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<document id="812016F61E626AE68BD1334063AC45FF" ID-CLB-Dataset="51708" ID-DOI="10.1002/ajp.20191" ID-GBIF-Dataset="3ded904d-e34f-4cd1-9903-79b8392a7f66" checkinTime="1436211515535" checkinUser="donat" docAuthor="Urs Thalmann &amp; Thomas Geissmann" docDate="2005" docId="03CB3A0C5A1F055BFF3C0965FDFFCD2E" docLanguage="en" docName="2005avahi_cleesei.pdf" docOrigin="AmerIcan Journal of PrImatology 67" docTitle="Avahi cleesei New, New SpecIes" docType="treatment" docVersion="7" lastPageNumber="375" masterDocId="FFF242745A1D055FFF9A0D61FFDFC83A" masterDocTitle="New Species of Woolly Lemur Avahi (Primates: Lemuriformes) in Bemaraha (Central Western Madagascar)" masterLastPageNumber="376" masterPageNumber="371" pageNumber="373" updateTime="1732846922826" updateUser="ExternalLinkService">
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<mods:title id="353882F39D4BD695FB2A183BB2FB3D5F">New Species of Woolly Lemur Avahi (Primates: Lemuriformes) in Bemaraha (Central Western Madagascar)</mods:title>
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<mods:name id="DB7C5974E1B702E3047D3EB5C067380F" type="personal">
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<mods:namePart id="F6489964228CB81DA7BC7122D30ECE09">Urs Thalmann</mods:namePart>
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<mods:namePart id="2A3CE7EF3980B8D6936244A47B50C4E8">Thomas Geissmann</mods:namePart>
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<mods:title id="177D2AF391EC9727DE1A25116B0D6649">AmerIcan Journal of PrImatology</mods:title>
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<mods:part id="97EA31C6A2F1DC91BDB92696905D2941">
<mods:date id="C3C2F044FFECC198FA69E6065F4272B6">2005</mods:date>
<mods:detail id="CF4EF02E6B0B9D3B62AE730833701298" type="volume">
<mods:number id="77B9A4834E045D26CA9DDFCF12A30501">67</mods:number>
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<mods:identifier id="10DCC8FFEB5D7E899C38B559E8E31C96" type="DOI">10.1002/ajp.20191</mods:identifier>
</mods:mods>
<treatment id="03CB3A0C5A1F055BFF3C0965FDFFCD2E" ID-DOI="http://doi.org/10.5281/zenodo.6266920" ID-GBIF-Taxon="114924054" ID-Zenodo-Dep="6266920" LSID="urn:lsid:plazi:treatment:03CB3A0C5A1F055BFF3C0965FDFFCD2E" httpUri="http://treatment.plazi.org/id/03CB3A0C5A1F055BFF3C0965FDFFCD2E" lastPageId="4" lastPageNumber="375" pageId="2" pageNumber="373" scope_infraOrder="Lemuriformes" scope_order="Primates">
<subSubSection id="C378D8915A1F055DFF3C0965FDD3CC26" box="[166,524,1028,1052]" pageId="2" pageNumber="373" type="nomenclature">
<paragraph id="8BDD8B1A5A1F055DFF3C0965FDD3CC26" box="[166,524,1028,1052]" pageId="2" pageNumber="373">
<heading id="D0953C765A1F055DFF3C0965FDD3CC26" bold="true" box="[166,524,1028,1052]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<taxonomicName id="4C62F0995A1F055DFF3C0965FE41CC26" ID-CoL="K29P" authority="New" authorityName="New" box="[166,414,1028,1052]" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="2" pageNumber="373" phylum="Chordata" rank="species" species="cleesei">
<emphasis id="B91657085A1F055DFF3C0965FE84CC26" bold="true" box="[166,347,1028,1052]" italics="true" pageId="2" pageNumber="373">
<taxonomicName id="4C62F0995A1F055DFF3C0965FF2BCC26" box="[166,244,1028,1052]" pageId="2" pageNumber="373">Avahi</taxonomicName>
cleesei:
</emphasis>
<taxonomicNameLabel id="A225EA735A1F055DFEFE0965FDD3CC26" box="[356,524,1028,1052]" pageId="2" pageNumber="373" rank="species">New SpecIes</taxonomicNameLabel>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378D8915A1F055DFF40094AFED1CCCA" pageId="2" pageNumber="373" type="description">
<paragraph id="8BDD8B1A5A1F055DFF40094AFCFBCC7E" box="[218,804,1067,1092]" pageId="2" pageNumber="373">
<heading id="D0953C765A1F055DFF40094AFCFBCC7E" box="[218,804,1067,1092]" fontSize="10" level="4" pageId="2" pageNumber="373" reason="1">Bemaraha form [Thalmann &amp; Geissmann, 2000].</heading>
</paragraph>
<paragraph id="8BDD8B1A5A1F055DFF3C0915FEADCCB6" box="[166,370,1140,1164]" pageId="2" pageNumber="373">
<heading id="D0953C765A1F055DFF3C0915FEADCCB6" bold="true" box="[166,370,1140,1164]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis id="B91657085A1F055DFF3C0915FEADCCB6" bold="true" box="[166,370,1140,1164]" italics="true" pageId="2" pageNumber="373">
<typeStatus id="54D935B85A1F055DFF3C0915FECFCCB6" box="[166,272,1140,1164]" pageId="2" pageNumber="373">Holotype</typeStatus>
(
<figureCitation id="1359979F5A1F055DFEBE0915FEB7CCB6" box="[292,360,1140,1164]" captionStart="Fig. 2" captionStartId="3.[166,201,1175,1195]" captionTargetBox="[175,1115,188,1155]" captionTargetId="figure@3.[173,1117,185,1159]" captionTargetPageId="3" captionText="Fig. 2. Type photographs (AIMZ 13854. b-d) of an adult male Avahi cleesei after it was captured on 3 October 1991 (a and b) and after it was released on 4 October 1991 (c). [Color figure can be viewed in the online issue, which is available at www. interscience. wiley. com.]" pageId="2" pageNumber="373">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A1F055DFF4009F0FBBCCC93" box="[218,1123,1168,1193]" pageId="2" pageNumber="373">
<materialsCitation id="3B0A81475A1F055DFF4009F0FB80CC93" ID-GBIF-Occurrence="1227780005" box="[218,1119,1168,1193]" collectingDate="1991-10-03" country="Afghanistan" location="Bemaraha" pageId="2" pageNumber="373" specimenCount="1">
<taxonomicName id="4C62F0995A1F055DFF4009F0FEC0CC93" box="[218,287,1169,1193]" pageId="2" pageNumber="373">
<emphasis id="B91657085A1F055DFF4009F0FEC0CC93" bold="true" box="[218,287,1169,1193]" italics="true" pageId="2" pageNumber="373">Avahi</emphasis>
</taxonomicName>
male (body weight =
<quantity id="4C9A26FF5A1F055DFDB309F0FDB7CC93" box="[553,616,1168,1193]" metricMagnitude="-1" metricUnit="kg" metricValue="8.3" pageId="2" pageNumber="373" unit="g" value="830.0">830 g</quantity>
) captured in
<location id="8EBDDDC15A1F055DFC9C09F1FCA0CC93" LSID="urn:lsid:plazi:treatment:03CB3A0C5A1F055BFF3C0965FDFFCD2E:8EBDDDC15A1F055DFC9C09F1FCA0CC93" box="[774,895,1168,1193]" country="Afghanistan" name="Bemaraha" pageId="2" pageNumber="373">Bemaraha</location>
on
<collectingDate id="EF9854325A1F055DFC3009F0FB80CC93" box="[938,1119,1168,1193]" pageId="2" pageNumber="373" value="1991-10-03">
<date id="FFDCADDA5A1F055DFC3009F0FB80CC93" box="[938,1119,1168,1193]" pageId="2" pageNumber="373" value="1991-10-03">3 October 1991</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD8B1A5A1F055DFF3C09B8FED1CCCA" box="[166,270,1241,1264]" pageId="2" pageNumber="373">
<heading id="D0953C765A1F055DFF3C09B8FED1CCCA" bold="true" box="[166,270,1241,1264]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis id="B91657085A1F055DFF3C09B8FED1CCCA" bold="true" box="[166,270,1241,1264]" italics="true" pageId="2" pageNumber="373">Material</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378D8915A1F055DFF400994FBBCCDF0" pageId="2" pageNumber="373" type="materials_examined">
<paragraph id="8BDD8B1A5A1F055DFF400994FD24CD5F" pageId="2" pageNumber="373">
<materialsCitation id="3B0A81475A1F055DFF400994FD28CD5F" ID-GBIF-Occurrence="1227780004" collectionCode="AIMZ" country="Madagascar" pageId="2" pageNumber="373" specimenCode="AIMZ 13854" specimenCount="1">
Hair samples (
<specimenCode id="DBC423615A1F055DFE1E0994FDC2CD34" box="[388,541,1269,1294]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.a), accompanying photographs (
<specimenCode id="DBC423615A1F055DFC080994FBF7CD34" box="[914,1064,1269,1294]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.b-n), and videotape (
<specimenCode id="DBC423615A1F055DFEC50873FE27CD11" box="[351,504,1298,1323]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.o) and audiotape recordings (
<specimenCode id="DBC423615A1F055DFCC70873FC28CD11" box="[861,1015,1298,1323]" pageId="2" pageNumber="373">AIMZ 13854</specimenCode>
.p) of the
<specimenCount id="9D6440935A1F055DFF3C0851FECBCD72" box="[166,276,1328,1352]" pageId="2" pageNumber="373" type="generic">specimen</specimenCount>
are stored in the Anthropological Institute and Museum of the University of Zurich (
<collectionCode id="ED7313DF5A1F055DFE33082CFE2CCD5F" box="[425,499,1357,1381]" collectionName="AIMZ" pageId="2" pageNumber="373">AIMZ</collectionCode>
), Zurich, Switzerland
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD8B1A5A1F055DFF3C08F7FE9ECD97" box="[166,321,1429,1453]" pageId="2" pageNumber="373">
<heading id="D0953C765A1F055DFF3C08F7FE9ECD97" bold="true" box="[166,321,1429,1453]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis id="B91657085A1F055DFF3C08F7FE9ECD97" bold="true" box="[166,321,1429,1453]" italics="true" pageId="2" pageNumber="373">
<typeStatus id="54D935B85A1F055DFF3C08F7FF3FCD97" box="[166,224,1430,1453]" pageId="2" pageNumber="373">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B0A81475A1F055DFF4008D3FBBCCDF0" ID-GBIF-Occurrence="1227780001" country="Madagascar" location="Madagascar" pageId="2" pageNumber="373" specimenCount="1">
<paragraph id="8BDD8B1A5A1F055DFF4008D3FD7FCDDD" pageId="2" pageNumber="373">
<geoCoordinate id="EE56EDDD5A1F055DFF4008D3FEECCDF0" box="[218,307,1455,1482]" pageId="2" pageNumber="373">18°59̸S</geoCoordinate>
,
<geoCoordinate id="EE56EDDD5A1F055DFED108D3FE78CDF0" box="[331,423,1455,1482]" pageId="2" pageNumber="373">44°45̸E</geoCoordinate>
, approximately
<locationDeviation id="9A6F92385A1F055DFDE308D3FD1FCDF0" box="[633,704,1458,1482]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="373" unit="km" value="3.0">
<quantity id="4C9A26FF5A1F055DFDE308D3FD1FCDF0" box="[633,704,1458,1482]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="373" unit="km" value="3.0">3 km</quantity>
</locationDeviation>
east-northeast of the village Ambalarano, western
<collectingCountry id="F375CB8A5A1F055DFE3708AEFDE5CDDD" box="[429,570,1487,1511]" name="Madagascar" pageId="2" pageNumber="373" value="Madagascar">Madagascar</collectingCountry>
(
<figureCitation id="1359979F5A1F055DFDD108AEFD51CDDD" box="[587,654,1487,1511]" captionStart="Fig. 1" captionStartId="1.[166,201,1119,1139]" captionTargetBox="[173,1116,187,1095]" captionTargetId="figure@1.[173,1117,185,1103]" captionTargetPageId="1" captionText="Fig. 1. Distribution map of western taxa of woolly lemurs." pageId="2" pageNumber="373">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BDD8B1A5A1F055DFBD708D3FBBCCDF0" box="[1101,1123,1458,1482]" pageId="2" pageNumber="373">of</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C378D8915A1F055DFF3C0B79FE3DCE9E" pageId="2" pageNumber="373" type="etymology">
<paragraph id="8BDD8B1A5A1F055DFF3C0B79FEFACE15" box="[166,293,1560,1583]" pageId="2" pageNumber="373">
<heading id="D0953C765A1F055DFF3C0B79FEFACE15" bold="true" box="[166,293,1560,1583]" fontSize="10" level="3" pageId="2" pageNumber="373" reason="0">
<emphasis id="B91657085A1F055DFF3C0B79FEFACE15" bold="true" box="[166,293,1560,1583]" italics="true" pageId="2" pageNumber="373">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A1F055DFF400B55FE3DCE9E" pageId="2" pageNumber="373">
Named in honor of British actor and comedian John Cleese for his promotion of conservation issues in movies such as
<emphasis id="B91657085A1F055DFDE50B33FCE1CE50" bold="true" box="[639,830,1618,1642]" italics="true" pageId="2" pageNumber="373">Fierce Creatures</emphasis>
[Chepisi &amp; Young, 1997] and documentaries such as
<emphasis id="B91657085A1F055DFE680B11FBBCCEBD" bold="true" box="[498,1123,1647,1671]" italics="true" pageId="2" pageNumber="373">Born to be Wild: Operation Lemur With John Cleese</emphasis>
[Kershaw &amp; Cleese, 1999].
</paragraph>
</subSubSection>
<caption id="DF1DDB925A1E055CFF3C09F6FC8DCCEC" pageId="3" pageNumber="374" targetBox="[175,1115,188,1155]" targetPageId="3">
<paragraph id="8BDD8B1A5A1E055CFF3C09F6FC8DCCEC" pageId="3" pageNumber="374">
Fig. 2. Type photographs (AIMZ 13854.b-d) of an adult male
<taxonomicName id="4C62F0995A1E055CFD6D09F9FCACCC91" box="[759,883,1176,1195]" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="cleesei">
<emphasis id="B91657085A1E055CFD6D09F9FCACCC91" bold="true" box="[759,883,1176,1195]" italics="true" pageId="3" pageNumber="374">
<taxonomicName id="4C62F0995A1E055CFD6D09F9FCEFCC91" box="[759,816,1176,1195]" pageId="3" pageNumber="374">Avahi</taxonomicName>
cleesei
</emphasis>
</taxonomicName>
after it was captured on 3 October 1991 (
<emphasis id="B91657085A1E055CFEDE09CFFE8FCCFB" bold="true" box="[324,336,1198,1217]" pageId="3" pageNumber="374">a</emphasis>
and
<emphasis id="B91657085A1E055CFEE509CFFE53CCFB" bold="true" box="[383,396,1198,1217]" pageId="3" pageNumber="374">b</emphasis>
) and after it was released on 4 October 1991 (
<emphasis id="B91657085A1E055CFCD809CFFC92CCFB" bold="true" box="[834,845,1198,1217]" pageId="3" pageNumber="374">c</emphasis>
). [Color figure can be viewed in the online issue, which is available at www.interscience.wiley.com.]
</paragraph>
</caption>
<subSubSection id="C378D8915A1E055CFF3C0877FCCACDBF" pageId="3" pageNumber="374" type="diagnosis">
<paragraph id="8BDD8B1A5A1E055CFF3C0877FEC0CD14" box="[166,287,1302,1326]" pageId="3" pageNumber="374">
<heading id="D0953C765A1E055CFF3C0877FEC0CD14" bold="true" box="[166,287,1302,1326]" fontSize="10" level="3" pageId="3" pageNumber="374" reason="0">
<emphasis id="B91657085A1E055CFF3C0877FEC0CD14" bold="true" box="[166,287,1302,1326]" italics="true" pageId="3" pageNumber="374">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A1E055CFF400853FCCACDBF" pageId="3" pageNumber="374">
Distinguished from
<taxonomicName id="4C62F0995A1E055CFE540852FDA1CD71" box="[462,638,1331,1355]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1340" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="B91657085A1E055CFE540852FDA1CD71" bold="true" box="[462,638,1331,1355]" italics="true" pageId="3" pageNumber="374">A. occidentalis</emphasis>
</taxonomicName>
by its lack of a white facial mask and broad dark eye-rings, and from both
<taxonomicName id="4C62F0995A1E055CFD180831FCEBCD52" box="[642,820,1360,1384]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1359" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="B91657085A1E055CFD180831FCEBCD52" bold="true" box="[642,820,1360,1384]" italics="true" pageId="3" pageNumber="374">A. occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C62F0995A1E055CFCE60831FBDBCD52" box="[892,1028,1360,1384]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1347" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B91657085A1E055CFCE60831FBDBCD52" bold="true" box="[892,1028,1360,1384]" italics="true" pageId="3" pageNumber="374">A. unicolor</emphasis>
</taxonomicName>
by the presence of a dark chevron pattern on the forehead.
</paragraph>
</subSubSection>
<subSubSection id="C378D8915A1E055BFF3C08A1FE31C95E" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374" type="description">
<paragraph id="8BDD8B1A5A1E055CFF3C08A1FE4DCDE2" box="[166,402,1472,1496]" pageId="3" pageNumber="374">
<heading id="D0953C765A1E055CFF3C08A1FE4DCDE2" bold="true" box="[166,402,1472,1496]" fontSize="10" level="3" pageId="3" pageNumber="374" reason="0">
<emphasis id="B91657085A1E055CFF3C08A1FE4DCDE2" bold="true" box="[166,402,1472,1496]" italics="true" pageId="3" pageNumber="374">
Description (
<figureCitation id="1359979F5A1E055CFEDE08A1FE57CDE2" box="[324,392,1472,1496]" captionStart="Fig. 2" captionStartId="3.[166,201,1175,1195]" captionTargetBox="[175,1115,188,1155]" captionTargetId="figure@3.[173,1117,185,1159]" captionTargetPageId="3" captionText="Fig. 2. Type photographs (AIMZ 13854. b-d) of an adult male Avahi cleesei after it was captured on 3 October 1991 (a and b) and after it was released on 4 October 1991 (c). [Color figure can be viewed in the online issue, which is available at www. interscience. wiley. com.]" pageId="3" pageNumber="374">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A1E055BFF4008BDFE31C95E" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374">
The face of
<taxonomicName id="4C62F0995A1E055CFEC508BCFE17CDCF" box="[351,456,1501,1525]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="300" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="cleesei">
<emphasis id="B91657085A1E055CFEC508BCFE17CDCF" bold="true" box="[351,456,1501,1525]" italics="true" pageId="3" pageNumber="374">A. cleesei</emphasis>
</taxonomicName>
is only slightly paler than the upper forehead and crown. The facial area above the nose extends upward toward the forehead (
<figureCitation id="1359979F5A1E055CFC53089BFBD2CE28" box="[969,1037,1530,1554]" captionStart="Fig. 2" captionStartId="3.[166,201,1175,1195]" captionTargetBox="[175,1115,188,1155]" captionTargetId="figure@3.[173,1117,185,1159]" captionTargetPageId="3" captionText="Fig. 2. Type photographs (AIMZ 13854. b-d) of an adult male Avahi cleesei after it was captured on 3 October 1991 (a and b) and after it was released on 4 October 1991 (c). [Color figure can be viewed in the online issue, which is available at www. interscience. wiley. com.]" pageId="3" pageNumber="374">Fig. 2</figureCitation>
c). This upward extension contrasts with the virtually opposite pattern created by the triangle of forehead pelage that invades the facial area in other western
<taxonomicName id="4C62F0995A1E055CFB870B54FBBCCE76" box="[1053,1123,1589,1612]" pageId="3" pageNumber="374">
<emphasis id="B91657085A1E055CFB870B54FBBCCE76" bold="true" box="[1053,1123,1589,1612]" italics="true" pageId="3" pageNumber="374">Avahi</emphasis>
</taxonomicName>
(e.g.,
<taxonomicName id="4C62F0995A1E055CFF7F0B33FE50CE50" box="[229,399,1618,1642]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1450" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="B91657085A1E055CFF7F0B33FE50CE50" bold="true" box="[229,399,1618,1642]" italics="true" pageId="3" pageNumber="374">A. occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C62F0995A1E055CFE5D0B33FD97CE50" box="[455,584,1618,1642]" class="Mammalia" family="Indriidae" genus="Avahi" genus.bestMatchDistance="1438" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="B91657085A1E055CFE5D0B33FD97CE50" bold="true" box="[455,584,1618,1642]" italics="true" pageId="3" pageNumber="374">A. unicolor</emphasis>
</taxonomicName>
). The forehead fur immediately bordering the facial area is blackish and forms a dark chevron pattern above the facial area. The eyes are maroon and have black, hairless eyelids. The snout is also black and hairless. The fur surrounding the corners of mouth is whitish. The fur on the head and body has a brown-gray coloration and a woolly (slightly curled) flecked appearance. The tail is beige or brown-gray, and is slightly reddish only on the dorsal side of the root. The inner dorsal surface of the lower limbs is white. The fur of the chest, belly, and inner surface of the upper limbs is relatively thin, downy, and very light gray.
</paragraph>
</subSubSection>
<subSubSection id="C378D8915A19055BFF3C0CF1FDB3CA4E" pageId="4" pageNumber="375" type="distribution">
<paragraph id="8BDD8B1A5A19055BFF3C0CF1FEE5C992" box="[166,314,400,424]" pageId="4" pageNumber="375">
<heading id="D0953C765A19055BFF3C0CF1FEE5C992" bold="true" box="[166,314,400,424]" fontSize="10" level="3" pageId="4" pageNumber="375" reason="0">
<emphasis id="B91657085A19055BFF3C0CF1FEE5C992" bold="true" box="[166,314,400,424]" italics="true" pageId="4" pageNumber="375">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A19055BFF400CCDFDB3CA4E" pageId="4" pageNumber="375">This species has been found only in the Tsingy de Bemaraha region north of the Manambolo River. The northern limit is unclear. There is no evidence from surveys and interviews that it is distributed in the wider surroundings of Bemaraha [Ausilio &amp; Raveloanrinoro, 1998; Sterling, 1998], the Namoroka Strict Nature Reserve (between the Sambao and Mahavavy rivers [Hawkins et al., 1998; Thalmann et al., 1999], or between the Mahavavy and Betsiboka rivers [Curtis, 1997; Thalmann &amp; Geissmann, 2000].</paragraph>
</subSubSection>
<subSubSection id="C378D8915A19055BFF3C0FC1FD29CB35" pageId="4" pageNumber="375" type="etymology">
<paragraph id="8BDD8B1A5A19055BFF3C0FC1FE5CCA82" box="[166,387,672,696]" pageId="4" pageNumber="375">
<heading id="D0953C765A19055BFF3C0FC1FE5CCA82" bold="true" box="[166,387,672,696]" fontSize="10" level="3" pageId="4" pageNumber="375" reason="0">
<emphasis id="B91657085A19055BFF3C0FC1FE5CCA82" bold="true" box="[166,387,672,696]" italics="true" pageId="4" pageNumber="375">Vernacular names</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A19055BFF400FDCFD29CB35" pageId="4" pageNumber="375">
Locally, these animals are called Dadintsifaky, which translates to grandparent of the sifaky (
<taxonomicName id="4C62F0995A19055BFE370FBAFDE8CAC8" box="[429,567,731,754]" class="Mammalia" family="Indriidae" genus="Propithecus" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="4" pageNumber="375" phylum="Chordata" rank="genus">
<emphasis id="B91657085A19055BFE370FBAFDE8CAC8" bold="true" box="[429,567,731,754]" italics="true" pageId="4" pageNumber="375">Propithecus</emphasis>
</taxonomicName>
). As a vernacular name in English, we propose Cleeses woolly lemur or Bemaraha woolly lemur.
</paragraph>
</subSubSection>
<subSubSection id="C378D8915A19055BFF3C0E5DFDFFCD2E" pageId="4" pageNumber="375" type="discussion">
<paragraph id="8BDD8B1A5A19055BFF3C0E5DFE4DCB69" box="[166,402,828,851]" pageId="4" pageNumber="375">
<heading id="D0953C765A19055BFF3C0E5DFE4DCB69" allCaps="true" bold="true" box="[166,402,828,851]" fontSize="10" level="1" pageId="4" pageNumber="375" reason="0">
<emphasis id="B91657085A19055BFF3C0E5DFE4DCB69" bold="true" box="[166,402,828,851]" pageId="4" pageNumber="375">CONSERVATION</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8B1A5A19055BFF400E02FC5DCC37" pageId="4" pageNumber="375">
<taxonomicName id="4C62F0995A19055BFF400E02FEB0CB41" box="[218,367,867,891]" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="cleesei">
<emphasis id="B91657085A19055BFF400E02FEB0CB41" bold="true" box="[218,367,867,891]" italics="true" pageId="4" pageNumber="375">
<taxonomicName id="4C62F0995A19055BFF400E02FEC0CB41" box="[218,287,867,891]" pageId="4" pageNumber="375">Avahi</taxonomicName>
cleesei
</emphasis>
</taxonomicName>
clearly falls into the category of Endangered (
<collectionCode id="ED7313DF5A19055BFC120E02FC6ECB41" box="[904,945,867,891]" pageId="4" pageNumber="375">EN</collectionCode>
) as established by the IUCN Red List [IUCN, 2001], according to the following criteria (B1ac(i, ii, iii, v)): the extent of occurrence is estimated to be less than
<quantity id="4C9A26FF5A19055BFCE30EFFFC38CB8F" box="[889,999,925,950]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="375" unit="km" value="5000.0">5,000 km</quantity>
2 (
<collectionCode id="ED7313DF5A19055BFB9E0EFCFBC5CB8F" box="[1028,1050,925,949]" pageId="4" pageNumber="375">B</collectionCode>
1), the species is known in just one location (
<collectionCode id="ED7313DF5A19055BFDC50EDBFDABCBE9" box="[607,628,954,979]" pageId="4" pageNumber="375">B</collectionCode>
1a), and the known population is declining (
<collectionCode id="ED7313DF5A19055BFF2A0EB9FF1ACBCA" box="[176,197,984,1008]" pageId="4" pageNumber="375">B</collectionCode>
1c) in the extent of occurrence (i); area of occupancy (ii); area, extent, and/or quality of habitat (iii); and number of mature individuals (v).
</paragraph>
<paragraph id="8BDD8B1A5A19055BFF400972FDFFCD2E" pageId="4" pageNumber="375">
However, more detailed analyses and surveys may reveal that the taxon should be moved to the Critically Endangered (
<collectionCode id="ED7313DF5A19055BFD7D094EFCD1CC7D" box="[743,782,1071,1095]" pageId="4" pageNumber="375">CR</collectionCode>
) category. For example, the disturbed forest close to the village of Ankinajao, which supported a substantial number of individuals in 1994 [Thalmann &amp; Geissmann, 2000], had been cut completely by 2003 (Thalmann, unpublished data). The subhumid forest at the base of the escarpment of the Tsingy de Bemaraha is under continuous pressure from bush fires that reduce it in many places every yearin some places to only a few meters in width. Such subhumid forests are the only habitat in which
<taxonomicName id="4C62F0995A19055BFB8709BEFF2FCD2E" class="Mammalia" family="Indriidae" genus="Avahi" higherTaxonomySource="CoL" kingdom="Animalia" order="Primates" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="cleesei">
<emphasis id="B91657085A19055BFB8709BEFF2FCD2E" bold="true" italics="true" pageId="4" pageNumber="375">
<taxonomicName id="4C62F0995A19055BFB8709BEFBBCCCCD" box="[1053,1123,1247,1271]" pageId="4" pageNumber="375">Avahi</taxonomicName>
cleesei
</emphasis>
</taxonomicName>
is known to occur so far.
</paragraph>
</subSubSection>
</treatment>
</document>