treatments-xml/data/03/FB/87/03FB87B7FFF6182BFEC030A8FE8A0B11.xml

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<mods:title id="6ADB16772BD27CBF03A6B9DAA60FAA18">Kapsulotaenia chisholmae n. sp. (Cestoda: Proteocephalidae), from Varanus spenceri (Reptilia: Varanidae) in Australia</mods:title>
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<mods:namePart id="E4B52514B37C9BA96ADE9E9505ED35C3">Hugh I. Jones</mods:namePart>
<mods:affiliation id="060D178D9E551D16CD7E7B8E4930F6C7">Discipline of Microbiology, M 502, University of Western Australia, Nedlands, WA 6009, Australia</mods:affiliation>
<mods:nameIdentifier id="AF959C55056B1858095A1A0594EB3A8C" type="email">alain.dechambrier@ville-ge.ch</mods:nameIdentifier>
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<mods:namePart id="1C3D5828859699C8B4EDED7ECDA121A2">Alain de Chambrier</mods:namePart>
<mods:affiliation id="F36F9D9919CBC6CBB547743D29978114">Muséum d’histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<emphasis id="B926EAB3FFF6182CFEC030A8FDCF0CB6" bold="true" box="[275,566,1187,1211]" italics="true" pageId="1" pageNumber="210">Kapsulotaenia chisholmae</emphasis>
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<figureCitation id="13692A24FFF6182CFE4030CFFE010CD0" box="[403,504,1220,1245]" captionStart-0="Figs 1 - 3" captionStart-1="Figs 4 - 7" captionStart-2="Figs 8 - 10" captionStart-3="Fig 11" captionStartId-0="2.[151,192,1938,1959]" captionStartId-1="3.[151,192,1879,1900]" captionStartId-2="4.[151,192,698,719]" captionStartId-3="4.[151,183,1424,1445]" captionTargetBox-0="[197,1395,268,1862]" captionTargetBox-1="[310,1254,266,1825]" captionTargetBox-2="[163,1433,257,644]" captionTargetBox-3="[151,1436,799,1390]" captionTargetId-0="figure@2.[151,1436,257,1862]" captionTargetId-1="figure@3.[204,1383,257,1826]" captionTargetId-2="figure@4.[151,1436,257,645]" captionTargetId-3="figure@4.[148,1440,799,1390]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." captionText-1="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." captionText-2="Figs 8 - 10. Kapsulotaenia chisholmae n. sp. Figs. 8 - 10, QM G 235019. Scanning electron micrographs. (8) Dorsoventral view. (9) Lateral view. (10) Apical view." captionText-3="Fig 11. Kapsulotaenia chisholmae n. sp. Schematic view of the scoleces, showing the different shape of contraction, with rostellum extended, partially and fully retracted." httpUri-0="https://zenodo.org/record/269243/files/figure.png" httpUri-1="https://zenodo.org/record/269244/files/figure.png" httpUri-2="https://zenodo.org/record/269245/files/figure.png" httpUri-3="https://zenodo.org/record/269246/files/figure.png" pageId="1" pageNumber="210">Figs 1-11</figureCitation>
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<emphasis id="B926EAB3FFF6182CFF4430F8FE960D07" bold="true" box="[151,367,1267,1290]" pageId="1" pageNumber="210">Material examined</emphasis>
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:
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, 
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.
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‒ 
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West of
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West of
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, collected 
<date id="FFEC1061FFF6182CFF2D32A6FE800EC8" box="[254,377,1709,1733]" pageId="1" pageNumber="210" value="2000-01-11">
<collectingDate id="EFA8E989FFF6182CFF2D32A6FE800EC8" box="[254,377,1709,1733]" pageId="1" pageNumber="210" value="2000-01-11">01.11.2000</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BED36A1FFF6182CFF4432C0FDE90F0E" blockId="1.[151,756,1267,1796]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFF4432C0FE720EEE" bold="true" box="[151,395,1739,1763]" italics="true" pageId="1" pageNumber="210">Additional specimens:</emphasis>
<materialsCitation id="3B3A3CFCFFF6182CFE4132C6FDF50F0E" collectionCode="QM" pageId="1" pageNumber="210" specimenCode="QM G235019, QM J15694" specimenCount="3">
<specimenCode id="DBF49EDAFFF6182CFE4132C6FDD30EEE" box="[402,554,1740,1765]" pageId="1" pageNumber="210">QM G235019</specimenCode>
; 
<specimenCount id="9D54FD28FFF6182CFDE532C6FD400EEE" box="[566,697,1740,1765]" pageId="1" pageNumber="210" type="generic">3 specimens</specimenCount>
from host 
<specimenCode id="DBF49EDAFFF6182CFF1932E7FEB40F0E" box="[202,333,1772,1796]" pageId="1" pageNumber="210">QM J15694</specimenCode>
; no date recorded
</materialsCitation>
.
</paragraph>
<paragraph id="8BED36A1FFF6182CFF443310FD430F3C" blockId="1.[151,698,1818,1843]" box="[151,698,1818,1843]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFF443310FECB0F3C" bold="true" box="[151,306,1818,1843]" pageId="1" pageNumber="210">
<typeStatus id="54E98803FFF6182CFF443310FF360F3E" box="[151,207,1819,1843]" pageId="1" pageNumber="210">Type</typeStatus>
locality:
</emphasis>
Central-west 
<collectingRegion id="4996F843FFF6182CFE143311FDB10F3C" box="[455,584,1818,1841]" country="Australia" name="Queensland" pageId="1" pageNumber="210">Queensland</collectingRegion>
, 
<collectingCountry id="F3457631FFF6182CFD823311FD4F0F3C" box="[593,694,1818,1841]" name="Australia" pageId="1" pageNumber="210">Australia</collectingCountry>
.
</paragraph>
<paragraph id="8BED36A1FFF6182CFF443341FF150FAC" blockId="1.[151,756,1866,1953]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFF443341FEEE0F6F" bold="true" box="[151,279,1866,1890]" pageId="1" pageNumber="210">
<typeStatus id="54E98803FFF6182CFF443341FF360F6F" box="[151,207,1866,1890]" pageId="1" pageNumber="210">Type</typeStatus>
host:
</emphasis>
<taxonomicName id="4C524D22FFF6182CFEF53341FD090F6F" authority="Lucas &amp; Frost, 1903" authorityName="Lucas &amp; Frost" authorityYear="1903" box="[294,752,1866,1890]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="B926EAB3FFF6182CFEF53341FE100F6C" box="[294,489,1866,1889]" italics="true" pageId="1" pageNumber="210">Varanus spenceri</emphasis>
<bibRefCitation id="EFC34B50FFF6182CFE243341FD090F6F" author="Lucas A. H. S. &amp; Frost C." box="[503,752,1866,1890]" firstAuthor="Lucas" journalOrPublisher="Proceedings of the Royal Society of Victoria" pageId="1" pageNumber="210" pagination="145 - 147" part="15" refId="ref5047" refString="Lucas A. H. S., Frost C. 1903. Description of two new Australian lizards, Varanus spenceri and Diplodactylus bilineatus. Proceedings of the Royal Society of Victoria 15: 145 - 147." title="Description of two new Australian lizards, Varanus spenceri and Diplodactylus bilineatus" type="journal article" year="1903">Lucas &amp; Frost, 1903</bibRefCitation>
</taxonomicName>
. Reptilia, 
<taxonomicName id="4C524D22FFF6182CFF283362FE900F8D" box="[251,361,1897,1920]" class="Reptilia" family="Varanidae" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="210" phylum="Chordata" rank="family">Varanidae</taxonomicName>
. 
<collectingRegion id="4996F843FFF6182CFEA53362FE0D0F8D" box="[374,500,1897,1920]" country="Australia" name="Queensland" pageId="1" pageNumber="210">Queensland</collectingRegion>
Museum accession no. J47127.
</paragraph>
<paragraph id="8BED36A1FFF6182CFF4433BCFDBD0FC3" blockId="1.[151,580,1975,1999]" box="[151,580,1975,1999]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFF4433BCFEAB0FC3" bold="true" box="[151,338,1975,1998]" pageId="1" pageNumber="210">Site of infection:</emphasis>
Upper small intestine.
</paragraph>
<paragraph id="8BED36A1FFF6182CFF4433EDFE810010" blockId="1.[151,756,2021,2078]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFF4433EDFE5C0FF1" bold="true" box="[151,421,2021,2046]" pageId="1" pageNumber="210">Prevalence of infection:</emphasis>
Five out of 14 examined hosts were infected (36%).
</paragraph>
</subSubSection>
<subSubSection id="C348652AFFF6182CFCEC3508FAB90936" pageId="1" pageNumber="210" type="etymology">
<paragraph id="8BED36A1FFF6182CFCEC3508FAB90936" blockId="1.[831,1346,258,315]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFCEC3508FC3E0916" bold="true" box="[831,967,259,283]" pageId="1" pageNumber="210">Etymology:</emphasis>
The species is dedicated to Leslie Chisholm, South Australian Museum, 
<collectingCountry id="F3457631FFF6182CFB043529FAC00934" box="[1239,1337,290,313]" name="Australia" pageId="1" pageNumber="210">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C348652AFFF61829FCEC3559FC350FAB" lastPageId="4" lastPageNumber="213" pageId="1" pageNumber="210" type="description">
<paragraph id="8BED36A1FFF6182CFCEC3559FC280AA0" blockId="1.[831,1437,338,2078]" pageId="1" pageNumber="210">
<emphasis id="B926EAB3FFF6182CFCEC3559FC3C0964" bold="true" box="[831,965,338,361]" pageId="1" pageNumber="210">Description</emphasis>
(based on 
<typeStatus id="54E98803FFF6182CFB803559FB4B0964" box="[1107,1202,338,361]" pageId="1" pageNumber="210">holotype</typeStatus>
and two 
<typeStatus id="54E98803FFF6182CFAFE3558FA610966" box="[1325,1432,339,363]" pageId="1" pageNumber="210">paratypes</typeStatus>
; measurements in micrometres unless otherwise stated): Strobila acraspedote, anapolytic, total length 
<quantity id="4CAA9B44FFF6182CFAB83598FC5D09C6" metricMagnitude="-1" metricUnit="m" metricValue="2.475" metricValueMax="3.15" metricValueMin="1.8" pageId="1" pageNumber="210" unit="mm" value="247.5" valueMax="315.0" valueMin="180.0">180- 315 mm</quantity>
, maximum width 840, body dorsoventrally flattened, early proglottides short and wide, immature proglottides wider than long to longer than wide, mature proglottides approximately twice as long as wide, 1.30 to 1.95 mm long, 715-840 wide, with pregravid and gravid proglottides longer than wide (1.8-2.3 mm long × 755-995 wide and 2.5-3.6 mm long × 875-1090 wide, respectively).
</paragraph>
<paragraph id="8BED36A1FFF6182CFCEC36BDFACB0CDE" blockId="1.[831,1437,338,2078]" pageId="1" pageNumber="210">
Scolex 340-450 wide, with pyramidal anterior retractile apex, almost twice as long as wide when totally everted, with apical organ 
<quantity id="4CAA9B44FFF6182CFBD436FCFB9D0B03" box="[1031,1124,759,783]" metricMagnitude="0" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="1" pageNumber="210" unit="in" value="80.0" valueMax="90.0" valueMin="70.0">70-90 in</quantity>
diameter, and width at base about 130 (
<figureCitation id="13692A24FFF6182CFC6D371CFBFD0B3D" box="[958,1028,791,816]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="1" pageNumber="210">Figs 1</figureCitation>
, 
<figureCitation id="13692A24FFF6182CFBC73713FBBE0B3D" box="[1044,1095,792,816]" captionStart-0="Figs 8 - 10" captionStart-1="Fig 11" captionStartId-0="4.[151,192,698,719]" captionStartId-1="4.[151,183,1424,1445]" captionTargetBox-0="[163,1433,257,644]" captionTargetBox-1="[151,1436,799,1390]" captionTargetId-0="figure@4.[151,1436,257,645]" captionTargetId-1="figure@4.[148,1440,799,1390]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="Figs 8 - 10. Kapsulotaenia chisholmae n. sp. Figs. 8 - 10, QM G 235019. Scanning electron micrographs. (8) Dorsoventral view. (9) Lateral view. (10) Apical view." captionText-1="Fig 11. Kapsulotaenia chisholmae n. sp. Schematic view of the scoleces, showing the different shape of contraction, with rostellum extended, partially and fully retracted." httpUri-0="https://zenodo.org/record/269245/files/figure.png" httpUri-1="https://zenodo.org/record/269246/files/figure.png" pageId="1" pageNumber="210">8-11</figureCitation>
). Four circular suckers, appear to face anteriorly, 
<quantity id="4CAA9B44FFF6182CFBC7373CFB7B0B43" box="[1044,1154,823,846]" metricMagnitude="0" metricUnit="m" metricValue="3.175" metricValueMax="4.064" metricValueMin="2.286" pageId="1" pageNumber="210" unit="in" value="125.0" valueMax="160.0" valueMin="90.0">90-160 in</quantity>
diameter. Rostellum-like retractable (
<figureCitation id="13692A24FFF6182CFC133753FBF70B7C" box="[960,1038,856,881]" captionStart="Fig 11" captionStartId="4.[151,183,1424,1445]" captionTargetBox="[151,1436,799,1390]" captionTargetId="figure@4.[148,1440,799,1390]" captionTargetPageId="4" captionText="Fig 11. Kapsulotaenia chisholmae n. sp. Schematic view of the scoleces, showing the different shape of contraction, with rostellum extended, partially and fully retracted." httpUri="https://zenodo.org/record/269246/files/figure.png" pageId="1" pageNumber="210">Fig. 11</figureCitation>
). Small gladiate spinitriches on neck and proglottides. Proliferation zone about 140 μm wide. Very fine layer of subtegumentary longitudinal muscles, with well-stained subtegumentary cells. Internal longitudinal muscles formed by 8-10 bundles of muscle fibres; bundles absent on lateral sides of proglottides (
<figureCitation id="13692A24FFF6182CFC943011FC5F0C3C" box="[839,934,1050,1073]" captionStart="Figs 4 - 7" captionStartId="3.[151,192,1879,1900]" captionTargetBox="[310,1254,266,1825]" captionTargetId="figure@3.[204,1383,257,1826]" captionTargetPageId="3" captionText="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269244/files/figure.png" pageId="1" pageNumber="210">Figs 4-6</figureCitation>
). Two pairs of osmoregulatory canals situated between testes and vitelline follicles, with dorsal canals slightly medial to ventral ones. Ventral canals thinwalled, 15-18 μm wide, sometimes overlapping vitelline follicles. Dorsal canals thick-walled (surrounded by thin muscle fibres), about 5-10 μm wide (
<figureCitation id="13692A24FFF6182CFB1B30B7FADC0CDE" box="[1224,1317,1212,1235]" captionStart="Figs 4 - 7" captionStartId="3.[151,192,1879,1900]" captionTargetBox="[310,1254,266,1825]" captionTargetId="figure@3.[204,1383,257,1826]" captionTargetPageId="3" captionText="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269244/files/figure.png" pageId="1" pageNumber="210">Figs 4-6</figureCitation>
).
</paragraph>
<paragraph id="8BED36A1FFF6182CFCEC30D6FB920E74" blockId="1.[831,1437,338,2078]" pageId="1" pageNumber="210">
Testes medullary (
<figureCitation id="13692A24FFF6182CFBDC30D7FB820CFE" box="[1039,1147,1244,1269]" captionStart="Figs 4 - 7" captionStartId="3.[151,192,1879,1900]" captionTargetBox="[310,1254,266,1825]" captionTargetId="figure@3.[204,1383,257,1826]" captionTargetPageId="3" captionText="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269244/files/figure.png" pageId="1" pageNumber="210">Figs 4, 5</figureCitation>
), spherical to oval, often slightly elongated (40 × 24), in one or two layers, in two almost separated lateral fields (
<figureCitation id="13692A24FFF6182CFB4B3116FB040D3B" box="[1176,1277,1309,1334]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="1" pageNumber="210">Figs 2, 3</figureCitation>
) total number 89-132, (x=108, n=10); 36-52 preporal testes, 5-12 postporal testes and 44-76 aporal testes. Vas deferens strongly coiled, directed anteriorly, reaching to midline of proglottis, often crossing it (
<figureCitation id="13692A24FFF6182CFB7B3195FAED0DBA" box="[1192,1300,1438,1463]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="1" pageNumber="210">Figs 2, 3</figureCitation>
). Cirrus-sac thick-walled, 175-200 long and 70-85 wide, representing 20-27% (x=24%, n=12) of proglottis width. Cirrus occupies about 40% of length of cirrus-sac. Genital atrium narrow, deep; genital pores irregularly alternating, post-equatorial, situated at 66-77% (x=71%, n=12) of proglottis length (
<figureCitation id="13692A24FFF6182CFC2E326AFBA40E74" box="[1021,1117,1633,1657]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="1" pageNumber="210">Figs 2, 3</figureCitation>
).
</paragraph>
<paragraph id="8BED36A1FFF61829FCEC3289FE7A0EB9" blockId="1.[831,1437,338,2078]" lastBlockId="4.[151,757,1560,2079]" lastPageId="4" lastPageNumber="213" pageId="1" pageNumber="210">
Ovary medullary, bilobed, with dorsal lobes penetrating to cortex (
<figureCitation id="13692A24FFF6182CFC6232AAFC0D0EB5" box="[945,1012,1697,1720]" captionStart="Figs 4 - 7" captionStartId="3.[151,192,1879,1900]" captionTargetBox="[310,1254,266,1825]" captionTargetId="figure@3.[204,1383,257,1826]" captionTargetPageId="3" captionText="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269244/files/figure.png" pageId="1" pageNumber="210">Fig. 6</figureCitation>
), with narrow isthmus and follicles on dorsal side; ovary 385-630 wide, occupying 57-65% (x=62%, n=12) of proglottis width (
<figureCitation id="13692A24FFF6182CFB1A32E9FAD40EF6" box="[1225,1325,1762,1787]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="1" pageNumber="210">Figs 2, 3</figureCitation>
). Relative ovarian size (ratio of ovarian size in relation to that of entire proglottis – see 
<bibRefCitation id="EFC34B50FFF6182CFB953328FA890F31" author="de Chambrier A. &amp; Binh T. T. &amp; Scholz T." box="[1094,1392,1827,1852]" etAl="et al." firstAuthor="de Chambrier" journalOrPublisher="Systematic Parasitology" pageId="1" pageNumber="210" pagination="39 - 50" part="81" refId="ref4401" refString="de Chambrier A., Binh T. T., Scholz T. 2012. Ophiotaenia bungari n. sp. (Cestoda), a parasite of Bungarus fasciatus (Schneider) (Ophidia: Elapidae) from Vietnam, with comments on relative ovarian size as a new and potentially useful diagnostic character for proteocephalidean tapeworms. Systematic Parasitology, 81: 39 - 50." title="Ophiotaenia bungari n. sp. (Cestoda), a parasite of Bungarus fasciatus (Schneider) (Ophidia: Elapidae) from Vietnam, with comments on relative ovarian size as a new and potentially useful diagnostic character for proteocephalidean tapeworms" type="journal article" year="2012">
de Chambrier 
<emphasis id="B926EAB3FFF6182CFB38332FFADE0F37" box="[1259,1319,1827,1851]" italics="true" pageId="1" pageNumber="210">et al.</emphasis>
, 2012
</bibRefCitation>
for methods of measuring) 6.9-7.5% of proglottis size. Vagina anterior (93%) or posterior (7%, n=55) to cirrussac, lined with stained cells in its terminal (distal) part, without observable vaginal sphincter. Mehlis’ gland 
<quantity id="4CAA9B44FFF6182CFAAA33AFFC810FD1" metricMagnitude="0" metricUnit="m" metricValue="1.8415" metricValueMax="2.413" metricValueMin="1.27" pageId="1" pageNumber="210" unit="in" value="72.5" valueMax="95.0" valueMin="50.0">50- 95 in</quantity>
diameter, representing 7-10% of proglottis width. Vitelline follicles oval, small, arranged in two lateral bands on each side of proglottis (
<figureCitation id="13692A24FFF6182CFB7E3C0DFAE90013" box="[1197,1296,2054,2078]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="1" pageNumber="210">Figs 2, 3</figureCitation>
), interrupted on poral side ventrally at level of terminal genitalia (cirrus-sac and vagina). Vitelline follicles do not reach anterior or posterior margin of proglottides, occupying 92-98% of proglottis length on poral side and 92-97% on aporal side (
<figureCitation id="13692A24FFF31829FEC93297FE8F0EB9" box="[282,374,1692,1716]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="4" pageNumber="213">Figs 2-3</figureCitation>
).
</paragraph>
<caption id="DF2D6629FFF5182FFF443399FBC20017" httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="2" pageNumber="211" targetBox="[197,1395,268,1862]" targetPageId="2">
<paragraph id="8BED36A1FFF5182FFF443399FBC20017" blockId="2.[151,1438,1938,2075]" pageId="2" pageNumber="211">
Figs 1-3. 
<taxonomicName id="4C524D22FFF5182FFF293399FE030FAA" authority="Hugh I. Jones &amp; Alain de Chambrier, 2016" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[250,506,1938,1959]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="2" pageNumber="211" phylum="Platyhelminthes" rank="species" species="chisholmae" status="n. sp.">
<emphasis id="B926EAB3FFF5182FFF293399FE030FAA" box="[250,506,1938,1959]" italics="true" pageId="2" pageNumber="211">Kapsulotaenia chisholmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557C8FFF5182FFDD23398FDCC0FA5" box="[513,565,1939,1960]" pageId="2" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
(1) Total lateral view of a scolex (holotype – 
<specimenCode id="DBF49EDAFFF5182FFC203398FB870FAA" box="[1011,1150,1938,1960]" pageId="2" pageNumber="211">QM G235015</specimenCode>
). (2) Dorsal view of a mature proglottis (paratype 
<specimenCode id="DBF49EDAFFF5182FFE1433A4FD610FC8" box="[455,664,1967,1989]" pageId="2" pageNumber="211">MHNG-PLAT-31201</specimenCode>
). (3) Dorsal view of a pregravid proglottis (paratype 
<specimenCode id="DBF49EDAFFF5182FFB1133A4FA6A0FC8" box="[1218,1427,1967,1989]" pageId="2" pageNumber="211">MHNG-PLAT-31201</specimenCode>
). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal.
</paragraph>
</caption>
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<paragraph id="8BED36A1FFF4182EFF44335CFE290014" blockId="3.[151,1439,1879,2073]" pageId="3" pageNumber="212">
Figs 4-7. 
<taxonomicName id="4C524D22FFF4182EFF29335CFE040F61" authority="Hugh I. Jones &amp; Alain de Chambrier, 2016" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[250,509,1879,1900]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="3" pageNumber="212" phylum="Platyhelminthes" rank="species" species="chisholmae" status="n. sp.">
<emphasis id="B926EAB3FFF4182EFF29335CFE040F61" box="[250,509,1879,1900]" italics="true" pageId="3" pageNumber="212">Kapsulotaenia chisholmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557C8FFF4182EFDDB3353FDB90F60" box="[520,576,1880,1901]" pageId="3" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
(4) Transversal section of a proglottis at level of its anterior part (paratype 
<specimenCode id="DBF49EDAFFF4182EFA9E3353FE830F86" pageId="3" pageNumber="212">MHNG-PLAT-31201</specimenCode>
). (5) Transversal section of a proglottis at level of cirrus-sac (paratype 
<specimenCode id="DBF49EDAFFF4182EFBF8337EFB040F86" box="[1067,1277,1909,1931]" pageId="3" pageNumber="212">MHNG-PLAT-31201</specimenCode>
). (6) Transversal section of a proglottis at level of the ovary (paratype 
<specimenCode id="DBF49EDAFFF4182EFD283399FC7F0FAB" box="[763,902,1937,1959]" pageId="3" pageNumber="212">QM G235018</specimenCode>
). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal.
</paragraph>
</caption>
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<paragraph id="8BED36A1FFF31829FF4436B1FDD50AE1" blockId="4.[151,1437,698,750]" pageId="4" pageNumber="213">
Figs 8-10. 
<taxonomicName id="4C524D22FFF31829FED136B1FDFD0AC2" authority="Hugh I. Jones &amp; Alain de Chambrier, 2016" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[258,516,698,719]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="4" pageNumber="213" phylum="Platyhelminthes" rank="species" species="chisholmae" status="n. sp.">
<emphasis id="B926EAB3FFF31829FED136B1FDFD0AC2" box="[258,516,698,719]" italics="true" pageId="4" pageNumber="213">Kapsulotaenia chisholmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557C8FFF31829FDDE36B0FDBC0ADD" box="[525,581,699,720]" pageId="4" pageNumber="213" rank="species">n. sp.</taxonomicNameLabel>
Figs. 8-10, 
<specimenCode id="DBF49EDAFFF31829FD1336B0FCB40AC2" box="[704,845,698,720]" pageId="4" pageNumber="213">QM G235019</specimenCode>
. Scanning electron micrographs. (8) Dorsoventral view. (9) Lateral view. (10) Apical view.
</paragraph>
</caption>
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<paragraph id="8BED36A1FFF31829FF44319BFDA30DCF" blockId="4.[151,1437,1424,1474]" pageId="4" pageNumber="213">
Fig 11. 
<taxonomicName id="4C524D22FFF31829FF39319BFE130DA8" authority="Hugh I. Jones &amp; Alain de Chambrier, 2016" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[234,490,1424,1445]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="4" pageNumber="213" phylum="Platyhelminthes" rank="species" species="chisholmae" status="n. sp.">
<emphasis id="B926EAB3FFF31829FF39319BFE130DA8" box="[234,490,1424,1445]" italics="true" pageId="4" pageNumber="213">Kapsulotaenia chisholmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557C8FFF31829FE22319AFDDF0DAB" box="[497,550,1425,1446]" pageId="4" pageNumber="213" rank="species">n. sp.</taxonomicNameLabel>
Schematic view of the scoleces, showing the different shape of contraction, with rostellum extended, partially and fully retracted.
</paragraph>
</caption>
<paragraph id="8BED36A1FFF31829FF4432B7FA940E7E" blockId="4.[151,757,1560,2079]" lastBlockId="4.[831,1437,1560,1958]" pageId="4" pageNumber="213">
Primordium of uterine stem medullary, already present in immature proglottides as an elongated structure formed by a thick layer of chromophilic cells. Formation of uterus of 
<typeStatus id="54E98803FFF31829FF2F332BFED30F35" box="[252,298,1824,1848]" pageId="4" pageNumber="213">type</typeStatus>
1 according to 
<bibRefCitation id="EFC34B50FFF31829FE033314FD080F35" author="de Chambrier A. &amp; Zehnder, M. &amp; Vaucher C. &amp; Mariaux J." box="[464,753,1823,1848]" etAl="et al." firstAuthor="de Chambrier" journalOrPublisher="Systematic Parasitology" pageId="4" pageNumber="213" pagination="159 - 171" part="57" refId="ref4524" refString="de Chambrier A., Zehnder, M., Vaucher C., Mariaux J. 2004. The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development. Systematic Parasitology 57: 159 - 171." title="The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development" type="journal article" year="2004">
de Chambrier 
<emphasis id="B926EAB3FFF31829FDBA332AFD660F3B" box="[617,671,1823,1848]" italics="true" pageId="4" pageNumber="213">et al.</emphasis>
(2004)
</bibRefCitation>
: uterine stem tipped with conspicuous concentration of numerous intensely stained cells (
<figureCitation id="13692A24FFF31829FD9A336AFD6D0F77" box="[585,660,1889,1914]" captionStart="Figs 1 - 3" captionStartId="2.[151,192,1938,1959]" captionTargetBox="[197,1395,268,1862]" captionTargetId="figure@2.[151,1436,257,1862]" captionTargetPageId="2" captionText="Figs 1 - 3. Kapsulotaenia chisholmae n. sp. (1) Total lateral view of a scolex (holotype – QM G 235015). (2) Dorsal view of a mature proglottis (paratype MHNG-PLAT- 31201). (3) Dorsal view of a pregravid proglottis (paratype MHNG-PLAT- 31201). Abbreviations: ao – apical organ; cc – chromophilic cells; ci – cirrus; cm – circular musculature; cs – cirrus-sac; do – dorsal osmoregulatory canal; du – uterine diverticula; mg – Mehlis’ glands; ov – ovary; te – testes; us – uterine stem; va – vas deferens; vc – vaginal canal; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269243/files/figure.png" pageId="4" pageNumber="213">Fig. 2</figureCitation>
); lumen appears in first mature proglottides. Formation of lateral diverticula begins before appearance of eggs in uterus. Thin-walled lateral diverticula grow in pregravid and gravid proglottides, occupying up to 75% of proglottis width. Gravid proglottides with 300- 
<specimenCount id="9D54FD28FFF31829FDE73C0DFD690012" box="[564,656,2054,2079]" pageId="4" pageNumber="213" type="egg">350 egg</specimenCount>
clusters. In gravid proglottides, ovaries, Mehlis’ gland, testes and vas deferens diminished in size, whereas cirrus-sac still visible. Lateral uterine branches difficult to observe.
</paragraph>
<paragraph id="8BED36A1FFF31829FCEC3275FC350FAB" blockId="4.[831,1437,1560,1958]" pageId="4" pageNumber="213">
8- 
<specimenCount id="9D54FD28FFF31829FC863275FC5E0E9A" box="[853,935,1662,1687]" pageId="4" pageNumber="213" type="egg">13 eggs</specimenCount>
in spherical to oval clusters, 100-125 
<date id="FFEC1061FFF31829FAEB3274FA9E0E98" box="[1336,1383,1662,1686]" pageId="4" pageNumber="213" value="1995-10">x 95</date>
-110, surrounded by smooth, thin membrane (outer envelope) closely apposed to eggs (
<figureCitation id="13692A24FFF31829FB8132CAFB6F0ED7" box="[1106,1174,1729,1754]" captionStart="Figs 4 - 7" captionStartId="3.[151,192,1879,1900]" captionTargetBox="[310,1254,266,1825]" captionTargetId="figure@3.[204,1383,257,1826]" captionTargetPageId="3" captionText="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269244/files/figure.png" pageId="4" pageNumber="213">Fig. 7</figureCitation>
). Eggs spherical, with a three-layered embryophore, with thick external layer, 
<quantity id="4CAA9B44FFF31829FAAA32EFFC800F11" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.143" metricValueMin="0.9398" pageId="4" pageNumber="213" unit="in" value="41.0" valueMax="45.0" valueMin="37.0">37- 45 in</quantity>
diameter, intermediate layer nucleate of irregular shape 
<quantity id="4CAA9B44FFF31829FC5B332CFC110F33" box="[904,1000,1831,1854]" metricMagnitude="-1" metricUnit="m" metricValue="9.525" metricValueMax="10.16" metricValueMin="8.89" pageId="4" pageNumber="213" unit="in" value="37.5" valueMax="40.0" valueMin="35.0">35-40 in</quantity>
diameter, internal layer envelope well developed, 
<quantity id="4CAA9B44FFF31829FC683341FBEC0F6D" box="[955,1045,1865,1890]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="7.112" metricValueMin="5.842" pageId="4" pageNumber="213" unit="in" value="25.5" valueMax="28.0" valueMin="23.0">23-28 in</quantity>
diameter. Oncosphere spherical, 
<quantity id="4CAA9B44FFF31829FAAA3341FC8F0F8F" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="4" pageNumber="213" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">12- 15 in</quantity>
diameter, with three pairs of embryonic hooks, 6-7 long (
<figureCitation id="13692A24FFF31829FCAD3386FC460FAB" box="[894,959,1933,1958]" captionStart="Figs 4 - 7" captionStartId="3.[151,192,1879,1900]" captionTargetBox="[310,1254,266,1825]" captionTargetId="figure@3.[204,1383,257,1826]" captionTargetPageId="3" captionText="Figs 4 - 7. Kapsulotaenia chisholmae n. sp. (4) Transversal section of a proglottis at level of its anterior part (paratype MHNG- PLAT- 31201). (5) Transversal section of a proglottis at level of cirrus-sac (paratype MHNG-PLAT- 31201). (6) Transversal section of a proglottis at level of the ovary (paratype QM G 235018). (7) Egg cluster. Abbreviations: ci – cirrus; cs – cirrussac; do – dorsal osmoregulatory canal; em – tri-layered embryophore; lm – internal longitudinal musculature; oe – outer envelope; on – oncosphere; ov – ovary; st – subtegumental muscle fibres; su – subtegumental cells; te – testes; tg – tegument; ud – uteroduct; ut – uterus; va – vas deferens; vc – vaginal canal; vd – vitelloduct; vi – vitelline follicles; vo – ventral osmoregulatory canal." httpUri="https://zenodo.org/record/269244/files/figure.png" pageId="4" pageNumber="213">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348652AFFF2182BFF44350AFE8A0B11" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214" type="discussion">
<paragraph id="8BED36A1FFF21828FF44350AFEE60915" blockId="5.[151,287,257,280]" box="[151,287,257,280]" pageId="5" pageNumber="214">
<heading id="D0A581CDFFF21828FF44350AFEE60915" allCaps="true" bold="true" box="[151,287,257,280]" fontSize="10" level="1" pageId="5" pageNumber="214" reason="2">
<emphasis id="B926EAB3FFF21828FF44350AFEE60915" bold="true" box="[151,287,257,280]" pageId="5" pageNumber="214">REMARKS</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36A1FFF21828FF443539FE580AA6" blockId="5.[151,757,305,1838]" pageId="5" pageNumber="214">
Due to the fact that the majority of cestodes were recovered from preserved lizards, most specimens of 
<taxonomicName id="4C524D22FFF21828FF443579FED40985" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[151,301,369,393]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="chisholmae">
<emphasis id="B926EAB3FFF21828FF443579FED40985" box="[151,301,369,393]" italics="true" pageId="5" pageNumber="214">K. chisholmae</emphasis>
</taxonomicName>
were not in optimal condition: longitudinal excretory canals were sometimes difficult to distinguish, the uterine lateral diverticula in gravid proglottides were unclear, and the irregular shape and size of the testes may have been an artifact of poor preservation. The specimen collected from a living host in Nelia is better preserved, but does not have the scolex. Nonetheless, several constant features could be identified which readily differentiate this species from the other species of 
<taxonomicName id="4C524D22FFF21828FF443699FECA0AA4" authorityName="Freze" authorityYear="1963" box="[151,307,658,681]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FF443699FECA0AA4" box="[151,307,658,681]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
(
<tableCitation id="C6D0031AFFF21828FE933699FE680AA6" box="[320,401,658,683]" captionStart="Table 1" captionStartId="7.[151,204,636,657]" captionTargetBox="[166,1327,876,2058]" captionText="Table 1. Summary of diagnostic traits in Kapsulotaenia species. Measurements in micrometres (μm) unless otherwise stated. PC = proportion of the length of cirrus-sac to the width of the proglottis; PP = position of the genital pore (cirrus pore) as % of the proglottis length; OV = percentage of the width of the ovary in relation to the width of the proglottis; SL = supplementary layer within the embryophore (see de Chambrier, 2006, p. 91);* see de Chambrier et al. (2012); D = taken from the original drawings; B = measurements taken from cotypes of Beddard (BMNH 1915.5.14.2); R = measurements taken from type specimens of Ratz (MHNG-PLAT-36037)." httpUri="http://table.plazi.org/id/DF2D6629FFF0182AFF443677FB3D0B2C" pageId="5" pageNumber="214" tableUuid="DF2D6629FFF0182AFF443677FB3D0B2C">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BED36A1FFF21828FF4436B9FE840CA1" blockId="5.[151,757,305,1838]" pageId="5" pageNumber="214">
<bibRefCitation id="EFC34B50FFF21828FF4436B9FE700AC4" author="de Chambrier A." box="[151,393,690,713]" firstAuthor="de Chambrier" journalOrPublisher="Systematic Parasitology" pageId="5" pageNumber="214" pagination="83 - 93" part="63" refId="ref4357" refString="de Chambrier A. 2006. Redescription of Kapsulotaenia sandgroundi (Carter, 1943) (Eucestoda: Proteocephalidae: Acanthotaeniinae), a parasite of Varanus komodoensis (Varanoidea: Varanidae) from Indonesia. Systematic Parasitology 63: 83 - 93." title="Redescription of Kapsulotaenia sandgroundi (Carter, 1943) (Eucestoda: Proteocephalidae: Acanthotaeniinae), a parasite of Varanus komodoensis (Varanoidea: Varanidae) from Indonesia" type="journal article" year="2006">de Chambrier (2006)</bibRefCitation>
recognized five species which have been formally described, viz. 
<taxonomicName id="4C524D22FFF21828FD8B36D9FE3C0B06" authority="(Johnston, 1909)" authorityName="Johnston" authorityYear="1909" baseAuthorityName="Johnston" baseAuthorityYear="1909" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">
<emphasis id="B926EAB3FFF21828FD8B36D9FF080B04" italics="true" pageId="5" pageNumber="214">Kapsulotaenia tidswelli</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FEDF36F9FE440B06" author="Johnston T. H." box="[268,445,754,779]" firstAuthor="Johnston" journalOrPublisher="Journal of the Royal Society of New South Wales" pageId="5" pageNumber="214" pagination="103 - 116" part="43" refId="ref4821" refString="Johnston T. H. 1909. On a new reptilian cestode. Journal of the Royal Society of New South Wales 43: 103 - 116." title="On a new reptilian cestode" type="journal article" year="1909">Johnston, 1909</bibRefCitation>
)
</taxonomicName>
from 
<taxonomicName id="4C524D22FFF21828FDCC36FFFD350B06" baseAuthorityName="White" baseAuthorityYear="1790" box="[543,716,756,779]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="varius">
<emphasis id="B926EAB3FFF21828FDCC36FFFD350B06" box="[543,716,756,779]" italics="true" pageId="5" pageNumber="214">Varanus varius</emphasis>
</taxonomicName>
in eastern 
<collectingCountry id="F3457631FFF21828FF273719FEAF0B24" box="[244,342,786,809]" name="Australia" pageId="5" pageNumber="214">Australia</collectingCountry>
(
<bibRefCitation id="EFC34B50FFF21828FEA33719FDDD0B26" author="Johnston T. H." box="[368,548,786,811]" firstAuthor="Johnston" journalOrPublisher="Journal of the Royal Society of New South Wales" pageId="5" pageNumber="214" pagination="103 - 116" part="43" refId="ref4821" refString="Johnston T. H. 1909. On a new reptilian cestode. Journal of the Royal Society of New South Wales 43: 103 - 116." title="On a new reptilian cestode" type="journal article" year="1909">Johnston, 1909</bibRefCitation>
), 
<taxonomicName id="4C524D22FFF21828FD90371FFED40B46" authority="(Carter, 1943)" authorityName="Carter" authorityYear="1943" baseAuthorityName="Carter" baseAuthorityYear="1943" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="sandgroundi">
<emphasis id="B926EAB3FFF21828FD90371FFD0D0B24" box="[579,756,786,811]" italics="true" pageId="5" pageNumber="214">K. sandgroundi</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FF4D3738FEDF0B46" author="Carter W. J." box="[158,294,819,843]" firstAuthor="Carter" journalOrPublisher="Transactions of the American Microscopical Society" pageId="5" pageNumber="214" pagination="301 - 305" part="62" refId="ref4195" refString="Carter W. J. 1943. Proteocephalus sandgroundi, a new tetraphyllidean cestode from an East Indian monitor lizard. Transactions of the American Microscopical Society 62: 301 - 305." title="Proteocephalus sandgroundi, a new tetraphyllidean cestode from an East Indian monitor lizard" type="journal article" year="1943">Carter, 1943</bibRefCitation>
)
</taxonomicName>
from the Komodo dragon, 
<taxonomicName id="4C524D22FFF21828FD83373FFECD0B66" authority="(Carter, 1943)" authorityName="Carter" authorityYear="1943" baseAuthorityName="Carter" baseAuthorityYear="1943" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="komodoensis">
<emphasis id="B926EAB3FFF21828FD83373FFD0D0B44" box="[592,756,818,843]" italics="true" pageId="5" pageNumber="214">V. komodoensis</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FF4D3758FED50B66" author="Carter W. J." box="[158,300,851,875]" firstAuthor="Carter" journalOrPublisher="Transactions of the American Microscopical Society" pageId="5" pageNumber="214" pagination="301 - 305" part="62" refId="ref4195" refString="Carter W. J. 1943. Proteocephalus sandgroundi, a new tetraphyllidean cestode from an East Indian monitor lizard. Transactions of the American Microscopical Society 62: 301 - 305." title="Proteocephalus sandgroundi, a new tetraphyllidean cestode from an East Indian monitor lizard" type="journal article" year="1943">Carter, 1943</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C524D22FFF21828FE90375FFD4C0B66" authority="Schmidt &amp; Kuntz, 1974" authorityName="Schmidt &amp; Kuntz" authorityYear="1974" box="[323,693,851,875]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="frezei">
<emphasis id="B926EAB3FFF21828FE90375FFE590B66" box="[323,416,851,875]" italics="true" pageId="5" pageNumber="214">K. frezei</emphasis>
<bibRefCitation id="EFC34B50FFF21828FE783758FD4C0B66" author="Schmidt G. D. &amp; Kuntz R. E." box="[427,693,851,875]" firstAuthor="Schmidt" journalOrPublisher="Proceedings of the Helminthological Society of Washington" pageId="5" pageNumber="214" pagination="195 - 199" part="41" refId="ref5255" refString="Schmidt G. D., Kuntz R. E. 1974. Tapeworms from Philippine reptiles, with two new species of Proteocephalata. Proceedings of the Helminthological Society of Washington 41: 195 - 199." title="Tapeworms from Philippine reptiles, with two new species of Proteocephalata" type="journal article" year="1974">Schmidt &amp; Kuntz, 1974</bibRefCitation>
</taxonomicName>
from 
<taxonomicName id="4C524D22FFF21828FF44377FFEF30B87" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[151,266,883,907]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="salvator">
<emphasis id="B926EAB3FFF21828FF44377FFEF30B87" box="[151,266,883,907]" italics="true" pageId="5" pageNumber="214">V. salvator</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C524D22FFF21828FEE1377FFF370BA6" authority="Koch, Gaulke &amp; Bohme, 2010" authorityName="Koch, Gaulke &amp; Bohme" authorityYear="2010" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="B926EAB3FFF21828FEE1377FFE270B87" box="[306,478,883,907]" italics="true" pageId="5" pageNumber="214">V. palawanensis</emphasis>
<bibRefCitation id="EFC34B50FFF21828FE353778FF370BA6" author="Koch A. &amp; Gaulke M. &amp; Bohme W." firstAuthor="Koch" journalOrPublisher="Zootaxa" pageId="5" pageNumber="214" pagination="1 - 54" part="2446" refId="ref4917" refString="Koch A., Gaulke M., Bohme W. 2010. Unravelling the underestimated diversity of Philippine water monitor lizards (Squamata: Varanus salvator complex), with the description of two new species and a new subspecies. Zootaxa 2446: 1 - 54." title="Unravelling the underestimated diversity of Philippine water monitor lizards (Squamata: Varanus salvator complex), with the description of two new species and a new subspecies" type="journal article" year="2010">Koch, Gaulke &amp; Böhme, 2010</bibRefCitation>
</taxonomicName>
) in 
<collectingRegion id="4996F843FFF21828FED23798FEA70BA7" box="[257,350,915,938]" country="Philippines" name="Palawan" pageId="5" pageNumber="214">Palawan</collectingRegion>
, 
<collectingCountry id="F3457631FFF21828FEBE3798FE1C0BA7" box="[365,485,915,938]" name="Philippines" pageId="5" pageNumber="214">Philippines</collectingCountry>
(see 
<bibRefCitation id="EFC34B50FFF21828FDF53798FD090BA6" author="Koch A. &amp; Gaulke M. &amp; Bohme W." box="[550,752,915,939]" etAl="et al." firstAuthor="Koch" journalOrPublisher="Zootaxa" pageId="5" pageNumber="214" pagination="1 - 54" part="2446" refId="ref4917" refString="Koch A., Gaulke M., Bohme W. 2010. Unravelling the underestimated diversity of Philippine water monitor lizards (Squamata: Varanus salvator complex), with the description of two new species and a new subspecies. Zootaxa 2446: 1 - 54." title="Unravelling the underestimated diversity of Philippine water monitor lizards (Squamata: Varanus salvator complex), with the description of two new species and a new subspecies" type="journal article" year="2010">
Koch 
<emphasis id="B926EAB3FFF21828FDB8379FFD5F0BA7" box="[619,678,915,939]" italics="true" pageId="5" pageNumber="214">et al.</emphasis>
, 2010
</bibRefCitation>
; 
<bibRefCitation id="EFC34B50FFF21828FF4437B8FE490BC1" author="Schmidt G. D. &amp; Kuntz R. E." box="[151,432,947,972]" firstAuthor="Schmidt" journalOrPublisher="Proceedings of the Helminthological Society of Washington" pageId="5" pageNumber="214" pagination="195 - 199" part="41" refId="ref5255" refString="Schmidt G. D., Kuntz R. E. 1974. Tapeworms from Philippine reptiles, with two new species of Proteocephalata. Proceedings of the Helminthological Society of Washington 41: 195 - 199." title="Tapeworms from Philippine reptiles, with two new species of Proteocephalata" type="journal article" year="1974">Schmidt &amp; Kuntz, 1974</bibRefCitation>
), 
<taxonomicName id="4C524D22FFF21828FE1937BFFD090BC1" authority="(Ratz, 1900)" authorityName="Ratz" authorityYear="1900" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[458,752,948,972]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FE1937BFFDAA0BC1" box="[458,595,948,972]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FDBB37BFFD110BC1" author="Ratz I. von" box="[616,744,948,972]" firstAuthor="Ratz" journalOrPublisher="Comptes rendus des seances de la Societe de biologie et de ses filiales" pageId="5" pageNumber="214" pagination="980 - 981" part="52" refId="ref5188" refString="Ratz I. von. 1900 a. Trois nouveaux cestodes de reptiles. Comptes rendus des seances de la Societe de biologie et de ses filiales 52: 980 - 981." title="Trois nouveaux cestodes de reptiles" type="journal article" year="1900">Ratz, 1900</bibRefCitation>
)
</taxonomicName>
, from 
<taxonomicName id="4C524D22FFF21828FF0737DFFED20BE6" authorityName="Merrem" authorityYear="1820" box="[212,299,980,1003]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="genus">
<emphasis id="B926EAB3FFF21828FF0737DFFED20BE6" box="[212,299,980,1003]" italics="true" pageId="5" pageNumber="214">Varanus</emphasis>
</taxonomicName>
sp. in New 
<collectingCountry id="F3457631FFF21828FE6537D8FDFA0BE7" box="[438,515,979,1002]" name="Guinea-Bissau" pageId="5" pageNumber="214">Guinea</collectingCountry>
(
<bibRefCitation id="EFC34B50FFF21828FDC737DFFD630BE1" author="Ratz I. von" box="[532,666,980,1004]" firstAuthor="Ratz" journalOrPublisher="Comptes rendus des seances de la Societe de biologie et de ses filiales" pageId="5" pageNumber="214" pagination="980 - 981" part="52" refId="ref5188" refString="Ratz I. von. 1900 a. Trois nouveaux cestodes de reptiles. Comptes rendus des seances de la Societe de biologie et de ses filiales 52: 980 - 981." title="Trois nouveaux cestodes de reptiles" type="journal article" year="1900" yearSuffix="a">Ratz, 1900a</bibRefCitation>
, b), and 
<taxonomicName id="4C524D22FFF21828FF4437FFFE4F0C01" authority="(Beddard, 1913)" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[151,438,1011,1036]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FF4437FFFF0D0C06" box="[151,244,1012,1035]" italics="true" pageId="5" pageNumber="214">K. varia</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FEDB37F8FE560C01" author="Beddard F. E." box="[264,431,1011,1036]" firstAuthor="Beddard" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="5" pageNumber="214" pagination="4 - 36" part="1913" refId="ref4160" refString="Beddard F. E. 1913. Contributions to the anatomy and systematic arrangement of the Cestoidea. Proceedings of the Zoological Society of London 1913 (part 1): 4 - 36." title="Contributions to the anatomy and systematic arrangement of the Cestoidea" type="proceedings paper" year="1913">Beddard, 1913</bibRefCitation>
)
</taxonomicName>
from 
<taxonomicName id="4C524D22FFF21828FDD037FFFD9F0C06" baseAuthorityName="White" baseAuthorityYear="1790" box="[515,614,1012,1035]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="varius">
<emphasis id="B926EAB3FFF21828FDD037FFFD9F0C06" box="[515,614,1012,1035]" italics="true" pageId="5" pageNumber="214">V. varius</emphasis>
</taxonomicName>
in 
<collectingCountry id="F3457631FFF21828FD4137F8FD0D0C07" box="[658,756,1011,1034]" name="Australia" pageId="5" pageNumber="214">Australia</collectingCountry>
(
<bibRefCitation id="EFC34B50FFF21828FF4D3018FEBD0C21" author="Beddard F. E." box="[158,324,1043,1068]" firstAuthor="Beddard" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="5" pageNumber="214" pagination="4 - 36" part="1913" refId="ref4160" refString="Beddard F. E. 1913. Contributions to the anatomy and systematic arrangement of the Cestoidea. Proceedings of the Zoological Society of London 1913 (part 1): 4 - 36." title="Contributions to the anatomy and systematic arrangement of the Cestoidea" type="proceedings paper" year="1913">Beddard, 1913</bibRefCitation>
). 
<bibRefCitation id="EFC34B50FFF21828FE8A3018FDE80C27" author="Yamaguti S." bookContentInfo="860 pp." box="[345,529,1043,1066]" firstAuthor="Yamaguti" journalOrPublisher="Interscience Publishers, New York" pageId="5" pageNumber="214" refId="ref5852" refString="Yamaguti S. 1959. Systema Helminthum. Vol. 3, The Cestodes of Vertebrates. Interscience Publishers, New York, 860 pp." title="Systema Helminthum. Vol. 3, The Cestodes of Vertebrates" type="book" year="1959">Yamaguti (1959)</bibRefCitation>
however considered 
<taxonomicName id="4C524D22FFF21828FF44303EFF160C41" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[151,239,1077,1100]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FF44303EFF160C41" box="[151,239,1077,1100]" italics="true" pageId="5" pageNumber="214">K. varia</emphasis>
</taxonomicName>
to be a synonym of 
<taxonomicName id="4C524D22FFF21828FE1D303EFD8D0C47" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[462,628,1075,1100]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="sandgroundi">
<emphasis id="B926EAB3FFF21828FE1D303EFD8D0C47" box="[462,628,1075,1100]" italics="true" pageId="5" pageNumber="214">K. sandgroundi</emphasis>
</taxonomicName>
, an opinion that the present authors do not share. 
<bibRefCitation id="EFC34B50FFF21828FD903058FD0D0C61" author="Johnston T. H." box="[579,756,1107,1132]" firstAuthor="Johnston" journalOrPublisher="Journal of the Royal Society of New South Wales" pageId="5" pageNumber="214" pagination="103 - 116" part="43" refId="ref4821" refString="Johnston T. H. 1909. On a new reptilian cestode. Journal of the Royal Society of New South Wales 43: 103 - 116." title="On a new reptilian cestode" type="journal article" year="1909">Johnston (1909)</bibRefCitation>
commented that the scolex in 
<taxonomicName id="4C524D22FFF21828FE06307EFDAF0C81" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[469,598,1140,1164]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FE06307EFDAF0C81" box="[469,598,1140,1164]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
is very similar to that in 
<taxonomicName id="4C524D22FFF21828FF2E309EFE8F0CA6" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[253,374,1172,1196]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">
<emphasis id="B926EAB3FFF21828FF2E309EFE8F0CA6" box="[253,374,1172,1196]" italics="true" pageId="5" pageNumber="214">K. tidswelli</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BED36A1FFF21828FF4430BFFDF20F23" blockId="5.[151,757,305,1838]" pageId="5" pageNumber="214">
The cestodes recovered from five out of fourteen 
<taxonomicName id="4C524D22FFF21828FF4430DEFEF30CE1" authorityName="Lucas &amp; Frost" authorityYear="1903" box="[151,266,1237,1260]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="B926EAB3FFF21828FF4430DEFEF30CE1" box="[151,266,1237,1260]" italics="true" pageId="5" pageNumber="214">V. spenceri</emphasis>
</taxonomicName>
in the present study resemble 
<taxonomicName id="4C524D22FFF21828FD9A30DEFD140CE6" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[585,749,1236,1260]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="sandgroundi">
<emphasis id="B926EAB3FFF21828FD9A30DEFD140CE6" box="[585,749,1236,1260]" italics="true" pageId="5" pageNumber="214">K. sandgroundi</emphasis>
</taxonomicName>
, as described by 
<bibRefCitation id="EFC34B50FFF21828FE8630FEFE150D00" author="Carter W. J." box="[341,492,1269,1293]" firstAuthor="Carter" journalOrPublisher="Transactions of the American Microscopical Society" pageId="5" pageNumber="214" pagination="301 - 305" part="62" refId="ref4195" refString="Carter W. J. 1943. Proteocephalus sandgroundi, a new tetraphyllidean cestode from an East Indian monitor lizard. Transactions of the American Microscopical Society 62: 301 - 305." title="Proteocephalus sandgroundi, a new tetraphyllidean cestode from an East Indian monitor lizard" type="journal article" year="1943">Carter (1943)</bibRefCitation>
and redescribed by 
<bibRefCitation id="EFC34B50FFF21828FD0830FFFEA50D26" author="de Chambrier A." firstAuthor="de Chambrier" journalOrPublisher="Systematic Parasitology" pageId="5" pageNumber="214" pagination="83 - 93" part="63" refId="ref4357" refString="de Chambrier A. 2006. Redescription of Kapsulotaenia sandgroundi (Carter, 1943) (Eucestoda: Proteocephalidae: Acanthotaeniinae), a parasite of Varanus komodoensis (Varanoidea: Varanidae) from Indonesia. Systematic Parasitology 63: 83 - 93." title="Redescription of Kapsulotaenia sandgroundi (Carter, 1943) (Eucestoda: Proteocephalidae: Acanthotaeniinae), a parasite of Varanus komodoensis (Varanoidea: Varanidae) from Indonesia" type="journal article" year="2006">de Chambrier (2006)</bibRefCitation>
. However, 
<taxonomicName id="4C524D22FFF21828FE04311FFD0A0D26" authority="Hugh I. Jones &amp; Alain de Chambrier, 2016" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[471,755,1300,1323]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="chisholmae" status="n. sp.">
<emphasis id="B926EAB3FFF21828FE04311FFD0A0D26" box="[471,755,1300,1323]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia chisholmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557C8FFF21828FF44313EFF2C0D41" box="[151,213,1333,1356]" pageId="5" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
differs from this species and from 
<taxonomicName id="4C524D22FFF21828FDB4313DFD3B0D41" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[615,706,1333,1357]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FDB4313DFD3B0D41" box="[615,706,1333,1357]" italics="true" pageId="5" pageNumber="214">K. varia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FF44315DFEEF0D66" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[151,278,1364,1389]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">
<emphasis id="B926EAB3FFF21828FF44315DFEEF0D66" box="[151,278,1364,1389]" italics="true" pageId="5" pageNumber="214">K. tidswelli</emphasis>
</taxonomicName>
by the anterior position of the vagina to cirrus-sac. It also differs from 
<taxonomicName id="4C524D22FFF21828FE27317DFDA80D80" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[500,593,1398,1421]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FE27317DFDA80D80" box="[500,593,1398,1421]" italics="true" pageId="5" pageNumber="214">K. varia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C524D22FFF21828FDB0317DFD380D80" authorityName="Schmidt &amp; Kuntz" authorityYear="1974" box="[611,705,1397,1421]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="frezei">
<emphasis id="B926EAB3FFF21828FDB0317DFD380D80" box="[611,705,1397,1421]" italics="true" pageId="5" pageNumber="214">K. frezei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FF44319DFEE50DA0" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[151,284,1429,1453]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FF44319DFEE50DA0" box="[151,284,1429,1453]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
by a different egg number in each cluster (
<quantity id="4CAA9B44FFF21828FF7331BEFF160DC1" box="[160,239,1461,1485]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.302" metricValueMin="2.032" pageId="5" pageNumber="214" unit="in" value="10.5" valueMax="13.0" valueMin="8.0">8-13 in</quantity>
<taxonomicName id="4C524D22FFF21828FF2B31BDFE6A0DC1" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[248,403,1461,1485]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="chisholmae">
<emphasis id="B926EAB3FFF21828FF2B31BDFE6A0DC1" box="[248,403,1461,1485]" italics="true" pageId="5" pageNumber="214">K. chisholmae</emphasis>
</taxonomicName>
versus 12-20, 90-100 and more than 100, respectively), from 
<taxonomicName id="4C524D22FFF21828FE0F31DDFDC10DE0" authorityName="Schmidt &amp; Kuntz" authorityYear="1974" box="[476,568,1493,1517]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="frezei">
<emphasis id="B926EAB3FFF21828FE0F31DDFDC10DE0" box="[476,568,1493,1517]" italics="true" pageId="5" pageNumber="214">K. frezei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FDA231DDFD0D0DE0" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[625,756,1493,1517]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FDA231DDFD0D0DE0" box="[625,756,1493,1517]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
by a different cluster shape (spherical to oval versus banana-shaped in the two latter), from 
<taxonomicName id="4C524D22FFF21828FD98321DFD0D0E21" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[587,756,1557,1581]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="sandgroundi">
<emphasis id="B926EAB3FFF21828FD98321DFD0D0E21" box="[587,756,1557,1581]" italics="true" pageId="5" pageNumber="214">K. sandgroundi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FF10323DFEC20E41" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[195,315,1589,1613]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">
<emphasis id="B926EAB3FFF21828FF10323DFEC20E41" box="[195,315,1589,1613]" italics="true" pageId="5" pageNumber="214">K. tidswelli</emphasis>
</taxonomicName>
by a greater diameter of the embryophore (37-45 μm versus respectively 25-30 μm and 19-32 μm); from 
<taxonomicName id="4C524D22FFF21828FF00327CFE800E81" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[211,377,1653,1678]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="sandgroundi">
<emphasis id="B926EAB3FFF21828FF00327CFE800E81" box="[211,377,1653,1678]" italics="true" pageId="5" pageNumber="214">K. sandgroundi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FE7D327CFDFE0E83" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[430,519,1655,1678]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FE7D327CFDFE0E83" box="[430,519,1655,1678]" italics="true" pageId="5" pageNumber="214">K. varia</emphasis>
</taxonomicName>
(
<emphasis id="B926EAB3FFF21828FDC4327CFDAB0E83" box="[535,594,1655,1678]" italics="true" pageId="5" pageNumber="214">sensu</emphasis>
<bibRefCitation id="EFC34B50FFF21828FD89327EFD0D0E83" author="Nybelin O." box="[602,756,1653,1678]" firstAuthor="Nybelin" journalOrPublisher="Kungliga Svenska Vetenskapsakademiens Handlingar" pageId="5" pageNumber="214" pagination="1 - 48" part="52" refId="ref5120" refString="Nybelin O. 1917. Australischen Cestoden. Results of D. E. Mjoeberg's Swedish scientific expeditions to Australia, 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar 52 (14): 1 - 48." title="Australischen Cestoden. Results of D. E. Mjoeberg's Swedish scientific expeditions to Australia, 1910 - 1913" type="journal article" year="1917">Nybelin, 1917</bibRefCitation>
– see below), by the absence of a vaginal sphincter. Finally, 
<taxonomicName id="4C524D22FFF21828FF3D32BCFE7C0EC1" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[238,389,1717,1742]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="chisholmae">
<emphasis id="B926EAB3FFF21828FF3D32BCFE7C0EC1" box="[238,389,1717,1742]" italics="true" pageId="5" pageNumber="214">K. chisholmae</emphasis>
</taxonomicName>
differs from 
<emphasis id="B926EAB3FFF21828FDC232BCFF160EE3" italics="true" pageId="5" pageNumber="214">
<taxonomicName id="4C524D22FFF21828FDC232BCFD930EC3" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[529,618,1719,1742]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">K. varia</taxonomicName>
, 
<taxonomicName id="4C524D22FFF21828FDA732BCFD090EC1" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[628,752,1717,1742]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">K. tidswelli</taxonomicName>
, 
<taxonomicName id="4C524D22FFF21828FF4432DCFF160EE3" authorityName="Schmidt &amp; Kuntz" authorityYear="1974" box="[151,239,1750,1774]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="frezei">K. frezei</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C524D22FFF21828FEF232DCFE580EE3" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[289,417,1750,1774]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FEF232DCFE580EE3" box="[289,417,1750,1774]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
by a larger length of the strobila (up to 
<quantity id="4CAA9B44FFF21828FF3132FDFEC50F03" box="[226,316,1782,1806]" metricMagnitude="-1" metricUnit="m" metricValue="3.15" pageId="5" pageNumber="214" unit="mm" value="315.0">315 mm</quantity>
versus 
<quantity id="4CAA9B44FFF21828FE4032FDFE1B0F03" box="[403,482,1782,1806]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="214" unit="mm" value="30.0">30 mm</quantity>
, 
<quantity id="4CAA9B44FFF21828FE2332FDFD9A0F03" box="[496,611,1782,1806]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="5" pageNumber="214" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27-30 mm</quantity>
, 
<quantity id="4CAA9B44FFF21828FDA332FDFD390F03" box="[624,704,1782,1806]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="214" unit="mm" value="40.0">40 mm</quantity>
, and 
<quantity id="4CAA9B44FFF21828FF44331DFEF10F23" box="[151,264,1814,1838]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="5" pageNumber="214" unit="mm" value="25.0" valueMax="40.0" valueMin="10.0">10-40 mm</quantity>
, respectively) (
<tableCitation id="C6D0031AFFF21828FE79331DFE020F23" box="[426,507,1814,1838]" captionStart="Table 1" captionStartId="7.[151,204,636,657]" captionTargetBox="[166,1327,876,2058]" captionText="Table 1. Summary of diagnostic traits in Kapsulotaenia species. Measurements in micrometres (μm) unless otherwise stated. PC = proportion of the length of cirrus-sac to the width of the proglottis; PP = position of the genital pore (cirrus pore) as % of the proglottis length; OV = percentage of the width of the ovary in relation to the width of the proglottis; SL = supplementary layer within the embryophore (see de Chambrier, 2006, p. 91);* see de Chambrier et al. (2012); D = taken from the original drawings; B = measurements taken from cotypes of Beddard (BMNH 1915.5.14.2); R = measurements taken from type specimens of Ratz (MHNG-PLAT-36037)." httpUri="http://table.plazi.org/id/DF2D6629FFF0182AFF443677FB3D0B2C" pageId="5" pageNumber="214" tableUuid="DF2D6629FFF0182AFF443677FB3D0B2C">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BED36A1FFF21828FF44337DFEC30F80" blockId="5.[151,314,1910,1933]" box="[151,314,1910,1933]" pageId="5" pageNumber="214">
<heading id="D0A581CDFFF21828FF44337DFEC30F80" allCaps="true" bold="true" box="[151,314,1910,1933]" fontSize="10" level="1" pageId="5" pageNumber="214" reason="2">
<emphasis id="B926EAB3FFF21828FF44337DFEC30F80" bold="true" box="[151,314,1910,1933]" pageId="5" pageNumber="214">DISCUSSION</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36A1FFF21828FF4433AEFBF70C91" blockId="5.[151,756,1957,2078]" lastBlockId="5.[831,1437,258,2078]" pageId="5" pageNumber="214">
Within the cestode order 
<taxonomicName id="4C524D22FFF21828FE7633AEFD0D0FB1" authority=", Freze (1963)" authorityName="Freze" authorityYear="1963" box="[421,756,1957,1981]" class="Cestoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="order">
Proteocephalidea, 
<bibRefCitation id="EFC34B50FFF21828FDB833ADFD0D0FB1" author="Freze V. I." box="[619,756,1957,1981]" firstAuthor="Freze" journalOrPublisher="Tezisy Konferentsii VOG" pageId="5" pageNumber="214" pagination="155 - 157" refId="ref4697" refString="Freze V. I. 1963. Brief analysis of the cestode system of suborder Proteocephalata. Tezisy Konferentsii VOG 2: 155 - 157. [in Russian]" title="Brief analysis of the cestode system of subor- der Proteocephalata" type="book chapter" year="1963">Freze (1963)</bibRefCitation>
</taxonomicName>
erected the subfamilyAcanthotaeniinae, with three genera 
<taxonomicName id="4C524D22FFF21828FF4433EEFE170FF3" authority="Linstow, 1903" authorityName="Linstow" authorityYear="1903" box="[151,494,2021,2046]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FF4433EEFECA0FF1" box="[151,307,2021,2044]" italics="true" pageId="5" pageNumber="214">Acanthotaenia</emphasis>
<bibRefCitation id="EFC34B50FFF21828FE9533EEFE170FF3" author="Linstow O." box="[326,494,2021,2046]" firstAuthor="Linstow" journalOrPublisher="Zentralblatt fur Bakteriologie und Parasitologie" pageId="5" pageNumber="214" pagination="532 - 535" part="33" refId="ref5025" refString="Linstow O. 1903. Drei neue Tanien aus Ceylon. Zentralblatt fur Bakteriologie und Parasitologie 33: 532 - 535." title="Drei neue Tanien aus Ceylon" type="journal article" year="1903">Linstow, 1903</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C524D22FFF21828FDD633EEFF350010" authority="Freze, 1963" authorityName="Freze" authorityYear="1963" class="Cestoda" family="Proteocephalidae" genus="Rostellotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FDD633EEFD660FF1" box="[517,671,2021,2044]" italics="true" pageId="5" pageNumber="214">Rostellotaenia</emphasis>
<bibRefCitation id="EFC34B50FFF21828FD6133ECFF350010" author="Freze V. I." firstAuthor="Freze" journalOrPublisher="Tezisy Konferentsii VOG" pageId="5" pageNumber="214" pagination="155 - 157" refId="ref4697" refString="Freze V. I. 1963. Brief analysis of the cestode system of suborder Proteocephalata. Tezisy Konferentsii VOG 2: 155 - 157. [in Russian]" title="Brief analysis of the cestode system of subor- der Proteocephalata" type="book chapter" year="1963">Freze, 1963</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FED13C0DFDDF0010" authority="Freze, 1963" authorityName="Freze" authorityYear="1963" box="[258,550,2054,2078]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FED13C0DFE670010" box="[258,414,2054,2077]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
<bibRefCitation id="EFC34B50FFF21828FE773C0CFDDF0010" author="Freze V. I." box="[420,550,2054,2078]" firstAuthor="Freze" journalOrPublisher="Tezisy Konferentsii VOG" pageId="5" pageNumber="214" pagination="155 - 157" refId="ref4697" refString="Freze V. I. 1963. Brief analysis of the cestode system of suborder Proteocephalata. Tezisy Konferentsii VOG 2: 155 - 157. [in Russian]" title="Brief analysis of the cestode system of subor- der Proteocephalata" type="book chapter" year="1963">Freze, 1963</bibRefCitation>
</taxonomicName>
, all of which occur in monitors (
<taxonomicName id="4C524D22FFF21828FC1F3508FBDA0917" authorityName="Merrem" authorityYear="1820" box="[972,1059,259,282]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="genus">
<emphasis id="B926EAB3FFF21828FC1F3508FBDA0917" box="[972,1059,259,282]" italics="true" pageId="5" pageNumber="214">Varanus</emphasis>
</taxonomicName>
spp.) (
<bibRefCitation id="EFC34B50FFF21828FBA33508FB0C0914" author="Freze V. I." bookContentInfo="597 pp." box="[1136,1269,258,283]" firstAuthor="Freze" journalOrPublisher="Israel Program for Scientific Translation" pageId="5" pageNumber="214" refId="ref4728" refString="Freze V. I. 1965. Proteocephalata in fish, amphibians and reptiles. Essentials of Cestodology, vol. 5, ed: K. I. Skrjabin. Izdatel'stvo ' Nauka', Moscow 1965, 597 pp. Israel Program for Scientific Translation, 1969. [Translated from Russian]" title="Proteocephalata in fish, amphibians and reptiles" type="book" volumeTitle="Essentials of Cestodology, vol. 5, ed: K. I. Skrjabin. Izdatel'stvo ' Nauka', Moscow 1965" year="1965">Freze, 1965</bibRefCitation>
). 
<bibRefCitation id="EFC34B50FFF21828FADA3509FBCF0936" author="de Chambrier S. &amp; de Chambrier A." firstAuthor="de Chambrier" journalOrPublisher="Folia Parasitologica" pageId="5" pageNumber="214" pagination="263 - 279" part="57" refId="ref4575" refString="de Chambrier S., de Chambrier A. 2010. Two new genera and two new species of proteocephalidean tapeworms (Eucestoda) from reptiles and amphibians in Australia. Folia Parasitologica, 57: 263 - 279." title="Two new genera and two new species of proteocephalidean tapeworms (Eucestoda) from reptiles and amphibians in Australia" type="journal article" year="2010">de Chambrier &amp; de Chambrier (2010)</bibRefCitation>
described another acanthotaenine genus, 
<emphasis id="B926EAB3FFF21828FC5F3549FBD90954" box="[908,1056,322,345]" italics="true" pageId="5" pageNumber="214">Vandiermenia</emphasis>
, a parasite of the red-bellied snake 
<taxonomicName id="4C524D22FFF21828FCEC3569FBB70974" baseAuthorityName="Shaw" baseAuthorityYear="1794" box="[831,1102,354,377]" class="Reptilia" family="Elapidae" genus="Pseudechis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="porphyriacus">
<emphasis id="B926EAB3FFF21828FCEC3569FBB70974" box="[831,1102,354,377]" italics="true" pageId="5" pageNumber="214">Pseudechis porphyriacus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F3457631FFF21828FB433569FB0C0974" box="[1168,1269,354,377]" name="Australia" pageId="5" pageNumber="214">Australia</collectingCountry>
. 
<taxonomicName id="4C524D22FFF21828FAD23569FA640974" authorityName="Freze" authorityYear="1963" box="[1281,1437,354,377]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FAD23569FA640974" box="[1281,1437,354,377]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
is characterized principally by the presence of eggs in clusters in the gravid proglottides. Based on molecular data, 
<bibRefCitation id="EFC34B50FFF21828FCAF35C9FB5B09D6" author="de Chambrier A. &amp; Zehnder, M. &amp; Vaucher C. &amp; Mariaux J." box="[892,1186,450,475]" etAl="et al." firstAuthor="de Chambrier" journalOrPublisher="Systematic Parasitology" pageId="5" pageNumber="214" pagination="159 - 171" part="57" refId="ref4524" refString="de Chambrier A., Zehnder, M., Vaucher C., Mariaux J. 2004. The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development. Systematic Parasitology 57: 159 - 171." title="The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development" type="journal article" year="2004">
de Chambrier 
<emphasis id="B926EAB3FFF21828FBC935CFFBB209D4" box="[1050,1099,450,475]" italics="true" pageId="5" pageNumber="214">et al</emphasis>
. (2004)
</bibRefCitation>
separated four species in this genus in 
<collectingCountry id="F3457631FFF21828FC2335E9FBAC09F4" box="[1008,1109,482,505]" name="Australia" pageId="5" pageNumber="214">Australia</collectingCountry>
, with one species in 
<taxonomicName id="4C524D22FFF21828FAEE35EFFC220A16" authority="(White, 1790)" authorityName="White" authorityYear="1790" baseAuthorityName="White" baseAuthorityYear="1790" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="varius">
<emphasis id="B926EAB3FFF21828FAEE35EFFA6409F6" box="[1341,1437,484,507]" italics="true" pageId="5" pageNumber="214">V. varius</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FC943609FC2D0A16" author="White J." bookContentInfo="229 pp." box="[839,980,514,539]" firstAuthor="White" journalOrPublisher="Debrett, London" pageId="5" pageNumber="214" refId="ref5785" refString="White J. 1790. Journal of a voyage to New South Wales, with sixty-five plates of non-descript animals, birds, lizards, serpents, curious cones of trees and other natural productions. Debrett, London, 229 pp." title="Journal of a voyage to New South Wales, with sixty-five plates of non-descript animals, birds, lizards, serpents, curious cones of trees and other natural productions" type="book" year="1790">White, 1790</bibRefCitation>
)
</taxonomicName>
, two in 
<taxonomicName id="4C524D22FFF21828FBE8360FFAC20A16" authority="(Gray, 1838)" authorityName="Gray" authorityYear="1838" baseAuthorityName="Gray" baseAuthorityYear="1838" box="[1083,1339,514,539]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="gouldii">
<emphasis id="B926EAB3FFF21828FBE8360FFB5C0A14" box="[1083,1189,514,539]" italics="true" pageId="5" pageNumber="214">V. gouldii</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FB653608FACA0A16" author="Gray J. E." box="[1206,1331,515,539]" firstAuthor="Gray" journalOrPublisher="Annals and Magazine of Natural History, London" pageId="5" pageNumber="214" pagination="388 - 394" part="1" refId="ref4782" refString="Gray J. E. 1838. Catalogue of the slender-tongued saurians, with descriptions of many new genera and species. Part 3. Annals and Magazine of Natural History, London 1: 388 - 394." title="Catalogue of the slender-tongued saurians, with descriptions of many new genera and species. Part 3" type="journal article" year="1838">Gray, 1838</bibRefCitation>
)
</taxonomicName>
and one in 
<taxonomicName id="4C524D22FFF21828FCB6362FFB4D0A37" authority="Mertens, 1957" authorityName="Mertens" authorityYear="1957" box="[869,1204,547,571]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="rosenbergi">
<emphasis id="B926EAB3FFF21828FCB6362FFC050A37" box="[869,1020,547,571]" italics="true" pageId="5" pageNumber="214">V. rosenbergi</emphasis>
<bibRefCitation id="EFC34B50FFF21828FBDF362FFB4D0A37" author="Mertens R." box="[1036,1204,547,571]" firstAuthor="Mertens" journalOrPublisher="Zoologischer Anzeiger" pageId="5" pageNumber="214" pagination="17 - 20" part="159" refId="ref5086" refString="Mertens R. 1957. Ein neuer melanistischer Waran aus dem sudlichen Australien. (V. gouldii rosenbergi, subsp. nov.). Zoologischer Anzeiger 159: 17 - 20." title="Ein neuer melanistischer Waran aus dem sudlichen Australien. (V. gouldii rosenbergi, subsp. nov.)" type="journal article" year="1957">Mertens, 1957</bibRefCitation>
</taxonomicName>
. There is also one species in 
<taxonomicName id="4C524D22FFF21828FC67364FFB340A56" authority="(Daudin, 1802)" authorityName="Daudin" authorityYear="1802" baseAuthorityName="Daudin" baseAuthorityYear="1802" box="[948,1229,579,603]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="indicus">
<emphasis id="B926EAB3FFF21828FC67364FFBE70A57" box="[948,1054,579,603]" italics="true" pageId="5" pageNumber="214">V. indicus</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FBFD3648FB3C0A56" author="Daudin F. M." box="[1070,1221,579,603]" firstAuthor="Daudin" journalOrPublisher="Sonnini" pageId="5" pageNumber="214" part="3" refId="ref4615" refString="Daudin F. M. 1802. Histoire naturelle, generale et particuliere des reptiles, ouvrage faisant suite a l'histoire naturelle, generale et particuliere composee par Leclerc de Buffon, et redigee par C. S. Sonnini, vol. 3. F. Dufart, Paris." title="Histoire naturelle, generale et particuliere des reptiles, ouvrage faisant suite a l'histoire naturelle, generale et particuliere composee par Leclerc de Buffon, et redigee par C. S" type="journal volume" year="1802">Daudin, 1802</bibRefCitation>
)
</taxonomicName>
(A. de Chambrier, unpublished). Species of 
<taxonomicName id="4C524D22FFF21828FB863668FB080A77" authorityName="Freze" authorityYear="1963" box="[1109,1265,611,634]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FB863668FB080A77" box="[1109,1265,611,634]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
are believed to exhibit oioxenous specificity for their host (
<emphasis id="B926EAB3FFF21828FACF368FFAAE0A96" box="[1308,1367,644,667]" italics="true" pageId="5" pageNumber="214">sensu</emphasis>
<bibRefCitation id="EFC34B50FFF21828FAB3368FFBFD0AB1" author="Euzet L. &amp; Combes C." firstAuthor="Euzet" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="5" pageNumber="214" pagination="239 - 285" part="40" refId="ref4666" refString="Euzet L., Combes C. 1980. Les problemes de l'espece chez les animaux parasites. Memoires de la Societe Zoologique de France 40: 239 - 285." title="Les problemes de l'espece chez les animaux parasites" type="journal article" year="1980">Euzet &amp; Combes, 1980</bibRefCitation>
). The main problem in the study of the 
<taxonomicName id="4C524D22FFF21828FCB936C8FBFF0AD7" authorityName="Freze" authorityYear="1963" box="[874,1030,707,730]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FCB936C8FBFF0AD7" box="[874,1030,707,730]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
species is that most descriptions are incomplete. 
<bibRefCitation id="EFC34B50FFF21828FC1736EFFBB10AF1" author="Ratz I. von" box="[964,1096,740,764]" firstAuthor="Ratz" journalOrPublisher="Comptes rendus des seances de la Societe de biologie et de ses filiales" pageId="5" pageNumber="214" pagination="980 - 981" part="52" refId="ref5188" refString="Ratz I. von. 1900 a. Trois nouveaux cestodes de reptiles. Comptes rendus des seances de la Societe de biologie et de ses filiales 52: 980 - 981." title="Trois nouveaux cestodes de reptiles" type="journal article" year="1900" yearSuffix="a">Ratz (1900a</bibRefCitation>
, b) gave an identical, very poor description of 
<taxonomicName id="4C524D22FFF21828FC0A3708FB2D0B11" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[985,1236,771,796]" class="Cestoda" family="Proteocephalidae" genus="Ichthyotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FC0A3708FB2D0B11" box="[985,1236,771,796]" italics="true" pageId="5" pageNumber="214">Ichthyotaenia saccifera</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C524D22FFF21828FB2B370FFA800B11" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[1272,1401,772,796]" class="Cestoda" family="Proteocephalidae" genus="Ichthyotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FB2B370FFA800B11" box="[1272,1401,772,796]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
) in two different publications, one in French and the other in German. 
<bibRefCitation id="EFC34B50FFF21828FC713748FBB20B51" author="Schwarz R." bookContentInfo="52 pp." box="[930,1099,835,860]" firstAuthor="Schwarz" journalOrPublisher="Inaugural dissertation, University of Basel" pageId="5" pageNumber="214" refId="ref5293" refString="Schwarz R. 1908. Die Ichthyotaenien der Reptilien und Beitrage zur Kenntnis der Bothriocephalen. Inaugural dissertation, University of Basel, 52 pp." title="Die Ichthyotaenien der Reptilien und Beitrage zur Kenntnis der Bothriocephalen" type="book" year="1908">Schwarz (1908)</bibRefCitation>
redescribed this taxon, but both descriptions are incomplete (missing a description of the scolex, number of testes, cirrus-sac ratio, testis field, uterine branches number, etc.) and no illustration was presented. Furthermore, the host was not identified to species. But 
<bibRefCitation id="EFC34B50FFF21828FC1F37E8FB810BF1" author="Schwarz R." bookContentInfo="52 pp." box="[972,1144,995,1020]" firstAuthor="Schwarz" journalOrPublisher="Inaugural dissertation, University of Basel" pageId="5" pageNumber="214" refId="ref5293" refString="Schwarz R. 1908. Die Ichthyotaenien der Reptilien und Beitrage zur Kenntnis der Bothriocephalen. Inaugural dissertation, University of Basel, 52 pp." title="Die Ichthyotaenien der Reptilien und Beitrage zur Kenntnis der Bothriocephalen" type="book" year="1908">Schwarz (1908)</bibRefCitation>
described for 
<taxonomicName id="4C524D22FFF21828FACA37EEFA650BF1" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[1305,1436,996,1020]" class="Cestoda" family="Proteocephalidae" genus="Ichthyotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FACA37EEFA650BF1" box="[1305,1436,996,1020]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
egg clusters elongated, bag-shaped (banana-shaped), each proglottis containing 2 to 11 clusters, which allows us to differentiate this taxa from our new species. Pending a revision of the genus, we consider 
<taxonomicName id="4C524D22FFF21828FB33306EFA9C0C71" baseAuthorityName="Ratz" baseAuthorityYear="1900" box="[1248,1381,1124,1148]" class="Cestoda" family="Proteocephalidae" genus="Ichthyotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="saccifera">
<emphasis id="B926EAB3FFF21828FB33306EFA9C0C71" box="[1248,1381,1124,1148]" italics="true" pageId="5" pageNumber="214">K. saccifera</emphasis>
</taxonomicName>
as a 
<emphasis id="B926EAB3FFF21828FCEC308EFBFE0C96" box="[831,1031,1156,1180]" italics="true" pageId="5" pageNumber="214">species inquirenda</emphasis>
.
</paragraph>
<paragraph id="8BED36A1FFF21828FCEC30AFFAD20FB1" blockId="5.[831,1437,258,2078]" pageId="5" pageNumber="214">
<bibRefCitation id="EFC34B50FFF21828FCEC30AFFBF80CB0" author="Beddard F. E." box="[831,1025,1188,1213]" firstAuthor="Beddard" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="5" pageNumber="214" pagination="4 - 36" part="1913" refId="ref4160" refString="Beddard F. E. 1913. Contributions to the anatomy and systematic arrangement of the Cestoidea. Proceedings of the Zoological Society of London 1913 (part 1): 4 - 36." title="Contributions to the anatomy and systematic arrangement of the Cestoidea" type="proceedings paper" year="1913">Beddard (1913)</bibRefCitation>
described 
<taxonomicName id="4C524D22FFF21828FB7B30AFFA650CB1" authorityName="Beddard" authorityYear="1913" box="[1192,1436,1188,1212]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FB7B30AFFA650CB1" box="[1192,1436,1188,1212]" italics="true" pageId="5" pageNumber="214">Acanthotaenia varia</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C524D22FFF21828FCBD30CFFBAE0CD1" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[878,1111,1220,1244]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FCBD30CFFBAE0CD1" box="[878,1111,1220,1244]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia varia</emphasis>
</taxonomicName>
) from 
<taxonomicName id="4C524D22FFF21828FB1230CEFA8B0CD1" baseAuthorityName="White" baseAuthorityYear="1790" box="[1217,1394,1221,1244]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="varius">
<emphasis id="B926EAB3FFF21828FB1230CEFA8B0CD1" box="[1217,1394,1221,1244]" italics="true" pageId="5" pageNumber="214">Varanus varius</emphasis>
</taxonomicName>
in a “Zoological Garden”. 
<bibRefCitation id="EFC34B50FFF21828FBA230EFFADC0CF0" author="Nybelin O." box="[1137,1317,1252,1277]" firstAuthor="Nybelin" journalOrPublisher="Kungliga Svenska Vetenskapsakademiens Handlingar" pageId="5" pageNumber="214" pagination="1 - 48" part="52" refId="ref5120" refString="Nybelin O. 1917. Australischen Cestoden. Results of D. E. Mjoeberg's Swedish scientific expeditions to Australia, 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar 52 (14): 1 - 48." title="Australischen Cestoden. Results of D. E. Mjoeberg's Swedish scientific expeditions to Australia, 1910 - 1913" type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
collected some cestode specimens from 
<taxonomicName id="4C524D22FFF21828FB7C310EFA9D0D16" baseAuthorityName="Gray" baseAuthorityYear="1838" box="[1199,1380,1284,1308]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="gouldii">
<emphasis id="B926EAB3FFF21828FB7C310EFA9D0D16" box="[1199,1380,1284,1308]" italics="true" pageId="5" pageNumber="214">Varanus gouldii</emphasis>
</taxonomicName>
that he identified as 
<taxonomicName id="4C524D22FFF21828FBD3312FFA600D30" authority="Beddard, 1913" authorityName="Beddard" authorityYear="1913" box="[1024,1433,1316,1341]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FBD3312FFB1B0D31" box="[1024,1250,1316,1340]" italics="true" pageId="5" pageNumber="214">Acanthotaenia varia</emphasis>
<bibRefCitation id="EFC34B50FFF21828FB22312FFA600D30" author="Beddard F. E." box="[1265,1433,1316,1341]" firstAuthor="Beddard" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="5" pageNumber="214" pagination="4 - 36" part="1913" refId="ref4160" refString="Beddard F. E. 1913. Contributions to the anatomy and systematic arrangement of the Cestoidea. Proceedings of the Zoological Society of London 1913 (part 1): 4 - 36." title="Contributions to the anatomy and systematic arrangement of the Cestoidea" type="proceedings paper" year="1913">Beddard, 1913</bibRefCitation>
</taxonomicName>
. However, the description of these worms differs from the ones incompletely described by 
<bibRefCitation id="EFC34B50FFF21828FB38316FFA650D70" author="Beddard F. E." box="[1259,1436,1380,1405]" firstAuthor="Beddard" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="5" pageNumber="214" pagination="4 - 36" part="1913" refId="ref4160" refString="Beddard F. E. 1913. Contributions to the anatomy and systematic arrangement of the Cestoidea. Proceedings of the Zoological Society of London 1913 (part 1): 4 - 36." title="Contributions to the anatomy and systematic arrangement of the Cestoidea" type="proceedings paper" year="1913">Beddard (1913)</bibRefCitation>
in the position of the vagina relative to the cirrussac (posterior in Beddard’s description and anterior/ posterior in Nybelin’s) and in the absence of a vaginal sphincter in the former specimens but present in the latter (
<tableCitation id="C6D0031AFFF21828FC5D320EFC110E10" box="[910,1000,1541,1565]" captionStart="Table 1" captionStartId="7.[151,204,636,657]" captionTargetBox="[166,1327,876,2058]" captionText="Table 1. Summary of diagnostic traits in Kapsulotaenia species. Measurements in micrometres (μm) unless otherwise stated. PC = proportion of the length of cirrus-sac to the width of the proglottis; PP = position of the genital pore (cirrus pore) as % of the proglottis length; OV = percentage of the width of the ovary in relation to the width of the proglottis; SL = supplementary layer within the embryophore (see de Chambrier, 2006, p. 91);* see de Chambrier et al. (2012); D = taken from the original drawings; B = measurements taken from cotypes of Beddard (BMNH 1915.5.14.2); R = measurements taken from type specimens of Ratz (MHNG-PLAT-36037)." httpUri="http://table.plazi.org/id/DF2D6629FFF0182AFF443677FB3D0B2C" pageId="5" pageNumber="214" tableUuid="DF2D6629FFF0182AFF443677FB3D0B2C">Table 1</tableCitation>
). Therefore, the specimens identified as 
<taxonomicName id="4C524D22FFF21828FCB1322EFAFA0E30" authority="(Beddard, 1913)" authorityName="sensu Nybelin" authorityYear="1917" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[866,1283,1573,1597]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="varia">
<emphasis id="B926EAB3FFF21828FCB1322EFBBB0E30" box="[866,1090,1573,1597]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia varia</emphasis>
(
<bibRefCitation id="EFC34B50FFF21828FB85322EFB050E30" author="Beddard F. E." box="[1110,1276,1573,1597]" firstAuthor="Beddard" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="5" pageNumber="214" pagination="4 - 36" part="1913" refId="ref4160" refString="Beddard F. E. 1913. Contributions to the anatomy and systematic arrangement of the Cestoidea. Proceedings of the Zoological Society of London 1913 (part 1): 4 - 36." title="Contributions to the anatomy and systematic arrangement of the Cestoidea" type="proceedings paper" year="1913">Beddard, 1913</bibRefCitation>
)
</taxonomicName>
described by 
<bibRefCitation id="EFC34B50FFF21828FCEC324EFC150E50" author="Nybelin O." box="[831,1004,1605,1629]" firstAuthor="Nybelin" journalOrPublisher="Kungliga Svenska Vetenskapsakademiens Handlingar" pageId="5" pageNumber="214" pagination="1 - 48" part="52" refId="ref5120" refString="Nybelin O. 1917. Australischen Cestoden. Results of D. E. Mjoeberg's Swedish scientific expeditions to Australia, 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar 52 (14): 1 - 48." title="Australischen Cestoden. Results of D. E. Mjoeberg's Swedish scientific expeditions to Australia, 1910 - 1913" type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
are considered as 
<taxonomicName id="4C524D22FFF21828FB00324EFA960E51" authorityName="Freze" authorityYear="1963" box="[1235,1391,1605,1628]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FB00324EFA960E51" box="[1235,1391,1605,1628]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
sp. Furthermore, 
<bibRefCitation id="EFC34B50FFF21828FC00326EFB120E70" author="de Chambrier A. &amp; Zehnder, M. &amp; Vaucher C. &amp; Mariaux J." box="[979,1259,1637,1661]" etAl="et al." firstAuthor="de Chambrier" journalOrPublisher="Systematic Parasitology" pageId="5" pageNumber="214" pagination="159 - 171" part="57" refId="ref4524" refString="de Chambrier A., Zehnder, M., Vaucher C., Mariaux J. 2004. The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development. Systematic Parasitology 57: 159 - 171." title="The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development" type="journal article" year="2004">
de Chambrier 
<emphasis id="B926EAB3FFF21828FBBE326DFB650E71" box="[1133,1180,1637,1661]" italics="true" pageId="5" pageNumber="214">et al</emphasis>
. (2004
</bibRefCitation>
, 
<bibRefCitation id="EFC34B50FFF21828FB25326EFAD40E71" author="de Chambrier A. &amp; Waeschenbach A. &amp; Fisseha M. &amp; Scholz T. &amp; Mariaux J." box="[1270,1325,1637,1660]" firstAuthor="de Chambrier" journalOrPublisher="ZooKeys" pageId="5" pageNumber="214" pagination="25 - 59" part="500" refId="ref4470" refString="de Chambrier A., Waeschenbach A., Fisseha M., Scholz T., Mariaux J. 2015. A large 28 S rDNA-based phylogeny confirms the limitations of the established morphological characters for classification of proteocephalidean tapeworms (Platyhelminthes, Cestoda). ZooKeys 500: 25 - 59." title="A large 28 S rDNA-based phylogeny confirms the limitations of the established morphological characters for classification of proteocephalidean tapeworms (Platyhelminthes, Cestoda)" type="journal article" year="2015">2015</bibRefCitation>
) observed a strict (oioxenous) specificity for the members of the monophyletic 
<taxonomicName id="4C524D22FFF21828FC0E32AEFB800EB1" authorityName="Freze" authorityYear="1963" box="[989,1145,1701,1724]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FC0E32AEFB800EB1" box="[989,1145,1701,1724]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
and described differences between 
<taxonomicName id="4C524D22FFF21828FC7232CEFBC40ED1" authorityName="Freze" authorityYear="1963" box="[929,1085,1733,1756]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FC7232CEFBC40ED1" box="[929,1085,1733,1756]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
sp. 2 from 
<taxonomicName id="4C524D22FFF21828FB1232CDFA940ED1" baseAuthorityName="Gray" baseAuthorityYear="1838" box="[1217,1389,1733,1757]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="gouldii">
<emphasis id="B926EAB3FFF21828FB1232CDFA940ED1" box="[1217,1389,1733,1757]" italics="true" pageId="5" pageNumber="214">Varanus gouldii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C524D22FFF21828FCEC32EEFC220EF1" authorityName="Freze" authorityYear="1963" box="[831,987,1765,1788]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FCEC32EEFC220EF1" box="[831,987,1765,1788]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
sp. 4 from 
<taxonomicName id="4C524D22FFF21828FBA232EDFB2C0EF0" baseAuthorityName="White" baseAuthorityYear="1790" box="[1137,1237,1766,1789]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="varius">
<emphasis id="B926EAB3FFF21828FBA232EDFB2C0EF0" box="[1137,1237,1766,1789]" italics="true" pageId="5" pageNumber="214">V. varius</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFC34B50FFF21828FB3A32EEFA650EF3" author="Johnston T. H." box="[1257,1436,1765,1790]" firstAuthor="Johnston" journalOrPublisher="Journal of the Royal Society of New South Wales" pageId="5" pageNumber="214" pagination="103 - 116" part="43" refId="ref4821" refString="Johnston T. H. 1909. On a new reptilian cestode. Journal of the Royal Society of New South Wales 43: 103 - 116." title="On a new reptilian cestode" type="journal article" year="1909">Johnston (1909)</bibRefCitation>
described 
<taxonomicName id="4C524D22FFF21828FC7D330DFBD00F11" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[942,1065,1797,1821]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">
<emphasis id="B926EAB3FFF21828FC7D330DFBD00F11" box="[942,1065,1797,1821]" italics="true" pageId="5" pageNumber="214">K. tidswelli</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4C524D22FFF21828FB8A330EFAAE0F11" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[1113,1367,1797,1820]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="tidswelli">
<emphasis id="B926EAB3FFF21828FB8A330EFAAE0F11" box="[1113,1367,1797,1820]" italics="true" pageId="5" pageNumber="214">Acanthotaenia tidswelli</emphasis>
</taxonomicName>
) from 
<taxonomicName id="4C524D22FFF21828FCEC332CFC1F0F30" baseAuthorityName="White" baseAuthorityYear="1790" box="[831,998,1830,1854]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="varius">
<emphasis id="B926EAB3FFF21828FCEC332CFC1F0F30" box="[831,998,1830,1854]" italics="true" pageId="5" pageNumber="214">Varanus varius</emphasis>
</taxonomicName>
, but the same author (
<bibRefCitation id="EFC34B50FFF21828FB38332EFA6C0F33" author="Johnston T. H." box="[1259,1429,1829,1854]" firstAuthor="Johnston" journalOrPublisher="Proceedings of the Royal Society of Queensland" pageId="5" pageNumber="214" pagination="233 - 249" part="23" refId="ref4849" refString="Johnston T. H. 1912. A census of Australian reptilian Entozoa. Proceedings of the Royal Society of Queensland 23: 233 - 249." title="A census of Australian reptilian Entozoa" type="journal article" year="1912">Johnston, 1912</bibRefCitation>
) listed this species also as parasite of 
<taxonomicName id="4C524D22FFF21828FB34334CFA6F0F51" baseAuthorityName="Gray" baseAuthorityYear="1838" box="[1255,1430,1861,1886]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="gouldii">
<emphasis id="B926EAB3FFF21828FB34334CFA6F0F51" box="[1255,1430,1861,1886]" italics="true" pageId="5" pageNumber="214">Varanus gouldii</emphasis>
</taxonomicName>
. However, this host record should be confirmed (voucher specimens are not available). It is obvious that the genus 
<taxonomicName id="4C524D22FFF21828FCEC33AEFC220FB1" authorityName="Freze" authorityYear="1963" box="[831,987,1957,1980]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="genus">
<emphasis id="B926EAB3FFF21828FCEC33AEFC220FB1" box="[831,987,1957,1980]" italics="true" pageId="5" pageNumber="214">Kapsulotaenia</emphasis>
</taxonomicName>
is pending taxonomic revision.
</paragraph>
<paragraph id="8BED36A1FFF2182BFCEC33CEFE8A0B11" blockId="5.[831,1437,258,2078]" lastBlockId="6.[151,756,258,797]" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214">
The ecology and distribution of 
<taxonomicName id="4C524D22FFF21828FB7C33CCFA650FD1" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[1199,1436,1989,2014]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="komodoensis">
<emphasis id="B926EAB3FFF21828FB7C33CCFA650FD1" box="[1199,1436,1989,2014]" italics="true" pageId="5" pageNumber="214">Varanus komodoensis</emphasis>
</taxonomicName>
(host of 
<taxonomicName id="4C524D22FFF21828FC7D33ECFBA40FF0" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[942,1117,2022,2046]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="sandgroundi">
<emphasis id="B926EAB3FFF21828FC7D33ECFBA40FF0" box="[942,1117,2022,2046]" italics="true" pageId="5" pageNumber="214">K. sandgroundi</emphasis>
</taxonomicName>
) and 
<taxonomicName id="4C524D22FFF21828FB7C33ECFAD70FF3" authorityName="Lucas &amp; Frost" authorityYear="1903" box="[1199,1326,2023,2046]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="B926EAB3FFF21828FB7C33ECFAD70FF3" box="[1199,1326,2023,2046]" italics="true" pageId="5" pageNumber="214">V. spenceri</emphasis>
</taxonomicName>
(host of 
<taxonomicName id="4C524D22FFF21828FCEC3C0CFC210010" authorityName="Hugh I. Jones &amp; Alain de Chambrier" authorityYear="2016" box="[831,984,2054,2078]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="5" pageNumber="214" phylum="Platyhelminthes" rank="species" species="chisholmae">
<emphasis id="B926EAB3FFF21828FCEC3C0CFC210010" box="[831,984,2054,2078]" italics="true" pageId="5" pageNumber="214">K. chisholmae</emphasis>
</taxonomicName>
) differ. 
<taxonomicName id="4C524D22FFF21828FBE33C0CFB2C0010" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[1072,1237,2054,2078]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="komodoensis">
<emphasis id="B926EAB3FFF21828FBE33C0CFB2C0010" box="[1072,1237,2054,2078]" italics="true" pageId="5" pageNumber="214">V. komodoensis</emphasis>
</taxonomicName>
only occurs in the 
<collectingRegion id="4996F843FFF1182BFF443508FE710914" box="[151,392,258,282]" country="Indonesia" name="Nusa Tenggara Timur" pageId="6" pageNumber="215">Lesser Sunda islands</collectingRegion>
, 
<collectingCountry id="F3457631FFF1182BFE4A3509FDFA0914" box="[409,515,258,281]" name="Indonesia" pageId="6" pageNumber="215">Indonesia</collectingCountry>
, principally Komodo and 
<collectingCountry id="F3457631FFF1182BFF1A3529FEF70934" box="[201,270,290,313]" name="Indonesia" pageId="6" pageNumber="215">Flores</collectingCountry>
, where it inhabits deciduous monsoon rain forest, savannah and mangrove (
<bibRefCitation id="EFC34B50FFF1182BFE273549FD8A0956" author="Ciofi C." bookContentInfo="588 pp." box="[500,627,322,347]" editor="Pianka E." firstAuthor="Ciofi" journalOrPublisher="Indiana University Press" pageId="6" pageNumber="215" pagination="197 - 204" refId="ref4228" refString="Ciofi C. 2004. Varanus komodoensis. [pp. 197 - 204]. In: Pianka E., King D. R., King R. A. (Eds), Varanoid Lizards of the World. Indiana University Press, 588 pp." title="Varanus komodoensis" type="book chapter" volumeTitle="Varanoid Lizards of the World" year="2004">Ciofi, 2004</bibRefCitation>
). As well as feeding on invertebrates and smaller vertebrates, adult 
<taxonomicName id="4C524D22FFF1182BFF44358FFEC60997" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[151,319,387,411]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="komodoensis">
<emphasis id="B926EAB3FFF1182BFF44358FFEC60997" box="[151,319,387,411]" italics="true" pageId="6" pageNumber="215">V. komodoensis</emphasis>
</taxonomicName>
can kill and consume large vertebrates such as deer, wild boar and goats; decomposing carcasses form a significant part of its diet (
<bibRefCitation id="EFC34B50FFF1182BFDD535C8FD3309D1" author="Auffenberg W." bookContentInfo="406 pp." box="[518,714,451,476]" firstAuthor="Auffenberg" journalOrPublisher="University Press of Florida, Gainesville" pageId="6" pageNumber="215" refId="ref4137" refString="Auffenberg W. 1981. The behavioral ecology of the Komodo monitor. University Press of Florida, Gainesville. 406 pp." title="The behavioral ecology of the Komodo monitor" type="book" year="1981">Auffenberg, 1981</bibRefCitation>
). In contrast, 
<taxonomicName id="4C524D22FFF1182BFED335EEFE8209F1" authorityName="Lucas &amp; Frost" authorityYear="1903" box="[256,379,485,508]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="B926EAB3FFF1182BFED335EEFE8209F1" box="[256,379,485,508]" italics="true" pageId="6" pageNumber="215">V. spenceri</emphasis>
</taxonomicName>
is a burrowing species confined to relatively treeless grasslands with 
<taxonomicName id="4C524D22FFF1182BFD93360FFD600A16" box="[576,665,516,539]" class="Liliopsida" family="Poaceae" genus="Astrebla" kingdom="Plantae" order="Poales" pageId="6" pageNumber="215" phylum="Tracheophyta" rank="genus">
<emphasis id="B926EAB3FFF1182BFD93360FFD600A16" box="[576,665,516,539]" italics="true" pageId="6" pageNumber="215">Astrebla</emphasis>
</taxonomicName>
spp. in central 
<collectingRegion id="4996F843FFF1182BFF3B362FFE9F0A36" box="[232,358,548,571]" country="Australia" name="Queensland" pageId="6" pageNumber="215">Queensland</collectingRegion>
and the adjacent 
<collectingRegion id="4996F843FFF1182BFDFB362FFD080A36" box="[552,753,548,571]" country="Australia" name="Northern Territory" pageId="6" pageNumber="215">
<collectingRegion id="4996F843FFF1182BFDFB362FFD710A36" box="[552,648,548,571]" country="Sri Lanka" name="Northern" pageId="6" pageNumber="215">Northern</collectingRegion>
Territory
</collectingRegion>
, and feeds on other smaller reptiles, small mammals such as 
<emphasis id="B926EAB3FFF1182BFF24366DFE350A76" box="[247,460,612,637]" italics="true" pageId="6" pageNumber="215">
<taxonomicName id="4C524D22FFF1182BFF24366DFE310A76" baseAuthorityName="Waite" baseAuthorityYear="1898" box="[247,456,612,637]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="villosisimus">Rattus villosisimus</taxonomicName>
,
</emphasis>
and invertebrates such as orthopterans including plague locust species (
<bibRefCitation id="EFC34B50FFF1182BFD5A368FFF360AB0" author="Pengilley R." firstAuthor="Pengilley" journalOrPublisher="Australian Journal of Herpetology" pageId="6" pageNumber="215" pagination="23 - 26" part="1" refId="ref5158" refString="Pengilley R. 1981. Notes on the biology of Varanus spenceri and V. gouldii, Barkly Tablelands, Northern Territory. Australian Journal of Herpetology 1: 23 - 26." title="Notes on the biology of Varanus spenceri and V. gouldii, Barkly Tablelands, Northern Territory" type="journal article" year="1981">Pengilley, 1981</bibRefCitation>
; 
<bibRefCitation id="EFC34B50FFF1182BFF3236AEFE020AB0" author="Valentic R. A. &amp; Turner G." box="[225,507,677,701]" firstAuthor="Valentic" journalOrPublisher="Herpetofauna" pageId="6" pageNumber="215" pagination="43 - 45" part="27" refId="ref5755" refString="Valentic R. A., Turner G. 1997. Diet and reproductive status of a road-killed Spencer's monitor Varanus spenceri. Herpetofauna 27: 43 - 45." title="Diet and reproductive status of a road-killed Spencer's monitor Varanus spenceri" type="journal article" year="1997">Valentic &amp; Turner, 1997</bibRefCitation>
; 
<bibRefCitation id="EFC34B50FFF1182BFDDE36AEFD550AB0" author="Wooley P." box="[525,684,677,701]" firstAuthor="Wooley" journalOrPublisher="Biawak" pageId="6" pageNumber="215" pagination="24 - 25" part="4" refId="ref5828" refString="Wooley P. 2010. Varanus spenceri: a last supper of grasshoppers (Acrididae). Biawak 4: 24 - 25." title="Varanus spenceri: a last supper of grasshoppers (Acrididae)" type="journal article" year="2010">Wooley, 2010</bibRefCitation>
). It is interesting to note that no cestodes were recovered from 
<date id="FFEC1061FFF1182BFF4436EDFF2B0AF0" box="[151,210,742,766]" pageId="6" pageNumber="215" value="2012-05">
12 
<taxonomicName id="4C524D22FFF1182BFF6D36EDFF2B0AF0" baseAuthorityName="Gray" baseAuthorityYear="1838" box="[190,210,742,765]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="gouldii">
<emphasis id="B926EAB3FFF1182BFF6D36EDFF2B0AF0" box="[190,210,742,765]" italics="true" pageId="6" pageNumber="215">V.</emphasis>
</taxonomicName>
</date>
gouldii and 
<date id="FFEC1061FFF1182BFEB536EDFE580AF0" box="[358,417,742,766]" pageId="6" pageNumber="215" value="2010-05">
10 
<taxonomicName id="4C524D22FFF1182BFE5F36EDFE580AF0" authorityName="Storr" authorityYear="1980" box="[396,417,742,765]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="panoptes">
<emphasis id="B926EAB3FFF1182BFE5F36EDFE580AF0" box="[396,417,742,765]" italics="true" pageId="6" pageNumber="215">V.</emphasis>
</taxonomicName>
</date>
panoptes in 
<collectingRegion id="4996F843FFF1182BFDEB36EEFD4F0AF1" box="[568,694,741,764]" country="Australia" name="Queensland" pageId="6" pageNumber="215">Queensland</collectingRegion>
(H.I. Jones, unpublished).
</paragraph>
</subSubSection>
</treatment>
</document>