203 lines
34 KiB
XML
203 lines
34 KiB
XML
<document id="FC17809DAE98A577CD8F2B768167622E" ID-DOI="10.5281/zenodo.175869" ID-GBIF-Dataset="ef14a2a6-57bf-4a25-94ff-5a94baeec224" ID-ISSN="1175-5326" ID-Zenodo-Dep="175869" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459822160839" checkinUser="plazi" docAuthor="Bartsch, Ilse" docDate="2007" docId="03FB87B1CE2CB637088BFBFCB47DA142" docLanguage="en" docName="zt01435p049.pdf" docOrigin="Zootaxa 1435" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Copidognathus parvulus Bartsch, 2007, spec. nov." docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="FFC2FFC9CE28B630081CFFABB54EA508" masterDocTitle="Halacarid mites (Acari: Halacaridae) from Esperance (Western Australia), description of two new Copidognathus species" masterLastPageNumber="49" masterPageNumber="41" pageNumber="45" updateTime="1698221939282" updateUser="plazi">
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<mods:titleInfo id="980260EAAD105F66D1C94A16AA00F0AC">
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<mods:title id="C96A10C957465DD9AC37DC72CDB74500">Halacarid mites (Acari: Halacaridae) from Esperance (Western Australia), description of two new Copidognathus species</mods:title>
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<mods:name id="1DD2387C94CD150D409BB56846B5E61F" type="personal">
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<mods:namePart id="3134451813213BA9BE0D09759A95C6E7">Bartsch, Ilse</mods:namePart>
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<mods:typeOfResource id="A5397693E42DA05B2DABC79774120B5E">text</mods:typeOfResource>
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<mods:title id="D5FC48672D73EB3AF3A91F5FA62B4F10">Zootaxa</mods:title>
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<mods:part id="79164EF61EA74F09766FEF50E185FD65">
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<mods:date id="10B869F03474B0174EF350103632743E">2007</mods:date>
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<mods:detail id="CCF09B7D4DB7FE4799F58AA18C980368" type="volume">
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<mods:number id="0DBF9FAC78FCE7E6294E0B8BE639B5AE">1435</mods:number>
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<mods:classification id="C815479838A37CE04DC3D78F9E764787">journal article</mods:classification>
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<mods:identifier id="B28E267CDB14DB45805EA4EE256507D5" type="DOI">10.5281/zenodo.175869</mods:identifier>
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<mods:identifier id="D33CDF18F5EC9D3919663BEFA2011B8B" type="GBIF-Dataset">ef14a2a6-57bf-4a25-94ff-5a94baeec224</mods:identifier>
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<mods:identifier id="EE7D0D794E621BAA82D9A500C1438BED" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BFC72F247BA3BF10D390FAF115583BF7" type="Zenodo-Dep">175869</mods:identifier>
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<treatment id="03FB87B1CE2CB637088BFBFCB47DA142" ID-DOI="http://doi.org/10.5281/zenodo.6247703" ID-GBIF-Taxon="119356014" ID-Zenodo-Dep="6247703" LSID="urn:lsid:plazi:treatment:03FB87B1CE2CB637088BFBFCB47DA142" httpUri="http://treatment.plazi.org/id/03FB87B1CE2CB637088BFBFCB47DA142" lastPageId="7" lastPageNumber="48" pageId="4" pageNumber="45">
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<subSubSection id="C348652CCE2CB634088BFBFCB457A19C" pageId="4" pageNumber="45" type="nomenclature">
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<paragraph id="8BED36A7CE2CB634088BFBFCB770A179" blockId="4.[151,574,1111,1172]" box="[151,574,1111,1137]" pageId="4" pageNumber="45">
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<heading id="D0A581CBCE2CB634088BFBFCB770A179" bold="true" box="[151,574,1111,1137]" fontSize="11" level="1" pageId="4" pageNumber="45" reason="1">
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<emphasis id="B926EAB5CE2CB634088BFBFCB770A179" bold="true" box="[151,574,1111,1137]" pageId="4" pageNumber="45">
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<taxonomicName id="4C524D24CE2CB634088BFBFCB48EA179" box="[151,448,1111,1137]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
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<emphasis id="B926EAB5CE2CB634088BFBFCB48EA179" bold="true" box="[151,448,1111,1137]" italics="true" pageId="4" pageNumber="45">Copidognathus parvulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21557CECE2CB63409DBFBFCB770A179" box="[455,574,1111,1137]" pageId="4" pageNumber="45" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BED36A7CE2CB634088BFBD1B457A19C" blockId="4.[151,574,1111,1172]" box="[151,281,1146,1172]" pageId="4" pageNumber="45">
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<figureCitation id="13692A22CE2CB634088BFBD1B457A19C" box="[151,281,1146,1172]" captionStart-0="FIGURES 14 – 21" captionStart-1="FIGURES 22 – 29" captionStartId-0="5.[151,269,1925,1949]" captionStartId-1="6.[151,269,1517,1541]" captionTargetBox-0="[152,1411,827,1924]" captionTargetBox-1="[213,1390,228,1442]" captionTargetId-0="figure@5.[151,1436,794,1973]" captionTargetId-1="figure@6.[151,1436,194,1494]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." captionText-1="FIGURES 22 – 29. Copidognathus parvulus spec. nov., male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm." httpUri-0="https://zenodo.org/record/175872/files/figure.png" httpUri-1="https://zenodo.org/record/175873/files/figure.png" pageId="4" pageNumber="45">Figs 14–28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C348652CCE2CB634088BFB61B19AA004" pageId="4" pageNumber="45" type="materials_examined">
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<paragraph id="8BED36A7CE2CB634088BFB61B19AA004" blockId="4.[151,1437,1226,2012]" pageId="4" pageNumber="45">
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Material examined.
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<typeStatus id="54E98805CE2CB6340990FB61B4B7A1EC" box="[396,505,1226,1252]" pageId="4" pageNumber="45" type="holotype">Holotype</typeStatus>
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male,
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<collectionCode id="ED43AE62CE2CB6340A4FFB61B7D5A1EC" LSID="urn:lsid:biocol.org:col:34981" box="[595,667,1226,1252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="4" pageNumber="45">WAM</collectionCode>
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T78347, Western
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<collectingCountry id="F3457637CE2CB6340B9FFB61B6BCA1EC" box="[899,1010,1226,1252]" name="Australia" pageId="4" pageNumber="45">Australia</collectingCountry>
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, Esperance, Duke of Orleans Bay,
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<geoCoordinate id="EE665060CE2CB634088BFB59B5B4A004" box="[151,250,1266,1292]" direction="south" orientation="latitude" pageId="4" pageNumber="45" precision="925" value="-33.916668">33°55’S</geoCoordinate>
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,
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<geoCoordinate id="EE665060CE2CB634091AFB59B437A004" box="[262,377,1266,1292]" direction="east" orientation="longitude" pageId="4" pageNumber="45" precision="925" value="122.583336">122°35’E</geoCoordinate>
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, corallines on demosponge, just below low water line,
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<date id="FFEC1067CE2CB6340BE0FB59B19EA004" box="[1020,1232,1266,1292]" pageId="4" pageNumber="45" value="2003-02-17">17 February 2003</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348652CCE2CB63408DAFAB1B74BA32C" pageId="4" pageNumber="45" type="diagnosis">
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<paragraph id="8BED36A7CE2CB63408DAFAB1B74BA32C" blockId="4.[151,1437,1226,2012]" pageId="4" pageNumber="45">
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<emphasis id="B926EAB5CE2CB63408DAFAB1B408A03C" bold="true" box="[198,326,1306,1332]" pageId="4" pageNumber="45">Diagnosis.</emphasis>
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Length 290 µm. On dorsal plates areolae with rosette pores distinctly delimited from rather smooth remainder of plates. AD with three small porose areolae; on PD pair of marginal costae separated from transverse porose areola in posterior margin. Ventral plates with small, porose areolae. Gnathosoma slender; rostrum long, extending to about middle of P-4. Legs slender, telofemora, genua and tibiae with articular membranes, tarsi with fossa membranes. Telofemora III and IV with 2/1 and 2/0 dorsal/ventral setae. Tibiae I to IV with 2, 2, 1, 0 bipectinate setae and 1, 1, 1, 2 slender ventral setae. Tarsi III and IV with four and three dorsal setae, respectively.
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</paragraph>
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</subSubSection>
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<subSubSection id="C348652CCE2CB63708DAF999B486A712" lastPageId="7" lastPageNumber="48" pageId="4" pageNumber="45" type="description">
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<paragraph id="8BED36A7CE2CB63508DAF999B184A472" blockId="4.[151,1437,1226,2012]" lastBlockId="5.[151,1437,152,738]" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45">
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<emphasis id="B926EAB5CE2CB63408DAF999B411A344" bold="true" box="[198,351,1586,1612]" pageId="4" pageNumber="45">Description.</emphasis>
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Length 290 µm, width 164 µm. Idiosoma heavily armoured (
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<figureCitation id="13692A22CE2CB6340C5EF999B1D4A344" box="[1090,1178,1586,1612]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="4" pageNumber="45">Fig. 14</figureCitation>
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), with narrow, coarse striae between the plates. Most parts of dorsal plates smooth, except for clearly demarcated porose areolae with rosette pores and scattered shallow foveae in anterior part of AD and within reticulate lateral margin of OC. Rosette pores with small surficial ostium and four to six canaliculi in deeper integumental layers. AD longer than wide, length 92 µm, width 70 µm; shape octagonal, anterior margin with truncate lamella, posterior margin truncate. Anterior porose areola transversely oblong, two posterior areolae round and immediately adjacent; scattered shallow foveae between anterior and paired areolae (
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<figureCitation id="13692A22CE2CB6340BFBF889B10EA234" box="[999,1088,1826,1852]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="4" pageNumber="45">Fig. 16</figureCitation>
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). Pair of porose areolae close together at 0.54. Length of OC 82 µm, width 52 µm, posteriorly with small ‘tails’, each plate with two large corneae, gland pore level with posterior cornea, pore canaliculus slightly posterior to gland pore; a porose areola medial to corneae; integument along lateral edge of OC raised and with foveae but without canaliculi (
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<figureCitation id="13692A22CE2CB6340D71F831B5F9A2D4" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="4" pageNumber="45">Fig. 17</figureCitation>
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). Length of PD 180 µm, width 106 µm; anterior margin ovate, extending between OC. PD with pair of porose costae near lateral margin of plate, costae far from reaching anterior margin of PD. Rosette pores present from level of 0.24 to 0.75; costae two rosette pores wide in anterior half, a single rosette pore wide in posterior half. Area between costae and posterior porose areola slightly raised. One pair of gland pores in lateral margin of costae halfway between ds-4 and ds-5 and somewhat anterior to the level of insertion of leg IV. Pair of ds-1 on AD in margin of pair of porose areolae; ds-
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<quantity id="4CAA9B42CE2DB6350B5FFE93B621A45A" box="[835,879,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="46" unit="in" value="2.0">2 in</quantity>
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striated integument; ds-3, ds-4 and ds-5 on PD, ds-3 anterior to costae, ds-4 and ds-5 within costae. Adanal setae in posterior margin of PD.
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</paragraph>
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<paragraph id="8BED36A7CE2DB63508DAFE23B789A7EA" blockId="5.[151,1437,152,738]" pageId="5" pageNumber="46">
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Ventral plates with small, distinctly delimited porose areolae, otherwise smooth. On AE porose areolae anterior to insertion of leg II. PE with two marginal areolae, one each anterior to insertions of legs III and IV, and one ventral areola medial to insertion of leg III. On GA areolae on either side of GO (
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<figureCitation id="13692A22CE2DB6350CA7FE73B05DA4FA" box="[1211,1299,472,498]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="5" pageNumber="46">Fig. 15</figureCitation>
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). Length of AE 110 µm, width 139 µm; posterior margin truncate; AE with large epimeral processes lamelliform flanking bases of trochanters I and II (
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<figureCitation id="13692A22CE2DB63509F3FD83B708A74A" box="[495,582,552,578]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="5" pageNumber="46">Fig. 18</figureCitation>
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). Length of PE 139 µm, extending beyond insertion of leg IV. Length of GA 147 µm, width 100 µm, anterior margin truncate. GO in posterior half of GA. Length of GO 37 µm, width 27 µm. Distance between anterior margin of GO and that of GA 1.9 times length of GO. Twenty-six perigenital setae arranged in a ring around GO. Genital sclerites with four pairs of subgenital setae, two anterior pairs setiform, two posterior pairs spur-like (
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<figureCitation id="13692A22CE2DB6350A7EFD63B7F7A7EA" box="[610,697,712,738]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="5" pageNumber="46">Fig. 19</figureCitation>
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).
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</paragraph>
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<caption id="DF2D662FCE2DB635088BF82EB02CA2EA" httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="5" pageNumber="46" targetBox="[152,1411,827,1924]" targetPageId="5">
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<paragraph id="8BED36A7CE2DB635088BF82EB02CA2EA" blockId="5.[151,1436,1925,2018]" pageId="5" pageNumber="46">
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<emphasis id="B926EAB5CE2DB635088BF82EB411A295" bold="true" box="[151,351,1925,1949]" pageId="5" pageNumber="46">FIGURES 14–21.</emphasis>
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<taxonomicName id="4C524D24CE2DB6350974F82EB73FA294" box="[360,625,1925,1948]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
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<emphasis id="B926EAB5CE2DB6350974F82EB73FA294" box="[360,625,1925,1948]" italics="true" pageId="5" pageNumber="46">Copidognathus parvulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21557CECE2DB6350A66F82EB7A4A295" box="[634,746,1925,1949]" pageId="5" pageNumber="46" rank="species">
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<emphasis id="B926EAB5CE2DB6350A66F82EB7A4A295" bold="true" box="[634,746,1925,1949]" pageId="5" pageNumber="46">spec. nov.</emphasis>
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</taxonomicNameLabel>
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, male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds-1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm.
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</paragraph>
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</caption>
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<caption id="DF2D662FCE2EB636088BFA46B642A365" httpUri="https://zenodo.org/record/175873/files/figure.png" pageId="6" pageNumber="47" targetBox="[213,1390,228,1442]" targetPageId="6">
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<paragraph id="8BED36A7CE2EB636088BFA46B642A365" blockId="6.[151,1437,1517,1645]" pageId="6" pageNumber="47">
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<emphasis id="B926EAB5CE2EB636088BFA46B413A30D" bold="true" box="[151,349,1517,1541]" pageId="6" pageNumber="47">FIGURES 22–29.</emphasis>
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<taxonomicName id="4C524D24CE2EB6360978FA45B724A30D" box="[356,618,1518,1541]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
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<emphasis id="B926EAB5CE2EB6360978FA45B724A30D" box="[356,618,1518,1541]" italics="true" pageId="6" pageNumber="47">Copidognathus parvulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21557CECE2EB6360A6DFA46B790A30D" box="[625,734,1517,1541]" pageId="6" pageNumber="47" rank="species">
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<emphasis id="B926EAB5CE2EB6360A6DFA46B790A30D" bold="true" box="[625,734,1517,1541]" pageId="6" pageNumber="47">spec. nov.</emphasis>
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</taxonomicNameLabel>
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, male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm.
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</paragraph>
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</caption>
|
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<paragraph id="8BED36A7CE2EB63608DAF90AB02CA23B" blockId="6.[151,1437,1697,2003]" pageId="6" pageNumber="47">
|
||
Gnathosoma long and slender; length 145 µm, width 45 µm. Rostrum extending to about middle of P-4 (
|
||
<figureCitation id="13692A22CE2EB6360883F962B467A3EB" box="[159,297,1737,1763]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="6" pageNumber="47">Figs 20, 21</figureCitation>
|
||
), its length about twice that of gnathosomal base. P-2 longer than P-4; P-2 with one dorsal seta, P-4 with three setae in basal whorl, one setula and two spurs at apex. Tectum triangular, its margin dentate (
|
||
<figureCitation id="13692A22CE2EB6360883F8B2B5B9A23B" box="[159,247,1817,1843]" captionStart="FIGURES 14 – 21" captionStartId="5.[151,269,1925,1949]" captionTargetBox="[152,1411,827,1924]" captionTargetId="figure@5.[151,1436,794,1973]" captionTargetPageId="5" captionText="FIGURES 14 – 21. Copidognathus parvulus spec. nov., male, 14, idiosoma, dorsal; 15, idiosoma, ventral; 16, anterior part of idiosoma with AD, dorsal; 17, ocular plate; 18, part of AE with epimeral pore; 19, genital opening; 20, gnathosoma, dorsal; 21, gnathosoma, ventral. (ds- 1, first dorsal seta; glp, gland pore, pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175872/files/figure.png" pageId="6" pageNumber="47">Fig. 20</figureCitation>
|
||
). Lateral flank of gnathosomal base with rosette pores, dorsal flank with scattered, small foveae.
|
||
</paragraph>
|
||
<paragraph id="8BED36A7CE2EB63708DAF8EAB41EA4C2" blockId="6.[151,1437,1697,2003]" lastBlockId="7.[151,1437,152,1098]" lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47">
|
||
Legs slender, heavily sclerotized, ventral integument markedly thicker than dorsal one (
|
||
<figureCitation id="13692A22CE2EB6360CC8F8EAB063A253" box="[1236,1325,1857,1883]" captionStart="FIGURES 22 – 29" captionStartId="6.[151,269,1517,1541]" captionTargetBox="[213,1390,228,1442]" captionTargetId="figure@6.[151,1436,194,1494]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Copidognathus parvulus spec. nov., male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm." httpUri="https://zenodo.org/record/175873/files/figure.png" pageId="6" pageNumber="47">Fig. 26</figureCitation>
|
||
). Surface of telofemora and tibiae covered with small cuticular droplets, that of telofemora also minutely reticulate. Ventral margin of telofemora apparently delicately crenulate, due to small cuticular droplets on the surface. Telofemora rather short, on all legs shorter than tibiae. Telofemora I to IV 2.5, 2.3, 2.0 and 2.2 times longer than high, respectively. Telofemora, genua and tibiae with large articular membranes, tarsi with fossa membranes (
|
||
<figureCitation id="13692A22CE2FB63708E9FF6BB4CFA5D2" box="[245,385,192,218]" captionStart="FIGURES 22 – 29" captionStartId="6.[151,269,1517,1541]" captionTargetBox="[213,1390,228,1442]" captionTargetId="figure@6.[151,1436,194,1494]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Copidognathus parvulus spec. nov., male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm." httpUri="https://zenodo.org/record/175873/files/figure.png" pageId="7" pageNumber="48">Figs 22–25</figureCitation>
|
||
). Trochanters III and IV each with small medial lamella (
|
||
<figureCitation id="13692A22CE2FB6370C29FF6BB18CA5D2" box="[1077,1218,192,218]" captionStart="FIGURES 22 – 29" captionStartId="6.[151,269,1517,1541]" captionTargetBox="[213,1390,228,1442]" captionTargetId="figure@6.[151,1436,194,1494]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Copidognathus parvulus spec. nov., male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm." httpUri="https://zenodo.org/record/175873/files/figure.png" pageId="7" pageNumber="48">Figs 24, 25</figureCitation>
|
||
). Distal margin of telofemora I and II with minute spiniform dorsal lamella. Leg chaetotaxy: leg I, 1, 2, 5, 4, 7, 7; leg II, 1, 2, 5, 4, 7, 4; leg III, 1, 2, 2, 3, 5, 4; leg IV, 0, 2, 3, 3, 5, 3. Tibiae I to IV with 2, 2, 1, 0 bipectinate and 1, 1, 1, 2 smooth ventral setae. Solenidion on both tarsus
|
||
<date id="FFEC1067CE2FB6370ADBFE93B60FA45A" box="[711,833,312,338]" pageId="7" pageNumber="48" value="1916-01" valueMax="1916-02-31" valueMin="1916-01-16">I and II 16</date>
|
||
µm long and on dorsolateral fossa membrane (
|
||
<figureCitation id="13692A22CE2FB6370D74FE93B5ABA472" captionStart="FIGURES 22 – 29" captionStartId="6.[151,269,1517,1541]" captionTargetBox="[213,1390,228,1442]" captionTargetId="figure@6.[151,1436,194,1494]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Copidognathus parvulus spec. nov., male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm." httpUri="https://zenodo.org/record/175873/files/figure.png" pageId="7" pageNumber="48">Figs 27, 28</figureCitation>
|
||
). Basal seta on tarsus III removed from fossary seta by height of tarsus (
|
||
<figureCitation id="13692A22CE2FB6370C22FECBB1D8A472" box="[1086,1174,352,378]" captionStart="FIGURES 22 – 29" captionStartId="6.[151,269,1517,1541]" captionTargetBox="[213,1390,228,1442]" captionTargetId="figure@6.[151,1436,194,1494]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Copidognathus parvulus spec. nov., male, 22, leg I, medial; 23, leg II, medial; 24, leg III, medial; 25, leg IV, medial; 26, telofemur to tibia IV, lateral (optical section in dotted line); 27, part of tibia and tarsus I, lateral (medial setae and claw omitted); 28, tarsus II, lateral (medial setae and claw omitted); 29, part of tibia and tarsus III, lateral (medial setae and claw in broken line). Scale = 50 µm." httpUri="https://zenodo.org/record/175873/files/figure.png" pageId="7" pageNumber="48">Fig. 29</figureCitation>
|
||
). Tarsus I with pair of doubled pas, tarsus II with single pas on either side; tarsi III and IV with setiform lateral pas and short, bristlelike medial pas.
|
||
</paragraph>
|
||
<paragraph id="8BED36A7CE2FB63708DAFE73B486A712" blockId="7.[151,1437,152,1098]" pageId="7" pageNumber="48">Claws of all tarsi with accessory process and pecten, the latter with numerous delicate tines. Pecten not extending to base of claw.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348652CCE2FB63708DAFD8CB1B9A74A" box="[198,1271,551,578]" pageId="7" pageNumber="48" type="etymology">
|
||
<paragraph id="8BED36A7CE2FB63708DAFD8CB1B9A74A" blockId="7.[151,1437,152,1098]" box="[198,1271,551,578]" pageId="7" pageNumber="48">
|
||
<emphasis id="B926EAB5CE2FB63708DAFD8CB41DA749" bold="true" box="[198,339,551,577]" pageId="7" pageNumber="48">Etymology.</emphasis>
|
||
It is a rather small and slender species, hence the name
|
||
<taxonomicName id="4C524D24CE2FB6370BFFFD83B1EDA74A" authority="Latin" authorityName="Latin" box="[995,1187,552,578]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="parvulus">parvulus (Latin)</taxonomicName>
|
||
, small.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348652CCE2FB63708DAFDE4B138A7B2" pageId="7" pageNumber="48" type="discussion">
|
||
<paragraph id="8BED36A7CE2FB63708DAFDE4B138A7B2" blockId="7.[151,1437,152,1098]" pageId="7" pageNumber="48">
|
||
<emphasis id="B926EAB5CE2FB63708DAFDE4B40EA761" bold="true" box="[198,320,591,617]" pageId="7" pageNumber="48">Remarks.</emphasis>
|
||
The most obvious combination of characters is: rostrum long and slender; porose areolae of dorsal and ventral plates distinctly delimited; AD with three small areolae and pair of glp-1 removed from lateral margin; PD with pair of rather short costae not fused with large posterior areola.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348652CCE2FB63708DAFD63B1C5A6F2" pageId="7" pageNumber="48" type="reference_group">
|
||
<paragraph id="8BED36A7CE2FB63708DAFD63B1C5A6F2" blockId="7.[151,1437,152,1098]" pageId="7" pageNumber="48">
|
||
<taxonomicName id="4C524D24CE2FB63708DAFD63B735A7EA" authority="Bartsch, 1999" authorityName="Bartsch" authorityYear="1999" box="[198,635,712,738]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="attalus">
|
||
<emphasis id="B926EAB5CE2FB63708DAFD63B49FA7E9" box="[198,465,712,737]" italics="true" pageId="7" pageNumber="48">Copidognathus attalus</emphasis>
|
||
<bibRefCitation id="EFC34B56CE2FB63709C4FD63B735A7EA" author="Bartsch" box="[472,635,712,738]" pageId="7" pageNumber="49" refString="Bartsch, I. (1999) Halacaridae (Acari) from Western Australia. Four species of Copidognathus. In D. I. Walker and F. E. Wells (eds), Proceedings of the Ninth International Marine Biological Workshop: The Seagrass Flora and Fauna of Rottnest Island, Western Australia, Western Australian Museum, Perth, 333 - 357." type="book chapter" year="1999">Bartsch, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
is strikingly similar to
|
||
<taxonomicName id="4C524D24CE2FB6370B90FD63B15AA7E9" box="[908,1044,712,737]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B926EAB5CE2FB6370B90FD63B15AA7E9" box="[908,1044,712,737]" italics="true" pageId="7" pageNumber="48">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
and shares characters such as (1) porose areolae of dorsal and ventral plates distinctly delimited, (2) three small areolae on AD and (3) glp-1 and ds-1 immediately adjacent. Marked differences are in the ornamentation of the plates and the shape of the rostrum. The porose areolae of the dorsal plates are wider in
|
||
<taxonomicName id="4C524D24CE2FB6370B7CFCEBB6A7A651" box="[864,1001,832,857]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B926EAB5CE2FB6370B7CFCEBB6A7A651" box="[864,1001,832,857]" italics="true" pageId="7" pageNumber="48">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C524D24CE2FB6370C54FCEBB1F4A651" box="[1096,1210,832,857]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="attalus">
|
||
<emphasis id="B926EAB5CE2FB6370C54FCEBB1F4A651" box="[1096,1210,832,857]" italics="true" pageId="7" pageNumber="48">C. attalus</emphasis>
|
||
</taxonomicName>
|
||
; in
|
||
<taxonomicName id="4C524D24CE2FB6370CF4FCEBB03EA651" box="[1256,1392,832,857]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B926EAB5CE2FB6370CF4FCEBB03EA651" box="[1256,1392,832,857]" italics="true" pageId="7" pageNumber="48">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
the integument of the dorsal and ventral plates is smooth outside the porose areolae, in
|
||
<taxonomicName id="4C524D24CE2FB6370C79FCC3B199A689" box="[1125,1239,872,897]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="attalus">
|
||
<emphasis id="B926EAB5CE2FB6370C79FCC3B199A689" box="[1125,1239,872,897]" italics="true" pageId="7" pageNumber="48">C. attalus</emphasis>
|
||
</taxonomicName>
|
||
the dorsal plates are reticulate and the ventral plates bear a delicately foveate ornamentation. The rostrum of
|
||
<taxonomicName id="4C524D24CE2FB6370CEFFC3BB031A6A1" box="[1267,1407,912,937]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B926EAB5CE2FB6370CEFFC3BB031A6A1" box="[1267,1407,912,937]" italics="true" pageId="7" pageNumber="48">C. parvulus</emphasis>
|
||
</taxonomicName>
|
||
is very long, about twice the length of the gnathosomal base, and extends far beyond the base of P-4, the rostrum of
|
||
<taxonomicName id="4C524D24CE2FB63708AAFC4BB467A6F1" box="[182,297,992,1017]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="attalus">
|
||
<emphasis id="B926EAB5CE2FB63708AAFC4BB467A6F1" box="[182,297,992,1017]" italics="true" pageId="7" pageNumber="48">C. attalus</emphasis>
|
||
</taxonomicName>
|
||
is slightly longer than the gnathosomal base, just reaching the end of P-3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348652CCE2FB63708DAFBACB47DA142" pageId="7" pageNumber="48" type="distribution">
|
||
<paragraph id="8BED36A7CE2FB63708DAFBACB47DA142" blockId="7.[151,1437,152,1098]" pageId="7" pageNumber="48">
|
||
<emphasis id="B926EAB5CE2FB63708DAFBACB429A129" bold="true" box="[198,359,1031,1057]" pageId="7" pageNumber="48">Distribution.</emphasis>
|
||
Western
|
||
<collectingCountry id="F3457637CE2FB63709C4FBA3B709A12A" box="[472,583,1032,1058]" name="Australia" pageId="7" pageNumber="48">Australia</collectingCountry>
|
||
, Esperance. The only specimen collected lived amongst corallines on a demosponge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |