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<taxonomicName id="4C634D765C49FFAA6EADFEF4FD02FE9B" ID-CoL="FGNF" authority="Balthasar, 1945" authorityName="Balthasar" authorityYear="1945" box="[227,748,318,340]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="botulus" subGenus="Phalacronothus">
<emphasis id="B917EAE75C49FFAA6EADFEF4FD02FE9B" bold="true" box="[227,748,318,340]" pageId="1" pageNumber="8">
<emphasis id="B917EAE75C49FFAA6EADFEF4FEACFE9C" bold="true" box="[227,322,318,340]" italics="true" pageId="1" pageNumber="8">Aphodius</emphasis>
(
<emphasis id="B917EAE75C49FFAA6F1EFEF4FE01FE9C" bold="true" box="[336,495,318,340]" italics="true" pageId="1" pageNumber="8">Phalacronothus</emphasis>
)
<emphasis id="B917EAE75C49FFAA6FB2FEF4FDABFE9C" bold="true" box="[508,581,318,340]" italics="true" pageId="1" pageNumber="8">botulus</emphasis>
<bibRefCitation id="EFF24B045C49FFAA6C05FEF5FD02FE9B" author="Balthasar, V." box="[587,748,318,340]" pageId="1" pageNumber="8" pagination="40 - 44" refId="ref1577" refString="Balthasar, V. 1945. Quator Generis Aphodius Illig. Subgenera nova. Contributio 82. ad congnitionem Scarabaeidarum (Col.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 42: 40 - 44" type="book chapter" year="1945">Balthasar, 1945</bibRefCitation>
</emphasis>
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</paragraph>
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<treatmentCitation id="0AC210E45C49FFAA6EADFE4AFD0BFE5D" author="Balthasar, V." box="[227,741,384,406]" page="43" pageId="1" pageNumber="8" year="1945">
<taxonomicName id="4C634D765C49FFAA6EADFE4AFD0BFE5D" ID-CoL="FGNF" authority="Balthasar, 1945: 43" authorityName="Balthasar" authorityPageNumber="43" authorityYear="1945" box="[227,741,384,406]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="botulus" subGenus="Paragolius">
<emphasis id="B917EAE75C49FFAA6EADFE4AFED0FE5D" box="[227,318,384,405]" italics="true" pageId="1" pageNumber="8">Aphodius</emphasis>
(
<emphasis id="B917EAE75C49FFAA6F1EFE4AFE55FE5D" box="[336,443,384,405]" italics="true" pageId="1" pageNumber="8">Paragolius</emphasis>
)
<emphasis id="B917EAE75C49FFAA6F83FE4AFDFAFE5D" box="[461,532,384,405]" italics="true" pageId="1" pageNumber="8">botulus</emphasis>
<bibRefCitation id="EFF24B045C49FFAA6C50FE4AFD0BFE5D" author="Balthasar, V." box="[542,741,384,406]" pageId="1" pageNumber="8" pagination="40 - 44" refId="ref1577" refString="Balthasar, V. 1945. Quator Generis Aphodius Illig. Subgenera nova. Contributio 82. ad congnitionem Scarabaeidarum (Col.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 42: 40 - 44" type="book chapter" year="1945">Balthasar, 1945: 43</bibRefCitation>
</taxonomicName>
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.
<typeStatus id="54D888575C49FFAA6CBDFE4AFCCAFE5D" box="[755,804,384,405]" pageId="1" pageNumber="8">Type</typeStatus>
locality “Mandschukuo, circum oppidium Charbin” [=
<collectingCountry id="F37476655C49FFAA6F92FE51FDF5FE79" box="[476,539,411,433]" name="China" pageId="1" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="49A7F8175C49FFAA6C6AFE51FD49FE79" box="[548,679,411,433]" country="China" name="Heilongjiang" pageId="1" pageNumber="8">Heilongjiang</collectingRegion>
];
<treatmentCitation id="0AC210E45C49FFAA6CF8FE51FC6FFE79" author="Balthasar, V." box="[694,897,411,433]" page="281" pageId="1" pageNumber="8" year="1964">
<bibRefCitation id="EFF24B045C49FFAA6CF8FE51FC6FFE79" author="Balthasar, V." box="[694,897,411,433]" pageId="1" pageNumber="8" refId="ref1612" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 3. Aphodiidae. Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag. 652 pp." type="book" year="1964">Balthasar, 1964: 281</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="AB7311DB5C49FFAA6EADFE7DFCDCFE21" pageId="1" pageNumber="8">
<taxonomicName id="4C634D765C49FFAA6EADFE7DFDD2FE04" ID-CoL="FGNF" authorityName="Balthasar" authorityYear="1945" box="[227,572,439,460]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="botulus" subGenus="Phalacronothus">
<emphasis id="B917EAE75C49FFAA6EADFE7DFED0FE04" box="[227,318,439,460]" italics="true" pageId="1" pageNumber="8">Aphodius</emphasis>
(
<emphasis id="B917EAE75C49FFAA6F03FE7DFE08FE04" box="[333,486,439,460]" italics="true" pageId="1" pageNumber="8">Phalacronothus</emphasis>
)
<emphasis id="B917EAE75C49FFAA6FBBFE7DFDD2FE04" box="[501,572,439,460]" italics="true" pageId="1" pageNumber="8">botulus</emphasis>
</taxonomicName>
: Kim, 1989: 3;
<treatmentCitation id="0AC210E45C49FFAA6C97FE7DFC4CFE04" author="Takahashi, T." box="[729,930,439,461]" page="34" pageId="1" pageNumber="8" year="1999">
<bibRefCitation id="EFF24B045C49FFAA6C97FE7DFC4CFE04" author="Takahashi, T." box="[729,930,439,461]" pageId="1" pageNumber="8" refId="ref2324" refString="Takahashi, T. 1999. Systematic Catalog of Scarabaeoidea from Japan (1 st. ed. Supplement)." type="book" year="1999">Takahashi, 1999: 34</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6DE3FE7DFE4CFE20" author="Dellacasa, M. &amp; Dellacasa, G." page="134" pageId="1" pageNumber="8" year="2006">
<bibRefCitation id="EFF24B045C49FFAA6DE3FE7DFE4CFE20" author="Dellacasa, M. &amp; Dellacasa, G." pageId="1" pageNumber="8" pagination="105 - 143" refId="ref1649" refString="Dellacasa, M. &amp; Dellacasa, G. 2006. Tribe Aphodiini Leach, 1815. P. 105 - 143. In: Lobl, I., Smetana, A. (Ed.). Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup." type="book chapter" year="2006">Dellacasa &amp; Dellacasa, 2006: 134</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6FE2FE19FD64FE20" author="Murata, K. &amp; Tsuchiya, K. &amp; Suzuki, H. &amp; Tsukada, T." box="[428,650,467,488]" page="84" pageId="1" pageNumber="8" year="2011">
<bibRefCitation id="EFF24B045C49FFAA6FE2FE19FD64FE20" author="Murata, K. &amp; Tsuchiya, K. &amp; Suzuki, H. &amp; Tsukada, T." box="[428,650,467,488]" pageId="1" pageNumber="8" pagination="79 - 92" refId="ref1877" refString="Murata, K., Tsuchiya, K., Suzuki, H. &amp; Tsukada, T. 2011. Species composition of dung beetles in the grasslands of the Aso region and conservation of their habitats. Insect (New Series), 14 (2): 79 - 92. [In Japanese]. DOI: 10.20848 / kontyu. 14.2 _ 79" type="journal article" year="2011">
Murata
<emphasis id="B917EAE75C49FFAA6FB9FE19FDCFFE20" box="[503,545,467,488]" italics="true" pageId="1" pageNumber="8">et al</emphasis>
., 2011: 84
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6CDAFE19FCC1FE21" author="Kim, J. I." box="[660,815,467,489]" page="126" pageId="1" pageNumber="8" year="2012">
<bibRefCitation id="EFF24B045C49FFAA6CDAFE19FCC1FE21" author="Kim, J. I." box="[660,815,467,489]" pageId="1" pageNumber="8" pagination="1 - 209" refId="ref1837" refString="Kim, J. I. 2012. Insect Fauna of Korea. Arthropoda: Insecta: Coleoptera: Scarabaeoidea (Laparosticti). The Flora and Fauna of Korea, 12 (3): 1 - 209." type="journal article" year="2012">Kim, 2012: 126</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BDC36F55C49FFAA6EADFE24FCCDFDCC" blockId="1.[227,1128,384,709]" box="[227,803,494,516]" pageId="1" pageNumber="8">
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<taxonomicName id="4C634D765C49FFAA6EADFE24FE27FDCB" ID-CoL="4FQS9" baseAuthorityName="Balthasar" baseAuthorityYear="1945" box="[227,457,494,515]" class="Insecta" family="Scarabaeidae" genus="Phalacronothus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="botulus">
<emphasis id="B917EAE75C49FFAA6EADFE24FE27FDCB" box="[227,457,494,515]" italics="true" pageId="1" pageNumber="8">Phalacronothus botulus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AC210E45C49FFAA6F98FE24FCF1FDCC" author="Dellacasa, M. &amp; Dellacasa, G. &amp; Kral, D. &amp; Bezdek, A." box="[470,799,494,516]" page="145" pageId="1" pageNumber="8" year="2016">
<bibRefCitation id="EFF24B045C49FFAA6F98FE24FCF1FDCC" author="Dellacasa, M. &amp; Dellacasa, G. &amp; Kral, D. &amp; Bezdek, A." box="[470,799,494,516]" pageId="1" pageNumber="8" pagination="98 - 155" refId="ref1712" refString="Dellacasa, M., Dellacasa, G., Kral, D. &amp; Bezdek, A. 2016. Tribe Aphodiini Leach, 1815. P. 98 - 155. In: Lobl, I., Lobl, D. (Ed.). Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. Brill, Leiden, Boston." type="book chapter" year="2016">Dellacasa &amp; Dellacasa, 2016: 145</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36F55C49FFAA6EADFDC0FE8AFD9E" blockId="1.[227,1128,384,709]" pageId="1" pageNumber="8">
<treatmentCitationGroup id="AB7311DB5C49FFAA6EADFDC0FE8AFD9E" pageId="1" pageNumber="8">
<treatmentCitation id="0AC210E45C49FFAA6EADFDC0FD32FDD7" author="Nakane, T." box="[227,732,522,544]" page="120" pageId="1" pageNumber="8" year="1956">
<taxonomicName id="4C634D765C49FFAA6EADFDC0FD32FDD7" ID-CoL="FGM9" authority="Nakane, 1956: 120" authorityName="Nakane" authorityPageNumber="120" authorityYear="1956" box="[227,732,522,544]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="naraensis" subGenus="Orodalus">
<emphasis id="B917EAE75C49FFAA6EADFDC0FED0FDD7" box="[227,318,522,543]" italics="true" pageId="1" pageNumber="8">Aphodius</emphasis>
(
<emphasis id="B917EAE75C49FFAA6F00FDC0FE47FDD7" box="[334,425,522,543]" italics="true" pageId="1" pageNumber="8">Orodalus</emphasis>
)
<emphasis id="B917EAE75C49FFAA6FF6FDC1FDFBFDE8" box="[440,533,523,544]" italics="true" pageId="1" pageNumber="8">naraensis</emphasis>
<bibRefCitation id="EFF24B045C49FFAA6C50FDC0FD32FDD7" author="Nakane, T." box="[542,732,522,544]" pageId="1" pageNumber="8" pagination="119 - 124" refId="ref1945" refString="Nakane, T. 1956. Neue Aphodiinen aus Japan (Coleoptera, Scarabaeidae). Insecta matsumurana, 20: 119 - 124." type="journal article" year="1956">Nakane, 1956: 120</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D888575C49FFAA6CA9FDC0FCF6FDD7" box="[743,792,522,543]" pageId="1" pageNumber="8">Type</typeStatus>
locality “
<collectingRegion id="49A7F8175C49FFAA6D37FDC0FC42FDD7" box="[889,940,522,543]" country="Japan" name="Nara" pageId="1" pageNumber="8">Nara</collectingRegion>
, Yamato, Honshu” [=
<collectingCountry id="F37476655C49FFAA6F52FDECFEB9FDF3" box="[284,343,550,571]" name="Japan" pageId="1" pageNumber="8">Japan</collectingCountry>
: Honshu];
<treatmentCitation id="0AC210E45C49FFAA6FF3FDECFD6DFDF3" author="Balthasar, V." box="[445,643,550,572]" page="201" pageId="1" pageNumber="8" year="1964">
<bibRefCitation id="EFF24B045C49FFAA6FF3FDECFD6DFDF3" author="Balthasar, V." box="[445,643,550,572]" pageId="1" pageNumber="8" refId="ref1612" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 3. Aphodiidae. Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag. 652 pp." type="book" year="1964">Balthasar, 1964: 201</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6CC3FDECFCB9FDF3" author="Stebnicka, Z." box="[653,855,550,572]" page="239" pageId="1" pageNumber="8" year="1980">
<bibRefCitation id="EFF24B045C49FFAA6CC3FDECFCB9FDF3" author="Stebnicka, Z." box="[653,855,550,572]" pageId="1" pageNumber="8" pagination="191 - 298" refId="ref2255" refString="Stebnicka, Z. 1980. Scarabaeoidea (Coleoptera) of the Democratic People's Republic of Korea. Acta zoologica cracoviensia, 24 (5): 191 - 298." type="journal article" year="1980">Stebnicka, 1980: 239</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0AC210E45C49FFAA6A4BFDECFE8FFD9E" author="Stebnicka, Z." page="78" pageId="1" pageNumber="8" year="1982">
<bibRefCitation id="EFF24B045C49FFAA6A4BFDECFE8FFD9E" author="Stebnicka, Z." pageId="1" pageNumber="8" pagination="75 - 80" refId="ref2286" refString="Stebnicka, Z. 1982. New and little known species of Aphodius Illig. (Coleoptera, Scarabaeidae, Aphodiinae). Archives des Sciences de Geneve, 35 (1): 75 - 80." type="journal article" year="1982">Stebnicka, 1982: 78</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36F55C49FFAA6EADFD97FC24FD45" blockId="1.[227,1128,384,709]" pageId="1" pageNumber="8">
<treatmentCitationGroup id="AB7311DB5C49FFAA6EADFD97FC24FD45" pageId="1" pageNumber="8">
<treatmentCitation id="0AC210E45C49FFAA6EADFD97FCD2FDBA" author="Petrovitz, R. von" box="[227,828,605,627]" page="138" pageId="1" pageNumber="8" year="1958">
<taxonomicName id="4C634D765C49FFAA6EADFD97FCD2FDBA" ID-CoL="FFZK" authority="Petrovitz, 1958: 138" authorityName="Petrovitz" authorityPageNumber="138" authorityYear="1958" box="[227,828,605,627]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="manschuriensis" subGenus="Amidorus">
<emphasis id="B917EAE75C49FFAA6EADFD97FED0FDBA" box="[227,318,605,626]" italics="true" pageId="1" pageNumber="8">Aphodius</emphasis>
(
<emphasis id="B917EAE75C49FFAA6F1CFD97FE41FDBA" box="[338,431,605,626]" italics="true" pageId="1" pageNumber="8">Amidorus</emphasis>
)
<emphasis id="B917EAE75C49FFAA6F8DFD97FDB7FDBA" box="[451,601,605,626]" italics="true" pageId="1" pageNumber="8">manschuriensis</emphasis>
<bibRefCitation id="EFF24B045C49FFAA6C2AFD97FCD2FDBA" author="Petrovitz, R. von" box="[612,828,605,627]" pageId="1" pageNumber="8" pagination="131 - 139" refId="ref1970" refString="Petrovitz, R. von. 1958. Neue Asiatische Aphodiusarten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 9: 131 - 139" type="journal article" year="1958">Petrovitz, 1958: 138</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D888575C49FFAA6D05FD97FC93FDBA" box="[843,893,605,626]" pageId="1" pageNumber="8">Type</typeStatus>
locality ”Manschurei: Charbin” [=
<collectingCountry id="F37476655C49FFAA6F34FDB2FE57FD46" box="[378,441,632,654]" name="China" pageId="1" pageNumber="8">China</collectingCountry>
:
<collectingRegion id="49A7F8175C49FFAA6F8CFDB2FDABFD46" box="[450,581,632,654]" country="China" name="Heilongjiang" pageId="1" pageNumber="8">Heilongjiang</collectingRegion>
]. Synonymized by
<treatmentCitation id="0AC210E45C49FFAA6CB2FDB2FC28FD45" author="Balthasar, V." box="[764,966,632,654]" page="281" pageId="1" pageNumber="8" year="1964">
<bibRefCitation id="EFF24B045C49FFAA6CB2FDB2FC28FD45" author="Balthasar, V." box="[764,966,632,654]" pageId="1" pageNumber="8" refId="ref1612" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Band 3. Aphodiidae. Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag. 652 pp." type="book" year="1964">Balthasar, 1964: 281</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36F55C49FFAA6EADFD5EFD7CFD0D" blockId="1.[227,1128,384,709]" pageId="1" pageNumber="8">
<treatmentCitationGroup id="AB7311DB5C49FFAA6EADFD5EFD7CFD0D" pageId="1" pageNumber="8">
<taxonomicName id="4C634D765C49FFAA6EADFD5EFE4AFD61" ID-CoL="3V89F" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[227,420,660,681]" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="B917EAE75C49FFAA6EADFD5EFE4AFD61" box="[227,420,660,681]" italics="true" pageId="1" pageNumber="8">Liothorax plagiatus</emphasis>
</taxonomicName>
auct.:
<treatmentCitation id="0AC210E45C49FFAA6FA5FD5EFD5BFD61" author="Shabalin, S. A." box="[491,693,660,682]" page="188" pageId="1" pageNumber="8" year="2020">
<bibRefCitation id="EFF24B045C49FFAA6FA5FD5EFD5BFD61" author="Shabalin, S. A." box="[491,693,660,682]" pageId="1" pageNumber="8" pagination="185 - 198" refId="ref2002" refString="Shabalin, S. A. 2020. Assembly of dung beetles (Coleoptera: Scarabaeidae) - inhabitants of sheep dung in the south of Sikhote-Alin. A. I. Kurentsov's Annual Memorial Meetings, 31: 185 - 198. [In Russian]. DOI: 10.25221 / kurentzov. 31.16" type="journal article" year="2020">Shabalin, 2020: 188</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6C8CFD5EFC62FD62" author="Shabalin, S. A." box="[706,908,660,682]" page="625" pageId="1" pageNumber="8" year="2021">
<bibRefCitation id="EFF24B045C49FFAA6C8CFD5EFC62FD62" author="Shabalin, S. A." box="[706,908,660,682]" pageId="1" pageNumber="8" pagination="500 - 514" refId="ref2057" refString="Shabalin, S. A. 2021. Assembly of dung beetles (Coleoptera, Scarabaeoidea) inhabiting sheep droppings in the northern spurs of the Manchurian-Korean Mountains (Russian Far East). Contemporary Problems of Ecology, 14 (5): 500 - 514. DOI: 10.1134 / S 1995425521050140" type="journal article" year="2021">Shabalin, 2021: 625</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6DD7FD5EFB8DFD61" author="Shabalin, S. A." box="[921,1123,660,682]" page="477" pageId="1" pageNumber="8" year="2022">
<bibRefCitation id="EFF24B045C49FFAA6DD7FD5EFB8DFD61" author="Shabalin, S. A." box="[921,1123,660,682]" pageId="1" pageNumber="8" pagination="475 - 483" refId="ref2111" refString="Shabalin, S. A. 2022. Microstatial distribution of dung beetles (Coleoptera: Scarabaeoidea) in horse excrement in the southern Sikhote-Alina, Russia. Contemporary Problems of Ecology, 15 (5): 475 - 483. DOI: 10.1134 / S 1995425522050110" type="journal article" year="2022">Shabalin, 2022: 477</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6F47FD65FE2FFD0D" author="Shabalin, S. A." box="[265,449,687,709]" page="20" pageId="1" pageNumber="8" year="2023">
<bibRefCitation id="EFF24B045C49FFAA6F47FD65FE2FFD0D" author="Shabalin, S. A." box="[265,449,687,709]" pageId="1" pageNumber="8" pagination="17 - 30" refId="ref2159" refString="Shabalin, S. A. 2023. Estimation of biomass of dung beetles (Coleoptera: Scarabaeoidea) from the Russian Far East. Far Eastern Entomologist, 475: 17 - 30. DOI: 10.25221 / fee. 475.3" type="journal article" year="2023">Shabalin, 2023: 20</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210E45C49FFAA6F84FD65FD60FD0D" author="Shabalin, S. A." box="[458,654,687,709]" page="256" pageId="1" pageNumber="8" year="2024">
<bibRefCitation id="EFF24B045C49FFAA6F84FD65FD60FD0D" author="Shabalin, S. A." box="[458,654,687,709]" pageId="1" pageNumber="8" pagination="227 - 240" refId="ref2201" refString="Shabalin, S. A. 2024. Assembly of dung beetles (Coleoptera, Scarabaeoidea) inhabiting cow dung in the northern spurs of the Manchurian-Korean Mountains (Russian Far East). Contemporary Problems of Ecology, 17 (2): 227 - 240. DOI: 10.1134 / S 1995425524020100" type="journal article" year="2024">Shabalin, 2024: 256</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C379657E5C49FFAA6F47FD2DFE81FAED" pageId="1" pageNumber="8" type="materials_examined">
<paragraph id="8BDC36F55C49FFAA6F47FD2DFE81FAED" blockId="1.[227,1127,742,1593]" pageId="1" pageNumber="8">
<materialsCitation id="3B0B3CA85C49FFAA6F47FD2DFCF2FCD0" collectingDate="2018-05-29" collectionCode="V" collectorName="S. Shabalin" country="Russia" location="Ryazanovka River" longitude="131.38333" municipality="Primorskii" pageId="1" pageNumber="8" specimenCount="19">
MATERIAL.
<emphasis id="B917EAE75C49FFAA6FDCFD2CFE38FD34" bold="true" box="[402,470,742,764]" pageId="1" pageNumber="8">
<collectingCountry id="F37476655C49FFAA6FDCFD2CFE38FD34" box="[402,470,742,764]" name="Russia" pageId="1" pageNumber="8">Russia</collectingCountry>
</emphasis>
:
<collectingMunicipality id="6BB8AC8F5C49FFAA6FAAFD2DFDA4FD35" box="[484,586,743,765]" pageId="1" pageNumber="8">Primorskii</collectingMunicipality>
krai: valley of
<location id="8EBC602E5C49FFAA6C90FD2DFC62FD35" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6C90FD2DFC62FD35" box="[734,908,743,765]" country="Russia" longitude="131.38333" municipality="Primorskii" name="Ryazanovka River" pageId="1" pageNumber="8">Ryazanovka River</location>
(42°82' N,
<geoCoordinate id="EE5750325C49FFAA6DB2FD2DFBB2FD34" box="[1020,1116,743,764]" degrees="131" direction="east" minutes="23" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="131.38333">131°23' E</geoCoordinate>
),
<date id="FFDD10355C49FFAA6EADFCC9FEA3FCD0" box="[227,333,771,792]" pageId="1" pageNumber="8" value="2018-05-29">
<collectingDate id="EF99E9DD5C49FFAA6EADFCC9FEA3FCD0" box="[227,333,771,792]" pageId="1" pageNumber="8" value="2018-05-29">
29.
<collectionCode id="ED72AE305C49FFAA6EB1FCC9FEFCFCD0" box="[255,274,771,792]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2018
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6F17FCC9FE33FCD0" box="[345,477,770,792]" count="19" pageId="1" pageNumber="8" type="generic">19 specimens</specimenCount>
in sheep dung, leg.
<collectorName id="269653235C49FFAA6CE7FCC9FCF2FCD0" box="[681,796,770,792]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6D66FCC9FD53FCFC" collectingDate="2018-06-07" collectionCode="VI" collectorName="S. Shabalin" country="Russia" location="Ryazanovka River" longitude="131.38333" municipality="Primorskii" pageId="1" pageNumber="8" specimenCount="9">
same locality,
<date id="FFDD10355C49FFAA6DF6FCC9FBF0FCD0" box="[952,1054,771,792]" pageId="1" pageNumber="8" value="2018-06-07">
<collectingDate id="EF99E9DD5C49FFAA6DF6FCC9FBF0FCD0" box="[952,1054,771,792]" pageId="1" pageNumber="8" value="2018-06-07">
7.
<collectionCode id="ED72AE305C49FFAA6D86FCC9FC0AFCD0" box="[968,996,771,792]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="8">VI</collectionCode>
2018
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6A64FCC9FEC9FCFC" count="9" pageId="1" pageNumber="8" type="generic">9 specimens</specimenCount>
in sheep and cow dung, leg.
<collectorName id="269653235C49FFAA6C05FCD4FD53FCFC" box="[587,701,798,820]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6C87FCD4FDBDFC98" collectingDate="2018-06-15" collectionCode="VI" collectorName="S. Shabalin" country="Russia" location="Ryazanovka River" longitude="131.38333" municipality="Primorskii" pageId="1" pageNumber="8" specimenCount="5">
same locality,
<date id="FFDD10355C49FFAA6D16FCD4FC27FCFB" box="[856,969,798,820]" pageId="1" pageNumber="8" value="2018-06-15">
<collectingDate id="EF99E9DD5C49FFAA6D16FCD4FC27FCFB" box="[856,969,798,820]" pageId="1" pageNumber="8" value="2018-06-15">
15.
<collectionCode id="ED72AE305C49FFAA6D3CFCD4FC61FCFC" box="[882,911,798,820]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="8">VI</collectionCode>
2018
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D9AFCD4FBA2FCFC" box="[980,1100,798,820]" count="5" pageId="1" pageNumber="8" type="generic">5 specimens</specimenCount>
in sheep and cow dung, leg.
<collectorName id="269653235C49FFAA6FACFCF0FDBDFC98" box="[482,595,826,848]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6C10FCF0FE93FCA3" collectingDate="2018-07-06" collectionCode="VI" collectorName="S. Shabalin" country="Russia" location="Ryazanovka River" longitude="131.38333" municipality="Primorskii" pageId="1" pageNumber="8" specimenCount="1">
same locality,
<date id="FFDD10355C49FFAA6CA2FCF0FCB9FC87" box="[748,855,826,848]" pageId="1" pageNumber="8" value="2018-07-06">
<collectingDate id="EF99E9DD5C49FFAA6CA2FCF0FCB9FC87" box="[748,855,826,848]" pageId="1" pageNumber="8" value="2018-07-06">6.VII 2018</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D2AFCF0FC3CFC98" box="[868,978,826,848]" count="1" pageId="1" pageNumber="8" type="generic">1 specimen</specimenCount>
in sheep dung, leg.
<collectorName id="269653235C49FFAA6F42FC9FFE93FCA3" box="[268,381,853,875]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6FC8FC9FFEBDFC4F" collectingDate="2020-06-08" collectionCode="VI" collectorName="S. Shabalin" country="Russia" latitude="43.533333" location="Safari-Park" longLatPrecision="1143" longitude="132.66667" pageId="1" pageNumber="8" specimenCount="3">
<location id="8EBC602E5C49FFAA6FC8FC9FFE1DFCA3" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6FC8FC9FFE1DFCA3" box="[390,499,853,875]" country="Russia" latitude="43.533333" longLatPrecision="1143" longitude="132.66667" name="Safari-Park" pageId="1" pageNumber="8">Safari-Park</location>
(
<geoCoordinate id="EE5750325C49FFAA6C4FFC9FFDB7FCA3" box="[513,601,853,875]" degrees="43" direction="north" minutes="32" orientation="latitude" pageId="1" pageNumber="8" precision="925" value="43.533333">43°32' N</geoCoordinate>
,
<geoCoordinate id="EE5750325C49FFAA6C2DFC9FFD2AFCA2" box="[611,708,853,874]" degrees="132" direction="east" minutes="40" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="132.66667">132°40' E</geoCoordinate>
),
<date id="FFDD10355C49FFAA6C98FC9FFCD7FCA2" box="[726,825,853,874]" pageId="1" pageNumber="8" value="2020-06-08">
<collectingDate id="EF99E9DD5C49FFAA6C98FC9FFCD7FCA2" box="[726,825,853,874]" pageId="1" pageNumber="8" value="2020-06-08">
8.
<collectionCode id="ED72AE305C49FFAA6CAAFC9FFCEFFCA2" box="[740,769,853,874]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="8">VI</collectionCode>
2020
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D0DFC9FFC57FCA3" box="[835,953,853,875]" count="3" pageId="1" pageNumber="8" type="generic">3 specimens</specimenCount>
in deer dung, leg.
<collectorName id="269653235C49FFAA6EADFCBBFEBDFC4F" box="[227,339,881,903]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6F13FCBAFE93FC6A" collectingDate="2019-05-15" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="43.383335" location="Romanovka" longLatPrecision="1143" longitude="132.86667" pageId="1" pageNumber="8" specimenCount="6">
<quantity id="4C9B9B105C49FFAA6F13FCBAFE60FC4F" box="[349,398,880,903]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="8" unit="km" value="5.0">5 km</quantity>
E
<location id="8EBC602E5C49FFAA6FE7FCBBFDF3FC4F" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6FE7FCBBFDF3FC4F" box="[425,541,881,903]" country="Russia" latitude="43.383335" longLatPrecision="1143" longitude="132.86667" name="Romanovka" pageId="1" pageNumber="8">Romanovka</location>
(
<geoCoordinate id="EE5750325C49FFAA6C65FCBBFD6DFC4E" box="[555,643,881,902]" degrees="43" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="8" precision="925" value="43.383335">43°23' N</geoCoordinate>
,
<geoCoordinate id="EE5750325C49FFAA6CC0FCBAFD00FC4E" box="[654,750,880,902]" degrees="132" direction="east" minutes="52" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="132.86667">132°52' E</geoCoordinate>
)
<date id="FFDD10355C49FFAA6CB5FCBAFC8DFC4E" box="[763,867,880,902]" pageId="1" pageNumber="8" value="2019-05-15">
<collectingDate id="EF99E9DD5C49FFAA6CB5FCBAFC8DFC4E" box="[763,867,880,902]" pageId="1" pageNumber="8" value="2019-05-15">
15.
<collectionCode id="ED72AE305C49FFAA6D58FCBAFCC4FC4E" box="[790,810,880,902]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D23FCBBFC0DFC4F" box="[877,995,881,903]" count="6" pageId="1" pageNumber="8" type="generic">6 specimens</specimenCount>
in cow dung, leg.
<collectorName id="269653235C49FFAA6F42FC46FE93FC6A" box="[268,381,908,930]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6FC8FC46FE90FC76" collectingDate="2019-05-15" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="43.0" location="Peretino" longLatPrecision="1146" longitude="133.23334" pageId="1" pageNumber="8" specimenCount="1">
vicinity
<location id="8EBC602E5C49FFAA6F98FC46FDC8FC6A" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6F98FC46FDC8FC6A" box="[470,550,908,930]" country="Russia" latitude="43.0" longLatPrecision="1146" longitude="133.23334" name="Peretino" pageId="1" pageNumber="8">Peretino</location>
(
<geoCoordinate id="EE5750325C49FFAA6C7DFC46FD65FC6A" box="[563,651,908,930]" degrees="43" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="8" precision="925" value="43.0">43°00' N</geoCoordinate>
,
<geoCoordinate id="EE5750325C49FFAA6CDBFC46FD18FC69" box="[661,758,908,929]" degrees="133" direction="east" minutes="14" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="133.23334">133°14' E</geoCoordinate>
),
<date id="FFDD10355C49FFAA6D46FC46FC81FC69" box="[776,879,908,930]" pageId="1" pageNumber="8" value="2019-05-15">
<collectingDate id="EF99E9DD5C49FFAA6D46FC46FC81FC69" box="[776,879,908,930]" pageId="1" pageNumber="8" value="2019-05-15">
15.
<collectionCode id="ED72AE305C49FFAA6D6DFC46FCD9FC6A" box="[803,823,908,930]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D37FC46FC08FC6A" box="[889,998,908,930]" count="1" pageId="1" pageNumber="8" type="generic">1 specimen</specimenCount>
in cow dung, leg.
<collectorName id="269653235C49FFAA6F40FC62FE90FC76" box="[270,382,936,958]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6FC4FC62FE17FC11" collectingDate="2019-05-15" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="43.1" location="Partizansk" longLatPrecision="1145" longitude="133.3" pageId="1" pageNumber="8" specimenCount="19">
<quantity id="4C9B9B105C49FFAA6FC4FC62FE26FC76" box="[394,456,936,958]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="8" unit="km" value="16.0">16 km</quantity>
SSE
<location id="8EBC602E5C49FFAA6C4FFC62FD8CFC76" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6C4FFC62FD8CFC76" box="[513,610,936,958]" country="Russia" latitude="43.1" longLatPrecision="1145" longitude="133.3" name="Partizansk" pageId="1" pageNumber="8">Partizansk</location>
(
<geoCoordinate id="EE5750325C49FFAA6C3CFC62FD23FC75" box="[626,717,936,958]" degrees="43" direction="north" minutes="06" orientation="latitude" pageId="1" pageNumber="8" precision="925" value="43.1">43°06' N</geoCoordinate>
,
<geoCoordinate id="EE5750325C49FFAA6C94FC62FCD0FC75" box="[730,830,936,957]" degrees="133" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="133.3">133°18' E</geoCoordinate>
),
<date id="FFDD10355C49FFAA6D1CFC62FC52FC75" box="[850,956,936,958]" pageId="1" pageNumber="8" value="2019-05-15">
<collectingDate id="EF99E9DD5C49FFAA6D1CFC62FC52FC75" box="[850,956,936,958]" pageId="1" pageNumber="8" value="2019-05-15">
15.
<collectionCode id="ED72AE305C49FFAA6D23FC62FC6FFC76" box="[877,897,936,958]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D87FC62FBA2FC76" box="[969,1100,936,958]" count="19" pageId="1" pageNumber="8" type="generic">19 specimens</specimenCount>
in sheep dung, leg.
<collectorName id="269653235C49FFAA6FC7FC0EFE17FC11" box="[393,505,963,985]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6C4BFC0EFED4FC3D" collectingDate="2019-05-24" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="43.1" location="Partizansk" longLatPrecision="1145" longitude="133.3" pageId="1" pageNumber="8" specimenCount="11">
same locality,
<date id="FFDD10355C49FFAA6CDCFC0EFD14FC11" box="[658,762,964,985]" pageId="1" pageNumber="8" value="2019-05-24">
<collectingDate id="EF99E9DD5C49FFAA6CDCFC0EFD14FC11" box="[658,762,964,985]" pageId="1" pageNumber="8" value="2019-05-24">
24.
<collectionCode id="ED72AE305C49FFAA6CE1FC0EFD2FFC11" box="[687,705,964,985]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D48FC0EFC67FC11" box="[774,905,963,985]" count="11" pageId="1" pageNumber="8" type="generic">11 specimens</specimenCount>
in sheep dung, leg.
<collectorName id="269653235C49FFAA6A1DFC0EFED4FC3D" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6F0AFC15FC37FC3D" box="[324,985,991,1013]" collectingDate="2019-06-03" collectionCode="VI" collectorName="S. Shabalin" country="Russia" latitude="43.1" location="Partizansk" longLatPrecision="1145" longitude="133.3" pageId="1" pageNumber="8" specimenCount="8">
same locality,
<date id="FFDD10355C49FFAA6F9EFC15FDDFFC3C" box="[464,561,991,1012]" pageId="1" pageNumber="8" value="2019-06-03">
<collectingDate id="EF99E9DD5C49FFAA6F9EFC15FDDFFC3C" box="[464,561,991,1012]" pageId="1" pageNumber="8" value="2019-06-03">
3.
<collectionCode id="ED72AE305C49FFAA6F90FC15FE14FC3C" box="[478,506,991,1012]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="8">VI</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6C75FC15FD41FC3D" box="[571,687,991,1013]" count="8" pageId="1" pageNumber="8" type="generic">8 specimens</specimenCount>
in sheep dung, leg.
<collectorName id="269653235C49FFAA6D25FC15FC37FC3D" box="[875,985,991,1013]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6DACFC15FCE0FBD9" collectingDate="2019-06-12" collectionCode="VI" collectorName="S. Shabalin" country="Russia" latitude="43.1" location="Partizansk" longLatPrecision="1145" longitude="133.3" pageId="1" pageNumber="8" specimenCount="2">
same locality,
<date id="FFDD10355C49FFAA6EADFC31FEBDFBD8" box="[227,339,1019,1040]" pageId="1" pageNumber="8" value="2019-06-12">
<collectingDate id="EF99E9DD5C49FFAA6EADFC31FEBDFBD8" box="[227,339,1019,1040]" pageId="1" pageNumber="8" value="2019-06-12">
12.
<collectionCode id="ED72AE305C49FFAA6EB0FC31FEF4FBD8" box="[254,282,1019,1040]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="8">VI</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6F11FC31FE38FBD9" box="[351,470,1019,1041]" count="2" pageId="1" pageNumber="8" type="generic">2 specimens</specimenCount>
in sheep dung, leg.
<collectorName id="269653235C49FFAA6CD3FC31FCE0FBD9" box="[669,782,1019,1041]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6D57FC31FBC8FBE4" collectingDate="2019-05-15" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="43.433334" location="Partizansk" longLatPrecision="1143" longitude="133.28334" pageId="1" pageNumber="8" specimenCount="202">
<quantity id="4C9B9B105C49FFAA6D57FC31FCB9FBD9" box="[793,855,1019,1041]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="8" unit="km" value="15.0">15 km</quantity>
NNE
<location id="8EBC602E5C49FFAA6DD9FC31FC15FBD9" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6DD9FC31FC15FBD9" box="[919,1019,1019,1041]" country="Russia" latitude="43.433334" longLatPrecision="1143" longitude="133.28334" name="Partizansk" pageId="1" pageNumber="8">Partizansk</location>
(
<geoCoordinate id="EE5750325C49FFAA6A47FC31FB8DFBD8" box="[1033,1123,1019,1041]" degrees="43" direction="north" minutes="26" orientation="latitude" pageId="1" pageNumber="8" precision="925" value="43.433334">43°26' N</geoCoordinate>
,
<geoCoordinate id="EE5750325C49FFAA6EADFBDCFEA4FBE3" box="[227,330,1046,1067]" degrees="133" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="133.28334">133°17' E</geoCoordinate>
),
<date id="FFDD10355C49FFAA6F2FFBDCFE20FBE3" box="[353,462,1046,1068]" pageId="1" pageNumber="8" value="2019-05-15">
<collectingDate id="EF99E9DD5C49FFAA6F2FFBDCFE20FBE3" box="[353,462,1046,1068]" pageId="1" pageNumber="8" value="2019-05-15">
15.
<collectionCode id="ED72AE305C49FFAA6F33FBDCFE7FFBE4" box="[381,401,1046,1068]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6F90FBDCFD9CFBE4" box="[478,626,1046,1068]" count="202" pageId="1" pageNumber="8" type="generic">202 specimens</specimenCount>
in horse and cow dung, leg.
<collectorName id="269653235C49FFAA6DE1FBDCFBC8FBE4" box="[943,1062,1046,1068]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6A7BFBDDFC48FB80" collectingDate="2019-05-24" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="43.433334" location="Partizansk" longLatPrecision="1143" longitude="133.28334" pageId="1" pageNumber="8" specimenCount="8">
same locality,
<date id="FFDD10355C49FFAA6F79FBF8FE71FB8F" box="[311,415,1074,1095]" pageId="1" pageNumber="8" value="2019-05-24">
<collectingDate id="EF99E9DD5C49FFAA6F79FBF8FE71FB8F" box="[311,415,1074,1095]" pageId="1" pageNumber="8" value="2019-05-24">
24.
<collectionCode id="ED72AE305C49FFAA6F1AFBF8FE89FB8F" box="[340,359,1074,1095]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6FE7FBF8FDF1FB80" box="[425,543,1074,1096]" count="8" pageId="1" pageNumber="8" type="generic">8 specimens</specimenCount>
in horse and cow dung, leg.
<collectorName id="269653235C49FFAA6D7BFBF8FC48FB80" box="[821,934,1074,1096]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6DFEFBF8FCA5FBAB" collectingDate="2019-06-03" collectionCode="VI" collectorName="S. Shabalin" country="Russia" latitude="43.433334" location="Partizansk" longLatPrecision="1143" longitude="133.28334" pageId="1" pageNumber="8" specimenCount="7">
same locality,
<date id="FFDD10355C49FFAA6A75FBF8FEFBFBAB" pageId="1" pageNumber="8" value="2019-06-03">
<collectingDate id="EF99E9DD5C49FFAA6A75FBF8FEFBFBAB" pageId="1" pageNumber="8" value="2019-06-03">
3.
<collectionCode id="ED72AE305C49FFAA6A04FBF8FB89FB8F" box="[1098,1127,1074,1095]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="8">VI</collectionCode>
2019
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6F6AFB84FE71FBAB" box="[292,415,1101,1123]" count="7" pageId="1" pageNumber="8" type="generic">7 specimens</specimenCount>
in horse and cow dung, leg.
<collectorName id="269653235C49FFAA6C98FB84FCA5FBAB" box="[726,843,1101,1123]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6D14FB87FC8BFBB7" collectingDate="2020-05-15" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="44.05" location="Novogordeevka" longLatPrecision="1139" longitude="133.23334" pageId="1" pageNumber="8" specimenCount="1">
<location id="8EBC602E5C49FFAA6D14FB87FC1DFBAB" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6D14FB87FC1DFBAB" box="[858,1011,1101,1123]" country="Russia" latitude="44.05" longLatPrecision="1139" longitude="133.23334" name="Novogordeevka" pageId="1" pageNumber="8">Novogordeevka</location>
(
<geoCoordinate id="EE5750325C49FFAA6A4BFB84FB8DFBAB" box="[1029,1123,1102,1123]" degrees="44" direction="north" minutes="03" orientation="latitude" pageId="1" pageNumber="8" precision="925" value="44.05">44°03' N</geoCoordinate>
,
<geoCoordinate id="EE5750325C49FFAA6EADFBA3FEAAFBB6" box="[227,324,1129,1150]" degrees="133" direction="east" minutes="14" orientation="longitude" pageId="1" pageNumber="8" precision="925" value="133.23334">133°14' E</geoCoordinate>
),
<date id="FFDD10355C49FFAA6F18FBA3FE50FBB6" box="[342,446,1129,1151]" pageId="1" pageNumber="8" value="2020-05-15">
<collectingDate id="EF99E9DD5C49FFAA6F18FBA3FE50FBB6" box="[342,446,1129,1151]" pageId="1" pageNumber="8" value="2020-05-15">
15.
<collectionCode id="ED72AE305C49FFAA6F3CFBA3FE68FBB7" box="[370,390,1129,1151]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2020
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6F86FBA3FDDBFBB7" box="[456,565,1129,1151]" count="1" pageId="1" pageNumber="8" type="generic">1 specimen</specimenCount>
in horse dung, leg.
<collectorName id="269653235C49FFAA6CBBFBA3FC8BFBB7" box="[757,869,1129,1151]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6D21FBA0FD66FB53" collectingDate="2020-05-23" collectionCode="V" collectorName="S. Shabalin" country="Russia" latitude="44.05" location="Novogordeevka" longLatPrecision="1139" longitude="133.23334" pageId="1" pageNumber="8" specimenCount="1">
same locality,
<date id="FFDD10355C49FFAA6DB5FBA3FB8CFBB6" box="[1019,1122,1129,1150]" pageId="1" pageNumber="8" value="2020-05-23">
<collectingDate id="EF99E9DD5C49FFAA6DB5FBA3FB8CFBB6" box="[1019,1122,1129,1150]" pageId="1" pageNumber="8" value="2020-05-23">
23.
<collectionCode id="ED72AE305C49FFAA6A59FBA3FBC4FBB6" box="[1047,1066,1129,1150]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2020
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6EADFB4FFEBFFB53" box="[227,337,1157,1179]" count="1" pageId="1" pageNumber="8" type="generic">1 specimen</specimenCount>
in horse dung, leg.
<collectorName id="269653235C49FFAA6C58FB4FFD66FB53" box="[534,648,1157,1179]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6CDDFB4FFDF9FB7E" collectingDate="2020-05-15" collectionCode="V" collectorName="S. Shabalin" country="Russia" location="Kornilovka" pageId="1" pageNumber="8" specimenCount="2">
vicinity of
<location id="8EBC602E5C49FFAA6D4EFB4FFC9EFB53" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6D4EFB4FFC9EFB53" box="[768,880,1157,1179]" country="Russia" name="Kornilovka" pageId="1" pageNumber="8">Kornilovka</location>
,
<date id="FFDD10355C49FFAA6D35FB4EFC0AFB52" box="[891,996,1156,1178]" pageId="1" pageNumber="8" value="2020-05-15">
<collectingDate id="EF99E9DD5C49FFAA6D35FB4EFC0AFB52" box="[891,996,1156,1178]" pageId="1" pageNumber="8" value="2020-05-15">
15.
<collectionCode id="ED72AE305C49FFAA6DD8FB4EFC44FB52" box="[918,938,1156,1178]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2020
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6DA1FB4FFB88FB53" box="[1007,1126,1157,1179]" count="2" pageId="1" pageNumber="8" type="generic">2 specimens</specimenCount>
in cow dung, leg.
<collectorName id="269653235C49FFAA6FECFB6AFDF9FB7E" box="[418,535,1184,1206]" pageId="1" pageNumber="8">S. Shabalin</collectorName>
</materialsCitation>
;
<materialsCitation id="3B0B3CA85C49FFAA6C6BFB6BFED0FB1A" collectingDate="2020-05-23" collectionCode="V" collectorName="S. Shabalin." country="Russia" location="Kornilovka" pageId="1" pageNumber="8" specimenCount="1">
same locality,
<date id="FFDD10355C49FFAA6CF5FB6AFCC8FB7D" box="[699,806,1184,1205]" pageId="1" pageNumber="8" value="2020-05-23">
<collectingDate id="EF99E9DD5C49FFAA6CF5FB6AFCC8FB7D" box="[699,806,1184,1205]" pageId="1" pageNumber="8" value="2020-05-23">
23.
<collectionCode id="ED72AE305C49FFAA6C99FB6AFD04FB7D" box="[727,746,1184,1205]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="8" type="Museum">V</collectionCode>
2020
</collectingDate>
</date>
,
<specimenCount id="9D65FD7C5C49FFAA6D7BFB6AFC49FB7E" box="[821,935,1184,1206]" count="1" pageId="1" pageNumber="8" type="generic">1 specimen</specimenCount>
cow dung, leg.
<collectorName id="269653235C49FFAA6A1DFB6AFED0FB1A" pageId="1" pageNumber="8">S. Shabalin.</collectorName>
</materialsCitation>
<materialsCitation id="3B0B3CA85C49FFAA6F05FB76FE82FAED" collectorName="Zoological Institute &amp; Federal Scientific Center &amp; Vladivostok" country="Russia" location="East Asia Terrestrial Biodiversity" municipality="Ryazanovka River" pageId="1" pageNumber="8" specimenCount="4" stateProvince="Saint Petersburg">
<specimenCount id="9D65FD7C5C49FFAA6F05FB76FE04FB1A" box="[331,490,1212,1234]" count="4" pageId="1" pageNumber="8" type="generic">Four specimens</specimenCount>
from the valley of
<collectingMunicipality id="6BB8AC8F5C49FFAA6C85FB76FC6BFB1A" box="[715,901,1212,1234]" pageId="1" pageNumber="8">Ryazanovka River</collectingMunicipality>
are deposited in the collection of the
<collectorName id="269653235C49FFAA6FC9FB1DFDAAFB25" box="[391,580,1239,1261]" pageId="1" pageNumber="8">Zoological Institute</collectorName>
(
<collectingRegion id="49A7F8175C49FFAA6C1FFB1DFD1BFB25" box="[593,757,1239,1261]" country="Russia" name="Saint Petersburg" pageId="1" pageNumber="8">Saint Petersburg</collectingRegion>
), but the rest material deposited in the collection of the
<collectorName id="269653235C49FFAA6FEDFB39FD48FAC0" box="[419,678,1267,1289]" pageId="1" pageNumber="8">Federal Scientific Center</collectorName>
of the
<location id="8EBC602E5C49FFAA6D49FB39FB89FAC1" LSID="urn:lsid:plazi:treatment:03CA87E35C49FFA96EADFEF4FBA0FEF0:8EBC602E5C49FFAA6D49FB39FB89FAC1" box="[775,1127,1267,1289]" country="Russia" municipality="Ryazanovka River" name="East Asia Terrestrial Biodiversity" pageId="1" pageNumber="8" stateProvince="Saint Petersburg">East Asia Terrestrial Biodiversity</location>
(
<collectorName id="269653235C49FFAA6EA7FAC5FE8AFAED" box="[233,356,1295,1317]" pageId="1" pageNumber="8">Vladivostok</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C379657E5C49FFA96F47FAE0FBA0FEF0" lastPageId="2" lastPageNumber="9" pageId="1" pageNumber="8" type="distribution">
<paragraph id="8BDC36F55C49FFAA6F47FAE0FDDCFA94" blockId="1.[227,1127,742,1593]" pageId="1" pageNumber="8">
DISTRIBUTION.
<collectingCountry id="F37476655C49FFAA6FF3FAE0FE13FA88" box="[445,509,1322,1344]" name="Russia" pageId="1" pageNumber="8">Russia</collectingCountry>
(first record, Primorskii krai),
<collectingCountry id="F37476655C49FFAA6D68FAE1FCB2FA88" box="[806,860,1323,1344]" name="Japan" pageId="1" pageNumber="8">Japan</collectingCountry>
(Honshu, Kyushu),
<collectingCountry id="F37476655C49FFAA6A6EFAE0FECEFA93" name="North Korea" pageId="1" pageNumber="8">North Korea</collectingCountry>
,
<collectingCountry id="F37476655C49FFAA6F64FA8CFE8DFA94" box="[298,355,1350,1372]" name="China" pageId="1" pageNumber="8">China</collectingCountry>
(
<collectingRegion id="49A7F8175C49FFAA6F21FA8CFE1EFA94" box="[367,496,1350,1372]" country="China" name="Heilongjiang" pageId="1" pageNumber="8">Heilongjiang</collectingRegion>
,
<collectingRegion id="49A7F8175C49FFAA6FB4FA8CFDC9FA94" box="[506,551,1350,1372]" country="China" name="Jilin" pageId="1" pageNumber="8">Jilin</collectingRegion>
).
</paragraph>
<paragraph id="8BDC36F55C49FFA96F47FAA8FBA0FEF0" blockId="1.[227,1127,742,1593]" lastBlockId="2.[227,1124,263,313]" lastPageId="2" lastPageNumber="9" pageId="1" pageNumber="8">
NOTES. Adults of
<taxonomicName id="4C634D765C49FFAA6F82FAABFD99FABE" authorityName="Balthasar" authorityYear="1945" box="[460,631,1377,1398]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="botulus">
<emphasis id="B917EAE75C49FFAA6F82FAABFD99FABE" box="[460,631,1377,1398]" italics="true" pageId="1" pageNumber="8">Aphodius botulus</emphasis>
</taxonomicName>
can be separated from other species of subgenus
<taxonomicName id="4C634D765C49FFAA6EADFAB7FE92FA5A" box="[227,380,1405,1426]" pageId="1" pageNumber="8" rank="subGenus" subGenus="Phalacronothus">
<emphasis id="B917EAE75C49FFAA6EADFAB7FE92FA5A" box="[227,380,1405,1426]" italics="true" pageId="1" pageNumber="8">Phalacronothus</emphasis>
</taxonomicName>
recorded from
<collectingCountry id="F37476655C49FFAA6C51FAB7FD8CFA5B" box="[543,610,1405,1427]" name="Russia" pageId="1" pageNumber="8">Russia</collectingCountry>
, namely
<taxonomicName id="4C634D765C49FFAA6C8EFAB7FC38FA5A" authority="Germar, 1824" authorityName="Germar" authorityYear="1824" box="[704,982,1405,1426]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="biguttatus">
<emphasis id="B917EAE75C49FFAA6C8EFAB7FCD1FA5A" box="[704,831,1405,1426]" italics="true" pageId="1" pageNumber="8">A. biguttatus</emphasis>
Germar, 1824
</taxonomicName>
,
<taxonomicName id="4C634D765C49FFAA6DABFAB7FDEDFA66" authority="Semenov et Medvedev, 1928" authorityName="Semenov et Medvedev" authorityYear="1928" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="citellorum">
<emphasis id="B917EAE75C49FFAA6DABFAB7FB89FA5A" box="[997,1127,1405,1426]" italics="true" pageId="1" pageNumber="8">A. citellorum</emphasis>
Semenov et Medvedev, 1928
</taxonomicName>
, and
<taxonomicName id="4C634D765C49FFAA6C77FA53FD17FA66" baseAuthorityName="Linnaeus" baseAuthorityYear="1760" box="[569,761,1433,1454]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="B917EAE75C49FFAA6C77FA53FD17FA66" box="[569,761,1433,1454]" italics="true" pageId="1" pageNumber="8">A. quadrimaculatus</emphasis>
</taxonomicName>
, by denser and coarser punctation of head, pronotum and elytra, and by the shape of the parameres. In Primorskii krai,
<taxonomicName id="4C634D765C49FFAA6A4AFA7FFB89FA01" authorityName="Balthasar" authorityYear="1945" box="[1028,1127,1460,1482]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="botulus">
<emphasis id="B917EAE75C49FFAA6A4AFA7FFB89FA01" box="[1028,1127,1460,1482]" italics="true" pageId="1" pageNumber="8">A. botulus</emphasis>
</taxonomicName>
occurs along with the more abundant
<taxonomicName id="4C634D765C49FFAA6C1FFA1AFD56FA2D" baseAuthorityName="Herbst" baseAuthorityYear="1789" box="[593,696,1488,1509]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B917EAE75C49FFAA6C1FFA1AFD56FA2D" box="[593,696,1488,1509]" italics="true" pageId="1" pageNumber="8">A. pusillus</emphasis>
</taxonomicName>
, and can be confused with it. It differs from the latter, apart from the characters mentioned above, also in having a hooked protibial spur in males and in a smaller average size. In
<collectingCountry id="F37476655C49FFAA6CCFF9CDFD13F9D4" box="[641,765,1543,1565]" name="North Korea" pageId="1" pageNumber="8">North Korea</collectingCountry>
, the beetles were found in sheep and cow dung (
<bibRefCitation id="EFF24B045C49FFAA6F1EF9E9FE1AF9F0" author="Stebnicka, Z." box="[336,500,1571,1593]" pageId="1" pageNumber="8" pagination="191 - 298" refId="ref2255" refString="Stebnicka, Z. 1980. Scarabaeoidea (Coleoptera) of the Democratic People's Republic of Korea. Acta zoologica cracoviensia, 24 (5): 191 - 298." type="journal article" year="1980">Stebnicka, 1980</bibRefCitation>
). In addition to the dung of abovementioned animals, in
<collectingCountry id="F37476655C49FFAA6A54F9E9FBB4F9F1" box="[1050,1114,1571,1593]" name="Russia" pageId="1" pageNumber="8">Russia</collectingCountry>
the beetles were also found in horse and deer dung. Our data suggest that
<taxonomicName id="4C634D765C4AFFA96D9BFECDFBD4FED4" authorityName="Balthasar" authorityYear="1945" box="[981,1082,263,284]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="9" phylum="Arthropoda" rank="species" species="botulus">
<emphasis id="B917EAE75C4AFFA96D9BFECDFBD4FED4" box="[981,1082,263,284]" italics="true" pageId="2" pageNumber="9">A. botulus</emphasis>
</taxonomicName>
is a generalist coprophagous species with imago active mostly in late spring and early summer.
</paragraph>
</subSubSection>
</treatment>
</document>