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<document id="4CBCFE2AD62200B539B49F36E4281580" ID-DOI="10.11646/zootaxa.3941.3.1" ID-GBIF-Dataset="67bc3c9d-ec99-4b47-81df-3fca7bd584ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="243860" ID-ZooBank="9907D081-A413-44F1-ABF6-76CC13DC8813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026017375" checkinUser="plazi" docAuthor="Wang, Cheng-Bin &amp; Zhou, Hong-Zhang" docDate="2015" docId="03EC87E6FF9A883B9DBF1AF3FE650CA3" docLanguage="en" docName="zt03941p338.pdf" docOrigin="Zootaxa 3941 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptomaphaginus wuzhishanicus Wang &amp; Zhou, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="334" masterDocId="FFD5FF9EFF85881A9D281927FFF50E4C" masterDocTitle="Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species" masterLastPageNumber="338" masterPageNumber="301" pageNumber="332" updateTime="1698572131055" updateUser="plazi">
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<mods:title id="11C59AB999730CA6157A02EE56CD27E2">Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species</mods:title>
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<mods:namePart id="FC8AE4041E02830B50B9F060C22F9A7F">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="03EC87E6FF9A883B9DBF1AF3FE650CA3" ID-DOI="http://doi.org/10.5281/zenodo.5691686" ID-GBIF-Taxon="119606327" ID-Zenodo-Dep="5691686" LSID="urn:lsid:plazi:treatment:03EC87E6FF9A883B9DBF1AF3FE650CA3" httpUri="http://treatment.plazi.org/id/03EC87E6FF9A883B9DBF1AF3FE650CA3" lastPageId="33" lastPageNumber="334" pageId="31" pageNumber="332">
<subSubSection id="C35F657BFF9A88059DBF1AF3FEC40A43" pageId="31" pageNumber="332" type="nomenclature">
<paragraph id="8BFA36F0FF9A88059DBF1AF3FD800DA2" blockId="31.[151,629,980,1039]" box="[151,629,980,1006]" pageId="31" pageNumber="332">
<heading id="D0B2819CFF9A88059DBF1AF3FD800DA2" bold="true" box="[151,629,980,1006]" fontSize="11" level="1" pageId="31" pageNumber="332" reason="1">
<emphasis id="B931EAE2FF9A88059DBF1AF3FD800DA2" bold="true" box="[151,629,980,1006]" pageId="31" pageNumber="332">
<taxonomicName id="4C454D73FF9A88059DBF1AF3FDE50DA2" ID-CoL="6WLQ5" box="[151,528,980,1006]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="wuzhishanicus" status="sp. nov.">
<emphasis id="B931EAE2FF9A88059DBF1AF3FDE50DA2" bold="true" box="[151,528,980,1006]" italics="true" pageId="31" pageNumber="332">Ptomaphaginus wuzhishanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FF9A88059F3F1AF3FD800DA2" box="[535,629,980,1006]" pageId="31" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA36F0FF9A88059DBF1AD1FEC40A43" blockId="31.[151,629,980,1039]" box="[151,305,1014,1039]" pageId="31" pageNumber="332">
(
<figureCitation id="137E2A75FF9A88059DB71AD1FF0D0A43" box="[159,248,1014,1039]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Figs. 18</figureCitation>
AJ)
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF9A88059DBF1D19FD0B0A37" pageId="31" pageNumber="332" type="materials_examined">
<paragraph id="8BFA36F0FF9A88059DBF1D19FD0B0A37" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
<emphasis id="B931EAE2FF9A88059DBF1D19FD560A1B" bold="true" box="[151,675,1086,1111]" pageId="31" pageNumber="332">
<typeStatus id="54FE8852FF9A88059DBF1D19FF270A1B" box="[151,210,1086,1111]" pageId="31" pageNumber="332">Type</typeStatus>
material.
<typeStatus id="54FE8852FF9A88059C7A1D19FE360A1B" box="[338,451,1086,1111]" pageId="31" pageNumber="332" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3527660FF9A88059CFB1D19FDC70A1B" box="[467,562,1086,1111]" name="China" pageId="31" pageNumber="332">CHINA</collectingCountry>
, Hainan:
</emphasis>
♂, Wuzhishan [=Mt. Wuzhi],
<quantity id="4CBD9B15FF9A880599221D18FBA30A1A" box="[1034,1110,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="31" pageNumber="332" unit="m" value="790.0">790 m</quantity>
, undergrowth, window trap,
<date id="FFFB1030FF9A88059DBF1D44FED70A37" box="[151,290,1123,1147]" pageId="31" pageNumber="332" value="2004-07-10">10.VII.2004</date>
, Jie Wu &amp; Yong-Jie Chen legg. (IZ-CAS).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF9A883B9DEF1DA1FE6B0EFC" lastPageId="33" lastPageNumber="334" pageId="31" pageNumber="332" type="description">
<paragraph id="8BFA36F0FF9A88059DEF1DA1FF010B47" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
<emphasis id="B931EAE2FF9A88059DEF1DA1FEAC0AD3" bold="true" box="[199,345,1158,1183]" pageId="31" pageNumber="332">Description.</emphasis>
<emphasis id="B931EAE2FF9A88059C481DA1FE6F0AD3" box="[352,410,1158,1183]" italics="true" pageId="31" pageNumber="332">Male</emphasis>
. EBL:
<quantity id="4CBD9B15FF9A88059CC51DA1FDA00AD2" box="[493,597,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="2.62" pageId="31" pageNumber="332" unit="mm" value="2.62">2.62 mm</quantity>
. Length of different body parts: HL: AL: PL: ELL = 0.43: 0.68: 0.65:
<quantity id="4CBD9B15FF9A88059DBF1D8CFF080A8E" box="[151,253,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="332" unit="mm" value="1.4">1.40 mm</quantity>
; width: HW: EW: PW: ELW = 0.70: 0.10: 1.03:
<quantity id="4CBD9B15FF9A88059E681D8CFC530A8E" box="[832,934,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="31" pageNumber="332" unit="mm" value="1.19">1.19 mm</quantity>
. Proportion of antennomeres from base to tip in µm (length × width): 124 × 35, 103 × 44, 63 × 42, 34 × 43, 31 × 49, 21 × 54, 39 × 67, 16 × 65, 51 × 79, 53 × 77, 99 × 80.
</paragraph>
<paragraph id="8BFA36F0FF9A88059DEF1C31FB520BD7" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">Habitus elongated oval, relatively convex and lustreless. Moderately pigmented: mostly chestnut brown; mouthparts, basal five antennomeres and apical half of ultimate antennomere, base of pronotum, and tarsi yellowish. Dorsum continually clothed with fine, recumbent and sallow pubescence. Insertions of pubescence on dorsal surfaces of pronotum, scutellum, elytra and femora align along transverse striolations.</paragraph>
<paragraph id="8BFA36F0FF9A88059DEF1C81FB850846" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
Head moderately short and gently convex, finely punctured, without visible transverse striolations, HW/HL = 1.65; anterior margin round. Compound eyes rather developed, EW/HW = 0.15. Antennae slender (
<figureCitation id="137E2A75FF9A880599DB1CEDFAB60BAF" box="[1267,1347,1482,1507]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
A), AL/ HW = 0.96; 5th shorter than 4th; 6th transverse; 9th and 10th wider than long; 11th pear-like.
</paragraph>
<paragraph id="8BFA36F0FF9A88059DEF1F31FA73083A" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
Pronotum not rather transverse and convex (
<figureCitation id="137E2A75FF9A88059FC31F32FCB60862" box="[747,835,1557,1582]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
D), widest at base, PW/PL = 1.57. Sides almost rectilinearly narrowing from posterior to anterior; hind corners drawn out and blunt. Posterior margin with distinct postero-lateral emargination. Surface transversely striolated, SP: 4143, interspaces narrower than that on elytra.
</paragraph>
<paragraph id="8BFA36F0FF9A88059DFC1FA6FEE7094A" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
Elytra relatively short and convex, widest near base, ELL/EW = 1.18. Sides feebly curved, gradually narrowing from base to apices; apices obliquely truncated and with a tuft of strong hairs (ca. six) different to pubescence background (
<figureCitation id="137E2A75FF9A88059CEF1FEEFDD608AE" box="[455,547,1737,1762]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
E). Surface transversely striolated, SE: 7172. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8BFA36F0FF9A88059DEF1E36FC0D093D" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
Prolegs robust, with basal three protarsomeres only slightly expanded (
<figureCitation id="137E2A75FF9A88059ED41E36FBA70966" box="[1020,1106,1809,1834]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
B): TW/BTW = 1.70. Spinal arrangement on ventral side of protibia as shown in
<figureCitation id="137E2A75FF9A88059FF71E12FCC50902" box="[735,816,1845,1870]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
C. Profemora wider than protibiae, and hairs on ventral side of profemora similar to that of protibiae. Mesotibiae moderately arcuate.
</paragraph>
<paragraph id="8BFA36F0FF9A88059DEF1E5AFADB09F6" blockId="31.[151,1437,1086,2014]" pageId="31" pageNumber="332">
Ventrite VII simple. Ventrite VIII also simple, gently emarginate posteriorly (
<figureCitation id="137E2A75FF9A880599071E5AFB7509DA" box="[1071,1152,1917,1942]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
F). Segment IX: uncinate tooth of genital plate much small and spiculum gastrale bottle-like, truncated at posterior end (
<figureCitation id="137E2A75FF9A880599901E86FAFF09F6" box="[1208,1290,1953,1978]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
G).
</paragraph>
<paragraph id="8BFA36F0FF9A883A9DEF1EE2FDBD0F51" blockId="31.[151,1437,1086,2014]" lastBlockId="32.[151,1437,151,285]" lastPageId="32" lastPageNumber="333" pageId="31" pageNumber="332">
Aedeagus piriform in dorsal view (
<figureCitation id="137E2A75FF9A88059F721EE2FD5B0992" box="[602,686,1989,2014]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="31" pageNumber="332">Fig. 18</figureCitation>
H): median lobe widest at about basal 1/3, sides of preapical part parallel; opening of genital orifice trended to cut right margin of median lobe; parameres relatively narrow, firmly attached to median lobe. Ventral operculum prominently and deeply bisected into long, narrow lobes, each rounded apically (
<figureCitation id="137E2A75FFA5883A9C2919F8FEA00EB4" box="[257,341,223,248]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="32" pageNumber="333">
Fig.
<date id="FFFB1030FFA5883A9C1F19C7FEA00EB4" box="[311,341,224,248]" pageId="32" pageNumber="333" value="1918-01">18</date>
</figureCitation>
I). Aedeagus like a tobacco pipe in lateral view, median lobe very arched, forming a right angle (
<figureCitation id="137E2A75FFA5883A9DB71823FF040F50" box="[159,241,260,285]" captionStart="FIGURE 18" captionStartId="32.[151,250,1882,1904]" captionTargetBox="[158,1426,361,1851]" captionTargetId="figure@32.[151,1436,337,1861]" captionTargetPageId="32" captionText="FIGURE 18. Ptomaphaginus wuzhishanicus sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VIII (ventral view); G, abdominal segment IX (ventral view); H, aedeagus (dorsal view); I, aedeagus (ventral view); J, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="32" pageNumber="333">Fig. 18</figureCitation>
J). Internal stylus very slender.
</paragraph>
<caption id="DF3A6678FFA5883A9DBF1E7DFA7B09E2" httpUri="https://zenodo.org/record/243878/files/figure.png" pageId="32" pageNumber="333" targetBox="[158,1426,361,1851]" targetPageId="32">
<paragraph id="8BFA36F0FFA5883A9DBF1E7DFA7B09E2" blockId="32.[151,1436,1882,1966]" pageId="32" pageNumber="333">
<emphasis id="B931EAE2FFA5883A9DBF1E7DFCEF093C" bold="true" box="[151,794,1882,1904]" pageId="32" pageNumber="333">
FIGURE 18.
<taxonomicName id="4C454D73FFA5883A9C061E7DFD8D093C" box="[302,632,1882,1904]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="wuzhishanicus" status="sp. nov.">
<emphasis id="B931EAE2FFA5883A9C061E7DFD8D093C" bold="true" box="[302,632,1882,1904]" italics="true" pageId="32" pageNumber="333">Ptomaphaginus wuzhishanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FFA5883A9FAB1E7DFD22093C" box="[643,727,1882,1904]" pageId="32" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
♂: A
</emphasis>
, antenna (dorsal view);
<emphasis id="B931EAE2FFA5883A99091E7DFBC7093C" bold="true" box="[1057,1074,1882,1904]" pageId="32" pageNumber="333">B</emphasis>
, proleg (dorsal view);
<emphasis id="B931EAE2FFA5883A98021E7DFAC9093C" bold="true" box="[1322,1340,1882,1904]" pageId="32" pageNumber="333">C</emphasis>
, protibia (ventral view);
<emphasis id="B931EAE2FFA5883A9C1F1E5EFEBC09C3" bold="true" box="[311,329,1913,1935]" pageId="32" pageNumber="333">D</emphasis>
, pronotum (dorsal view);
<emphasis id="B931EAE2FFA5883A9F731E5EFD9909C3" bold="true" box="[603,620,1913,1935]" pageId="32" pageNumber="333">E</emphasis>
, elytral apex (dorsoapical view);
<emphasis id="B931EAE2FFA5883A9EE31E5EFC2D09C3" bold="true" box="[971,984,1913,1935]" pageId="32" pageNumber="333">F</emphasis>
, ventrite VIII (ventral view);
<emphasis id="B931EAE2FFA5883A983A1E5EFAD609C3" bold="true" box="[1298,1315,1913,1935]" pageId="32" pageNumber="333">G</emphasis>
, abdominal segment IX (ventral view);
<emphasis id="B931EAE2FFA5883A9C861EBFFE3709E2" bold="true" box="[430,450,1944,1966]" pageId="32" pageNumber="333">H</emphasis>
, aedeagus (dorsal view);
<emphasis id="B931EAE2FFA5883A9FED1EBFFD3A09E2" bold="true" box="[709,719,1944,1966]" pageId="32" pageNumber="333">I</emphasis>
, aedeagus (ventral view);
<emphasis id="B931EAE2FFA5883A9EF31EBFFC1209E2" bold="true" box="[987,999,1944,1966]" pageId="32" pageNumber="333">J</emphasis>
, aedeagus (lateral view). Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BFA36F0FFA4883B9DEF19B0FE6B0EFC" blockId="33.[151,1437,151,752]" box="[199,414,151,176]" pageId="33" pageNumber="334">
<emphasis id="B931EAE2FFA4883B9DEF19B0FEEE0EFC" box="[199,283,151,176]" italics="true" pageId="33" pageNumber="334">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFFA4883B9DEF199BFDE90E99" box="[199,540,188,213]" pageId="33" pageNumber="334" type="distribution">
<paragraph id="8BFA36F0FFA4883B9DEF199BFDE90E99" blockId="33.[151,1437,151,752]" box="[199,540,188,213]" pageId="33" pageNumber="334">
<emphasis id="B931EAE2FFA4883B9DEF199BFE950E99" bold="true" box="[199,352,188,213]" pageId="33" pageNumber="334">Distribution.</emphasis>
<collectingCountry id="F3527660FFA4883B9C4F199BFE5F0E99" box="[359,426,188,213]" name="China" pageId="33" pageNumber="334">China</collectingCountry>
(Hainan).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFFA4883B9DEF19F8FEA40F51" pageId="33" pageNumber="334" type="etymology">
<paragraph id="8BFA36F0FFA4883B9DEF19F8FEA40F51" blockId="33.[151,1437,151,752]" pageId="33" pageNumber="334">
<emphasis id="B931EAE2FFA4883B9DEF19F8FEBB0EB4" bold="true" box="[199,334,223,248]" pageId="33" pageNumber="334">Etymology.</emphasis>
The specific epithet is from the Chinese name (Pinyin) of the
<typeStatus id="54FE8852FFA4883B992919C7FBC40EB4" box="[1025,1073,224,248]" pageId="33" pageNumber="334">type</typeStatus>
locality “Wuzhishan” in Hainan province,
<collectingCountry id="F3527660FFA4883B9C2F1823FEBB0F51" box="[263,334,260,285]" name="China" pageId="33" pageNumber="334">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFFA4883B9DEF1800FE650CA3" pageId="33" pageNumber="334" type="discussion">
<paragraph id="8BFA36F0FFA4883B9DEF1800FE650CA3" blockId="33.[151,1437,151,752]" pageId="33" pageNumber="334">
<emphasis id="B931EAE2FFA4883B9DEF1800FECE0F0C" bold="true" box="[199,315,295,320]" pageId="33" pageNumber="334">Remarks.</emphasis>
This new species is sympatric with
<taxonomicName id="4C454D73FFA4883B9FFE180EFCB90F0C" box="[726,844,296,320]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="newtoni" status="sp. nov.">
<emphasis id="B931EAE2FFA4883B9FFE180EFD110F0C" box="[726,740,297,320]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9FDB180EFCB90F0C" box="[755,844,297,320]" italics="true" pageId="33" pageNumber="334">newtoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FFA4883B9E7C1800FC5B0F0C" box="[852,942,295,320]" pageId="33" pageNumber="334" rank="species">
<emphasis id="B931EAE2FFA4883B9E7C1800FC5B0F0C" bold="true" box="[852,942,295,320]" pageId="33" pageNumber="334">sp. nov.</emphasis>
</taxonomicNameLabel>
, but it is easily to be distinguished from all other congener species from
<collectingCountry id="F3527660FFA4883B9CC3186BFDDB0F29" box="[491,558,332,357]" name="China" pageId="33" pageNumber="334">China</collectingCountry>
by elytral apices with a tuft of strong hairs different from the pubescence background and its piriform aedeagus. Interestingly, this new species should belong to a group from Indian fauna with common character of piriform aedeagus, incluing
<taxonomicName id="4C454D73FFA4883B9E2F18B2FB550FE1" authority="Szymczakowski, 1975" authorityName="Szymczakowski" authorityYear="1975" box="[775,1184,404,429]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="apiculatus">
<emphasis id="B931EAE2FFA4883B9E2F18B2FCE00FE0" box="[775,789,405,428]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9E0C18B3FC6C0FE1" box="[804,921,404,429]" italics="true" pageId="33" pageNumber="334">apiculatus</emphasis>
<bibRefCitation id="EFD44B01FFA4883B9E8818B3FB550FE1" author="Szymczakowski" box="[928,1184,404,429]" pageId="33" pageNumber="334" refString="Szymczakowski, W. (1975) Catopidae (Coleoptera) recoltes dans le sud de l'lnde par l'expedition du Muselum d'Histoire Naturelle de Geneve en 1972. Acta Zoologica Cracoviensia, 20 (4), 121 - 149." type="journal article" year="1975">Szymczakowski, 1975</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C454D73FFA4883B998418B2FF240F9C" authority="Perreau, 1991" authorityName="Perreau" authorityYear="1991" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="pilipennis">
<emphasis id="B931EAE2FFA4883B998418B2FB4F0FE0" box="[1196,1210,405,428]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B99E118B3FAC20FE1" box="[1225,1335,404,429]" italics="true" pageId="33" pageNumber="334">pilipennis</emphasis>
<bibRefCitation id="EFD44B01FFA4883B981718B2FF240F9C" author="Perreau" pageId="33" pageNumber="334" refString="Perreau, M. (1991) Contribution a la connaissance des Cholevidae de l'Inde du Nord et du Pakistan (Coleoptera). Revue Suisse de Zoologie, 98 (3), 555 - 566." type="journal article" year="1991">Perreau, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C454D73FFA4883B9DF6189EFDC20F9C" authority="Perreau, 1991" authorityName="Perreau" authorityYear="1991" box="[222,567,439,464]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="pilipennoides">
<emphasis id="B931EAE2FFA4883B9DF6189EFF190F9C" box="[222,236,441,464]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9DD31890FE670F9C" box="[251,402,439,464]" italics="true" pageId="33" pageNumber="334">pilipennoides</emphasis>
<bibRefCitation id="EFD44B01FFA4883B9CB1189FFDC20F9C" author="Perreau" box="[409,567,440,464]" pageId="33" pageNumber="334" refString="Perreau, M. (1991) Contribution a la connaissance des Cholevidae de l'Inde du Nord et du Pakistan (Coleoptera). Revue Suisse de Zoologie, 98 (3), 555 - 566." type="journal article" year="1991">Perreau, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C454D73FFA4883B9F6D189EFC370F9C" authority="Szymczakowski, 1975" authorityName="Szymczakowski" authorityYear="1975" box="[581,962,439,464]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="piraster">
<emphasis id="B931EAE2FFA4883B9F6D189EFDA60F9C" box="[581,595,441,464]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9F4A189EFD4E0F9C" box="[610,699,441,464]" italics="true" pageId="33" pageNumber="334">piraster</emphasis>
<bibRefCitation id="EFD44B01FFA4883B9FEA1890FC370F9C" author="Szymczakowski" box="[706,962,439,464]" pageId="33" pageNumber="334" refString="Szymczakowski, W. (1975) Catopidae (Coleoptera) recoltes dans le sud de l'lnde par l'expedition du Muselum d'Histoire Naturelle de Geneve en 1972. Acta Zoologica Cracoviensia, 20 (4), 121 - 149." type="journal article" year="1975">Szymczakowski, 1975</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C454D73FFA4883B9ED6189EFAF90F9C" authority="Perreau, 1995" authorityName="Perreau" authorityYear="1995" box="[1022,1292,439,464]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="B931EAE2FFA4883B9ED6189EFBF90F9C" box="[1022,1036,441,464]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B99341890FB900F9C" box="[1052,1125,439,464]" italics="true" pageId="33" pageNumber="334">riedeli</emphasis>
<bibRefCitation id="EFD44B01FFA4883B9944189FFAF90F9C" author="Perreau" box="[1132,1292,439,464]" pageId="33" pageNumber="334" refString="Perreau, M. (1995) Especes nouvelles ou mal connues de Cholevidae (Coleoptera). Revue Suisse de Zoologie, 102 (4), 969 - 978." type="journal article" year="1995">Perreau, 1995</bibRefCitation>
</taxonomicName>
. However, it differs from
<taxonomicName id="4C454D73FFA4883B9C0418FAFD710FB9" authority="Szymczakowski" authorityName="Szymczakowski" box="[300,644,476,501]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="apiculatus">
<emphasis id="B931EAE2FFA4883B9C0418FAFECF0FB8" box="[300,314,477,500]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9C6318FBFE350FB9" box="[331,448,476,501]" italics="true" pageId="33" pageNumber="334">apiculatus</emphasis>
Szymczakowski
</taxonomicName>
,
<taxonomicName id="4C454D73FFA4883B9FBB18FAFC590FB8" authority="Perreau" authorityName="Perreau" box="[659,940,476,501]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="pilipennoides">
<emphasis id="B931EAE2FFA4883B9FBB18FAFD540FB8" box="[659,673,477,500]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9F9B18FBFCBF0FB9" box="[691,842,476,501]" italics="true" pageId="33" pageNumber="334">pilipennoides</emphasis>
Perreau
</taxonomicName>
,
<taxonomicName id="4C454D73FFA4883B9E9418FAFB000FB9" authority="Szymczakowski" authorityName="Szymczakowski" box="[956,1269,476,501]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="piraster">
<emphasis id="B931EAE2FFA4883B9E9418FAFC3F0FB8" box="[956,970,477,500]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9EF318FAFBC10FB8" box="[987,1076,477,500]" italics="true" pageId="33" pageNumber="334">piraster</emphasis>
Szymczakowski
</taxonomicName>
and
<taxonomicName id="4C454D73FFA4883B981B18FAFF190C5B" authority="Perreau" authorityName="Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="B931EAE2FFA4883B981B18FAFAB40FB8" box="[1331,1345,477,500]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B987B18FBFA690FB9" box="[1363,1436,476,501]" italics="true" pageId="33" pageNumber="334">riedeli</emphasis>
Perreau
</taxonomicName>
by its very arched median lobe, forming a right angle when viewed laterally; and in the new species, the aedeagus is relatively wider at basal half, and basal protarsomere obviously narrower than protibial apex, ELL/EW = 1.18, while in
<taxonomicName id="4C454D73FFA4883B9C7C1B6EFDCB0C13" authority="Perreau" authorityName="Perreau" box="[340,574,583,608]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="pilipennis">
<emphasis id="B931EAE2FFA4883B9C7C1B6EFE970C2C" box="[340,354,585,608]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9C5A1B60FE150C2C" box="[370,480,583,608]" italics="true" pageId="33" pageNumber="334">pilipennis</emphasis>
Perreau
</taxonomicName>
the aedeagus is relatively narrower at basal half, and basal protersomere as wide as protibial apex, ELL/EW = 1.3. This new species is also similar to
<taxonomicName id="4C454D73FFA4883B9EE51B4AFAAA0CC8" authority="Szymczakowski, 1972" authorityName="Szymczakowski" authorityYear="1972" box="[973,1375,620,645]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="scaphaner">
<emphasis id="B931EAE2FFA4883B9EE51B4AFC2E0CC8" box="[973,987,621,644]" italics="true" pageId="33" pageNumber="334">P</emphasis>
.
<emphasis id="B931EAE2FFA4883B9EC11B4BFBAB0CC9" box="[1001,1118,620,645]" italics="true" pageId="33" pageNumber="334">scaphaner</emphasis>
<bibRefCitation id="EFD44B01FFA4883B994C1B4BFAAA0CC8" author="Szymczakowski" box="[1124,1375,620,645]" pageId="33" pageNumber="334" refString="Szymczakowski, W. (1972) Catopidae (Coleoptera) recoltes au Viet-nam par Gy. Topal. Acta Zoologica Cracoviensia, 17 (12), 289 - 304." type="journal article" year="1972">Szymczakowski, 1972</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3527660FFA4883B9DBF1BA8FF0C0CE4" box="[151,249,655,680]" name="Vietnam" pageId="33" pageNumber="334">Vietnam</collectingCountry>
,
<collectingCountry id="F3527660FFA4883B9C2E1BA8FE980CE4" box="[262,365,655,680]" name="Malaysia" pageId="33" pageNumber="334">Malaysia</collectingCountry>
and
<collectingCountry id="F3527660FFA4883B9C811BA8FDEC0CE4" box="[425,537,655,680]" name="Indonesia" pageId="33" pageNumber="334">Indonesia</collectingCountry>
, but they have differences in aedeagal apex shape: the latter with right apical expansion of aedeagus triangular and somewhat dorsally upturned; while the new species without apical expansion of aedeagus stuck out.
</paragraph>
</subSubSection>
</treatment>
</document>