261 lines
40 KiB
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261 lines
40 KiB
XML
<document id="90ACE825352EE2AF5137D5AF07E19604" ID-CLB-Dataset="8080" ID-DOI="10.11646/zootaxa.4927.4.2" ID-GBIF-Dataset="0aa851eb-552a-43c4-b4dd-1dcc0a1297df" ID-ISSN="1175-5326" ID-Zenodo-Dep="4543045" ID-ZooBank="77145FDD-2036-4794-8336-9735F017E1BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613473405370" checkinUser="plazi" docAuthor="Curler, Gregory R. & Skibińska, Kornelia" docDate="2021" docId="03FB87F5FFA0FB44FF08FB517B4B89C0" docLanguage="en" docName="zootaxa.4927.4.2.pdf" docOrigin="Zootaxa 4927 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paleotelmatoscopus madrizi Curler & Skibinska 2021, sp. nov." docType="treatment" docUuid="3137EB10-98D9-4940-A214-6677AA4BE228" docUuidSource="ZooBank" docVersion="8" lastPageNumber="515" masterDocId="FFC2FF8DFFA4FB4EFF9FFFD77E118B31" masterDocTitle="Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species" masterLastPageNumber="524" masterPageNumber="505" pageNumber="509" updateTime="1698929717423" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1FE522A9935727F18B4AB07FBDF21DDA">Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species</mods:title>
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<mods:namePart id="BBAC30CF37C479C743029A2B9DCFF8D2">Curler, Gregory R.</mods:namePart>
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<mods:nameIdentifier id="A12F4D6AE3CC729D39EB1283E8E96E92" type="ORCID">0000-0002-6052-4960</mods:nameIdentifier>
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<mods:affiliation id="FECD28EFA3D14F9CBCC7BCA025447BBB">Mississippi Entomological Museum, Mississippi State University, 100 Old Highway 12, P. O. Drawer 9775, Mississippi State, MS 39762 - 9775, USA. gcurler @ gmail. com; https: // orcid. org / 0000 - 0002 - 6052 - 4960 & Research Collaborator, Smithsonian Institution</mods:affiliation>
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<mods:nameIdentifier id="8B309BCA081CD71D2AD35E8D6F636673" type="email">gcurler@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C103C6791D73180F9A156402AE3BBFF6">Skibińska, Kornelia</mods:namePart>
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<mods:nameIdentifier id="B8606826AD154882A65BF304F095EF43" type="ORCID">0000-0002-5971-9373</mods:nameIdentifier>
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<mods:affiliation id="B0438A64F8ACF62E82AB2AD4AD15AA11">Institute of Systematics and Evolution of Animals Polish Academy of Sciences, Sławkowska street 17, 31 - 016 Kraków, Poland. yukisiak @ gmail. com; https: // orcid. org / 0000 - 0002 - 5971 - 9373</mods:affiliation>
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<mods:nameIdentifier id="255312A5A09FADEB915485887B80A705" type="email">yukisiak@gmail.com</mods:nameIdentifier>
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<mods:title id="46B0458294BC5C4833EB77BA50AD3367">Zootaxa</mods:title>
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<mods:date id="F4C982AB001D91BBD75B946F0B473349">2021</mods:date>
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<mods:number id="D95BA9F4A3C91992C6EF33A51DDB7D0E">2021-02-16</mods:number>
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<mods:number id="916A6050F001712BC8D91C301C032CAE">4927</mods:number>
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<treatment id="03FB87F5FFA0FB44FF08FB517B4B89C0" ID-DOI="http://doi.org/10.5281/zenodo.4543049" ID-GBIF-Taxon="179904898" ID-Zenodo-Dep="4543049" LSID="urn:lsid:zoobank.org:act:3137EB10-98D9-4940-A214-6677AA4BE228" httpUri="http://treatment.plazi.org/id/03FB87F5FFA0FB44FF08FB517B4B89C0" lastPageId="10" lastPageNumber="515" pageId="4" pageNumber="509">
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<emphasis id="B926EAF1FFA0FB4AFF08FB517D5C8F90" bold="true" box="[151,845,1158,1185]" pageId="4" pageNumber="509">
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<taxonomicName id="4C524D60FFA0FB4AFF08FB517CF98F90" ID-CoL="9HM58" authority="Curler & Skibinska" authorityName="Curler & Skibinska" authorityYear="2021" box="[151,744,1158,1185]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="madrizi" status="sp. nov.">
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<emphasis id="B926EAF1FFA0FB4AFF08FB517FF88F90" bold="true" box="[151,489,1158,1185]" italics="true" pageId="4" pageNumber="509">Paleotelmatoscopus madrizi</emphasis>
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Curler & Skibińska
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</taxonomicName>
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<taxonomicNameLabel id="A215578AFFA0FB4AFD70FB507D5C8F90" box="[751,845,1159,1185]" pageId="4" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BED36E3FFA0FB4AFF08FB7C7DAC8FF4" blockId="4.[151,957,1158,1221]" box="[151,957,1195,1221]" pageId="4" pageNumber="509">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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<subSubSection id="C3486568FFA0FB4AFF08FB247D148E3C" box="[151,773,1266,1293]" pageId="4" pageNumber="509" type="reference_group">
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<paragraph id="8BED36E3FFA0FB4AFF08FB247D148E3C" blockId="4.[151,773,1266,1293]" box="[151,773,1266,1293]" pageId="4" pageNumber="509">
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<taxonomicName id="4C524D60FFA0FB4AFF08FB247C048E3C" ID-CoL="89NLW" authority="Meunier, 1905" authorityName="Meunier" authorityYear="1905" box="[151,533,1266,1293]" class="Insecta" family="Psychodidae" genus="Pericoma" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="formosa">
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<emphasis id="B926EAF1FFA0FB4AFF08FB247F768E3C" box="[151,359,1267,1293]" italics="true" pageId="4" pageNumber="509">Pericoma formosa</emphasis>
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<bibRefCitation id="EFC34B12FFA0FB4AFEF1FB247C048E3C" author="Meunier, F." box="[366,533,1266,1293]" pageId="4" pageNumber="509" pagination="235 - 255" refId="ref9212" refString="Meunier, F. (1905) Monographie des Psychodidae de l'Ambere de la Baltique. Annales Musei Nationalis Hungarici, 3, 235 - 255, tabs. VI + VII. [table legends interchanged]" type="journal article" year="1905">Meunier, 1905</bibRefCitation>
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</taxonomicName>
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, in part (see
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<tableCitation id="C6D00358FFA0FB4AFD3DFB247CE58E3C" box="[674,756,1267,1293]" captionStart="TABLE 1" captionStartId="13.[150,238,473,499]" captionTargetPageId="13" captionText="TABLE 1. Data pertaining to specimens included in this study." httpUri="http://table.plazi.org/id/DF2D666BFFA9FB43FF09FE0E7D268AC2" pageId="4" pageNumber="509" tableUuid="DF2D666BFFA9FB43FF09FE0E7D268AC2">Table 1</tableCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3486568FFA0FB4AFF08FAED7CDB8EF0" pageId="4" pageNumber="509" type="diagnosis">
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<paragraph id="8BED36E3FFA0FB4AFF08FAED7CDB8EF0" blockId="4.[151,1437,1338,2019]" pageId="4" pageNumber="509">
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<emphasis id="B926EAF1FFA0FB4AFF08FAED7F1B8E65" bold="true" box="[151,266,1338,1364]" pageId="4" pageNumber="509">Diagnosis</emphasis>
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. Species known only from Eocene Baltic amber. Male with the following combination of character states: head capsule with postocciput lacking corniculi (
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<figureCitation id="13692A66FFA0FB4AFD5BFA887D358E48" box="[708,804,1375,1401]" captionStart="FIGURES 13–14" captionStartId="7.[151,264,1754,1778]" captionTargetBox="[158,1427,465,1727]" captionTargetId="figure-135@7.[151,1436,453,1731]" captionTargetPageId="7" captionText="FIGURES 13–14. Interpretive line drawings of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. 13. Holotype male, frontolateral view [CCHH 1800-3]. 14. Paratype male terminalia, dorsolateral view [CCHH 911-1]. Abbreviations:A1 = first anal vein; asc = ascoid; crn = corniculum; Cu =cubital vein;epd = epandrium; f1 = flagellomere 1; f2 = flagellomere 2; gcx = gonocoxite; gst = gonostylus; hpd = hypandrium; hpr = hypoproct; M1 = first branch of media; M2 = second branch of media; M3 = third branch of media;M4 = fourth branch of media; Mfk = medial fork;ped = pedicel; prm = paramere; R1 = anterior branch of radius; R2 = upper branch of second branch of radius; R3 = lower branch of second branch of radius; R4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius; Rfk = radial fork; Sc, subcostal vein; scp = scape; sur = surstylus; ten = tenacula." figureDoi="http://doi.org/10.5281/zenodo.4543057" httpUri="https://zenodo.org/record/4543057/files/figure.png" pageId="4" pageNumber="509">Figs. 13</figureCitation>
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;
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<figureCitation id="13692A66FFA0FB4AFCAEFA897D6D8E48" box="[817,892,1374,1401]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1867,1891]" captionTargetBox="[283,1304,188,1837]" captionTargetId="figure-19@8.[277,1311,181,1843]" captionTargetPageId="8" captionText="FIGURES 15–20. Photomicrographs of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. specimens. 15. Holotype male, frontolateral view [CCHH 1800-3]. 16. Holotype male, posterolateral view [CCHH 1800-3]. 17. Paratype male, frontolateral view [CCHH 914-1]. 18. Paratype male, dorsolateral view [CCHH 914-1]. 19. Paratype male, frontal view [MZ 14841]. 20. Paratype male, lateral view [MZ 17279].Abbreviations: f1 = flagellomere 1; f2 = flagellomere 2; ped = pedicel; scp = scape. Scale bars: 0.50mm" figureDoi="http://doi.org/10.5281/zenodo.4543059" httpUri="https://zenodo.org/record/4543059/files/figure.png" pageId="4" pageNumber="509">15; 19</figureCitation>
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); antennal scape without prominent mesial lobe at apex (
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<figureCitation id="13692A66FFA0FB4AFF69FA547F458EAC" box="[246,340,1411,1437]" captionStart="FIGURES 13–14" captionStartId="7.[151,264,1754,1778]" captionTargetBox="[158,1427,465,1727]" captionTargetId="figure-135@7.[151,1436,453,1731]" captionTargetPageId="7" captionText="FIGURES 13–14. Interpretive line drawings of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. 13. Holotype male, frontolateral view [CCHH 1800-3]. 14. Paratype male terminalia, dorsolateral view [CCHH 911-1]. Abbreviations:A1 = first anal vein; asc = ascoid; crn = corniculum; Cu =cubital vein;epd = epandrium; f1 = flagellomere 1; f2 = flagellomere 2; gcx = gonocoxite; gst = gonostylus; hpd = hypandrium; hpr = hypoproct; M1 = first branch of media; M2 = second branch of media; M3 = third branch of media;M4 = fourth branch of media; Mfk = medial fork;ped = pedicel; prm = paramere; R1 = anterior branch of radius; R2 = upper branch of second branch of radius; R3 = lower branch of second branch of radius; R4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius; Rfk = radial fork; Sc, subcostal vein; scp = scape; sur = surstylus; ten = tenacula." figureDoi="http://doi.org/10.5281/zenodo.4543057" httpUri="https://zenodo.org/record/4543057/files/figure.png" pageId="4" pageNumber="509">Figs. 13</figureCitation>
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;
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<figureCitation id="13692A66FFA0FB4AFEC0FA557FB88EAC" box="[351,425,1410,1437]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1867,1891]" captionTargetBox="[283,1304,188,1837]" captionTargetId="figure-19@8.[277,1311,181,1843]" captionTargetPageId="8" captionText="FIGURES 15–20. Photomicrographs of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. specimens. 15. Holotype male, frontolateral view [CCHH 1800-3]. 16. Holotype male, posterolateral view [CCHH 1800-3]. 17. Paratype male, frontolateral view [CCHH 914-1]. 18. Paratype male, dorsolateral view [CCHH 914-1]. 19. Paratype male, frontal view [MZ 14841]. 20. Paratype male, lateral view [MZ 17279].Abbreviations: f1 = flagellomere 1; f2 = flagellomere 2; ped = pedicel; scp = scape. Scale bars: 0.50mm" figureDoi="http://doi.org/10.5281/zenodo.4543059" httpUri="https://zenodo.org/record/4543059/files/figure.png" pageId="4" pageNumber="509">15; 17</figureCitation>
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;
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<figureCitation id="13692A66FFA0FB4AFE2AFA547FC58EAC" box="[437,468,1411,1437]" captionStart="FIGURES 21–24" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[172,1413,191,1909]" captionTargetId="figure-19@9.[163,1424,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–24. Photomicrographs of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. specimens. 21. Paratype male, dorsal view [CCHH 911-1]. 22. Paratype female, lateral view [CCHH 499-1]. 23. Paratype female, lateral view [MZ 13274]. 24. Paratype female, lateral view [MZ 19042]. Scale bars: 0.50mm" figureDoi="http://doi.org/10.5281/zenodo.4543063" httpUri="https://zenodo.org/record/4543063/files/figure.png" pageId="4" pageNumber="509">21</figureCitation>
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); flagellomeres approximately equal in width from base to apex; anterior and posterior wing margin evenly convex (
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<figureCitation id="13692A66FFA0FB4AFE41FA707C2D8EF0" box="[478,572,1447,1473]" captionStart="FIGURES 13–14" captionStartId="7.[151,264,1754,1778]" captionTargetBox="[158,1427,465,1727]" captionTargetId="figure-135@7.[151,1436,453,1731]" captionTargetPageId="7" captionText="FIGURES 13–14. Interpretive line drawings of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. 13. Holotype male, frontolateral view [CCHH 1800-3]. 14. Paratype male terminalia, dorsolateral view [CCHH 911-1]. Abbreviations:A1 = first anal vein; asc = ascoid; crn = corniculum; Cu =cubital vein;epd = epandrium; f1 = flagellomere 1; f2 = flagellomere 2; gcx = gonocoxite; gst = gonostylus; hpd = hypandrium; hpr = hypoproct; M1 = first branch of media; M2 = second branch of media; M3 = third branch of media;M4 = fourth branch of media; Mfk = medial fork;ped = pedicel; prm = paramere; R1 = anterior branch of radius; R2 = upper branch of second branch of radius; R3 = lower branch of second branch of radius; R4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius; Rfk = radial fork; Sc, subcostal vein; scp = scape; sur = surstylus; ten = tenacula." figureDoi="http://doi.org/10.5281/zenodo.4543057" httpUri="https://zenodo.org/record/4543057/files/figure.png" pageId="4" pageNumber="509">Figs. 13</figureCitation>
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;
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<figureCitation id="13692A66FFA0FB4AFDD7FA717C838EF0" box="[584,658,1446,1473]" captionStart="FIGURES 15–20" captionStartId="8.[151,264,1867,1891]" captionTargetBox="[283,1304,188,1837]" captionTargetId="figure-19@8.[277,1311,181,1843]" captionTargetPageId="8" captionText="FIGURES 15–20. Photomicrographs of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. specimens. 15. Holotype male, frontolateral view [CCHH 1800-3]. 16. Holotype male, posterolateral view [CCHH 1800-3]. 17. Paratype male, frontolateral view [CCHH 914-1]. 18. Paratype male, dorsolateral view [CCHH 914-1]. 19. Paratype male, frontal view [MZ 14841]. 20. Paratype male, lateral view [MZ 17279].Abbreviations: f1 = flagellomere 1; f2 = flagellomere 2; ped = pedicel; scp = scape. Scale bars: 0.50mm" figureDoi="http://doi.org/10.5281/zenodo.4543059" httpUri="https://zenodo.org/record/4543059/files/figure.png" pageId="4" pageNumber="509">15; 17</figureCitation>
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;
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<figureCitation id="13692A66FFA0FB4AFD02FA707CAD8EF0" box="[669,700,1447,1473]" captionStart="FIGURES 21–24" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[172,1413,191,1909]" captionTargetId="figure-19@9.[163,1424,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–24. Photomicrographs of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. specimens. 21. Paratype male, dorsal view [CCHH 911-1]. 22. Paratype female, lateral view [CCHH 499-1]. 23. Paratype female, lateral view [MZ 13274]. 24. Paratype female, lateral view [MZ 19042]. Scale bars: 0.50mm" figureDoi="http://doi.org/10.5281/zenodo.4543063" httpUri="https://zenodo.org/record/4543063/files/figure.png" pageId="4" pageNumber="509">21</figureCitation>
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).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3486568FFA0FB44FF58FA1D7D318BC8" lastPageId="10" lastPageNumber="515" pageId="4" pageNumber="509" type="description">
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<paragraph id="8BED36E3FFA0FB49FF58FA1D7D6C8A9C" blockId="4.[151,1437,1338,2019]" lastBlockId="7.[151,1436,151,429]" lastPageId="7" lastPageNumber="512" pageId="4" pageNumber="509">
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<emphasis id="B926EAF1FFA0FB4AFF58FA1D7FB28ED4" bold="true" box="[199,419,1482,1509]" pageId="4" pageNumber="509">Description. Male.</emphasis>
|
||
Outline of head round in frontal view, vertex broadly rounded, narrowing abruptly at postocciput; corniculi absent from postocciput; eyebridge comprised of four facet rows, separated by two facet diameters at median, with ventral-most row interrupted by base of scape; interocular suture arched toward vertex, broadly U-shaped; frons, vertex, occiput covered with setae alveoli; frons inconspicuous, bulging slightly between antennae; clypeus with ventral margin convex; labrum inconspicuous, triangular in frontal view; labellum somewhat laterally compressed, narrow compared to clypeus; palp with segments cylindrical, nondescript; apical palp segment elongate, approximately 2.5 times the length of preceding segment; scapus elongate, approximately 2.5 times longer than wide, without mesial lobe at apex; mesial surface of scape covered in dark, spatulate setae; pedicel with basal third cylindrical, apical 2/3 globular, broad compared to node of flagellomere 1; flagellomere 1 fusiform, symmetrical, with node approximately equal in width to those of remaining flagellomeres; flagellomeres 2–13 nodiform, with nodes approximately equal in width, internodes shorter at base and apex of antenna; apical flagellomere with node oblong, with minute subapical projection, apiculus digitiform, setose. Thorax with lateral sclerites poorly visible in most specimens; expandable lobes and other remarkable features absent; midleg with femur longer than that of fore- or hindleg, tibia shorter than that of hindleg; wing with C slightly greater in thickness compared to longitudinal veins; Sc not reaching base of R
|
||
<subScript id="17D634A6FFA0FB4AFD97F8027C308CD2" attach="left" box="[520,545,2005,2019]" fontSize="6" pageId="4" pageNumber="509">2+3</subScript>
|
||
; radial fork placed slightly apical to medial fork. Terminalia with hypandrium band-like, without increase in length at median, not extending beyond anterior margins of gonocoxites; gonopods at point of articulation between gonocoxite and gonostyli approximately equal in width to preceding abdominal segments; gonocoxite with apical 2/3 cylindrical in dorsal and lateral view; gonostylus longer than gonocoxite, acuminate, with pronounced bend at midlength, apical half conspicuously narrowed in comparison to base; parameres membranous, fused into sheath, partially surrounding aedeagus; epandrium wider than long, narrowing from base to apex in dorsoventral view, emarginate posteriorly, with paired aperture ventrally; surstyli elongate, curved dorsally, tapered from base to apex, with cluster of fewer than 10 tenacula inserted dorsoapically; hypoproct conspicuous, tongue-shaped in dorsoventral view, narrowing from base to apex.
|
||
</paragraph>
|
||
<caption id="DF2D666BFFA1FB4BFF08F8407ABE8CC6" ID-DOI="http://doi.org/10.5281/zenodo.4543051" ID-Zenodo-Dep="4543051" httpUri="https://zenodo.org/record/4543051/files/figure.png" pageId="5" pageNumber="510" startId="5.[151,264,1943,1967]" targetBox="[220,1365,192,1911]" targetPageId="5">
|
||
<paragraph id="8BED36E3FFA1FB4BFF08F8407ABE8CC6" blockId="5.[151,1436,1943,2040]" pageId="5" pageNumber="510">
|
||
<emphasis id="B926EAF1FFA1FB4BFF08F8407F2C8C9E" bold="true" box="[151,317,1943,1967]" pageId="5" pageNumber="510">FIGURES 3–6.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C524D60FFA1FB4BFDB8F84F7A118C9E" authority="(Meunier, 1905)" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[551,1024,1943,1968]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="formosa">
|
||
<emphasis id="B926EAF1FFA1FB4BFDB8F84F7D5E8C81" box="[551,847,1944,1968]" italics="true" pageId="5" pageNumber="510">Paleotelmatoscopus formosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC34B12FFA1FB4BFCC0F84F7DE88C9E" author="Meunier, F." box="[863,1017,1943,1968]" pageId="5" pageNumber="510" pagination="235 - 255" refId="ref9212" refString="Meunier, F. (1905) Monographie des Psychodidae de l'Ambere de la Baltique. Annales Musei Nationalis Hungarici, 3, 235 - 255, tabs. VI + VII. [table legends interchanged]" type="journal article" year="1905">Meunier, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
specimens. 3. Male, frontal and partial ventral view [CCHH 1800-1]. 4. Male, caudal and partial dorsal view [CCHH 1800-1]. 5. Male, frontolateral view [CCHH 1800-2]. 6. Male, frontolateral view [MP/3960/2]. Abbreviation: crn = corniculum. Scale bars: 0.50mm
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2D666BFFA2FB48FF08F8EB7AD38CD5" ID-DOI="http://doi.org/10.5281/zenodo.4543053" ID-Zenodo-Dep="4543053" httpUri="https://zenodo.org/record/4543053/files/figure.png" pageId="6" pageNumber="511" startId="6.[151,264,1852,1876]" targetBox="[164,1420,188,1821]" targetPageId="6">
|
||
<paragraph id="8BED36E3FFA2FB48FF08F8EB7AD38CD5" blockId="6.[151,1437,1852,2020]" pageId="6" pageNumber="511">
|
||
<emphasis id="B926EAF1FFA2FB48FF08F8EB7F588C65" bold="true" box="[151,329,1852,1876]" pageId="6" pageNumber="511">FIGURES 7–12.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C524D60FFA2FB48FDAFF8EB7A178C65" authority="(Meunier, 1905)" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[560,1030,1852,1876]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="formosa">
|
||
<emphasis id="B926EAF1FFA2FB48FDAFF8EB7D468C65" box="[560,855,1852,1876]" italics="true" pageId="6" pageNumber="511">Paleotelmatoscopus formosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC34B12FFA2FB48FCF9F8EB7DEE8C65" author="Meunier, F." box="[870,1023,1852,1876]" pageId="6" pageNumber="511" pagination="235 - 255" refId="ref9212" refString="Meunier, F. (1905) Monographie des Psychodidae de l'Ambere de la Baltique. Annales Musei Nationalis Hungarici, 3, 235 - 255, tabs. VI + VII. [table legends interchanged]" type="journal article" year="1905">Meunier, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
specimens. 7. Male, frontal and partial ventral view [MP/3960/1]. 8. Male head and prothorax, frontolateral view [CCHH 1800-2]. 9. Male, frontolateral and partial ventral view [LACM 340459]. 10. Male head, frontal view [MZ 15165]. 11. Female, frontolateral view [MZ 13195]. 12. Male head and thorax, frontolateral view [CCHH 69-12]. Abbreviations: asc = ascoid; crn = corniculum; f1 = flagellomere 1; f2 = flagellomere 2; ped = pedicel; scp = scape. Scale bars: 0.20mm (Figs. 7, 9, 11); 0.50mm (Figs. 8, 10, 12).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2D666BFFA3FB49FF08F90D7F3C8CDE" ID-DOI="http://doi.org/10.5281/zenodo.4543057" ID-Zenodo-Dep="4543057" httpUri="https://zenodo.org/record/4543057/files/figure.png" pageId="7" pageNumber="512" startId="7.[151,264,1754,1778]" targetBox="[158,1427,465,1727]" targetPageId="7">
|
||
<paragraph id="8BED36E3FFA3FB49FF08F90D7F3C8CDE" blockId="7.[151,1436,1754,2031]" pageId="7" pageNumber="512">
|
||
<emphasis id="B926EAF1FFA3FB49FF08F90D7F458DC3" bold="true" box="[151,340,1754,1778]" pageId="7" pageNumber="512">FIGURES 13–14.</emphasis>
|
||
Interpretive line drawings of
|
||
<taxonomicName id="4C524D60FFA3FB49FD1DF90C7A6E8DC2" authority="Curler and Skibinska" authorityName="Curler and Skibinska" authorityYear="2021" box="[642,1151,1755,1779]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="madrizi" status="sp. nov.">
|
||
<emphasis id="B926EAF1FFA3FB49FD1DF90C7DB38DC2" box="[642,930,1755,1779]" italics="true" pageId="7" pageNumber="512">Paleotelmatoscopus madrizi</emphasis>
|
||
Curler and Skibińska
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A215578AFFA3FB49FB1AF90C7AC58DC3" box="[1157,1236,1755,1778]" pageId="7" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
|
||
13. Holotype male, frontolateral view [CCHH 1800-3]. 14. Paratype male terminalia, dorsolateral view [CCHH 911-1]. Abbreviations:A
|
||
<subScript id="17D634A6FFA3FB49FACFF8D97B468C2D" attach="left" box="[1360,1367,1806,1820]" fontSize="6" pageId="7" pageNumber="512">1</subScript>
|
||
= first anal vein; asc = ascoid; crn = corniculum; Cu =cubital vein;epd = epandrium; f1 = flagellomere 1; f2 = flagellomere 2; gcx = gonocoxite; gst = gonostylus; hpd = hypandrium; hpr = hypoproct; M
|
||
<subScript id="17D634A6FFA3FB49FCD0F8817D478C55" attach="left" box="[847,854,1878,1892]" fontSize="6" pageId="7" pageNumber="512">1</subScript>
|
||
= first branch of media; M
|
||
<subScript id="17D634A6FFA3FB49FBFCF8817A7B8C55" attach="left" box="[1123,1130,1878,1892]" fontSize="6" pageId="7" pageNumber="512">2</subScript>
|
||
= second branch of media; M
|
||
<subScript id="17D634A6FFA3FB49FA0AF8817B8D8C55" attach="left" box="[1429,1436,1878,1892]" fontSize="6" pageId="7" pageNumber="512">3</subScript>
|
||
= third branch of media;M
|
||
<subScript id="17D634A6FFA3FB49FE3EF8AD7FB98CB9" attach="left" box="[417,424,1914,1928]" fontSize="6" pageId="7" pageNumber="512">4</subScript>
|
||
= fourth branch of media; Mfk = medial fork;ped = pedicel; prm = paramere; R
|
||
<subScript id="17D634A6FFA3FB49FB58F8AD7ADF8CB9" attach="left" box="[1223,1230,1914,1928]" fontSize="6" pageId="7" pageNumber="512">1</subScript>
|
||
= anterior branch of radius; R
|
||
<subScript id="17D634A6FFA3FB49FF6CF8497EEB8C9D" attach="left" box="[243,250,1950,1964]" fontSize="6" pageId="7" pageNumber="512">2</subScript>
|
||
= upper branch of second branch of radius; R
|
||
<subScript id="17D634A6FFA3FB49FD52F8497CC58C9D" attach="left" box="[717,724,1950,1964]" fontSize="6" pageId="7" pageNumber="512">3</subScript>
|
||
= lower branch of second branch of radius; R
|
||
<subScript id="17D634A6FFA3FB49FB3CF8497ABB8C9D" attach="left" box="[1187,1194,1950,1964]" fontSize="6" pageId="7" pageNumber="512">4</subScript>
|
||
= upper branch of third branch of radius; R
|
||
<subScript id="17D634A6FFA3FB49FEC6F8147F718CE0" attach="left" box="[345,352,1987,2001]" fontSize="6" pageId="7" pageNumber="512">5</subScript>
|
||
= lower branch of third branch of radius; Rfk = radial fork; Sc, subcostal vein; scp = scape; sur = surstylus; ten = tenacula.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2D666BFFACFB46FF08F89C7F718CC2" ID-DOI="http://doi.org/10.5281/zenodo.4543059" ID-Zenodo-Dep="4543059" httpUri="https://zenodo.org/record/4543059/files/figure.png" pageId="8" pageNumber="513" startId="8.[151,264,1867,1891]" targetBox="[283,1304,188,1837]" targetPageId="8">
|
||
<paragraph id="8BED36E3FFACFB46FF08F89C7F718CC2" blockId="8.[151,1437,1867,2035]" pageId="8" pageNumber="513">
|
||
<emphasis id="B926EAF1FFACFB46FF08F89C7F478C52" bold="true" box="[151,342,1867,1891]" pageId="8" pageNumber="513">FIGURES 15–20.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C524D60FFACFB46FDA3F89C7A2E8C52" authority="Curler and Skibinska" authorityName="Curler and Skibinska" authorityYear="2021" box="[572,1087,1867,1891]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="madrizi" status="sp. nov.">
|
||
<emphasis id="B926EAF1FFACFB46FDA3F89C7D4C8C52" box="[572,861,1867,1891]" italics="true" pageId="8" pageNumber="513">Paleotelmatoscopus madrizi</emphasis>
|
||
Curler and Skibińska
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A215578AFFACFB46FBD8F89C7A868C53" box="[1095,1175,1867,1890]" pageId="8" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
|
||
specimens. 15. Holotype male, frontolateral view [CCHH 1800-3]. 16. Holotype male, posterolateral view [CCHH 1800-3]. 17. Paratype male, frontolateral view [CCHH 914-1]. 18. Paratype male, dorsolateral view [CCHH 914-1]. 19. Paratype male, frontal view [MZ 14841]. 20. Paratype male, lateral view [MZ 17279]. Abbreviations: f1 = flagellomere 1; f2 = flagellomere 2; ped = pedicel; scp = scape. Scale bars: 0.50mm
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2D666BFFADFB47FF08F8407D808CC6" ID-DOI="http://doi.org/10.5281/zenodo.4543063" ID-Zenodo-Dep="4543063" httpUri="https://zenodo.org/record/4543063/files/figure.png" pageId="9" pageNumber="514" startId="9.[151,264,1943,1967]" targetBox="[172,1413,191,1909]" targetPageId="9">
|
||
<paragraph id="8BED36E3FFADFB47FF08F8407D808CC6" blockId="9.[151,1437,1943,2040]" pageId="9" pageNumber="514">
|
||
<emphasis id="B926EAF1FFADFB47FF08F8407F478C9E" bold="true" box="[151,342,1943,1967]" pageId="9" pageNumber="514">FIGURES 21–24.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C524D60FFADFB47FDA1F84F7A538C81" authority="Curler and Skibinska" authorityName="Curler and Skibinska" authorityYear="2021" box="[574,1090,1944,1968]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="madrizi" status="sp. nov.">
|
||
<emphasis id="B926EAF1FFADFB47FDA1F84F7D4E8C81" box="[574,863,1944,1968]" italics="true" pageId="9" pageNumber="514">Paleotelmatoscopus madrizi</emphasis>
|
||
Curler and Skibińska
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A215578AFFADFB47FBD4F84F7A8D8C9E" box="[1099,1180,1944,1967]" pageId="9" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
specimens. 21. Paratype male, dorsal view [CCHH 911-1]. 22. Paratype female, lateral view [CCHH 499-1]. 23. Paratype female, lateral view [MZ 13274]. 24. Paratype female, lateral view [MZ 19042]. Scale bars: 0.50mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BED36E3FFAEFB44FF58FF407D318BC8" blockId="10.[151,1437,151,753]" pageId="10" pageNumber="515">
|
||
<emphasis id="B926EAF1FFAEFB44FF58FF407F348B80" bold="true" box="[199,293,151,177]" pageId="10" pageNumber="515">Female.</emphasis>
|
||
Head with eye bridge comprised of four facet rows, with ventral-most row incomplete; scape conical, widening from base to apex, at least twice as long as wide, pedicel globular, flagellomere 1 of similar shape to flagellomeres 2–13; ascoids palmate, with three branches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3486568FFAEFB44FF58FED47D2E8A54" pageId="10" pageNumber="515" type="etymology">
|
||
<paragraph id="8BED36E3FFAEFB44FF58FED47D2E8A54" blockId="10.[151,1437,151,753]" pageId="10" pageNumber="515">
|
||
<emphasis id="B926EAF1FFAEFB44FF58FED47F5F8A2C" bold="true" box="[199,334,259,285]" pageId="10" pageNumber="515">Etymology.</emphasis>
|
||
The specific epithet is a patronym in honor of Isaí Madriz, our friend and colleague who expertly illustrated the male genitalia, revised the habitus illustration and provided many hours of guidance during the process of capturing and processing photomicrographs for this work.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3486568FFAEFB44FF58FEB87A6D8AE0" pageId="10" pageNumber="515" type="materials_examined">
|
||
<paragraph id="8BED36E3FFAEFB44FF58FEB87A438A9C" blockId="10.[151,1437,151,753]" pageId="10" pageNumber="515">
|
||
<emphasis id="B926EAF1FFAEFB44FF58FEB87F678AB9" bold="true" box="[199,374,366,393]" pageId="10" pageNumber="515">Type material.</emphasis>
|
||
<materialsCitation id="3B3A3CBEFFAEFB44FEE2FEB87A438A9C" ID-GBIF-Occurrence="3041468302" collectionCode="SDEI" pageId="10" pageNumber="515" specimenCode="CCHH 1800-3" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="54E98841FFAEFB44FEE2FEB87FFB8AB8" box="[381,490,367,393]" pageId="10" pageNumber="515" type="holotype">Holotype</typeStatus>
|
||
: male,
|
||
<specimenCode id="DBF49E98FFAEFB44FDA5FEB87CF08AB8" box="[570,737,367,393]" collectionCode="CCHH" pageId="10" pageNumber="515">CCHH 1800-3</specimenCode>
|
||
, deposited in
|
||
<collectionCode id="ED43AE26FFAEFB44FCE1FEB87DD18AB8" box="[894,960,367,393]" pageId="10" pageNumber="515">SDEI</collectionCode>
|
||
. For a list of
|
||
<typeStatus id="54E98841FFAEFB44FBC9FEB87AD58AB8" box="[1110,1220,367,393]" pageId="10" pageNumber="515" type="paratype">paratypes</typeStatus>
|
||
, please see
|
||
<tableCitation id="C6D00358FFAEFB44FAD6FEB87B8D8AB8" box="[1353,1436,367,393]" captionStart="TABLE 1" captionStartId="13.[150,238,473,499]" captionTargetPageId="13" captionText="TABLE 1. Data pertaining to specimens included in this study." httpUri="http://table.plazi.org/id/DF2D666BFFA9FB43FF09FE0E7D268AC2" pageId="10" pageNumber="515" tableUuid="DF2D666BFFA9FB43FF09FE0E7D268AC2">Table 1</tableCitation>
|
||
and the corresponding folio in Morphobank: http://morphobank.org/permalink/?F896
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BED36E3FFAEFB44FF58FE617A6D8AE0" blockId="10.[151,1437,151,753]" box="[199,1148,438,465]" pageId="10" pageNumber="515">
|
||
<emphasis id="B926EAF1FFAEFB44FF58FE617FED8AE1" bold="true" box="[199,508,438,464]" pageId="10" pageNumber="515">Other material examined.</emphasis>
|
||
See
|
||
<tableCitation id="C6D00358FFAEFB44FDAEFE607C958AE0" box="[561,644,439,465]" captionStart="TABLE 1" captionStartId="13.[150,238,473,499]" captionTargetPageId="13" captionText="TABLE 1. Data pertaining to specimens included in this study." httpUri="http://table.plazi.org/id/DF2D666BFFA9FB43FF09FE0E7D268AC2" pageId="10" pageNumber="515" tableUuid="DF2D666BFFA9FB43FF09FE0E7D268AC2">Table 1</tableCitation>
|
||
and the corresponding folio in Morphobank.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3486568FFAEFB44FF58FE0D7B4B89C0" pageId="10" pageNumber="515" type="discussion">
|
||
<paragraph id="8BED36E3FFAEFB44FF58FE0D7B4B89C0" blockId="10.[151,1437,151,753]" pageId="10" pageNumber="515">
|
||
<emphasis id="B926EAF1FFAEFB44FF58FE0D7F5E8AC5" bold="true" box="[199,335,474,500]" pageId="10" pageNumber="515">Comments.</emphasis>
|
||
Some specimens of
|
||
<taxonomicName id="4C524D60FFAEFB44FDABFE0C7CBB8AC4" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[564,682,475,501]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="formosa">
|
||
<emphasis id="B926EAF1FFAEFB44FDABFE0C7CBB8AC4" box="[564,682,475,501]" italics="true" pageId="10" pageNumber="515">P. formosa</emphasis>
|
||
</taxonomicName>
|
||
examined by
|
||
<bibRefCitation id="EFC34B12FFAEFB44FCDAFE0C7DFF8AC4" author="Meunier, F." box="[837,1006,474,501]" pageId="10" pageNumber="515" pagination="235 - 255" refId="ref9212" refString="Meunier, F. (1905) Monographie des Psychodidae de l'Ambere de la Baltique. Annales Musei Nationalis Hungarici, 3, 235 - 255, tabs. VI + VII. [table legends interchanged]" type="journal article" year="1905">Meunier (1905</bibRefCitation>
|
||
, p. 244) were described as “Var.”. This is based on the flagellomeres being roughly the same width rather than being progressively more narrow from the base to apex of the antenna. These specimens are regarded here as
|
||
<taxonomicName id="4C524D60FFAEFB44FCE2FDF47AAA890C" authorityName="Curler and Skibinska" authorityYear="2021" box="[893,1211,547,573]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="madrizi">
|
||
<emphasis id="B926EAF1FFAEFB44FCE2FDF47AAA890C" box="[893,1211,547,573]" italics="true" pageId="10" pageNumber="515">Paleotelmatoscopus madrizi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6D00358FFAEFB44FB57FDF47B0D890C" box="[1224,1308,547,573]" captionStart="TABLE 1" captionStartId="13.[150,238,473,499]" captionTargetPageId="13" captionText="TABLE 1. Data pertaining to specimens included in this study." httpUri="http://table.plazi.org/id/DF2D666BFFA9FB43FF09FE0E7D268AC2" pageId="10" pageNumber="515" tableUuid="DF2D666BFFA9FB43FF09FE0E7D268AC2">Table 1</tableCitation>
|
||
). Male and female specimens of
|
||
<taxonomicName id="4C524D60FFAEFB44FE15FD907FEC8950" authorityName="Curler and Skibinska" authorityYear="2021" box="[394,509,583,609]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="madrizi">
|
||
<emphasis id="B926EAF1FFAEFB44FE15FD907FEC8950" box="[394,509,583,609]" italics="true" pageId="10" pageNumber="515">P. madrizi</emphasis>
|
||
</taxonomicName>
|
||
have a shorter scape and smaller wing (i.e. shorter and more narrow), and males have a more narrow head capsule in comparison to
|
||
<taxonomicName id="4C524D60FFAEFB44FD44FDBC7D4289B4" baseAuthorityName="Meunier" baseAuthorityYear="1905" box="[731,851,619,645]" class="Insecta" family="Psychodidae" genus="Paleotelmatoscopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="formosa">
|
||
<emphasis id="B926EAF1FFAEFB44FD44FDBC7D4289B4" box="[731,851,619,645]" italics="true" pageId="10" pageNumber="515">P. formosa</emphasis>
|
||
</taxonomicName>
|
||
. This may be true for females as well, but measurement was not possible for this character. In the line illustration of the
|
||
<typeStatus id="54E98841FFAEFB44FC2CFD587A048998" box="[947,1045,655,681]" pageId="10" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
male (
|
||
<figureCitation id="13692A66FFAEFB44FBFAFD587AA88998" box="[1125,1209,655,681]" captionStart="FIGURES 13–14" captionStartId="7.[151,264,1754,1778]" captionTargetBox="[158,1427,465,1727]" captionTargetId="figure-135@7.[151,1436,453,1731]" captionTargetPageId="7" captionText="FIGURES 13–14. Interpretive line drawings of Paleotelmatoscopus madrizi Curler and Skibińska sp. nov. 13. Holotype male, frontolateral view [CCHH 1800-3]. 14. Paratype male terminalia, dorsolateral view [CCHH 911-1]. Abbreviations:A1 = first anal vein; asc = ascoid; crn = corniculum; Cu =cubital vein;epd = epandrium; f1 = flagellomere 1; f2 = flagellomere 2; gcx = gonocoxite; gst = gonostylus; hpd = hypandrium; hpr = hypoproct; M1 = first branch of media; M2 = second branch of media; M3 = third branch of media;M4 = fourth branch of media; Mfk = medial fork;ped = pedicel; prm = paramere; R1 = anterior branch of radius; R2 = upper branch of second branch of radius; R3 = lower branch of second branch of radius; R4 = upper branch of third branch of radius; R 5 = lower branch of third branch of radius; Rfk = radial fork; Sc, subcostal vein; scp = scape; sur = surstylus; ten = tenacula." figureDoi="http://doi.org/10.5281/zenodo.4543057" httpUri="https://zenodo.org/record/4543057/files/figure.png" pageId="10" pageNumber="515">Fig. 13</figureCitation>
|
||
), ascoids are absent from some flagellomeres; this is to indicate that, on this particular specimen, we did not see ascoids inserted on those flagellomeres. Presumably they were lost prior to, or during preservation of the specimen in the amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |