324 lines
41 KiB
XML
324 lines
41 KiB
XML
<document id="D6185B0AB3A4DAAB14B5DAE55244797F" ID-CLB-Dataset="29868" ID-DOI="10.11646/zootaxa.4434.3.10" ID-GBIF-Dataset="d7dee720-286b-47c2-9016-b4bd868d2654" ID-ISSN="1175-5326" ID-Zenodo-Dep="1292243" ID-ZooBank="E4DBB0C4-2C3F-4F23-B41A-4C40FEB448B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529351258412" checkinUser="plazi" docAuthor="Kodada, Jan, Hendrich, Lars & Balke, Michael" docDate="2018" docId="03CBED67FF92897AF9C511FC3CE546D0" docLanguage="en" docName="zootaxa.4434.3.10.pdf.imf" docOrigin="Zootaxa 4434 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tepuidessus grulai Kodada & Hendrich & Balke 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="566" masterDocId="FFF2951FFF90897FF95217673F114758" masterDocTitle="Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)" masterLastPageNumber="572" masterPageNumber="561" pageNumber="563" updateTime="1698549797483" updateUser="ExternalLinkService">
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<mods:mods id="224E09EC3B0BBEF4B2E1CB27620F0715" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5588F7AB5E92B9F0E8129F876A71F161">
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<mods:title id="089EF8631D46987F62CAA1C9416C8167">Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)</mods:title>
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<mods:name id="28F031B41490878AD32B178A01D703F2" type="personal">
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<mods:role id="47D97A7F6689F1B33C3CD85ACC777BB8">
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<mods:roleTerm id="FA0F85799CA4E643D05F5C7619F0FE26">Author</mods:roleTerm>
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<mods:namePart id="8561A5B944195239D421A35B78BC60EC">Kodada, Jan</mods:namePart>
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<mods:namePart id="A1FB2CE1C4EE6177020E5B81BDBCC6B1">Hendrich, Lars</mods:namePart>
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<mods:name id="36FB00EF3E932B25FC82CF2F0F995DC8" type="personal">
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<mods:roleTerm id="D0927CD554EF673B432B05532E777E63">Author</mods:roleTerm>
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<mods:namePart id="84F5459F8092601CD93C7D1F7509A5A7">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="2DAC9C02016213EAD50F99F6A4025181">text</mods:typeOfResource>
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<mods:title id="914B08A30E4657E53C279D117A47DDEE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DBAEF0BF7C43C810E826CE7C0F5E817F">
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<mods:date id="FF3C2A85593D250AF0524C357A213E0B">2018</mods:date>
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<mods:number id="9B1417EDF97B269998663EAF3F7C722A">2018-06-18</mods:number>
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<mods:number id="9588326A7C6A76A74C514D14B2101A2D">4434</mods:number>
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<subSubSection id="C3780FFAFF92897DF9C511FC3ECF41ED" box="[151,478,1691,1717]" pageId="2" pageNumber="563" type="nomenclature">
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<paragraph id="8BDD5C71FF92897DF9C511FC3ECF41ED" blockId="2.[151,478,1691,1717]" box="[151,478,1691,1717]" pageId="2" pageNumber="563">
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<heading id="D095EB1DFF92897DF9C511FC3ECF41ED" bold="true" box="[151,478,1691,1717]" fontSize="11" level="1" pageId="2" pageNumber="563" reason="1">
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<emphasis id="B9168063FF92897DF9C511FC3ECF41ED" bold="true" box="[151,478,1691,1717]" pageId="2" pageNumber="563">
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<taxonomicName id="4C6227F2FF92897DF9C511FC3E6841ED" ID-CoL="8LZQZ" authority="Kodada & Hendrich & Balke, 2018" authorityName="Kodada & Hendrich & Balke" authorityYear="2018" box="[151,377,1691,1717]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="grulai" status="sp. nov.">
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<emphasis id="B9168063FF92897DF9C511FC3E6841ED" bold="true" box="[151,377,1691,1717]" italics="true" pageId="2" pageNumber="563">Tepuidessus grulai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2253D18FF92897DF8D211FC3ECF41ED" box="[384,478,1691,1717]" pageId="2" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3780FFAFF92897DF9C511863B6E40D2" pageId="2" pageNumber="563" type="materials_examined">
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<paragraph id="8BDD5C71FF92897DF9C511863CD141A2" blockId="2.[151,1436,1761,2038]" box="[151,960,1761,1786]" pageId="2" pageNumber="563">
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<materialsCitation id="3B0A562CFF92897DF9C511863CD141A2" ID-GBIF-Occurrence="1849338247" box="[151,960,1761,1786]" country="Venezuela" elevation="2000" latitude="5.194" location="Venezuela" longLatPrecision="78" longitude="-62.045" pageId="2" pageNumber="563" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9168063FF92897DF9C511863E3C41A2" bold="true" box="[151,301,1761,1786]" pageId="2" pageNumber="563">
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<typeStatus id="54D9E2D3FF92897DF9C511863FC341A2" box="[151,210,1761,1786]" pageId="2" pageNumber="563">Type</typeStatus>
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locality
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</emphasis>
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. Acopán Tepui, ca.
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<quantity id="4C9AF194FF92897DFB5C11853D7D41A1" box="[526,620,1762,1786]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="563" unit="m" value="2000.0">
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<elevation id="004FBB42FF92897DFB5C11853D7D41A1" box="[526,620,1762,1786]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="563" unit="m" value="2000.0">2,000 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE563AB6FF92897DFB2A11863DDA41A2" box="[632,715,1761,1786]" direction="north" orientation="latitude" pageId="2" pageNumber="563" precision="55" value="5.194">5.194N</geoCoordinate>
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<geoCoordinate id="EE563AB6FF92897DFB8011863C2B41A2" box="[722,826,1761,1786]" direction="west" orientation="longitude" pageId="2" pageNumber="563" precision="55" value="-62.045">62.045W</geoCoordinate>
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,
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<collectingCountry id="F3751CE1FF92897DFA1711863CAD41A2" box="[837,956,1761,1786]" name="Venezuela" pageId="2" pageNumber="563">Venezuela</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BDD5C71FF92897DF99510623B6E40D2" blockId="2.[151,1436,1761,2038]" pageId="2" pageNumber="563">
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<materialsCitation id="3B0A562CFF92897DF99510623DD5403E" ID-GBIF-Occurrence="1849338082" collectingDate="2015-11-14" collectionCode="ZSM" country="Venezuela" elevation="2022" location="Acopantepui" municipality="Estado Boliver" pageId="2" pageNumber="563" specimenCode="VEN6" specimenCount="2" typeStatus="holotype">
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<emphasis id="B9168063FF92897DF99510623D7B4046" bold="true" box="[199,618,1797,1822]" pageId="2" pageNumber="563">
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<typeStatus id="54D9E2D3FF92897DF99510623E244046" box="[199,309,1797,1822]" pageId="2" pageNumber="563" type="holotype">Holotype</typeStatus>
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, male (
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<collectionCode id="ED73C4B4FF92897DF8D810623EDC4046" box="[394,461,1797,1822]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="563">ZSM</collectionCode>
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): "
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<collectingCountry id="F3751CE1FF92897DF8BF10623D7B4046" box="[493,618,1797,1822]" name="Venezuela" pageId="2" pageNumber="563">Venezuela</collectingCountry>
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</emphasis>
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:
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<collectingMunicipality id="6BB9C60BFF92897DFB2B10623C334046" box="[633,802,1797,1822]" pageId="2" pageNumber="563">Estado Boliver</collectingMunicipality>
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, puddles on rocks in
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<location id="8EBD0AAAFF92897DFD5D10623B8F4046" LSID="urn:lsid:plazi:treatment:03CBED67FF92897AF9C511FC3CE546D0:8EBD0AAAFF92897DFD5D10623B8F4046" box="[1039,1182,1797,1822]" country="Venezuela" municipality="Estado Boliver" name="Acopantepui" pageId="2" pageNumber="563">Acopantepui</location>
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base camp" / "
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<quantity id="4C9AF194FF92897DFC1710613A894046" box="[1349,1432,1798,1822]" metricMagnitude="3" metricUnit="m" metricValue="2.022" pageId="2" pageNumber="563" unit="m" value="2022.0">
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<elevation id="004FBB42FF92897DFC1710613A894046" box="[1349,1432,1798,1822]" metricMagnitude="3" metricUnit="m" metricValue="2.022" pageId="2" pageNumber="563" unit="m" value="2022.0">2022m</elevation>
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</quantity>
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,
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<date id="FFDC7AB1FF92897DF9C5104E3E00401A" box="[151,273,1833,1858]" pageId="2" pageNumber="563" value="2015-11-14">
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||
<collectingDate id="EF988359FF92897DF9C5104E3E00401A" box="[151,273,1833,1858]" pageId="2" pageNumber="563" value="2015-11-14">14.xi.2015</collectingDate>
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</date>
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||
, 5.19413°[N] -62.04478°[W], (
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||
<specimenCode id="DBC4F40AFF92897DFBD2104E3DD8401A" box="[640,713,1833,1858]" collectionCode="VEN" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15643" name="Fundaci<63>n Instituto Bot<6F>nico de Venezuela" pageId="2" pageNumber="563">
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||
VEN
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||
<date id="FFDC7AB1FF92897DFBEE104E3DD8401A" box="[700,713,1833,1858]" pageId="2" pageNumber="563" value="2015-11-06">6</date>
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||
</specimenCode>
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||
/Nov2015)" / "
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||
<typeStatus id="54D9E2D3FF92897DFA24104E3CF0401A" box="[886,993,1833,1858]" pageId="2" pageNumber="563" type="holotype">Holotype</typeStatus>
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<taxonomicName id="4C6227F2FF92897DFABF104E3BD2401A" authority="Kodada & Hendrich & Balke, 2018" authorityName="Kodada & Hendrich & Balke" authorityYear="2018" box="[1005,1219,1833,1858]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="grulai" status="sp. nov.">
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||
<emphasis id="B9168063FF92897DFABF104E3BD2401A" box="[1005,1219,1833,1858]" italics="true" pageId="2" pageNumber="563">Tepuidessus grulai</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A2253D18FF92897DFD82104D3A3F4019" box="[1232,1326,1834,1857]" pageId="2" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
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, Kodada, Hendrich & Balke des. 2018" [red printed label]
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</materialsCitation>
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||
.
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<emphasis id="B9168063FF92897DFB82102A3C40403E" bold="true" box="[720,849,1869,1894]" pageId="2" pageNumber="563">Paratypes:</emphasis>
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43 exs with the same label data as holotype (CHF, CKB, MIZA, NMPC, ZSM). Each paratype is provided with a red printed paratype label.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3780FFAFF92897BF99510F23CE84189" lastPageId="4" lastPageNumber="565" pageId="2" pageNumber="563" type="description">
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<paragraph id="8BDD5C71FF92897DF99510F23D6540AD" blockId="2.[151,1436,1761,2038]" pageId="2" pageNumber="563">
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<emphasis id="B9168063FF92897DF99510F23EF140F6" bold="true" box="[199,480,1941,1966]" pageId="2" pageNumber="563">
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||
Description of
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||
<typeStatus id="54D9E2D3FF92897DF82B10F23EF140F6" box="[377,480,1941,1966]" pageId="2" pageNumber="563" type="holotype">holotype</typeStatus>
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</emphasis>
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||
. Habitus slightly broad oval, outline with particularly distinct discontinuity between pronotum and elytra (
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<figureCitation id="135940F4FF92897DF8DC10DE3EE6408A" box="[398,503,1977,2002]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1228,1250]" captionStartId-1="5.[151,250,1981,2003]" captionTargetBox-0="[160,1427,194,1206]" captionTargetBox-1="[331,1256,447,1959]" captionTargetId-0="figure@4.[159,1427,193,1206]" captionTargetId-1="figure@5.[331,1256,447,1959]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Tepuidessus grulai sp. nov., habitus of male (A) and female (B). Scale bar = 1 mm." captionText-1="FIGURE 4. Tepuidessus grulai sp. nov., head and pronotum of male (A) and female (B)." httpUri-0="https://zenodo.org/record/1292247/files/figure.png" httpUri-1="https://zenodo.org/record/1292249/files/figure.png" pageId="2" pageNumber="563">Figs 3–4</figureCitation>
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). Pronotum broadest before its midlength. Elytra widest at about midlength. Total length:
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<quantity id="4C9AF194FF92897DF9BE10B93E5540AD" box="[236,324,2014,2038]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="563" unit="mm" value="1.8">1.8 mm</quantity>
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||
; maximum width:
|
||
<quantity id="4C9AF194FF92897DFB4510B93D6140AD" box="[535,624,2014,2038]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="563" unit="mm" value="0.9">0.9 mm</quantity>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF93897CF99517F03CEC47E8" blockId="3.[151,1437,151,392]" box="[199,1021,151,176]" pageId="3" pageNumber="564">
|
||
<emphasis id="B9168063FF93897CF99517F03E4947E8" bold="true" box="[199,344,151,176]" pageId="3" pageNumber="564">Colouration</emphasis>
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||
. Black dorsally, dark brown to black ventrally (
|
||
<figureCitation id="135940F4FF93897CFA3C17F03CA747E8" box="[878,950,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1228,1250]" captionTargetBox="[160,1427,194,1206]" captionTargetId="figure@4.[159,1427,193,1206]" captionTargetPageId="4" captionText="FIGURE 3. Tepuidessus grulai sp. nov., habitus of male (A) and female (B). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1292247/files/figure.png" pageId="3" pageNumber="564">Figs 3</figureCitation>
|
||
,
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||
<figureCitation id="135940F4FF93897CFA9117FF3CC247E8" box="[963,979,152,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1981,2003]" captionTargetBox="[331,1256,447,1959]" captionTargetId="figure@5.[331,1256,447,1959]" captionTargetPageId="5" captionText="FIGURE 4. Tepuidessus grulai sp. nov., head and pronotum of male (A) and female (B)." httpUri="https://zenodo.org/record/1292249/files/figure.png" pageId="3" pageNumber="564">4</figureCitation>
|
||
,
|
||
<figureCitation id="135940F4FF93897CFA8D17F03CFE47E8" box="[991,1007,151,176]" captionStart="FIGURE 5" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[180,1421,195,1939]" captionTargetId="figure@6.[180,1421,195,1940]" captionTargetPageId="6" captionText="FIGURE 5. Tepuidessus grulai sp. nov., ventral surface." httpUri="https://zenodo.org/record/1292251/files/figure.png" pageId="3" pageNumber="564">5</figureCitation>
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).
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF93897CF99517DB3E5B46D0" blockId="3.[151,1437,151,392]" pageId="3" pageNumber="564">
|
||
<emphasis id="B9168063FF93897CF99517DB3E89478D" bold="true" box="[199,408,188,213]" pageId="3" pageNumber="564">Surface sculpture</emphasis>
|
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. Head with distinct microreticulation and few setiferous punctures (
|
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<figureCitation id="135940F4FF93897CFDC317DB3BFD478C" box="[1169,1260,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1981,2003]" captionTargetBox="[331,1256,447,1959]" captionTargetId="figure@5.[331,1256,447,1959]" captionTargetPageId="5" captionText="FIGURE 4. Tepuidessus grulai sp. nov., head and pronotum of male (A) and female (B)." httpUri="https://zenodo.org/record/1292249/files/figure.png" pageId="3" pageNumber="564">Fig. 4A</figureCitation>
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). Pronotum and elytron shiny (
|
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<figureCitation id="135940F4FF93897CF86917B83E8447A0" box="[315,405,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1228,1250]" captionTargetBox="[160,1427,194,1206]" captionTargetId="figure@4.[159,1427,193,1206]" captionTargetPageId="4" captionText="FIGURE 3. Tepuidessus grulai sp. nov., habitus of male (A) and female (B). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1292247/files/figure.png" pageId="3" pageNumber="564">Fig. 3A</figureCitation>
|
||
), mostly polished and distinct microreticulation only visible along pronotal margins; whole surface with dense and coarse setiferous punctation, exceptionally coarse and deep on disc (
|
||
<figureCitation id="135940F4FF93897CFD8416633A224645" box="[1238,1331,260,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1181,1203]" captionTargetBox="[151,1435,196,1158]" captionTargetId="figure@7.[151,1435,193,1160]" captionTargetPageId="7" captionText="FIGURE 6. Tepuidessus grulai sp. nov., reduced metathoracic wing (A), lateral view of last ventrite in female (B) and pronotum, punctation and basal stria as well as basal stria on elytron (C)." httpUri="https://zenodo.org/record/1292253/files/figure.png" pageId="3" pageNumber="564">Fig. 6C</figureCitation>
|
||
). Ventral surface microreticulate, abdominal ventrites 3–6 shinier; venter with distinct setiferous punctation; posterior margins of ventrites 3–5 without denticles (present in Neotropical genus
|
||
<taxonomicName id="4C6227F2FF93897CFA90162B3A15463D" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" box="[962,1284,332,357]" class="Insecta" family="Dytiscidae" genus="Bidessodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9168063FF93897CFA90162B3B2E463D" box="[962,1087,332,357]" italics="true" pageId="3" pageNumber="564">Bidessodes</emphasis>
|
||
Régimbart, 1895
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="EFF32180FF93897CFC68162B3E2F46D0" pageId="3" pageNumber="564" refId="ref2940" refString="Miller, K. B. & Bergsten, J. (2016) Diving Beetles of the World. Systematics and Biology of the Dytiscidae. Johns Hopkins University Press, Baltimore, 320 pp." type="book">Miller & Bergsten 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1D0CF9FF93897CF9C510CB3CC6409A" box="[151,983,1964,1986]" pageId="3" pageNumber="564">
|
||
<paragraph id="8BDD5C71FF93897CF9C510CB3CC6409A" blockId="3.[151,983,1964,1986]" box="[151,983,1964,1986]" pageId="3" pageNumber="564">
|
||
<emphasis id="B9168063FF93897CF9C510CB3E03409A" bold="true" box="[151,274,1964,1986]" pageId="3" pageNumber="564">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6227F2FF93897CF84B10CB3EF6409A" authorityName="Spangler" authorityYear="1981" box="[281,487,1964,1986]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="breweri">
|
||
<emphasis id="B9168063FF93897CF84B10CB3EF6409A" box="[281,487,1964,1986]" italics="true" pageId="3" pageNumber="564">Tepuidessus breweri</emphasis>
|
||
</taxonomicName>
|
||
, head and pronotum of male (A) and female (B).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1D0CF9FF94897BF9C513AB3B7443BA" box="[151,1125,1228,1250]" httpUri="https://zenodo.org/record/1292247/files/figure.png" pageId="4" pageNumber="565" startId="4.[151,250,1228,1250]" targetBox="[160,1427,194,1206]" targetPageId="4">
|
||
<paragraph id="8BDD5C71FF94897BF9C513AB3B7443BA" blockId="4.[151,1125,1228,1250]" box="[151,1125,1228,1250]" pageId="4" pageNumber="565">
|
||
<emphasis id="B9168063FF94897BF9C513AB3E0343BA" bold="true" box="[151,274,1228,1250]" pageId="4" pageNumber="565">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6227F2FF94897BF84B13AB3EC643BA" authority="Kodada & Hendrich & Balke, 2018" authorityName="Kodada & Hendrich & Balke" authorityYear="2018" box="[281,471,1228,1250]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="grulai" status="sp. nov.">
|
||
<emphasis id="B9168063FF94897BF84B13AB3EC643BA" box="[281,471,1228,1250]" italics="true" pageId="4" pageNumber="565">Tepuidessus grulai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9168063FF94897BF88F13AB3D3C43BA" bold="true" box="[477,557,1228,1250]" pageId="4" pageNumber="565">
|
||
<taxonomicNameLabel id="A2253D18FF94897BF88F13AB3D3C43BA" box="[477,557,1228,1250]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus of male (A) and female (B). Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDD5C71FF94897BF995126E3CE1428E" blockId="4.[151,1437,1289,2034]" pageId="4" pageNumber="565">
|
||
<emphasis id="B9168063FF94897BF995126E3E52427A" bold="true" box="[199,323,1289,1314]" pageId="4" pageNumber="565">Structures</emphasis>
|
||
. Antenna stout, moniliform. Head without occipital line and with rounded clypeus (as in
|
||
<figureCitation id="135940F4FF94897BFC1B126E3A9E4279" box="[1353,1423,1289,1314]" captionStart="FIGURE 4" captionStartId="5.[151,250,1981,2003]" captionTargetBox="[331,1256,447,1959]" captionTargetId="figure@5.[331,1256,447,1959]" captionTargetPageId="5" captionText="FIGURE 4. Tepuidessus grulai sp. nov., head and pronotum of male (A) and female (B)." httpUri="https://zenodo.org/record/1292249/files/figure.png" pageId="4" pageNumber="565">Fig. 4</figureCitation>
|
||
). Pronotum without lateral bead; with distinct and very deep basal striae (as in
|
||
<figureCitation id="135940F4FF94897BFD5A124B3B99421D" box="[1032,1160,1324,1349]" captionStart="FIGURE 6" captionStartId="7.[151,250,1181,1203]" captionTargetBox="[151,1435,196,1158]" captionTargetId="figure@7.[151,1435,193,1160]" captionTargetPageId="7" captionText="FIGURE 6. Tepuidessus grulai sp. nov., reduced metathoracic wing (A), lateral view of last ventrite in female (B) and pronotum, punctation and basal stria as well as basal stria on elytron (C)." httpUri="https://zenodo.org/record/1292253/files/figure.png" pageId="4" pageNumber="565">Figs 6A, C</figureCitation>
|
||
). Elytron with short and deep basal striae, but without sutural line, with distinct longitudinal depression on elytral disc roughly in middle of each elytron (as in
|
||
<figureCitation id="135940F4FF94897BF83B12123EAE42D5" box="[361,447,1397,1422]" captionStart="FIGURE 3" captionStartId="4.[151,250,1228,1250]" captionTargetBox="[160,1427,194,1206]" captionTargetId="figure@4.[159,1427,193,1206]" captionTargetPageId="4" captionText="FIGURE 3. Tepuidessus grulai sp. nov., habitus of male (A) and female (B). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1292247/files/figure.png" pageId="4" pageNumber="565">Fig. 3B</figureCitation>
|
||
, arrow). Basal epipleural transverse carina absent. Metathoracic wings vestigial, reduced to short membranous vestiges without any sign of venation (
|
||
<figureCitation id="135940F4FF94897BFA1D12FE3C8442EA" box="[847,917,1433,1458]" captionStart="FIGURE 6" captionStartId="7.[151,250,1181,1203]" captionTargetBox="[151,1435,196,1158]" captionTargetId="figure@7.[151,1435,193,1160]" captionTargetPageId="7" captionText="FIGURE 6. Tepuidessus grulai sp. nov., reduced metathoracic wing (A), lateral view of last ventrite in female (B) and pronotum, punctation and basal stria as well as basal stria on elytron (C)." httpUri="https://zenodo.org/record/1292253/files/figure.png" pageId="4" pageNumber="565">Fig. 6</figureCitation>
|
||
). Pro- and mesotarsi appearing stout because tarsomeres 1–3 distinctly dilated laterally; metatrochanter moderately offset.
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF94897BF99512863D26413E" blockId="4.[151,1437,1289,2034]" pageId="4" pageNumber="565">
|
||
<emphasis id="B9168063FF94897BF99512863E6542A2" bold="true" box="[199,372,1505,1530]" pageId="4" pageNumber="565">Male genitalia</emphasis>
|
||
. Median lobe of aedeagus simply curved, long in lateral view; in ventral view, slender and gently narrowed towards tip (
|
||
<figureCitation id="135940F4FF94897BF8BA11623D7B4145" box="[488,618,1541,1566]" captionStart="FIGURE 7" captionStartId="8.[151,250,1124,1146]" captionTargetBox="[463,1146,216,1088]" captionTargetId="figure@8.[419,1168,195,1102]" captionTargetPageId="8" captionText="FIGURE 7. Tepuidessus grulai sp. nov., median lobe of aedeagus and paramere in lateral view (A) and median lobe in ventral view (B)" httpUri="https://zenodo.org/record/1292255/files/figure.png" pageId="4" pageNumber="565">Figs 7A, B</figureCitation>
|
||
); lateral lobes (parameres) bisegmented and of a general
|
||
<taxonomicName id="4C6227F2FF94897BFDA911623A724146" authority="Kodada & Hendrich & Balke, 2018" authorityName="Kodada & Hendrich & Balke" authorityYear="2018" box="[1275,1379,1541,1566]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="561" phylum="Arthropoda" rank="genus" status="sp. nov.">Bidessini</taxonomicName>
|
||
<typeStatus id="54D9E2D3FF94897BFC3E11613A8D4146" box="[1388,1436,1542,1566]" pageId="4" pageNumber="565">type</typeStatus>
|
||
(see
|
||
<bibRefCitation id="EFF32180FF94897BF999114E3E764119" box="[203,359,1577,1602]" pageId="4" pageNumber="565" refId="ref2809" refString="Bistrom, O. (1988) Generic review of the Bidessini (Coleoptera, Dytiscidae). Acta Zoologica Fennica, 184, 1 - 41." type="journal article">Biström 1988</bibRefCitation>
|
||
), tip of distal joint rounded (
|
||
<figureCitation id="135940F4FF94897BFBF5114E3C114119" box="[679,768,1577,1602]" captionStart="FIGURE 7" captionStartId="8.[151,250,1124,1146]" captionTargetBox="[463,1146,216,1088]" captionTargetId="figure@8.[419,1168,195,1102]" captionTargetPageId="8" captionText="FIGURE 7. Tepuidessus grulai sp. nov., median lobe of aedeagus and paramere in lateral view (A) and median lobe in ventral view (B)" httpUri="https://zenodo.org/record/1292255/files/figure.png" pageId="4" pageNumber="565">Fig. 7A</figureCitation>
|
||
) and with no obvious "nose" or hook (as in e.g.
|
||
<bibRefCitation id="EFF32180FF94897BFC4E114E3FC2413E" pageId="4" pageNumber="565" refId="ref2668" refString="Balke, M., Ruthensteiner, B., Warikar, E. L., Neven, K. & Hendrich, L. (2015) Two new species of Limbodessus diving beetles from New Guinea - short verbal descriptions flanked by online content (digital photography, µCT scans, drawings and DNA sequence data). Biodiversity Data Journal, 3, e 7096. https: // doi. org / 10.3897 / bdj. 3. e 7096" type="journal article">
|
||
Balke
|
||
<emphasis id="B9168063FF94897BFC36114D3A84411A" box="[1380,1429,1577,1602]" italics="true" pageId="4" pageNumber="565">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 31;
|
||
<bibRefCitation id="EFF32180FF94897BF867112A3EC2413D" box="[309,467,1613,1638]" pageId="4" pageNumber="565" refId="ref2809" refString="Bistrom, O. (1988) Generic review of the Bidessini (Coleoptera, Dytiscidae). Acta Zoologica Fennica, 184, 1 - 41." type="journal article">Biström 1988</bibRefCitation>
|
||
: fig. 15).
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF94897BF99511163A4241F6" blockId="4.[151,1437,1289,2034]" pageId="4" pageNumber="565">
|
||
<emphasis id="B9168063FF94897BF99511163E0F41D2" bold="true" box="[199,286,1649,1674]" pageId="4" pageNumber="565">Female</emphasis>
|
||
. Dorsal and ventral surface dull due to well impressed microreticulation between surface punctation (
|
||
<figureCitation id="135940F4FF94897BF9CD11F23FEB41F5" box="[159,250,1685,1710]" captionStart="FIGURE 3" captionStartId="4.[151,250,1228,1250]" captionTargetBox="[160,1427,194,1206]" captionTargetId="figure@4.[159,1427,193,1206]" captionTargetPageId="4" captionText="FIGURE 3. Tepuidessus grulai sp. nov., habitus of male (A) and female (B). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1292247/files/figure.png" pageId="4" pageNumber="565">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="135940F4FF94897BF85411F23E3841F5" box="[262,297,1685,1709]" captionStart="FIGURE 4" captionStartId="5.[151,250,1981,2003]" captionTargetBox="[331,1256,447,1959]" captionTargetId="figure@5.[331,1256,447,1959]" captionTargetPageId="5" captionText="FIGURE 4. Tepuidessus grulai sp. nov., head and pronotum of male (A) and female (B)." httpUri="https://zenodo.org/record/1292249/files/figure.png" pageId="4" pageNumber="565">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135940F4FF94897BF86711F23E5441F6" box="[309,325,1685,1710]" captionStart="FIGURE 5" captionStartId="6.[151,250,1971,1993]" captionTargetBox="[180,1421,195,1939]" captionTargetId="figure@6.[180,1421,195,1940]" captionTargetPageId="6" captionText="FIGURE 5. Tepuidessus grulai sp. nov., ventral surface." httpUri="https://zenodo.org/record/1292251/files/figure.png" pageId="4" pageNumber="565">5</figureCitation>
|
||
). Apical ventrite flattened before posterior margin, tip in lateral view beak-shaped (
|
||
<figureCitation id="135940F4FF94897BFDBC11F23A5741F6" box="[1262,1350,1685,1710]" captionStart="FIGURE 6" captionStartId="7.[151,250,1181,1203]" captionTargetBox="[151,1435,196,1158]" captionTargetId="figure@7.[151,1435,193,1160]" captionTargetPageId="7" captionText="FIGURE 6. Tepuidessus grulai sp. nov., reduced metathoracic wing (A), lateral view of last ventrite in female (B) and pronotum, punctation and basal stria as well as basal stria on elytron (C)." httpUri="https://zenodo.org/record/1292253/files/figure.png" pageId="4" pageNumber="565">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF94897BF99511DE3CE84189" blockId="4.[151,1437,1289,2034]" box="[199,1017,1721,1746]" pageId="4" pageNumber="565">
|
||
<emphasis id="B9168063FF94897BF99511DE3E7A418A" bold="true" box="[199,363,1721,1746]" pageId="4" pageNumber="565">Size variation</emphasis>
|
||
. Total length:
|
||
<quantity id="4C9AF194FF94897BFB5C11DE3D894189" box="[526,664,1721,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="4" pageNumber="565" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
|
||
; maximum width:
|
||
<quantity id="4C9AF194FF94897BFA3811DE3CE54189" box="[874,1012,1721,1745]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="4" pageNumber="565" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3780FFAFF94897AF99511BA3E2C478D" lastPageId="5" lastPageNumber="566" pageId="4" pageNumber="565" type="diagnosis">
|
||
<paragraph id="8BDD5C71FF94897BF99511BA3C5840F1" blockId="4.[151,1437,1289,2034]" pageId="4" pageNumber="565">
|
||
<emphasis id="B9168063FF94897BF99511BA3ED641AE" bold="true" box="[199,455,1757,1782]" pageId="4" pageNumber="565">Differential diagnosis</emphasis>
|
||
. Using the key of
|
||
<bibRefCitation id="EFF32180FF94897BFBC911BA3CE841AD" box="[667,1017,1757,1782]" pageId="4" pageNumber="565" refId="ref2940" refString="Miller, K. B. & Bergsten, J. (2016) Diving Beetles of the World. Systematics and Biology of the Dytiscidae. Johns Hopkins University Press, Baltimore, 320 pp." type="book">Miller & Bergsten (2016: 220)</bibRefCitation>
|
||
, the new species from Acopán Tepui does not key out with
|
||
<taxonomicName id="4C6227F2FF94897BF8C210663D074042" authorityName="Spangler" authorityYear="1981" box="[400,534,1793,1818]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9168063FF94897BF8C210663D074042" box="[400,534,1793,1818]" italics="true" pageId="4" pageNumber="565">Tepuidessus</emphasis>
|
||
</taxonomicName>
|
||
, but with
|
||
<taxonomicName id="4C6227F2FF94897BFBDB10663CBD4041" authority="Balke, 2001" authorityName="Balke" authorityYear="2001" box="[649,940,1793,1818]" class="Insecta" family="Dytiscidae" genus="Papuadessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9168063FF94897BFBDB10663C0A4042" box="[649,795,1793,1818]" italics="true" pageId="4" pageNumber="565">Papuadessus</emphasis>
|
||
<bibRefCitation id="EFF32180FF94897BFA7010663CBD4041" box="[802,940,1793,1818]" pageId="4" pageNumber="565" refId="ref2476" refString="Balke, M. (2001) Papuadessus pakdjoko - a new genus and species of rheobiont diving beetle from New Guinea of potential use for environmental impact assessments (Coleoptera, Dytiscidae). Hydrobiologia, 464 (1), 107 - 112. https: // doi. org / 10.1023 / A: 1013931032342" type="journal article">Balke, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, a genus endemic to the Papuan region. The two
|
||
<taxonomicName id="4C6227F2FF94897BF99A10423E4B4066" authorityName="Balke" authorityYear="2001" box="[200,346,1829,1854]" class="Insecta" family="Dytiscidae" genus="Papuadessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9168063FF94897BF99A10423E4B4066" box="[200,346,1829,1854]" italics="true" pageId="4" pageNumber="565">Papuadessus</emphasis>
|
||
</taxonomicName>
|
||
species are morphologically rather divergent and considered members of one lineage only based on DNA sequence data (
|
||
<bibRefCitation id="EFF32180FF94897BF8E2102E3D264039" box="[432,567,1865,1890]" pageId="4" pageNumber="565" refId="ref2476" refString="Balke, M. (2001) Papuadessus pakdjoko - a new genus and species of rheobiont diving beetle from New Guinea of potential use for environmental impact assessments (Coleoptera, Dytiscidae). Hydrobiologia, 464 (1), 107 - 112. https: // doi. org / 10.1023 / A: 1013931032342" type="journal article">Balke 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF32180FF94897BFB17102E3C1C4039" box="[581,781,1865,1890]" pageId="4" pageNumber="565" refId="ref2753" refString="Balke, M., Warikar, E., Toussaint, E. F. A. & Hendrich, L. (2013) Papuadessus baueri spec. nov. from Biak Island, Papua (Coleoptera, Dytiscidae, Hydroporinae). Spixiana, 36 (2), 283 - 288." type="journal article">
|
||
Balke
|
||
<emphasis id="B9168063FF94897BFBDD102D3DD3403A" box="[655,706,1865,1890]" italics="true" pageId="4" pageNumber="565">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF32180FF94897BFA48102E3B20403A" box="[794,1073,1865,1890]" pageId="4" pageNumber="565" refId="ref2940" refString="Miller, K. B. & Bergsten, J. (2016) Diving Beetles of the World. Systematics and Biology of the Dytiscidae. Johns Hopkins University Press, Baltimore, 320 pp." type="book">Miller & Bergsten 2016</bibRefCitation>
|
||
). In the absence of any obvious morphological apomorphy that would group the species treated here with
|
||
<taxonomicName id="4C6227F2FF94897BFA81100A3B7440DE" authorityName="Balke" authorityYear="2001" box="[979,1125,1901,1926]" class="Insecta" family="Dytiscidae" genus="Papuadessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9168063FF94897BFA81100A3B7440DE" box="[979,1125,1901,1926]" italics="true" pageId="4" pageNumber="565">Papuadessus</emphasis>
|
||
</taxonomicName>
|
||
, or indeed any other genus, we opt for the pragmatic approach to assign it to
|
||
<taxonomicName id="4C6227F2FF94897BFBEE10F63C5340F2" authorityName="Spangler" authorityYear="1981" box="[700,834,1937,1962]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9168063FF94897BFBEE10F63C5340F2" box="[700,834,1937,1962]" italics="true" pageId="4" pageNumber="565">Tepuidessus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF94897AF99510D23E2C478D" blockId="4.[151,1437,1289,2034]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="566" pageId="4" pageNumber="565">
|
||
<taxonomicName id="4C6227F2FF94897BF99510D23EB14096" authority="Kodada & Hendrich & Balke, 2018" authorityName="Kodada & Hendrich & Balke" authorityYear="2018" box="[199,416,1973,1998]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="grulai" status="sp. nov.">
|
||
<emphasis id="B9168063FF94897BF99510D23EB14096" box="[199,416,1973,1998]" italics="true" pageId="4" pageNumber="565">Tepuidessus grulai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9168063FF94897BF8FD10D23D014096" bold="true" box="[431,528,1973,1998]" pageId="4" pageNumber="565">
|
||
<taxonomicNameLabel id="A2253D18FF94897BF8FD10D23D014096" box="[431,528,1973,1998]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C6227F2FF94897BFBE910D13C254096" authorityName="Spangler" authorityYear="1981" box="[699,820,1973,1998]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="breweri">
|
||
<emphasis id="B9168063FF94897BFBE910D13C254096" box="[699,820,1973,1998]" italics="true" pageId="4" pageNumber="565">T. breweri</emphasis>
|
||
</taxonomicName>
|
||
by (1) much broader habitus with distinct outline discontinuity between pronotum and elytra, (2) well developed, long and deep basal striae on pronotum and elytra and (3) by unbordered apical abdominal ventrite. The median lobe of aedeagus is more narrowed than in
|
||
<taxonomicName id="4C6227F2FF95897AFC7917FE3A8D47E8" authorityName="Spangler" authorityYear="1981" box="[1323,1436,151,176]" class="Insecta" family="Dytiscidae" genus="Tepuidessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="breweri">
|
||
<emphasis id="B9168063FF95897AFC7917FE3A8D47E8" box="[1323,1436,151,176]" italics="true" pageId="5" pageNumber="566">T. breweri</emphasis>
|
||
</taxonomicName>
|
||
in lateral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3780FFAFF95897AF99517B83FE64645" pageId="5" pageNumber="566" type="etymology">
|
||
<paragraph id="8BDD5C71FF95897AF99517B83FE64645" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="566">
|
||
<emphasis id="B9168063FF95897AF99517B83E5747A0" bold="true" box="[199,326,223,248]" pageId="5" pageNumber="566">Etymology</emphasis>
|
||
. Named after Daniel Gruľa, one of the collectors of the new species. It is a noun in the genitive singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3780FFAFF95897AF99516403CE546D0" pageId="5" pageNumber="566" type="distribution">
|
||
<paragraph id="8BDD5C71FF95897AF99516403BDC463D" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="566">
|
||
<emphasis id="B9168063FF95897AF99516403E484618" bold="true" box="[199,345,295,320]" pageId="5" pageNumber="566">Distribution</emphasis>
|
||
. Only known from the
|
||
<typeStatus id="54D9E2D3FF95897AFB23164F3DB04618" box="[625,673,296,320]" pageId="5" pageNumber="566">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="135940F4FF95897AFA4016403C4B4618" box="[786,858,295,320]" captionStart="FIGURE 8" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[178,1415,1226,1961]" captionTargetId="figure@8.[178,1415,1226,1961]" captionTargetPageId="8" captionText="FIGURE 8. Distribution of Tepuidessus breweri (black dot) and Tepuidessus grulai sp. nov. (red square) in southern Venezuela." httpUri="https://zenodo.org/record/1292257/files/figure.png" pageId="5" pageNumber="566">Fig. 8</figureCitation>
|
||
). This locality is situated around
|
||
<quantity id="4C9AF194FF95897AFDB6164F3A2C4618" box="[1252,1341,295,320]" metricMagnitude="5" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="566" unit="km" value="140.0">140 km</quantity>
|
||
west of Mount Roraima, the locality of the other
|
||
<taxonomicName id="4C6227F2FF95897AFB31162B3DDA463D" authority="Kodada & Hendrich & Balke, 2018" authorityName="Kodada & Hendrich & Balke" authorityYear="2018" box="[611,715,332,357]" class="Insecta" family="Dytiscidae" genus="Bidessini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="561" phylum="Arthropoda" rank="genus" status="sp. nov.">Bidessini</taxonomicName>
|
||
species described from the plateau of a tepui.
|
||
</paragraph>
|
||
<paragraph id="8BDD5C71FF95897AF99516083CE546D0" blockId="5.[151,1436,151,392]" box="[199,1012,367,392]" pageId="5" pageNumber="566">
|
||
<emphasis id="B9168063FF95897AF99516083E3246D0" bold="true" box="[199,291,367,392]" pageId="5" pageNumber="566">Habitat</emphasis>
|
||
. Puddles on rocks of summit plateau of Acopán Tepui (
|
||
<figureCitation id="135940F4FF95897AFAC116083CF746D0" box="[915,998,367,392]" captionStart="FIGURE 10" captionStartId="10.[151,250,1727,1749]" captionTargetBox="[278,1308,193,1705]" captionTargetId="figure@10.[278,1308,193,1705]" captionTargetPageId="10" captionText="FIGURE 10. Habitat of Tepuidessus grulai sp. nov. Puddles on rocks of summit plateau of Acopán Tepui (Foto: Peter Fenďa)." httpUri="https://zenodo.org/record/1292261/files/figure.png" pageId="5" pageNumber="566">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |