageId="26" pageNumber="27">escalerai-</emphasis>
|
||
and
|
||
<taxonomicName id="4C5246A7F803FFAEFE74FE67FDAC4901" authorityName="Heyden" authorityYear="1870" box="[387,513,511,536]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="vandalitiae">
|
||
<emphasis id="B926E136F803FFAEFE74FE67FDAC4901" box="[387,513,511,536]" italics="true" pageId="26" pageNumber="27">vandalitiae</emphasis>
|
||
</taxonomicName>
|
||
-groups. However, according to the characters used to define the
|
||
<taxonomicName id="4C5246A7F803FFAEFB24FE67FAFC4901" authorityName="Heyden" authorityYear="1870" box="[1235,1361,511,536]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="vandalitiae">
|
||
<emphasis id="B926E136F803FFAEFB24FE67FAFC4901" box="[1235,1361,511,536]" italics="true" pageId="26" pageNumber="27">vandalitiae</emphasis>
|
||
</taxonomicName>
|
||
-group the newly described
|
||
<taxonomicName id="4C5246A7F803FFAEFE66FDBDFDA44924" authority="Fresneda & Faille & Fery & Ribera, 2019" authorityName="Fresneda & Faille & Fery & Ribera" authorityYear="2019" box="[401,521,548,573]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
|
||
<emphasis id="B926E136F803FFAEFE66FDBDFDA44924" box="[401,521,548,573]" italics="true" pageId="26" pageNumber="27">S. brusteli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926E136F803FFAEFDE0FDBCFDF04924" bold="true" box="[535,605,548,573]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A2155C4DF803FFAEFDE0FDBCFDF04924" box="[535,605,548,573]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
should have been included in it, but we recovered it as sister to
|
||
<taxonomicName id="4C5246A7F803FFAEFA70FDBDFEB24979" baseAuthorityName="Jeannel" baseAuthorityYear="1936" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="maroccanus">
|
||
<emphasis id="B926E136F803FFAEFA70FDBDFEB24979" italics="true" pageId="26" pageNumber="27">S. maroccanus</emphasis>
|
||
</taxonomicName>
|
||
, which belongs to the
|
||
<taxonomicName id="4C5246A7F803FFAEFDDDFDDFFD314979" baseAuthorityName="Perris" baseAuthorityYear="1864" box="[554,668,583,608]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="clathratus">
|
||
<emphasis id="B926E136F803FFAEFDDDFDDFFD314979" box="[554,668,583,608]" italics="true" pageId="26" pageNumber="27">clathratus</emphasis>
|
||
</taxonomicName>
|
||
-group. Some of the characters used to define species groups were reconstructed as having been independently gained in different species. The thick and blunt setae (the "spine") on the ventral side of the apex of the parameres (
|
||
<figureCitation id="136921A1F803FFAEFD56FD17FC8F49B1" box="[673,802,655,680]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="26" pageNumber="27">Figs 19–26</figureCitation>
|
||
) is a derived state of clade A, present in
|
||
<taxonomicName id="4C5246A7F803FFAEFB00FD08FA3149B1" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[1271,1436,656,680]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="maroccanus">
|
||
<emphasis id="B926E136F803FFAEFB00FD08FAA849BE" box="[1271,1285,656,679]" italics="true" pageId="26" pageNumber="27">S</emphasis>
|
||
.
|
||
<emphasis id="B926E136F803FFAEFAE3FD09FA3149B1" box="[1300,1436,657,680]" italics="true" pageId="26" pageNumber="27">maroccanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136921A1F803FFAEFF68FD2CFF5949D4" box="[159,244,692,717]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="26" pageNumber="27">Fig. 26</figureCitation>
|
||
), in the species of the
|
||
<taxonomicName id="4C5246A7F803FFAEFE00FD2DFDD749D4" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[503,634,692,717]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
|
||
<emphasis id="B926E136F803FFAEFE00FD2DFDD749D4" box="[503,634,692,717]" italics="true" pageId="26" pageNumber="27">S. escalerai</emphasis>
|
||
</taxonomicName>
|
||
-group (
|
||
<figureCitation id="136921A1F803FFAEFD21FD2CFCF549D5" box="[726,856,692,717]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="26" pageNumber="27">Figs 19–23</figureCitation>
|
||
) including
|
||
<taxonomicName id="4C5246A7F803FFAEFC2CFD2DFB6E49D5" baseAuthorityName="Murray" baseAuthorityYear="1856" box="[987,1219,693,716]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="transversostriatus">
|
||
<emphasis id="B926E136F803FFAEFC2CFD2DFB6E49D5" box="[987,1219,693,716]" italics="true" pageId="26" pageNumber="27">S. transversostriatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136921A1F803FFAEFB23FD2CFA8A49D5" box="[1236,1319,692,717]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="26" pageNumber="27">Fig. 24</figureCitation>
|
||
) and in
|
||
<taxonomicName id="4C5246A7F803FFAEFA70FD2DFEA449E9" baseAuthorityName="Perris" baseAuthorityYear="1864" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="clathratus">
|
||
<emphasis id="B926E136F803FFAEFA70FD2DFEA449E9" italics="true" pageId="26" pageNumber="27">S. clathratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136921A1F803FFAEFEEEFD4FFEC349E9" box="[281,366,727,752]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="26" pageNumber="27">Fig. 25</figureCitation>
|
||
). According to our reconstruction the "spine" was secondarily lost in
|
||
<taxonomicName id="4C5246A7F803FFAEFB7CFD40FB5349E9" authority="Fresneda & Faille & Fery & Ribera, 2019" authorityName="Fresneda & Faille & Fery & Ribera" authorityYear="2019" box="[1163,1278,727,752]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
|
||
<emphasis id="B926E136F803FFAEFB7CFD40FB5349E9" box="[1163,1278,727,752]" italics="true" pageId="26" pageNumber="27">S. brusteli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926E136F803FFAEFAF0FD4FFAE549E9" bold="true" box="[1287,1352,727,752]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A2155C4DF803FFAEFAF0FD4FFAE549E9" box="[1287,1352,727,752]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and its likely sister
|
||
<taxonomicName id="4C5246A7F803FFAEFED7FD65FE65480C" box="[288,456,764,789]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="orchesioides">
|
||
<emphasis id="B926E136F803FFAEFED7FD65FE65480C" box="[288,456,764,789]" italics="true" pageId="26" pageNumber="27">S. orchesioides</emphasis>
|
||
</taxonomicName>
|
||
, and it is primarily absent in clade B and in
|
||
<taxonomicName id="4C5246A7F803FFAEFC4FFD64FB80480C" authorityName="Reitter" authorityYear="1885" box="[952,1069,764,789]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B926E136F803FFAEFC4FFD64FB80480C" box="[952,1069,764,789]" italics="true" pageId="26" pageNumber="27">Anemadus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136921A1F803FFAEFBCBFD64FB3C480D" box="[1084,1169,764,789]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="26" pageNumber="27">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BED3D24F803FFAEFF30FC87FBE948D1" blockId="26.[151,1436,151,968]" pageId="26" pageNumber="27">
|
||
The shortening of the parameres from the apparent plesiomorphic condition of being much longer than the median lobe has occurred independently in
|
||
<taxonomicName id="4C5246A7F803FFAEFD65FCDDFC944845" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[658,825,837,860]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="maroccanus">
|
||
<emphasis id="B926E136F803FFAEFD65FCDDFC944845" box="[658,825,837,860]" italics="true" pageId="26" pageNumber="27">S. maroccanus</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="4C5246A7F803FFAEFC55FCDDFB854844" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[930,1064,836,861]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="escalerai">
|
||
<emphasis id="B926E136F803FFAEFC55FCDDFB854844" box="[930,1064,836,861]" italics="true" pageId="26" pageNumber="27">S. escalerai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5246A7F803FFAEFB92FCDDFB5B4844" baseAuthorityName="Perris" baseAuthorityYear="1864" box="[1125,1270,836,861]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="clathratus">
|
||
<emphasis id="B926E136F803FFAEFB92FCDDFB5B4844" box="[1125,1270,836,861]" italics="true" pageId="26" pageNumber="27">S. clathratus</emphasis>
|
||
</taxonomicName>
|
||
-groups. Other characters used by
|
||
<bibRefCitation id="EFC340D5F803FFAEFE98FCFFFD344899" author="Giachino, P. M. & Vailati, D." box="[367,665,871,896]" pageId="26" pageNumber="27" refId="ref13309" refString="Giachino, P. M. & Vailati, D. (1993) Revisione degli Anemadinae. Monografie di " Natura Bresciana " No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino & Vailati (1993)</bibRefCitation>
|
||
, such as the presence of more or less developed striae on the elytral disc, are difficult to define precisely. Despite their usefulness in identifying the species of the genus, our results suggest that the species groups defined within
|
||
<taxonomicName id="4C5246A7F803FFAEFD56FC37FC9648D1" authorityName="Jeannel" authorityYear="1922" box="[673,827,943,968]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B926E136F803FFAEFD56FC37FC9648D1" box="[673,827,943,968]" italics="true" pageId="26" pageNumber="27">Speonemadus</emphasis>
|
||
</taxonomicName>
|
||
are in need of revision.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |