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<mods:title id="385248161677C4DB3E697F6975A9C60B">Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera)</mods:title>
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<subSubSection id="C3486534FFC6FFA4FE5A6AE4FCDAFF35" box="[410,792,221,247]" pageId="33" pageNumber="26" type="nomenclature">
<paragraph id="8BED36BFFFC6FFA4FE5A6AE4FCDAFF35" blockId="33.[410,792,221,247]" box="[410,792,221,247]" pageId="33" pageNumber="26">
<heading id="D0A581D3FFC6FFA4FE5A6AE4FCDAFF35" bold="true" box="[410,792,221,247]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="26" reason="2">
<emphasis id="B926EAADFFC6FFA4FE5A6AE4FCDAFF35" bold="true" box="[410,792,221,247]" pageId="33" pageNumber="26">
Genus
<taxonomicName id="4C524D3CFFC6FFA4FE296AE5FCDAFF35" ID-CoL="6353W" ID-ENA="500309" authority="Rohacek, 1999" authorityName="Rohacek" authorityYear="1999" box="[489,792,221,247]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC6FFA4FE296AE5FDA9FF35" bold="true" box="[489,619,223,246]" italics="true" pageId="33" pageNumber="26">Fungomyza</emphasis>
Roháček, 1999
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3486534FFC6FFA4FFB16B3CFE51FE8E" pageId="33" pageNumber="26" type="reference_group">
<paragraph id="8BED36BFFFC6FFA4FFB16B3CFE51FE8E" blockId="33.[113,1090,260,333]" pageId="33" pageNumber="26">
<treatmentCitationGroup id="AB421191FFC6FFA4FFB16B3CFE51FE8E" pageId="33" pageNumber="26">
<treatmentCitation id="0AF310AEFFC6FFA4FFB16B3CFE4EFED4" author="ROHACEK J." box="[113,396,260,279]" page="391" pageId="33" pageNumber="26" year="1999">
<taxonomicName id="4C524D3CFFC6FFA4FFB16B3CFE4EFED4" ID-CoL="6353W" authority="Rohacek, 1999 d: 391" authorityName="Rohacek" authorityPageNumber="391" authorityYear="1999" box="[113,396,260,279]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC6FFA4FFB16B3CFF16FED4" box="[113,212,262,279]" italics="true" pageId="33" pageNumber="26">Fungomyza</emphasis>
<bibRefCitation id="EFC34B4EFFC6FFA4FF196B3EFE4EFED4" author="ROHACEK J." box="[217,396,260,279]" pageId="33" pageNumber="26" pagination="373 - 404" refId="ref334295" refString="ROHACEK J. 1999 d: A revision and re-classincation of the genus Paranthomyza Czerny, with description of a new genus of Anthomyzidae. Studia Dipterologica 6: 373 - 404." type="journal article" year="1999">Roháček, 1999d: 391</bibRefCitation>
</taxonomicName>
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[feminine];
<treatmentCitation id="0AF310AEFFC6FFA4FE346B3FFD3DFED4" author="ROHACEK J. &amp; BARBER K. N." box="[500,767,260,280]" page="132" pageId="33" pageNumber="26" year="2004">
<bibRefCitation id="EFC34B4EFFC6FFA4FE346B3FFD1AFED4" author="ROHACEK J. &amp; BARBER K. N." box="[500,728,260,280]" pageId="33" pageNumber="26" pagination="131 - 141" refId="ref335772" refString="ROHACEK J. &amp; BARBER K. N. 2004: A new species of the genus Fungomyza from the Nearctic Region (Diptera: Anthomyzidae). Casopis Slezskeho Zemskeho Muzea, Opava (A) 53: 131 - 141." type="journal article" year="2004">ROHÁĆEK &amp; BARBER (2004)</bibRefCitation>
: 132
</treatmentCitation>
133 (redennition);
<treatmentCitation id="0AF310AEFFC6FFA4FC686B3FFF72FEF1" author="ROHACEK J." page="214" pageId="33" pageNumber="26" year="2006">
<bibRefCitation id="EFC34B4EFFC6FFA4FC686B3FFBFCFED4" author="ROHACEK J." box="[936,1086,260,280]" pageId="33" pageNumber="26" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK (2006a)</bibRefCitation>
: 214
</treatmentCitation>
215 (redescription); ROHÁĆEK (2009): 8493, 110111 (redescription, phylogeny);
<treatmentCitation id="0AF310AEFFC6FFA4FC496B25FF31FE8E" author="ROHACEK J. &amp; TOTHOVA A." page="169" pageId="33" pageNumber="26" year="2014">
<bibRefCitation id="EFC34B4EFFC6FFA4FC496B25FF09FE8E" author="ROHACEK J. &amp; TOTHOVA A." pageId="33" pageNumber="26" pagination="165 - 176" refId="ref336515" refString="ROHACEK J. &amp; TOTHOVA A. 2014: Morphology versus DNA - what will bring clarity to the relationships of phylogenetically unclear genera of Anthomyzidae (Diptera)? Arthropod Systematics &amp; Phylogeny 72: 165 - 176." type="journal article" year="2014">ROHÁĆEK &amp; TÓTHOVÁ (2014)</bibRefCitation>
: 169
</treatmentCitation>
170 (phylogeny).
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</paragraph>
</subSubSection>
<subSubSection id="C3486534FFC6FFA4FFB16B5FFC63FEBE" box="[113,929,357,381]" pageId="33" pageNumber="26" type="type_taxon">
<paragraph id="8BED36BFFFC6FFA4FFB16B5FFC63FEBE" blockId="33.[113,929,357,381]" box="[113,929,357,381]" pageId="33" pageNumber="26">
<emphasis id="B926EAADFFC6FFA4FFB16B5FFEC7FEBE" bold="true" box="[113,261,357,381]" pageId="33" pageNumber="26">
<typeStatus id="54E9881DFFC6FFA4FFB16B5FFF6BFEBE" box="[113,169,357,381]" pageId="33" pageNumber="26">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C524D3CFFC6FFA4FECC6B5DFD6FFEBE" authority="Meigen, 1830: 107" authorityName="Meigen" authorityPageNumber="107" authorityYear="1830" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[268,685,357,381]" class="Insecta" family="Heleomyzidae" genus="Opomyza" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="26" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="B926EAADFFC6FFA4FECC6B5DFE1EFEBE" box="[268,476,359,381]" italics="true" pageId="33" pageNumber="26">Opomyza albimana</emphasis>
<bibRefCitation id="EFC34B4EFFC6FFA4FE226B5FFD6FFEBE" author="MEIGEN J. W." box="[482,685,357,381]" pageId="33" pageNumber="26" refId="ref329671" refString="MEIGEN J. W. 1830: Systematische Beschreibung der bekannten europaischen zweissugeligen Insekten. Vol. 6. Schulzische Buchhandlung, Hamm, xi + 401 pp." type="book" year="1830">Meigen, 1830: 107</bibRefCitation>
</taxonomicName>
(original designation).
</paragraph>
</subSubSection>
<subSubSection id="C3486534FFC6FFA7FFB16BAEFE17FC10" lastPageId="34" lastPageNumber="31" pageId="33" pageNumber="26" type="diagnosis">
<paragraph id="8BED36BFFFC6FFA4FFB16BAEFBDDFB8F" blockId="33.[113,1090,403,1548]" pageId="33" pageNumber="26">
<emphasis id="B926EAADFFC6FFA4FFB16BAEFF24FE6F" bold="true" box="[113,230,404,428]" pageId="33" pageNumber="26">Diagnosis.</emphasis>
(1)
<emphasis id="B926EAADFFC6FFA4FED66BA9FE93FE6F" bold="true" box="[278,337,403,428]" italics="true" pageId="33" pageNumber="26">Head</emphasis>
distinctly higher than long. (2) Eye large, broadly oval, with longest diameter almost vertical. (3) Frons moderately broad; frontal triangle variable in length, shining or sparsely microtomentose. (4) Frontal lunule small but always distinct. (5) Antenna geniculate between pedicel and 1st nagellomere, the latter strongly compressed laterally. (6) Arista short-ciliate. (7) Palpus yellow, slender but somewhat clavate, with 1 longer subapical seta. Cephalic chaetotaxy: (8) pvt relatively long, crossed or strongly convergent; (9) vte, vti and oc long; (10) 3 ors, anterior slightly or distinctly shorter; 1 microsetula in front of anterior ors; (11) postocular setulae short, in single row; (12) 1 relatively short vi, subvibrissa indistinguishable from peristomal setulae; (13) peristomal setulae small and variable in number. (14)
<emphasis id="B926EAADFFC6FFA4FF636889FF36FD0F" bold="true" box="[163,244,691,716]" italics="true" pageId="33" pageNumber="26">Thorax</emphasis>
slightly narrower than head, more or less shining, despite some microtomentum. Thoracic chaetotaxy: (15) 1 hu, 2 npl (anterior longer); (16) 1 medium-sized sa, 1 longer pa; (17) 1 distinct, relatively long prs; (18) 2 postsutural dc, both long and strong; (19) ac microsetae small and numerous, in 4 rows, ending in front of level of posterior dc; (20) 2 sc (apical strong, laterobasal short); (21) 1 minute upcurved ppl; (22) 2 distinct stpl (anterior shorter) and a few setulae in dorsal half of sternopleuron. (23) At least one pair of femora variegated (yellow and brown); (24) f
<subScript id="17D634FAFFC6FFA4FE3669BEFE3CFC50" attach="left" box="[502,510,900,915]" fontSize="6" pageId="33" pageNumber="26">1</subScript>
with relatively short ctenidial spine; (25) t
<subScript id="17D634FAFFC6FFA4FC7369BEFC79FC50" attach="left" box="[947,955,900,915]" fontSize="6" pageId="33" pageNumber="26">2</subScript>
with distinct ventroapical seta; (26) male f
<subScript id="17D634FAFFC6FFA4FE67699EFE6DFC70" attach="left" box="[423,431,932,947]" fontSize="6" pageId="33" pageNumber="26">3</subScript>
with posteroventral row of setae, distal of which are short and thick. (27) Wing long, moderately narrow; (28) wing membrane unicolourous. (29) C with inconspicuous spinulae among nne setulae on Cs
<subScript id="17D634FAFFC6FFA4FDB369DEFDB9FC30" attach="left" box="[627,635,996,1011]" fontSize="6" pageId="33" pageNumber="26">2</subScript>
; (30) R
<subScript id="17D634FAFFC6FFA4FD0B69DEFD21FC30" attach="left" box="[715,739,996,1011]" fontSize="6" pageId="33" pageNumber="26">2+3</subScript>
long, parallel to C, ending nearer than or at the same distance from apex of R
<subScript id="17D634FAFFC6FFA4FD886E3EFDA2FBD0" attach="left" box="[584,608,1028,1043]" fontSize="6" pageId="33" pageNumber="26">4+5</subScript>
as M; (31) R
<subScript id="17D634FAFFC6FFA4FD316E3EFCCBFBD0" attach="left" box="[753,777,1028,1043]" fontSize="6" pageId="33" pageNumber="26">4+5</subScript>
slightly bent to sinuate; (32) cell dm relatively long; cross-vein r-m situated distal to middle of dm cell. (33) CuA
<subScript id="17D634FAFFC6FFA4FC2A6E1EFC30FBF0" attach="left" box="[1002,1010,1060,1075]" fontSize="6" pageId="33" pageNumber="26">1</subScript>
ending near, A
<subScript id="17D634FAFFC6FFA4FF7D6E7EFF07FB90" attach="left" box="[189,197,1092,1107]" fontSize="6" pageId="33" pageNumber="26">1</subScript>
far from wing margin. (34) Anal lobe and alula well developed, relatively large.
</paragraph>
<paragraph id="8BED36BFFFC6FFA4FF506E69FC64FB2F" blockId="33.[113,1090,403,1548]" pageId="33" pageNumber="26">
<emphasis id="B926EAADFFC6FFA4FF506E69FEF8FBAF" bold="true" box="[144,314,1107,1132]" italics="true" pageId="33" pageNumber="26">Male abdomen.</emphasis>
(35) T1 separate from T2, at least dorsally; (36) T2T5 large and broad. (37) S1S5 of variable width, dark but slightly paler than associated terga. Male postabdomen: (38) T6 short, transverse, bare and well sclerotized although paler-pigmented than S6S8. (39) S6 and S7 strongly asymmetrical, partly fused and situated laterally, each with a few setulae. (40) S8 unusually long, less asymmetrical, more setose and situated dorsally.
</paragraph>
<paragraph id="8BED36BFFFC6FFA7FF506EC9FE7FFE10" blockId="33.[113,1090,403,1548]" lastBlockId="34.[151,1128,219,979]" lastPageId="34" lastPageNumber="31" pageId="33" pageNumber="26">
<emphasis id="B926EAADFFC6FFA4FF506EC9FEFAFACF" bold="true" box="[144,312,1267,1292]" italics="true" pageId="33" pageNumber="26">Male genitalia.</emphasis>
(41) Epandrium large, relatively densely setose, with 23 pairs longer. (42) Anal nssure very small, narrowly rounded triangular. (43) Medandrium high, slightly to conspicuously narrowed dorsally. (44) Cercus simple in shape, variable in size. (45) Gonostylus with 23 denticles on apex, and with external micropubescence partly or entirely reduced. (46) Hypandrium with anterior nat lobes not or slightly projecting dorsally; (47) transandrium with a forked or medially divided caudal process. (48) Pregonite large, nat, fused to hypandrium, with 2 groups of setae. (49) Postgonite simple, slender, long and relatively straight. (50) Phallapodeme with normal apex and shortly bifurcate symmetrical base. (51) Phallophore short, with more or less distinct ventral process; (52) distiphallus composed of distally membranous saccus and slender sclerotized nlum. (53) Saccus with relatively small membranous part, distinct basal and internal sclerites, and unarmed or with adpressed spinulae and/or setulae on surface; (54) nlum formed by 2 long, dark, band-like sclerites fused basally and apically, with simple apex. (55) Aedeagal part of folding apparatus with reduced connecting sclerite and its external wall provided with small grains, tubercles or short spines combined with dark striae. (56) Basal membrane below caudal process with dense small teeth or spines. (57) Ejacapodeme small to medium-sized, with subterminal digitiform process or modined (in some species).
</paragraph>
<paragraph id="8BED36BFFFC5FFA7FF756BE1FE17FC10" blockId="34.[151,1128,219,979]" pageId="34" pageNumber="31">
(58)
<emphasis id="B926EAADFFC5FFA7FF276BE0FE67FE30" bold="true" box="[231,421,474,499]" italics="true" pageId="34" pageNumber="31">Female abdomen</emphasis>
relatively shining, with broader terga (T2T6) and narrower sterna (S2S5). (59) Postabdomen long, basally broad, caudally strongly tapered, telescopically retractable from 7th segment, with less sclerotized and paler T8, S8, T10 and S10. (60) T6 broad, S6 relatively small, narrow. (61) T7 and S7 separate, S7 small, narrow, paler than T7. (62) T8 simple, relatively nat, pale-pigmented; (63) S8 narrow, medially longitudinally divided; its posterior bare parts dorsally curved and far invaginated into 8th segment. (64) Internal sclerites of female genital chamber (uterus) weakly sclerotized and reduced, formed by 12 pairs of partly fused posterior plates and (65) 1 anteroventral, broad but thin, incomplete and unpigmented annular sclerite. (66) Anterior part of uterus provided with an elongate, vesicular, weakly sclerotized ventral receptacle terminating in short projecting apex of various ending. (67) Accessory glands relatively small, on slightly dilated ringed ducts. (68) Spermathecae (1+1) roughly cup-like to pyriform, characterized by deep terminal invagination and narrowed basal part covered by nne spinulae; spermathecal ducts short. (69) T10 long and narrow, bare except for 1 pair of dorsal setae; (70) S10, simple, pale, slightly longer than T10, almost without micropubescence. (71) Cercus long and very slender, with a number of longer nne setae.
</paragraph>
</subSubSection>
<subSubSection id="C3486534FFC5FFA1FF5769D0FB8AFEB7" lastPageId="36" lastPageNumber="33" pageId="34" pageNumber="31" type="discussion">
<paragraph id="8BED36BFFFC5FFA6FF5769D0FD6CFEF5" blockId="34.[151,1128,1002,1539]" lastBlockId="35.[113,1089,222,1558]" lastPageId="35" lastPageNumber="28" pageId="34" pageNumber="31">
<emphasis id="B926EAADFFC5FFA7FF5769D0FEDAFBC1" bold="true" box="[151,280,1002,1026]" pageId="34" pageNumber="31">Discussion.</emphasis>
The above generic diagnosis follows
<bibRefCitation id="EFC34B4EFFC5FFA7FD0669D1FC4AFBC0" author="ROHACEK J." box="[710,904,1003,1027]" pageId="34" pageNumber="31" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK (2009a)</bibRefCitation>
, including the most ancestral E. Palaearctic relative,
<taxonomicName id="4C524D3CFFC5FFA7FE376E37FCCFFBE0" authority="Rohacek, 2009" authorityName="Rohacek" authorityYear="2009" box="[503,781,1035,1059]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="species" species="cercata">
<emphasis id="B926EAADFFC5FFA7FE376E37FDA3FBE0" box="[503,609,1037,1059]" italics="true" pageId="34" pageNumber="31">F. cercata</emphasis>
Roháček, 2009
</taxonomicName>
. Although outer appearance and several other features indicate the superncial similarity of
<taxonomicName id="4C524D3CFFC5FFA7FCE46E17FC63FB80" authorityName="Rohacek" authorityYear="1999" box="[804,929,1069,1091]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FCE46E17FC63FB80" box="[804,929,1069,1091]" italics="true" pageId="34" pageNumber="31">Fungomyza</emphasis>
</taxonomicName>
to
<taxonomicName id="4C524D3CFFC5FFA7FC0A6E17FEEBFBA0" authority="Czerny, 1902" authorityName="Czerny" authorityYear="1902" class="Insecta" family="Anthomyzidae" genus="Paranthomyza" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FC0A6E17FBA4FB80" box="[970,1126,1069,1091]" italics="true" pageId="34" pageNumber="31">Paranthomyza</emphasis>
<bibRefCitation id="EFC34B4EFFC5FFA7FF576E71FEEBFBA0" author="CZERNY L." box="[151,297,1099,1123]" pageId="34" pageNumber="31" pagination="249 - 256" refId="ref324724" refString="CZERNY L. 1902: Bemerkungen zu den Arten der Gattungen Anthomyza Fll. und Ischnomyia Lw. Wiener Entomologische Zeitung 21: 249 - 256." type="journal article" year="1902">Czerny, 1902</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C524D3CFFC5FFA7FEF46E77FE7AFBA0" authorityName="Rohacek" authorityYear="2009" box="[308,440,1101,1123]" class="Insecta" family="Anthomyzidae" genus="Carexomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FEF46E77FE7AFBA0" box="[308,440,1101,1123]" italics="true" pageId="34" pageNumber="31">Carexomyza</emphasis>
</taxonomicName>
or
<taxonomicName id="4C524D3CFFC5FFA7FE1E6E77FDA0FBA0" authorityName="Rohacek &amp; Barber" authorityYear="2011" box="[478,610,1101,1123]" class="Insecta" family="Anthomyzidae" genus="Quametopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FE1E6E77FDA0FBA0" box="[478,610,1101,1123]" italics="true" pageId="34" pageNumber="31">Quametopia</emphasis>
</taxonomicName>
(e.g. 1, 14, 53, 54, 58), previous morphological (
<bibRefCitation id="EFC34B4EFFC5FFA7FF5F6E51FE8FFB40" author="ROHACEK J." box="[159,333,1131,1155]" pageId="34" pageNumber="31" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
) and molecular (ROHÁĆEK et al. 2009) phylogenetic analyses connrmed its association with the
<taxonomicName id="4C524D3CFFC5FFA7FE566EB7FDCFFB60" authorityName="Fallen" authorityYear="1810" box="[406,525,1165,1187]" class="Insecta" family="Anthomyzidae" genus="Anthomyza" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FE566EB7FDCFFB60" box="[406,525,1165,1187]" italics="true" pageId="34" pageNumber="31">Anthomyza</emphasis>
</taxonomicName>
clade (see
<figureCitation id="13692A3AFFC5FFA7FD476EB1FD23FB60" box="[647,737,1163,1187]" captionStart="Figs 136138" captionStartId="91.[113,150,1509,1528]" captionTargetBox="[129,1048,198,1476]" captionTargetId="figure-64@91.[126,1075,197,1509]" captionTargetPageId="91" captionText="Figs 136138. Arganthomyza bivittata Roháček &amp; Barber, 2013, paratype female (Canada: Ontario). 136 postabdomen, dorsally; 137 the same, ventrally; 138 the same, laterally. Scale = 0.2 mm. For abbreviations see p. 11. Adapted from ROHÁĆEK &amp; BARBER (2013: Figs 9092)." figureDoi="http://doi.org/10.5281/zenodo.4272897" httpUri="https://zenodo.org/record/4272897/files/figure.png" pageId="34" pageNumber="31">
Fig.
<quantity id="4CAA9B5AFFC5FFA7FD796EB1FD23FB60" box="[697,737,1163,1187]" metricMagnitude="0" metricUnit="m" metricValue="3.4798" pageId="34" pageNumber="31" unit="in" value="137.0">137</quantity>
</figureCitation>
in
<bibRefCitation id="EFC34B4EFFC5FFA7FCC56EB1FC70FB60" author="ROHACEK J." box="[773,946,1163,1187]" pageId="34" pageNumber="31" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
). Based on these phylogenetic hypotheses, there is support for a sister-group relationship between
<taxonomicName id="4C524D3CFFC5FFA7FC2A6E97FBA4FB00" authorityName="Rohacek" authorityYear="1999" box="[1002,1126,1197,1219]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FC2A6E97FBA4FB00" box="[1002,1126,1197,1219]" italics="true" pageId="34" pageNumber="31">Fungomyza</emphasis>
</taxonomicName>
and a lineage consisting of
<taxonomicName id="4C524D3CFFC5FFA7FE726EF7FD8FFB20" authorityName="Rohacek" authorityYear="2009" box="[434,589,1229,1251]" class="Insecta" family="Anthomyzidae" genus="Arganthomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FE726EF7FD8FFB20" box="[434,589,1229,1251]" italics="true" pageId="34" pageNumber="31">Arganthomyza</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D3CFFC5FFA7FDBD6EF7FD3AFB20" authorityName="Loew" authorityYear="1863" box="[637,760,1229,1251]" class="Insecta" family="Anthomyzidae" genus="Ischnomyia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FDBD6EF7FD3AFB20" box="[637,760,1229,1251]" italics="true" pageId="34" pageNumber="31">Ischnomyia</emphasis>
</taxonomicName>
as here redenned. The relationship of
<taxonomicName id="4C524D3CFFC5FFA7FF766ED7FEF0FAC0" authorityName="Rohacek" authorityYear="1999" box="[182,306,1261,1283]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FF766ED7FEF0FAC0" box="[182,306,1261,1283]" italics="true" pageId="34" pageNumber="31">Fungomyza</emphasis>
</taxonomicName>
to this sister pair seems to be supported by the following synapomorphies: caudal process of transandrium forked or medially desclerotized; female S8 longitudinally divided but with posterior bare parts recurved and deeply invaginated into 8th abdominal segment (strongest synapomorphy); spermathecal ducts markedly shortened (only in
<taxonomicName id="4C524D3CFFC5FFA7FBF76F77FEC9FA40" authorityName="Rohacek" authorityYear="2009" class="Insecta" family="Anthomyzidae" genus="Arganthomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FBF76F77FEC9FA40" italics="true" pageId="34" pageNumber="31">Arganthomyza</emphasis>
</taxonomicName>
, not in
<taxonomicName id="4C524D3CFFC5FFA7FEA26F57FE1FFA40" authorityName="Loew" authorityYear="1863" box="[354,477,1389,1411]" class="Insecta" family="Anthomyzidae" genus="Ischnomyia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FEA26F57FE1FFA40" box="[354,477,1389,1411]" italics="true" pageId="34" pageNumber="31">Ischnomyia</emphasis>
</taxonomicName>
); well-sclerotized abdominal sterna; similar construction of ventral receptacle (see
<bibRefCitation id="EFC34B4EFFC5FFA7FE4E6FB1FDF9FA60" author="ROHACEK J." box="[398,571,1419,1443]" pageId="34" pageNumber="31" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
). The sister clade of
<taxonomicName id="4C524D3CFFC5FFA7FCDA6FB7FC77FA60" authorityName="Rohacek" authorityYear="2009" box="[794,949,1421,1443]" class="Insecta" family="Anthomyzidae" genus="Arganthomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FCDA6FB7FC77FA60" box="[794,949,1421,1443]" italics="true" pageId="34" pageNumber="31">Arganthomyza</emphasis>
</taxonomicName>
+
<taxonomicName id="4C524D3CFFC5FFA7FC126FB7FB8FFA60" authorityName="Loew" authorityYear="1863" box="[978,1101,1421,1443]" class="Insecta" family="Anthomyzidae" genus="Ischnomyia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FC126FB7FB8FFA60" box="[978,1101,1421,1443]" italics="true" pageId="34" pageNumber="31">Ischnomyia</emphasis>
</taxonomicName>
is supported by modincation of the nlum of the distiphallus (sclerites expanded and attached or partly fused) and partial to complete coalescence of the female T7 and S7 to form a compact syntergosternite. The poorly known genus
<taxonomicName id="4C524D3CFFC5FFA7FDA06FD7FD16F9C0" authorityName="Rohacek" authorityYear="2006" box="[608,724,1517,1539]" class="Insecta" family="Anthomyzidae" genus="Receptrixa" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC5FFA7FDA06FD7FD16F9C0" box="[608,724,1517,1539]" italics="true" pageId="34" pageNumber="31">Receptrixa</emphasis>
</taxonomicName>
could also be a candidate for a closer (but highly modined) relative of all the above-discussed genera (shared features: short spermathecal ducts, distally positioned cross-vein r-m), but because the male of its only species is unknown this hypothesis cannot be properly tested.
</paragraph>
<paragraph id="8BED36BFFFC4FFA6FF506B04FBDCFD35" blockId="35.[113,1089,222,1558]" pageId="35" pageNumber="28">
The most recent hypothesis of phylogenetic relationships of anthomyzid genera based on multigene analyses (
<bibRefCitation id="EFC34B4EFFC4FFA6FE8E6B64FDB3FEB5" author="ROHACEK J. &amp; TOTHOVA A." box="[334,625,350,374]" pageId="35" pageNumber="28" pagination="165 - 176" refId="ref336515" refString="ROHACEK J. &amp; TOTHOVA A. 2014: Morphology versus DNA - what will bring clarity to the relationships of phylogenetically unclear genera of Anthomyzidae (Diptera)? Arthropod Systematics &amp; Phylogeny 72: 165 - 176." type="journal article" year="2014">ROHÁĆEK &amp; TÓTHOVÁ 2014</bibRefCitation>
) surprisingly separated
<taxonomicName id="4C524D3CFFC4FFA6FCB46B5AFC32FEB5" authorityName="Rohacek" authorityYear="1999" box="[884,1008,352,374]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FCB46B5AFC32FEB5" box="[884,1008,352,374]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
(represented by its
<typeStatus id="54E9881DFFC4FFA6FEC36B45FEF3FE54" box="[259,305,383,407]" pageId="35" pageNumber="28">type</typeStatus>
species, the European
<taxonomicName id="4C524D3CFFC4FFA6FDEE6BBAFCAAFE55" authority="(Meigen, 1830))" authorityName="Meigen" authorityYear="1830" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[558,872,382,406]" class="Insecta" family="Heleomyzidae" genus="Opomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="B926EAADFFC4FFA6FDEE6BBAFD6DFE55" box="[558,687,384,406]" italics="true" pageId="35" pageNumber="28">F. albimana</emphasis>
(
<bibRefCitation id="EFC34B4EFFC4FFA6FD7F6B44FC95FE55" author="MEIGEN J. W." box="[703,855,382,406]" pageId="35" pageNumber="28" refId="ref329671" refString="MEIGEN J. W. 1830: Systematische Beschreibung der bekannten europaischen zweissugeligen Insekten. Vol. 6. Schulzische Buchhandlung, Hamm, xi + 401 pp." type="book" year="1830">Meigen, 1830</bibRefCitation>
))
</taxonomicName>
as a sister group to all remaining analysed taxa of the family, thus excluding it from the
<taxonomicName id="4C524D3CFFC4FFA6FC8E6B9AFC07FE75" authorityName="Fallen" authorityYear="1810" box="[846,965,416,438]" class="Insecta" family="Anthomyzidae" genus="Anthomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FC8E6B9AFC07FE75" box="[846,965,416,438]" italics="true" pageId="35" pageNumber="28">Anthomyza</emphasis>
</taxonomicName>
clade. This relatively well-supported nnding contradicts the earlier molecular study of ROHÁĆEK et al. (2009) and, particularly, the current morphological knowledge (see above). Hitherto, we do not know of any morphological synapomorphy supporting the monophyly of the clade with all these (remaining) taxa. They, however, differ from
<taxonomicName id="4C524D3CFFC4FFA6FD47681AFCC1FDF5" authorityName="Rohacek" authorityYear="1999" box="[647,771,544,566]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FD47681AFCC1FDF5" box="[647,771,544,566]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
in having microsaprophagous larvae, feeding on at least partly rotten tissues of plants, while those of
<taxonomicName id="4C524D3CFFC4FFA6FCB0687AFC2EFD95" authorityName="Rohacek" authorityYear="1999" box="[880,1004,576,598]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FCB0687AFC2EFD95" box="[880,1004,576,598]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
species are mycophagous, developing in sporocarps of macrofungi (see DELY- DRASKOVITS 1972, CHANDLER 1978,
<bibRefCitation id="EFC34B4EFFC4FFA6FEE46845FE0DFD55" author="ROHACEK J." box="[292,463,638,662]" pageId="35" pageNumber="28" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
). The other possibility is that
<taxonomicName id="4C524D3CFFC4FFA6FCC768BAFC41FD55" authorityName="Rohacek" authorityYear="1999" box="[775,899,640,662]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FCC768BAFC41FD55" box="[775,899,640,662]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
actually diverged basally from the common stem of the
<taxonomicName id="4C524D3CFFC4FFA6FDCB68A4FD6AFD75" authorityName="Czerny" authorityYear="1903" box="[523,680,670,694]" class="Insecta" family="Anthomyzidae" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="subFamily" subFamily="Anthomyzinae">Anthomyzinae</taxonomicName>
as the results of
<bibRefCitation id="EFC34B4EFFC4FFA6FC9F68A5FF7AFD15" author="ROHACEK J. &amp; TOTHOVA A." pageId="35" pageNumber="28" pagination="165 - 176" refId="ref336515" refString="ROHACEK J. &amp; TOTHOVA A. 2014: Morphology versus DNA - what will bring clarity to the relationships of phylogenetically unclear genera of Anthomyzidae (Diptera)? Arthropod Systematics &amp; Phylogeny 72: 165 - 176." type="journal article" year="2014">ROHÁĆEK &amp; TÓTHOVÁ (2014)</bibRefCitation>
indicate; in this case, however, the former hypotheses based on morphological data would have to be incorrect (probably due to misinterpreted polarity of some characters).
</paragraph>
<paragraph id="8BED36BFFFC4FFA6FF5068C4FEF0FB15" blockId="35.[113,1089,222,1558]" pageId="35" pageNumber="28">
The most diagnostic characters of
<taxonomicName id="4C524D3CFFC4FFA6FE32693AFDACFCD5" authorityName="Rohacek" authorityYear="1999" box="[498,622,768,790]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FE32693AFDACFCD5" box="[498,622,768,790]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
(those apomorphic are marked “A” in parentheses) are: (1) eye with longest diameter almost vertical; (12) subvibrissa not developed (A); (17) 1 long prs; (23) at least one pair of femora variegated (bicolourous) (A); (32) cross-vein r-m situated distal to middle of cell dm; (38) male T6 short but well sclerotized; (40) male S8 unusually long (A); (42) anal nssure very small, narrowly rounded triangular (A); (43) medandrium high, more or less narrowed dorsally; (51) phallophore short, with more or less distinct ventral process; (53) saccus of distiphallus with membranous part having adpressed surface spinulae (A); (63) S8 narrow, longitudinally divided and invaginated into 8th segment (A); (64) internal paired sclerites of female genital chamber (uterus) weakly developed; (65) internal looped structure (homologous to annular sclerite) nne, non-sclerotized (?A); (68) spermatheca with terminal invagination (A). Most of the apomorphic features listed above are not unique for this genus some of them are shared with related genera of the
<taxonomicName id="4C524D3CFFC4FFA6FC0A6E5AFB83FBB5" authorityName="Fallen" authorityYear="1810" box="[970,1089,1120,1142]" class="Insecta" family="Anthomyzidae" genus="Anthomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FC0A6E5AFB83FBB5" box="[970,1089,1120,1142]" italics="true" pageId="35" pageNumber="28">Anthomyza</emphasis>
</taxonomicName>
clade, especially
<taxonomicName id="4C524D3CFFC4FFA6FEE76EBAFE00FB55" authorityName="Rohacek" authorityYear="2009" box="[295,450,1152,1174]" class="Insecta" family="Anthomyzidae" genus="Arganthomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FEE76EBAFE00FB55" box="[295,450,1152,1174]" italics="true" pageId="35" pageNumber="28">Arganthomyza</emphasis>
</taxonomicName>
. The larval mycophagy of
<taxonomicName id="4C524D3CFFC4FFA6FD1E6EBAFC98FB55" authorityName="Rohacek" authorityYear="1999" box="[734,858,1152,1174]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FD1E6EBAFC98FB55" box="[734,858,1152,1174]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
(though unconnrmed in
<taxonomicName id="4C524D3CFFC4FFA6FF506E9AFF3CFB75" authorityName="Rohacek" authorityYear="2009" box="[144,254,1184,1206]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="cercata">
<emphasis id="B926EAADFFC4FFA6FF506E9AFF3CFB75" box="[144,254,1184,1206]" italics="true" pageId="35" pageNumber="28">F. cercata</emphasis>
</taxonomicName>
) can be considered a non-morphological autapotypic feature of this genus (
<bibRefCitation id="EFC34B4EFFC4FFA6FFB96E84FEE4FB15" author="ROHACEK J." box="[121,294,1214,1238]" pageId="35" pageNumber="28" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK 2006a</bibRefCitation>
).
</paragraph>
<paragraph id="8BED36BFFFC4FFA6FF506EDAFDDCF9D5" blockId="35.[113,1089,222,1558]" pageId="35" pageNumber="28">
<taxonomicName id="4C524D3CFFC4FFA6FF506EDAFECEFB35" authorityName="Rohacek" authorityYear="1999" box="[144,268,1248,1270]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FF506EDAFECEFB35" box="[144,268,1248,1270]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
species can appear externally similar to other darkly coloured and relatively shining
<taxonomicName id="4C524D3CFFC4FFA6FF036EC4FEA0FAD5" box="[195,354,1278,1302]" class="Insecta" family="Anthomyzidae" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="family">Anthomyzidae</taxonomicName>
, including some
<taxonomicName id="4C524D3CFFC4FFA6FDCF6F3AFD6BFAD5" authorityName="Rohacek" authorityYear="2009" box="[527,681,1280,1302]" class="Insecta" family="Anthomyzidae" genus="Arganthomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FDCF6F3AFD6BFAD5" box="[527,681,1280,1302]" italics="true" pageId="35" pageNumber="28">Arganthomyza</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D3CFFC4FFA6FD196F3AFC8DFAD5" authorityName="Fallen" authorityYear="1810" box="[729,847,1280,1302]" class="Insecta" family="Anthomyzidae" genus="Anthomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FD196F3AFC8DFAD5" box="[729,847,1280,1302]" italics="true" pageId="35" pageNumber="28">Anthomyza</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C524D3CFFC4FFA6FC6B6F3AFE37FAF5" authority="(Rohacek &amp; Freidberg, 1993)" baseAuthorityName="Rohacek &amp; Freidberg" baseAuthorityYear="1993" class="Insecta" family="Anthomyzidae" genus="Receptrixa" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="receptrix">
<emphasis id="B926EAADFFC4FFA6FC6B6F3AFF78FAF5" italics="true" pageId="35" pageNumber="28">Receptrixa receptrix</emphasis>
(
<bibRefCitation id="EFC34B4EFFC4FFA6FF076F24FE2FFAF5" author="ROHACEK J. &amp; FREIDBERG A." box="[199,493,1310,1334]" pageId="35" pageNumber="28" pagination="61 - 112" refId="ref336250" refString="ROHACEK J. &amp; FREIDBERG A. 1993: The Anthomyzidae (Diptera) of Israel, with revision of Anagnota Becker. Israel Journal of Entomology 27: 61 - 112." type="journal article" year="1993">Roháček &amp; Freidberg, 1993</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C524D3CFFC4FFA6FE3E6F1AFD19FAF5" authorityName="" baseAuthorityName="Meigen" baseAuthorityYear="1838" box="[510,731,1312,1334]" class="Insecta" family="Anthomyzidae" genus="Paranthomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B926EAADFFC4FFA6FE3E6F1AFD19FAF5" box="[510,731,1312,1334]" italics="true" pageId="35" pageNumber="28">Paranthomyza nitida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D3CFFC4FFA6FCCA6F1AFC4FFAF5" authorityName="Rohacek" authorityYear="2009" box="[778,909,1312,1334]" class="Insecta" family="Anthomyzidae" genus="Carexomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FCCA6F1AFC4FFAF5" box="[778,909,1312,1334]" italics="true" pageId="35" pageNumber="28">Carexomyza</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D3CFFC4FFA6FC7E6F1AFB83FAF5" authorityName="Rohacek &amp; Barber" authorityYear="2011" box="[958,1089,1312,1334]" class="Insecta" family="Anthomyzidae" genus="Quametopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FC7E6F1AFB83FAF5" box="[958,1089,1312,1334]" italics="true" pageId="35" pageNumber="28">Quametopia</emphasis>
</taxonomicName>
species.
<taxonomicName id="4C524D3CFFC4FFA6FF0B6F7AFE85FA95" authorityName="Rohacek" authorityYear="1999" box="[203,327,1344,1366]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FF0B6F7AFE85FA95" box="[203,327,1344,1366]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
can be differentiated from these taxa by having more or less variegated legs, a distinctively reduced subvibrissa, sparse ac microsetae and a distally positioned r-m (the latter only known in
<taxonomicName id="4C524D3CFFC4FFA6FE506FBAFDC6FA55" authorityName="Rohacek" authorityYear="2006" box="[400,516,1408,1430]" class="Insecta" family="Anthomyzidae" genus="Receptrixa" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FE506FBAFDC6FA55" box="[400,516,1408,1430]" italics="true" pageId="35" pageNumber="28">Receptrixa</emphasis>
</taxonomicName>
). The recently described
<taxonomicName id="4C524D3CFFC4FFA6FCE06FBAFBC1FA55" authorityName="Rohacek" authorityYear="2013" box="[800,1027,1408,1430]" class="Insecta" family="Anthomyzidae" genus="Reliquantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="variipes">
<emphasis id="B926EAADFFC4FFA6FCE06FBAFBC1FA55" box="[800,1027,1408,1430]" italics="true" pageId="35" pageNumber="28">Reliquantha variipes</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C524D3CFFC4FFA6FEF76F9AFE71FA75" authorityName="Rohacek" authorityYear="1999" box="[311,435,1440,1462]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC4FFA6FEF76F9AFE71FA75" box="[311,435,1440,1462]" italics="true" pageId="35" pageNumber="28">Fungomyza</emphasis>
</taxonomicName>
species, especially
<taxonomicName id="4C524D3CFFC4FFA6FD496F9AFD3BFA75" authorityName="Rohacek &amp; Barber" authorityYear="2004" box="[649,761,1440,1462]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="buccata">
<emphasis id="B926EAADFFC4FFA6FD496F9AFD3BFA75" box="[649,761,1440,1462]" italics="true" pageId="35" pageNumber="28">F. buccata</emphasis>
</taxonomicName>
(including similar variegation of legs) but
<taxonomicName id="4C524D3CFFC4FFA6FF2F6FFAFEA0FA15" authorityName="Rohacek" authorityYear="2013" box="[239,354,1472,1494]" class="Insecta" family="Anthomyzidae" genus="Reliquantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="variipes">
<emphasis id="B926EAADFFC4FFA6FF2F6FFAFEA0FA15" box="[239,354,1472,1494]" italics="true" pageId="35" pageNumber="28">R. variipes</emphasis>
</taxonomicName>
lacks a ctenidial spine on f
<subScript id="17D634FAFFC4FFA6FD406FF4FD4AFA1E" attach="left" box="[640,648,1486,1501]" fontSize="6" pageId="35" pageNumber="28">1</subScript>
and has a distinct subvibrissa, in addition to many other structural differences in the male and female terminalia (see
<bibRefCitation id="EFC34B4EFFC4FFA6FC596FE5FB82FA35" author="ROHACEK J." box="[921,1088,1502,1526]" pageId="35" pageNumber="28" pagination="793 - 814" refId="ref335356" refString="ROHACEK J. 2013 c: Reliquantha variipes gen. &amp; sp. nov., a peculiar new taxon of Anthomyzidae (Diptera) from Great Britain with uncertain relationships. Acta Entomologica Musei Nationalis Pragae 53: 793 - 814." type="journal article" year="2013">ROHÁĆEK 2013c</bibRefCitation>
and discussion under
<taxonomicName id="4C524D3CFFC4FFA6FE986C3AFE04F9D5" authorityName="Rohacek &amp; Barber" authorityYear="2004" box="[344,454,1536,1558]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="28" phylum="Arthropoda" rank="species" species="buccata">
<emphasis id="B926EAADFFC4FFA6FE986C3AFE04F9D5" box="[344,454,1536,1558]" italics="true" pageId="35" pageNumber="28">F. buccata</emphasis>
</taxonomicName>
below).
</paragraph>
<paragraph id="8BED36BFFFC3FFA1FF756AE1FB8AFEB7" blockId="36.[151,1127,219,372]" pageId="36" pageNumber="33">
Three species included:
<taxonomicName id="4C524D3CFFC3FFA1FE7B6AE4FCF4FF37" authority="Rohacek, 2009" authorityName="Rohacek" authorityYear="2009" box="[443,822,219,244]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="cercata">
<emphasis id="B926EAADFFC3FFA1FE7B6AE4FD4CFF37" box="[443,654,222,244]" italics="true" pageId="36" pageNumber="33">Fungomyza cercata</emphasis>
Roháček, 2009
</taxonomicName>
from the Far East of
<collectingCountry id="F345762FFFC3FFA1FBDF6AE1FBA4FF30" box="[1055,1126,219,243]" name="Russia" pageId="36" pageNumber="33">Russia</collectingCountry>
is considered a sister species of the remaining two, viz.
<taxonomicName id="4C524D3CFFC3FFA1FD266AC4FF38FEF0" authority="(Meigen, 1830) (W. Palaearctic)" authorityName="W. Palaearctic" baseAuthorityName="Meigen" baseAuthorityYear="1830" class="Insecta" family="Heleomyzidae" genus="Opomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="B926EAADFFC3FFA1FD266AC4FCA6FED7" box="[742,868,254,276]" italics="true" pageId="36" pageNumber="33">F. albimana</emphasis>
(
<bibRefCitation id="EFC34B4EFFC3FFA1FCB26AC1FBCBFED7" author="MEIGEN J. W." box="[882,1033,251,276]" pageId="36" pageNumber="33" refId="ref329671" refString="MEIGEN J. W. 1830: Systematische Beschreibung der bekannten europaischen zweissugeligen Insekten. Vol. 6. Schulzische Buchhandlung, Hamm, xi + 401 pp." type="book" year="1830">Meigen, 1830</bibRefCitation>
) (W. Palaearctic)
</taxonomicName>
and
<taxonomicName id="4C524D3CFFC3FFA1FEF06B24FC8AFEF0" authority="Rohacek &amp; Barber, 2004" authorityName="Rohacek &amp; Barber" authorityYear="2004" box="[304,840,283,308]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="buccata">
<emphasis id="B926EAADFFC3FFA1FEF06B24FE5DFEF7" box="[304,415,286,308]" italics="true" pageId="36" pageNumber="33">F. buccata</emphasis>
<bibRefCitation id="EFC34B4EFFC3FFA1FE676B21FD71FEF7" author="ROHACEK J. &amp; BARBER K. N." box="[423,691,283,308]" pageId="36" pageNumber="33" pagination="131 - 141" refId="ref335772" refString="ROHACEK J. &amp; BARBER K. N. 2004: A new species of the genus Fungomyza from the Nearctic Region (Diptera: Anthomyzidae). Casopis Slezskeho Zemskeho Muzea, Opava (A) 53: 131 - 141." type="journal article" year="2004">Roháček &amp; Barber, 2004</bibRefCitation>
(E. Nearctic)
</taxonomicName>
. The latter two species are distributed on opposite sides of the Atlantic Ocean. This is a clear example of a sister-pair transatlantic distribution, where the more ancestral species is Nearctic (
<bibRefCitation id="EFC34B4EFFC3FFA1FC4F6B66FBFEFEB7" author="ROHACEK J." box="[911,1084,348,372]" pageId="36" pageNumber="33" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3486534FFC3FFA1FEA06BA6FC5FFE70" box="[352,925,411,436]" pageId="36" pageNumber="33" type="discussion">
<paragraph id="8BED36BFFFC3FFA1FEA06BA6FC5FFE70" blockId="36.[352,925,411,436]" box="[352,925,411,436]" pageId="36" pageNumber="33">
<heading id="D0A581D3FFC3FFA1FEA06BA6FC5FFE70" bold="true" box="[352,925,411,436]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="33" reason="2">
<emphasis id="B926EAADFFC3FFA1FEA06BA6FC5FFE70" bold="true" box="[352,925,411,436]" pageId="36" pageNumber="33">
Key to identincation of
<taxonomicName id="4C524D3CFFC3FFA1FDAB6BA6FD2FFE70" authorityName="Rohacek" authorityYear="1999" box="[619,749,412,435]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAADFFC3FFA1FDAB6BA6FD2FFE70" bold="true" box="[619,749,412,435]" italics="true" pageId="36" pageNumber="33">Fungomyza</emphasis>
</taxonomicName>
species (world)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3486534FFC3FFA1FF576BF1FBA4FBA0" pageId="36" pageNumber="33" type="key">
<keyStep id="30A62E1AFFC3FFA1FF576BF1FBA4FCC0" pageId="36" pageNumber="33">
<paragraph id="8BED36BFFFC3FFA1FF576BF1FBA4FDA0" blockId="36.[151,1127,459,1123]" pageId="36" pageNumber="33">
<keyLead id="30A3958AFFC3FFA1FF576BF1FBA4FDA0" pageId="36" pageNumber="33">
1 Fore femur and tibia uniformly yellow. Gonostylus small, externally without micropubescence, and apex with 3 denticles (
<bibRefCitation id="EFC34B4EFFC3FFA1FD9C6BD1FCCAFDC0" author="ROHACEK J." box="[604,776,491,515]" pageId="36" pageNumber="33" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FCD26BD1FCAFFDC1" box="[786,877,490,515]" captionStart="Figs 125127" captionStartId="87.[115,152,1484,1503]" captionTargetBox="[117,1086,348,1455]" captionTargetId="figure-52@87.[117,1086,347,1473]" captionTargetPageId="87" captionText="Figs 125127. Wings of the Nearctic species ofArganthomyza. 125 A. acuticuspis Roháček &amp; Barber, 2013, paratype male, wing length 2.5 mm (USA: New Mexico); 126 A. bivittata Roháček &amp; Barber, 2013, paratype male, wing length 2.8 mm; 127 A. duplex Roháček &amp; Barber, 2013, paratype male, wing length 2.6 mm (both Canada: Ontario). Photo by K. N. Barber. Adapted from ROHÁĆEK &amp; BARBER (2013: Figs 167169)." figureDoi="http://doi.org/10.5281/zenodo.4272891" httpUri="https://zenodo.org/record/4272891/files/figure.png" pageId="36" pageNumber="33">Fig. 125</figureCitation>
); male cercus large and with thick dark setae (
<bibRefCitation id="EFC34B4EFFC3FFA1FE056831FDB1FDE0" author="ROHACEK J." box="[453,627,523,547]" pageId="36" pageNumber="33" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FDBE6831FD1EFDE0" box="[638,732,523,547]" captionStart="Figs 117124" captionStartId="85.[113,150,1456,1475]" captionTargetBox="[163,1039,223,1423]" captionTargetId="figure-14@85.[141,1058,195,1461]" captionTargetPageId="85" captionText="Figs 117124. Arganthomyza acuticuspis Roháček &amp; Barber, 2013, paratype female (USA: New Mexico). 117 spermatheca; 118 postabdomen, dorsally; 119 the same, ventrally; 120 spermatheca; 121 ventral receptacle, laterally; 122 internal sclerites, ventrally; 123 the same, laterally; 124 female genital chamber and apex of postabdomen, laterally. Scales = 0.2 mm (Figs 118, 119), 0.1 mm (Figs 121124) and 0.05 mm (others). Adapted from ROHÁĆEK &amp; BARBER (2013: Figs 7481)." figureDoi="http://doi.org/10.5281/zenodo.4272889" httpUri="https://zenodo.org/record/4272889/files/figure.png" pageId="36" pageNumber="33">Fig. 123</figureCitation>
). Aedeagal part of folding apparatus with dense dark-pigmented spines and striae (
<bibRefCitation id="EFC34B4EFFC3FFA1FD026811FCB2FD80" author="ROHACEK J." box="[706,880,555,579]" pageId="36" pageNumber="33" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FCBB6811FC1BFD80" box="[891,985,555,579]" captionStart="Figs 125127" captionStartId="87.[115,152,1484,1503]" captionTargetBox="[117,1086,348,1455]" captionTargetId="figure-52@87.[117,1086,347,1473]" captionTargetPageId="87" captionText="Figs 125127. Wings of the Nearctic species ofArganthomyza. 125 A. acuticuspis Roháček &amp; Barber, 2013, paratype male, wing length 2.5 mm (USA: New Mexico); 126 A. bivittata Roháček &amp; Barber, 2013, paratype male, wing length 2.8 mm; 127 A. duplex Roháček &amp; Barber, 2013, paratype male, wing length 2.6 mm (both Canada: Ontario). Photo by K. N. Barber. Adapted from ROHÁĆEK &amp; BARBER (2013: Figs 167169)." figureDoi="http://doi.org/10.5281/zenodo.4272891" httpUri="https://zenodo.org/record/4272891/files/figure.png" pageId="36" pageNumber="33">Fig. 127</figureCitation>
). Female unknown. .....................................................
<taxonomicName id="4C524D3CFFC3FFA1FD4E6871FC62FDA0" ID-CoL="8C7GZ" authority="Rohacek, 2009" authorityName="Rohacek" authorityYear="2009" box="[654,928,587,611]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="cercata">
<emphasis id="B926EAADFFC3FFA1FD4E6871FD38FDA1" bold="true" box="[654,762,587,610]" italics="true" pageId="36" pageNumber="33">F. cercata</emphasis>
Roháček, 2009
</taxonomicName>
(
<collectingCountry id="F345762FFFC3FFA1FC6E6871FC39FDA0" box="[942,1019,587,611]" name="Russia" pageId="36" pageNumber="33">Russia</collectingCountry>
: Far East)
</keyLead>
</paragraph>
<paragraph id="8BED36BFFFC3FFA1FF576851FBA4FCC0" blockId="36.[151,1127,459,1123]" pageId="36" pageNumber="33">
<keyLead id="30A3958AFFC3FFA1FF576851FBA4FCC0" pageId="36" pageNumber="33">
Fore femur and tibia yellow and dark brown variegated (
<bibRefCitation id="EFC34B4EFFC3FFA1FCE56851FC13FD40" author="ROHACEK J." box="[805,977,619,643]" pageId="36" pageNumber="33" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK 2006a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FC1A6851FBF6FD40" box="[986,1076,619,643]" captionStart="Figs 460466" captionStartId="293.[113,150,1514,1533]" captionTargetBox="[138,1058,225,1488]" captionTargetId="figure-14@293.[114,1089,195,1511]" captionTargetPageId="293" captionText="Figs 460466. Anthomyza shewelli sp. nov., paratype male, typical form (Canada: Ontario). 460 external genitalia, caudally; 461 apex of nlum, subventrally; 462 external genitalia, laterally; 463 hypandrial complex, laterally; 464 transandrium, caudally; 465 aedeagal complex, laterally; 466 gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 461, 466) and 0.1 mm (others)." figureDoi="http://doi.org/10.5281/zenodo.4273009" httpUri="https://zenodo.org/record/4273009/files/figure.png" pageId="36" pageNumber="33">Fig. 466</figureCitation>
; OHÁĆEK 2009a:
<figureCitation id="13692A3AFFC3FFA1FEAC68B1FE08FD60" box="[364,458,651,675]" captionStart="Figs 146149" captionStartId="94.[151,188,1489,1508]" captionTargetBox="[170,1106,214,1476]" captionTargetId="figure-11@94.[170,1108,214,1477]" captionTargetPageId="94" captionText="Figs 146149. Living Arganthomyza duplex Roháček &amp; Barber, 2013 and its habitat. 146 mating pair of A. duplex, laterally, body length ca. 2.32.9 mm; 147 male, laterally, body length 2.5 mm; 148 female, laterally, body length ca. 3 mm; 149 fern-dominated undergowth of mixed forest, habitat of A. duplex and A. carbo (all photos from Canada: Ontario: Dubreuilville). Photo by J. Roháček. Fig. 146 adapted from ROHÁĆEK &amp; BARBER (2013: Fig. 100)." figureDoi="http://doi.org/10.5281/zenodo.4272901" httpUri="https://zenodo.org/record/4272901/files/figure.png" pageId="36" pageNumber="33">Fig. 147</figureCitation>
). Gonostylus larger, externally at least partly micropubescent, and tapered apex with 2 denticles (
<figureCitation id="13692A3AFFC3FFA1FD966891FD6AFD00" box="[598,680,683,707]" captionStart="Figs 3538" captionStartId="39.[113,150,1225,1244]" captionTargetBox="[159,1053,233,1193]" captionTargetId="figure-14@39.[124,1078,198,1214]" captionTargetPageId="39" captionText="Figs 3538. Fungomyza buccata Roháček &amp; Barber, 2004, male. 35 head, laterally; 36 external genitalia, caudally; 37 gonostylus, posterolaterally (widest extension); 38 whole genitalia, laterally. Fig. 35 based on holotype, others on paratype. Scales = 0.3 mm (Fig. 35) and 0.1 mm (others). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 14). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272855" httpUri="https://zenodo.org/record/4272855/files/figure.png" pageId="36" pageNumber="33">Fig. 37</figureCitation>
;
<bibRefCitation id="EFC34B4EFFC3FFA1FD766891FCA6FD00" author="ROHACEK J." box="[694,868,683,707]" pageId="36" pageNumber="33" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK 2006a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FCB06891FC12FD00" box="[880,976,683,707]" captionStart="Figs 467473" captionStartId="295.[115,152,1496,1515]" captionTargetBox="[147,1067,217,1469]" captionTargetId="figure-14@295.[115,1089,195,1497]" captionTargetPageId="295" captionText="Figs 467473. Anthomyza shewelli sp. nov., paratype male, darkest form (USA: Idaho). 467 external genitalia, caudally; 468 apex of nlum, subventrally; 469 external genitalia, laterally; 470 hypandrial complex, laterally; 471 transandrium, caudally; 472 aedeagal complex, laterally; 473 gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 468, 473) and 0.1 mm (others)." figureDoi="http://doi.org/10.5281/zenodo.4273011" httpUri="https://zenodo.org/record/4273011/files/figure.png" pageId="36" pageNumber="33">Fig. 468</figureCitation>
); male cercus small, nnely and pale setose (
<figureCitation id="13692A3AFFC3FFA1FDC968F1FD9AFD20" box="[521,600,715,739]" captionStart="Figs 3538" captionStartId="39.[113,150,1225,1244]" captionTargetBox="[159,1053,233,1193]" captionTargetId="figure-14@39.[124,1078,198,1214]" captionTargetPageId="39" captionText="Figs 3538. Fungomyza buccata Roháček &amp; Barber, 2004, male. 35 head, laterally; 36 external genitalia, caudally; 37 gonostylus, posterolaterally (widest extension); 38 whole genitalia, laterally. Fig. 35 based on holotype, others on paratype. Scales = 0.3 mm (Fig. 35) and 0.1 mm (others). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 14). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272855" httpUri="https://zenodo.org/record/4272855/files/figure.png" pageId="36" pageNumber="33">Fig. 36</figureCitation>
;
<bibRefCitation id="EFC34B4EFFC3FFA1FDA268F1FCCFFD20" author="ROHACEK J." box="[610,781,715,739]" pageId="36" pageNumber="33" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK 2006a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FCD668F1FCB0FD20" box="[790,882,715,739]" captionStart="Figs 467473" captionStartId="295.[115,152,1496,1515]" captionTargetBox="[147,1067,217,1469]" captionTargetId="figure-14@295.[115,1089,195,1497]" captionTargetPageId="295" captionText="Figs 467473. Anthomyza shewelli sp. nov., paratype male, darkest form (USA: Idaho). 467 external genitalia, caudally; 468 apex of nlum, subventrally; 469 external genitalia, laterally; 470 hypandrial complex, laterally; 471 transandrium, caudally; 472 aedeagal complex, laterally; 473 gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 468, 473) and 0.1 mm (others)." figureDoi="http://doi.org/10.5281/zenodo.4273011" httpUri="https://zenodo.org/record/4273011/files/figure.png" pageId="36" pageNumber="33">Fig. 467</figureCitation>
). Armature of aedeagal part of folding apparatus pale-pigmented, inconspicuous. .......................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A62E1AFFC3FFA1FF576931FBA4FBA0" pageId="36" pageNumber="33">
<paragraph id="8BED36BFFFC3FFA1FF576931FBA4FC60" blockId="36.[151,1127,459,1123]" pageId="36" pageNumber="33">
<keyLead id="30A3958AFFC3FFA1FF576931FBA4FC60" pageId="36" pageNumber="33">
2(1) Anterior ors long and gena unusually high (
<figureCitation id="13692A3AFFC3FFA1FD6D6931FD3CFCE1" box="[685,766,778,803]" captionStart="Figs 3538" captionStartId="39.[113,150,1225,1244]" captionTargetBox="[159,1053,233,1193]" captionTargetId="figure-14@39.[124,1078,198,1214]" captionTargetPageId="39" captionText="Figs 3538. Fungomyza buccata Roháček &amp; Barber, 2004, male. 35 head, laterally; 36 external genitalia, caudally; 37 gonostylus, posterolaterally (widest extension); 38 whole genitalia, laterally. Fig. 35 based on holotype, others on paratype. Scales = 0.3 mm (Fig. 35) and 0.1 mm (others). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 14). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272855" httpUri="https://zenodo.org/record/4272855/files/figure.png" pageId="36" pageNumber="33">Fig. 35</figureCitation>
); frons, including most of frontal triangle, microtomentose and dull. Mid and hind femora darkened on distal half. Gonostylus with posteromedially projecting apex (
<figureCitation id="13692A3AFFC3FFA1FD146971FCE1FCA0" box="[724,803,843,867]" captionStart="Figs 3538" captionStartId="39.[113,150,1225,1244]" captionTargetBox="[159,1053,233,1193]" captionTargetId="figure-14@39.[124,1078,198,1214]" captionTargetPageId="39" captionText="Figs 3538. Fungomyza buccata Roháček &amp; Barber, 2004, male. 35 head, laterally; 36 external genitalia, caudally; 37 gonostylus, posterolaterally (widest extension); 38 whole genitalia, laterally. Fig. 35 based on holotype, others on paratype. Scales = 0.3 mm (Fig. 35) and 0.1 mm (others). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 14). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272855" httpUri="https://zenodo.org/record/4272855/files/figure.png" pageId="36" pageNumber="33">Fig. 37</figureCitation>
) and pregonite without posterior projection (
<figureCitation id="13692A3AFFC3FFA1FEBB6951FE08FC40" box="[379,458,875,899]" captionStart="Figs 3538" captionStartId="39.[113,150,1225,1244]" captionTargetBox="[159,1053,233,1193]" captionTargetId="figure-14@39.[124,1078,198,1214]" captionTargetPageId="39" captionText="Figs 3538. Fungomyza buccata Roháček &amp; Barber, 2004, male. 35 head, laterally; 36 external genitalia, caudally; 37 gonostylus, posterolaterally (widest extension); 38 whole genitalia, laterally. Fig. 35 based on holotype, others on paratype. Scales = 0.3 mm (Fig. 35) and 0.1 mm (others). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 14). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272855" httpUri="https://zenodo.org/record/4272855/files/figure.png" pageId="36" pageNumber="33">Fig. 38</figureCitation>
). Female T8 short and transverse (
<figureCitation id="13692A3AFFC3FFA1FCF66951FC44FC40" box="[822,902,875,899]" captionStart="Figs 4246" captionStartId="42.[150,187,1462,1481]" captionTargetBox="[160,1125,249,1430]" captionTargetId="figure-11@42.[148,1131,225,1484]" captionTargetPageId="42" captionText="Figs 4246. Fungomyza buccata Roháček &amp; Barber, 2004, paratype female. 42 postabdomen, dorsally; 43 the same, ventrally; 44 female genital chamber and 8th segment, laterally; 45 internal sclerites of female genital chamber, ventrally; 46 spermathecae. Scales = 0.2 mm (Figs 42, 43), 0.1 mm (Figs 44, 45) and 0.05 mm (Fig. 46). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 812). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272859" httpUri="https://zenodo.org/record/4272859/files/figure.png" pageId="36" pageNumber="33">Fig. 42</figureCitation>
); spermathecae more rounded (
<figureCitation id="13692A3AFFC3FFA1FEFA69B1FE48FC60" box="[314,394,907,931]" captionStart="Figs 4246" captionStartId="42.[150,187,1462,1481]" captionTargetBox="[160,1125,249,1430]" captionTargetId="figure-11@42.[148,1131,225,1484]" captionTargetPageId="42" captionText="Figs 4246. Fungomyza buccata Roháček &amp; Barber, 2004, paratype female. 42 postabdomen, dorsally; 43 the same, ventrally; 44 female genital chamber and 8th segment, laterally; 45 internal sclerites of female genital chamber, ventrally; 46 spermathecae. Scales = 0.2 mm (Figs 42, 43), 0.1 mm (Figs 44, 45) and 0.05 mm (Fig. 46). Adapted from ROHÁĆEK &amp; BARBER (2004: Figs 812). For abbreviations see p. 11." figureDoi="http://doi.org/10.5281/zenodo.4272859" httpUri="https://zenodo.org/record/4272859/files/figure.png" pageId="36" pageNumber="33">Fig. 46</figureCitation>
). ........................
<taxonomicName id="4C524D3CFFC3FFA1FD8469B1FC04FC60" ID-CoL="6JRB8" authority="Rohacek &amp; Barber, 2004" authorityName="Rohacek &amp; Barber" authorityYear="2004" box="[580,966,906,931]" class="Insecta" family="Anthomyzidae" genus="Fungomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="buccata">
<emphasis id="B926EAADFFC3FFA1FD8469B1FD74FC60" bold="true" box="[580,694,906,931]" italics="true" pageId="36" pageNumber="33">F. buccata</emphasis>
<bibRefCitation id="EFC34B4EFFC3FFA1FD7C69B1FC04FC60" author="ROHACEK J. &amp; BARBER K. N." box="[700,966,907,931]" pageId="36" pageNumber="33" pagination="131 - 141" refId="ref335772" refString="ROHACEK J. &amp; BARBER K. N. 2004: A new species of the genus Fungomyza from the Nearctic Region (Diptera: Anthomyzidae). Casopis Slezskeho Zemskeho Muzea, Opava (A) 53: 131 - 141." type="journal article" year="2004">Roháček &amp; Barber, 2004</bibRefCitation>
</taxonomicName>
(eastern
<collectingCountry id="F345762FFFC3FFA1FBE869B1FB9DFC60" box="[1064,1119,907,931]" name="United States of America" pageId="36" pageNumber="33">USA</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BED36BFFFC3FFA1FF576991FBA4FBA0" blockId="36.[151,1127,459,1123]" pageId="36" pageNumber="33">
<keyLead id="30A3958AFFC3FFA1FF576991FBA4FBA0" pageId="36" pageNumber="33">
Anterior ors short and gena very low (
<bibRefCitation id="EFC34B4EFFC3FFA1FDAF6991FCDFFC00" author="ROHACEK J." box="[623,797,939,963]" pageId="36" pageNumber="33" pagination="1 - 180" refId="ref334979" refString="ROHACEK J. 2009 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Casopis Slezskeho Zemskeho Muzea, Opava (A) 58 (supplement 1): 1 - 180." type="journal article" year="2009">ROHÁĆEK 2009a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FCE76991FC46FC00" box="[807,900,939,963]" captionStart="Figs 146149" captionStartId="94.[151,188,1489,1508]" captionTargetBox="[170,1106,214,1476]" captionTargetId="figure-11@94.[170,1108,214,1477]" captionTargetPageId="94" captionText="Figs 146149. Living Arganthomyza duplex Roháček &amp; Barber, 2013 and its habitat. 146 mating pair of A. duplex, laterally, body length ca. 2.32.9 mm; 147 male, laterally, body length 2.5 mm; 148 female, laterally, body length ca. 3 mm; 149 fern-dominated undergowth of mixed forest, habitat of A. duplex and A. carbo (all photos from Canada: Ontario: Dubreuilville). Photo by J. Roháček. Fig. 146 adapted from ROHÁĆEK &amp; BARBER (2013: Fig. 100)." figureDoi="http://doi.org/10.5281/zenodo.4272901" httpUri="https://zenodo.org/record/4272901/files/figure.png" pageId="36" pageNumber="33">Fig. 147</figureCitation>
); frontal triangle and posterior part of orbit bare and shining. Mid and hind femora uniformly yellow (OHÁĆEK 2009a:
<figureCitation id="13692A3AFFC3FFA1FE9169D1FE6DFBC0" box="[337,431,1003,1027]" captionStart="Figs 146149" captionStartId="94.[151,188,1489,1508]" captionTargetBox="[170,1106,214,1476]" captionTargetId="figure-11@94.[170,1108,214,1477]" captionTargetPageId="94" captionText="Figs 146149. Living Arganthomyza duplex Roháček &amp; Barber, 2013 and its habitat. 146 mating pair of A. duplex, laterally, body length ca. 2.32.9 mm; 147 male, laterally, body length 2.5 mm; 148 female, laterally, body length ca. 3 mm; 149 fern-dominated undergowth of mixed forest, habitat of A. duplex and A. carbo (all photos from Canada: Ontario: Dubreuilville). Photo by J. Roháček. Fig. 146 adapted from ROHÁĆEK &amp; BARBER (2013: Fig. 100)." figureDoi="http://doi.org/10.5281/zenodo.4272901" httpUri="https://zenodo.org/record/4272901/files/figure.png" pageId="36" pageNumber="33">Fig. 147</figureCitation>
). Gonostylus apically narrowed but not projecting and pregonite with distinct posterior projection (
<bibRefCitation id="EFC34B4EFFC3FFA1FD836E31FD33FBE0" author="ROHACEK J." box="[579,753,1035,1059]" pageId="36" pageNumber="33" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK 2006a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FD3C6E31FC56FBE0" box="[764,916,1035,1059]" captionStart="Figs 467473" captionStartId="295.[115,152,1496,1515]" captionTargetBox="[147,1067,217,1469]" captionTargetId="figure-14@295.[115,1089,195,1497]" captionTargetPageId="295" captionText="Figs 467473. Anthomyza shewelli sp. nov., paratype male, darkest form (USA: Idaho). 467 external genitalia, caudally; 468 apex of nlum, subventrally; 469 external genitalia, laterally; 470 hypandrial complex, laterally; 471 transandrium, caudally; 472 aedeagal complex, laterally; 473 gonostylus, ventrolaterocaudally (widest extension). Scales = 0.05 mm (Figs 468, 473) and 0.1 mm (others)." figureDoi="http://doi.org/10.5281/zenodo.4273011" httpUri="https://zenodo.org/record/4273011/files/figure.png" pageId="36" pageNumber="33">Figs 468, 471</figureCitation>
). Female T8 longer than broad; spermathecae roughly pyriform with narrowed basal part (
<bibRefCitation id="EFC34B4EFFC3FFA1FC786E11FBA1FB80" author="ROHACEK J." box="[952,1123,1067,1091]" pageId="36" pageNumber="33" pagination="1 - 328" refId="ref334646" refString="ROHACEK J. 2006 a: A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Casopis Slezskeho Zemskeho Muzea, Opava (A) 55 (supplement 1): 1 - 328." type="journal article" year="2006">ROHÁĆEK 2006a</bibRefCitation>
:
<figureCitation id="13692A3AFFC3FFA1FF146E71FEA8FBA0" box="[212,362,1099,1123]" captionStart="Figs 474480" captionStartId="296.[152,189,1484,1503]" captionTargetBox="[169,1104,226,1458]" captionTargetId="figure-11@296.[152,1125,204,1491]" captionTargetPageId="296" captionText="Figs 474480. Anthomyza shewelli sp. nov., paratype female, typical form (Canada: Ontario). 474 postabdomen, dorsally; 475 the same, ventrally; 476 distal part of female genital chamber, laterally; 477 end of postabdomen with internal sclerites, laterally; 478 internal sclerites and S8, ventrally (micropubescence omitted); 479 spermathecae; 480 spermatheca of another specimen. Scales = 0.2 mm (Figs 474, 475), 0.1 mm (Figs 476478) and 0.05 mm (Figs 479, 480)." figureDoi="http://doi.org/10.5281/zenodo.4273013" httpUri="https://zenodo.org/record/4273013/files/figure.png" pageId="36" pageNumber="33">Figs 474, 476</figureCitation>
). .....................................
<taxonomicName id="4C524D3CFFC3FFA1FDB86E71FC6AFBA0" authority="(Meigen, 1830)" authorityName="W. Palaearctic" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[632,936,1098,1123]" class="Insecta" family="Heleomyzidae" genus="Opomyza" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="33" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="B926EAADFFC3FFA1FDB86E71FD39FBA0" bold="true" box="[632,763,1098,1123]" italics="true" pageId="36" pageNumber="33">F. albimana</emphasis>
(
<bibRefCitation id="EFC34B4EFFC3FFA1FCC96E71FC62FBA0" author="MEIGEN J. W." box="[777,928,1099,1123]" pageId="36" pageNumber="33" refId="ref329671" refString="MEIGEN J. W. 1830: Systematische Beschreibung der bekannten europaischen zweissugeligen Insekten. Vol. 6. Schulzische Buchhandlung, Hamm, xi + 401 pp." type="book" year="1830">Meigen, 1830</bibRefCitation>
)
</taxonomicName>
(Europe,
<collectingCountry id="F345762FFFC3FFA1FBD16E71FB9DFBA0" box="[1041,1119,1099,1123]" name="Turkey" pageId="36" pageNumber="33">Turkey</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>