240 lines
31 KiB
XML
240 lines
31 KiB
XML
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<mods:title id="97581FCFA6C7002014A5D466ECA24173">Four new species of Australian velvet ants (Hymenoptera: Mutillidae, Aglaotilla) reared from bee and wasp nests, with a review of Australian mutillid host records</mods:title>
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<mods:namePart id="E55CB2D2E8FC17B17E922D67D5508E16">Taylor, Christopher K.</mods:namePart>
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<mods:namePart id="E3EAD782A8D8D9EDB5F97AE04B092778">Murphy, Mark V.</mods:namePart>
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<mods:namePart id="B83C457DFE31954A85019C694660C8D0">Hitchen, Yvette</mods:namePart>
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<mods:namePart id="4EAAE89569E7AF77A2B9BF2A702B076E">Brothers, Denis J.</mods:namePart>
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<mods:title id="089929794EDAC3D115A14E97F01DA650">Zootaxa</mods:title>
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<mods:date id="A850ADED4194896804A7250566F1C116">2019</mods:date>
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<subSubSection id="C37965054B4AC25716BAFA12FE0439EF" box="[151,461,1424,1451]" pageId="2" pageNumber="203" type="nomenclature">
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<paragraph id="8BDC368E4B4AC25716BAFA12FE0439EF" blockId="2.[151,461,1424,1451]" box="[151,461,1424,1451]" pageId="2" pageNumber="203">
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<heading id="D09481E24B4AC25716BAFA12FE0439EF" bold="true" box="[151,461,1424,1451]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
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<taxonomicName id="4C634D0D4B4AC25716BAFA12FE0439EF" authority="Brothers, 2018" authorityName="Brothers" authorityYear="2018" box="[151,461,1424,1451]" class="Insecta" family="Mutillidae" genus="Aglaotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA9C4B4AC25716BAFA12FE0439EF" bold="true" box="[151,461,1424,1451]" pageId="2" pageNumber="203">
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<emphasis id="B917EA9C4B4AC25716BAFA12FEC739EE" bold="true" box="[151,270,1424,1451]" italics="true" pageId="2" pageNumber="203">Aglaotilla</emphasis>
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Brothers, 2018
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C37965054B4AC25716BAFA57FDCE39A8" box="[151,519,1492,1517]" pageId="2" pageNumber="203" type="reference_group">
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<paragraph id="8BDC368E4B4AC25716BAFA57FDCE39A8" blockId="2.[151,519,1492,1517]" box="[151,519,1492,1517]" pageId="2" pageNumber="203">
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<taxonomicName id="4C634D0D4B4AC25716BAFA57FE0639A8" authority="Brothers, 2018: 358" authorityName="Brothers" authorityPageNumber="358" authorityYear="2018" box="[151,463,1492,1517]" class="Insecta" family="Mutillidae" genus="Aglaotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA9C4B4AC25716BAFA57FF3539A8" box="[151,252,1493,1517]" italics="true" pageId="2" pageNumber="203">Aglaotilla</emphasis>
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Brothers, 2018: 358
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</taxonomicName>
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–361.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37965054B4AC25716BAF990FCCE3A6F" box="[151,775,1553,1578]" pageId="2" pageNumber="203" type="materials_examined">
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<paragraph id="8BDC368E4B4AC25716BAF990FCCE3A6F" blockId="2.[151,775,1553,1578]" box="[151,775,1553,1578]" pageId="2" pageNumber="203">
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<emphasis id="B917EA9C4B4AC25716BAF990FEEA3A6F" bold="true" box="[151,291,1553,1578]" pageId="2" pageNumber="203">
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<typeStatus id="54D8882C4B4AC25716BAF990FF053A6F" box="[151,204,1554,1578]" pageId="2" pageNumber="203">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C634D0D4B4AC2571707F990FCCD3A6F" authority="Turner, 1916 a" authorityName="Turner" authorityYear="1916" box="[298,772,1553,1578]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="submetallescens">
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<emphasis id="B917EA9C4B4AC2571707F990FDA43A6F" box="[298,621,1554,1578]" italics="true" pageId="2" pageNumber="203">Ephutomorpha submetallescens</emphasis>
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Turner, 1916a
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37965054B4AC25616BAF9D5FD4B3F7E" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203" type="discussion">
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<paragraph id="8BDC368E4B4AC25716BAF9D5FA903B63" blockId="2.[151,1437,1623,2011]" pageId="2" pageNumber="203">
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<emphasis id="B917EA9C4B4AC25716BAF9D5FED63A37" bold="true" box="[151,287,1623,1650]" pageId="2" pageNumber="203">Comments.</emphasis>
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The taxonomic framework for Australian
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<taxonomicName id="4C634D0D4B4AC2571528F9DAFCB33A37" authorityName="Latreille" authorityYear="1802" box="[773,890,1624,1650]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="family">Mutillidae</taxonomicName>
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remains rudimentary, particularly regarding generic divisions. Keys to both males and females were provided by
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<bibRefCitation id="EFF24B7F4B4AC257155BF9FEFBCC3AD3" author="Andre, E." box="[886,1029,1659,1686]" pageId="2" pageNumber="203" pagination="475 - 517" refId="ref12746" refString="Andre, E. (1895) Mutillides d'Australie nouvelles ou imparfaitement connues. Memoires de la Societ Zoologique de France, 8, 475 - 517." type="journal article" year="1895">André (1895</bibRefCitation>
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,
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<bibRefCitation id="EFF24B7F4B4AC257123DF9FEFB823AD3" author="Andre, E." box="[1040,1099,1660,1686]" pageId="2" pageNumber="203" pagination="256 - 308" refId="ref12774" refString="Andre, E. (1898) Contribution a la connaissance des mutillides de l'Australie. Memoires de la Societ Zoologique de France, 11, 256 - 308." type="journal article" year="1898">1898</bibRefCitation>
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,
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<bibRefCitation id="EFF24B7F4B4AC257127BF9FEFB5B3AD3" author="Andre, E." box="[1110,1170,1660,1686]" pageId="2" pageNumber="203" pagination="467 - 513" refId="ref12804" refString="Andre, E. (1901) Nouvelle contribution a la connaissance des mutillides de l'Australie. Memoires de la Societ Zoologique de France, 14, 467 - 513." type="journal article" year="1901">1901</bibRefCitation>
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||
); although the
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<date id="FFDD104E4B4AC257131BF9F9FF063AFF" pageId="2" pageNumber="203" value="1895" valueMax="1898">1895 and 1898</date>
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keys included the majority of species described to those dates, the 1901 key primarily included only species described in that publication.
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<bibRefCitation id="EFF24B7F4B4AC25717CAF947FD4D3A9B" author="Turner, R. E." box="[487,644,1732,1759]" pageId="2" pageNumber="203" pagination="429 - 450" refId="ref15710" refString="Turner, R. E. (1914) Notes on fossorial Hymenoptera. XIV. Annals and Magazine of Natural History, Series 8, 14 (84), 429 - 450. https: // doi. org / 10.1080 / 00222931408693600" type="journal article" year="1914">Turner (1914)</bibRefCitation>
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reviewed selected species from
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<collectingRegion id="49A7F86C4B4AC25715D9F947FB753A9B" box="[1012,1212,1732,1759]" country="Australia" name="Western Australia" pageId="2" pageNumber="203">Western Australia</collectingRegion>
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and
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<collectingRegion id="49A7F86C4B4AC25712D9F946FAAB3A9B" box="[1268,1378,1732,1758]" country="Australia" name="Tasmania" pageId="2" pageNumber="203">Tasmania</collectingRegion>
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, providing a key to females only. Unfortunately, creating a detailed and up-to-date key to Australian
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<taxonomicName id="4C634D0D4B4AC25712EAF96AFAF63B47" authorityName="Latreille" authorityYear="1802" box="[1223,1343,1768,1794]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="family">Mutillidae</taxonomicName>
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, or even to the genus
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<taxonomicName id="4C634D0D4B4AC2571708F88EFE5C3B63" authorityName="Brothers" authorityYear="2018" box="[293,405,1804,1830]" class="Insecta" family="Mutillidae" genus="Aglaotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA9C4B4AC2571708F88EFE5C3B63" box="[293,405,1804,1830]" italics="true" pageId="2" pageNumber="203">Aglaotilla</emphasis>
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</taxonomicName>
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alone, would require detailed revisionary work beyond the scope of the current study.
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</paragraph>
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<paragraph id="8BDC368E4B4AC25616EAF8B2FD643EC2" blockId="2.[151,1437,1623,2011]" lastBlockId="3.[151,1437,153,827]" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
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The majority of species of Australian
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<taxonomicName id="4C634D0D4B4AC2571443F8B2FD2A3B0F" authorityName="Latreille" authorityYear="1802" box="[622,739,1840,1866]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="family">Mutillidae</taxonomicName>
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are currently placed in the genus
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<taxonomicName id="4C634D0D4B4AC2571271F8B2FA513B0F" authority="Andre, 1902" authorityName="Andre" authorityYear="1902" box="[1116,1432,1840,1866]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA9C4B4AC2571271F8B2FACD3B0F" box="[1116,1284,1840,1866]" italics="true" pageId="2" pageNumber="203">Ephutomorpha</emphasis>
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<bibRefCitation id="EFF24B7F4B4AC2571324F8B2FA513B0F" author="Andre, E." box="[1289,1432,1840,1866]" pageId="2" pageNumber="203" pagination="1 - 77" refId="ref12835" refString="Andre, E. (1902) Hymenoptera. Fam. Mutillidae. Genera Insectorum, Fasc. 11, 1 - 77, 3 pls." type="book chapter" year="1902">André, 1902</bibRefCitation>
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</taxonomicName>
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, explicitly established as a taxon of convenience to contain all Australasian species that would now be placed in the subfamily
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<taxonomicName id="4C634D0D4B4AC257173DF8FAFD613BD7" authority="(Brothers 1975)" authorityName="Brothers" authorityYear="1975" baseAuthorityName="Brothers" baseAuthorityYear="1975" box="[272,680,1911,1938]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="subFamily" subFamily="Sphaeropthalminae">
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Sphaeropthalminae (
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<bibRefCitation id="EFF24B7F4B4AC25717D5F8FAFD563BD7" author="Brothers, D. J." box="[504,671,1911,1938]" pageId="2" pageNumber="203" pagination="483 - 648" refId="ref13002" refString="Brothers, D. J. (1975) Phylogeny and classification of the aculeate Hymenoptera, with special reference to Mutillidae. University of Kansas Science Bulletin, 50, 483 - 648." type="journal article" year="1975">Brothers 1975</bibRefCitation>
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)
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</taxonomicName>
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.
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<bibRefCitation id="EFF24B7F4B4AC257149EF8FAFC853BD7" author="Andre, E." box="[691,844,1912,1938]" pageId="2" pageNumber="203" pagination="1 - 77" refId="ref12835" refString="Andre, E. (1902) Hymenoptera. Fam. Mutillidae. Genera Insectorum, Fasc. 11, 1 - 77, 3 pls." type="book chapter" year="1902">André (1902)</bibRefCitation>
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expressed the expectation that future studies would prove the value of dividing his ‘
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<taxonomicName id="4C634D0D4B4AC2571421F81EFD7D3BF3" authorityName="Andre" authorityYear="1902" box="[524,692,1948,1974]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA9C4B4AC2571421F81EFD7D3BF3" box="[524,692,1948,1974]" italics="true" pageId="2" pageNumber="203">Ephutomorpha</emphasis>
|
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</taxonomicName>
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’ into several genera. A handful of generic names have also been proposed for more restricted species groups, but relationships between these and the bulk of ‘
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<taxonomicName id="4C634D0D4B4AC257129AF842FA963B9F" authorityName="Andre" authorityYear="1902" box="[1207,1375,1984,2010]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4AC257129AF842FA963B9F" box="[1207,1375,1984,2010]" italics="true" pageId="2" pageNumber="203">Ephutomorpha</emphasis>
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</taxonomicName>
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’ spe- cies remain uninvestigated. The results of this study have thus proved particularly valuable in associating the sexes of several species which are clearly congeneric, and thus enabling the characterization of a striking new genus, described as
|
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<taxonomicName id="4C634D0D4B4BC2561703FF63FDC03CBE" authority="Brothers" authorityName="Brothers" authorityYear="2018" box="[302,521,225,251]" class="Insecta" family="Mutillidae" genus="Aglaotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA9C4B4BC2561703FF63FE573CBE" box="[302,414,225,251]" italics="true" pageId="3" pageNumber="204">Aglaotilla</emphasis>
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Brothers
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</taxonomicName>
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in an accompanying paper (Brothers, 2018). This genus differs from the other Australasian sphaeropthalmine genera (all placed in
|
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<taxonomicName id="4C634D0D4B4BC25614CEFE87FCB43D5A" authorityName="Brothers & Lelej" authorityYear="2017" box="[739,893,261,287]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="tribe" tribe="Dasymutillini">Dasymutillini</taxonomicName>
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by
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<bibRefCitation id="EFF24B7F4B4BC2561585FE87FB6E3D5A" author="Brothers, D. J. & Lelej, A. S." box="[936,1191,261,287]" pageId="3" pageNumber="204" pagination="1 - 97" refId="ref13258" refString="Brothers, D. J. & Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97. https: // doi. org / 10.3897 / jhr. 60.20091" type="journal article" year="2017">Brothers & Lelej 2017</bibRefCitation>
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) as follows: from
|
||
<taxonomicName id="4C634D0D4B4BC2561359FE87FE7E3D06" authority="Brothers, 2012" authorityName="Brothers" authorityYear="2012" class="Insecta" family="Mutillidae" genus="Ancistrotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC2561359FE87FECA3D06" italics="true" pageId="3" pageNumber="204">Ancistrotilla</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC2561721FEABFE7E3D06" author="Brothers, D. J." box="[268,439,297,323]" pageId="3" pageNumber="204" pagination="223 - 251" refId="ref13203" refString="Brothers, D. J. (2012) The new genus Ancistrotilla n. gen., with new species from Vanuatu and New Caledonia (Hymenoptera, Mutillidae). Zoosystema, 34, 223 - 251. https: // doi. org / 10.5252 / z 2012 n 2 a 2" type="journal article" year="2012">Brothers, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0D4B4BC25617EEFEABFD173D06" authority="Lelej, 1983" authorityName="Lelej" authorityYear="1983" box="[451,734,297,323]" class="Insecta" family="Mutillidae" genus="Australotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC25617EEFEABFD983D06" box="[451,593,297,323]" italics="true" pageId="3" pageNumber="204">Australotilla</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC2561474FEABFD173D06" author="Lelej, A. S." box="[601,734,297,323]" pageId="3" pageNumber="204" pagination="152 - 160" refId="ref14316" refString="Lelej, A. S. (1983) [On the velvet ants (Hymenoptera, Mutillidae) of Australia]. Entomologicheskoe Obozrenie, 62, 612 - 619. [in Russian, transl. (1984) Contributions to the knowledge of the velvet ants (Hymenoptera, Mutillidae) of Australia. Entomological Review, 62, 152 - 160]" type="journal article" year="1983">Lelej, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0D4B4BC25614C7FEABFB833D06" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[746,1098,297,323]" class="Insecta" family="Mutillidae" genus="Bothriomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC25614C7FEABFC463D06" box="[746,911,297,323]" italics="true" pageId="3" pageNumber="204">Bothriomutilla</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC25615BBFEABFB833D06" author="Ashmead, W. H." box="[918,1098,297,323]" pageId="3" pageNumber="204" pagination="45 - 60" refId="ref12862" refString="Ashmead, W. H. (1899) Super-families in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0D4B4BC256127AFEABFA513D06" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[1111,1432,297,323]" class="Insecta" genus="Eurymutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC256127AFEABFB143D06" box="[1111,1245,297,323]" italics="true" pageId="3" pageNumber="204">Eurymutilla</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC25612C9FEABFA513D06" author="Ashmead, W. H." box="[1252,1432,297,323]" pageId="3" pageNumber="204" pagination="45 - 60" refId="ref12862" refString="Ashmead, W. H. (1899) Super-families in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae and Mutillidae. Journal of the New York Entomological Society, 7, 45 - 60." type="journal article" year="1899">Ashmead, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0D4B4BC25616BAFECFFE0B3D22" authority="Lelej, 1983" authorityName="Lelej" authorityYear="1983" box="[151,450,333,359]" class="Insecta" family="Mutillidae" genus="Odontomyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC25616BAFECFFEFE3D22" box="[151,311,333,359]" italics="true" pageId="3" pageNumber="204">Odontomyrme</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC256176CFECFFE0B3D22" author="Lelej, A. S." box="[321,450,333,359]" pageId="3" pageNumber="204" pagination="152 - 160" refId="ref14316" refString="Lelej, A. S. (1983) [On the velvet ants (Hymenoptera, Mutillidae) of Australia]. Entomologicheskoe Obozrenie, 62, 612 - 619. [in Russian, transl. (1984) Contributions to the knowledge of the velvet ants (Hymenoptera, Mutillidae) of Australia. Entomological Review, 62, 152 - 160]" type="journal article" year="1983">Lelej, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D0D4B4BC25617D0FECFFCE73D22" authority="Brothers, 1994" authorityName="Brothers" authorityYear="1994" box="[509,814,333,359]" class="Insecta" genus="Ponerotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC25617D0FECFFDB53D22" box="[509,636,333,359]" italics="true" pageId="3" pageNumber="204">Ponerotilla</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC25614A8FECFFCE73D22" author="Brothers, D. J." box="[645,814,333,359]" pageId="3" pageNumber="204" pagination="143 - 152" refId="ref13138" refString="Brothers, D. J. (1994) A new genus and four new species of Mutillidae associated with Brachyponera lutea Mayr (Formicidae) in Western Australia (Hymenoptera). Journal of the Australian Entomological Society, 33, 143 - 152. https: // doi. org / 10.1111 / j. 1440 - 6055.1994. tb 00941. x" type="journal article" year="1994">Brothers, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
by the absence of a well-defined pygidial plate in the females (Brothers 2018: fig. 5) and the bright metallic coloration and hyaline wings of the males; from
|
||
<taxonomicName id="4C634D0D4B4BC2561309FEF3FEF43DEA" authority="Brothers, 1971" authorityName="Brothers" authorityYear="1971" class="Insecta" family="Mutillidae" genus="Ascetotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC2561309FEF3FA553DCE" box="[1316,1436,369,395]" italics="true" pageId="3" pageNumber="204">Ascetotilla</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC25616BAFE17FEF43DEA" author="Brothers, D. J." box="[151,317,405,431]" pageId="3" pageNumber="204" pagination="471 - 485" refId="ref12970" refString="Brothers, D. J. (1971) Ascetotilla, a new genus of Mutillidae from New Guinea (Hymenoptera). Pacific Insects, 13, 471 - 485." type="journal article" year="1971">Brothers, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
by the heavily punctate metapleuron and lateral face of the propodeum in the females and the bright metallic coloration and hyaline wings of the males; and from
|
||
<taxonomicName id="4C634D0D4B4BC2561579FE3BFB463D96" authority="Andre, 1902" authorityName="Taylor & Murphy & Hitchen & Brothers" authorityYear="2019" box="[852,1167,441,467]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B917EA9C4B4BC2561579FE3BFC353D96" box="[852,1020,441,467]" italics="true" pageId="3" pageNumber="204">Ephutomorpha</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC256122FFE3BFB463D96" author="Andre, E." box="[1026,1167,441,467]" pageId="3" pageNumber="204" pagination="1 - 77" refId="ref12835" refString="Andre, E. (1902) Hymenoptera. Fam. Mutillidae. Genera Insectorum, Fasc. 11, 1 - 77, 3 pls." type="book chapter" year="1902">André, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E74B4BC25612BAFE38FAEF3D96" box="[1175,1318,442,467]" pageId="3" pageNumber="204" sensu="stricto">
|
||
<emphasis id="B917EA9C4B4BC25612BAFE38FAEF3D96" box="[1175,1318,442,467]" italics="true" pageId="3" pageNumber="204">sensu stricto</emphasis>
|
||
</taxonomicNameLabel>
|
||
(i.e. those species grouped with the
|
||
<typeStatus id="54D8882C4B4BC2561798FE5FFE2C3DB2" box="[437,485,477,503]" pageId="3" pageNumber="204">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C634D0D4B4BC2561461FE5FFC603DB2" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[588,937,476,503]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="aurata">
|
||
<emphasis id="B917EA9C4B4BC2561461FE5FFD383DB2" box="[588,753,477,503]" italics="true" pageId="3" pageNumber="204">Mutilla aurata</emphasis>
|
||
<bibRefCitation id="EFF24B7F4B4BC25614D5FE5FFC603DB2" author="Fabricius, J. C." box="[760,937,476,503]" pageId="3" pageNumber="204" refId="ref13796" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus. Libraria Korti, Flensburgi [Flensburg] et Lipsiae [Copenhagen], [xvi] + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
|
||
</taxonomicName>
|
||
, as discussed by
|
||
<bibRefCitation id="EFF24B7F4B4BC2561244FE5FFADB3DB2" author="Brothers, D. J." box="[1129,1298,477,503]" pageId="3" pageNumber="204" pagination="223 - 251" refId="ref13203" refString="Brothers, D. J. (2012) The new genus Ancistrotilla n. gen., with new species from Vanuatu and New Caledonia (Hymenoptera, Mutillidae). Zoosystema, 34, 223 - 251. https: // doi. org / 10.5252 / z 2012 n 2 a 2" type="journal article" year="2012">Brothers, 2012</bibRefCitation>
|
||
) by the lack of a well-defined transverse groove on S2 (figs 3b, 7b, 9b) and the apically constricted T1 without an apical pale integumental band in the females (figs 1c, 3a) (males of
|
||
<emphasis id="B917EA9C4B4BC2561523FDA7FC293E7A" box="[782,992,549,575]" italics="true" pageId="3" pageNumber="204">
|
||
<taxonomicName id="4C634D0D4B4BC2561523FDA7FC7F3E7A" authority="Taylor & Murphy & Hitchen & Brothers, 2019" authorityName="Taylor & Murphy & Hitchen & Brothers" authorityYear="2019" box="[782,950,549,575]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus" sensu="stricto">Ephutomorpha</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E74B4BC2561591FDA4FC293E7A" box="[956,992,550,575]" pageId="3" pageNumber="204" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are unknown). Many other species currently assigned to
|
||
<taxonomicName id="4C634D0D4B4BC256174BFDCBFDC73E26" authorityName="Taylor & Murphy & Hitchen & Brothers" authorityYear="2019" box="[358,526,585,611]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA9C4B4BC256174BFDCBFDC73E26" box="[358,526,585,611]" italics="true" pageId="3" pageNumber="204">Ephutomorpha</emphasis>
|
||
</taxonomicName>
|
||
remain unrevised and may be transferred to new genera in the future; such revisions are beyond the scope of the current paper.
|
||
</paragraph>
|
||
<paragraph id="8BDC368E4B4BC25616EAFD13FD4B3F7E" blockId="3.[151,1437,153,827]" pageId="3" pageNumber="204">
|
||
Brothers (2018) provided the first detailed description and illustrations of male genitalia for a species of
|
||
<emphasis id="B917EA9C4B4BC256137DFD13FE603E8A" italics="true" pageId="3" pageNumber="204">
|
||
<taxonomicName id="4C634D0D4B4BC256137DFD13FF0E3E8A" authorityName="Brothers" authorityYear="2018" class="Insecta" family="Mutillidae" genus="Aglaotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">Aglaotilla</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D0D4B4BC25616FEFD37FE603E8A" authorityName="Turner" authorityYear="1916" box="[211,425,693,719]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="submetallescens">A. submetallescens</taxonomicName>
|
||
</emphasis>
|
||
. In the species described below (except possibly
|
||
<taxonomicName id="4C634D0D4B4BC25615F4FD37FBF33E8A" authority="Taylor & Murphy & Hitchen & Brothers, 2019" authorityName="Taylor & Murphy & Hitchen & Brothers" authorityYear="2019" box="[985,1082,693,719]" class="Insecta" family="Mutillidae" genus="Aglaotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="micra" status="sp. nov.">
|
||
<emphasis id="B917EA9C4B4BC25615F4FD37FBF33E8A" box="[985,1082,693,719]" italics="true" pageId="3" pageNumber="204">A. micra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457E74B4BC256126EFD37FB543E8A" box="[1091,1181,693,719]" pageId="3" pageNumber="204" rank="species">
|
||
<emphasis id="B917EA9C4B4BC256126EFD37FB543E8A" bold="true" box="[1091,1181,693,719]" pageId="3" pageNumber="204">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, for which the male is currently unknown) the male genitalia closely resemble those of
|
||
<taxonomicName id="4C634D0D4B4BC256154FFD5BFBFF3EB6" authorityName="Turner" authorityYear="1916" box="[866,1078,729,755]" class="Insecta" family="Mutillidae" genus="Ephutomorpha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="submetallescens">
|
||
<emphasis id="B917EA9C4B4BC256154FFD5BFBFF3EB6" box="[866,1078,729,755]" italics="true" pageId="3" pageNumber="204">A. submetallescens</emphasis>
|
||
</taxonomicName>
|
||
in overall appearance with most variation involving length and pilosity of the parameres. Arrangements of the parapenial lobes, cuspides and penis valves vary less obviously between species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |