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<document id="3FF1741E71921734D220B0C77054C0CA" ID-DOI="10.1649/0010-065X-78.2.255" ID-ISSN="1938-4394" ID-Zenodo-Dep="13243970" ID-ZooBank="urn:lsid:zoobank.org:pub:96AA091D-040B-4C64-AAB8-CD8FC550C4F9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722969630902" checkinUser="felipe" docAuthor="Minkina, Łukasz, Gama, Emanuel R. R. &amp; Skelley, Paul E." docDate="2024" docId="03CA8783FFB2D65E56FBFAE6F82AF94E" docLanguage="en" docName="TheColeopBull.78.2.255-264.pdf" docOrigin="The Coleopterists Bulletin 78 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-78.2.255" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Ataenius kudrnai Minkina, Gama, and SkelleY 2024, new species" docType="treatment" docUuid="BF9E525F-5BF9-48A6-A1BF-15934B944798" docUuidSource="ZooBank" docVersion="2" lastPageNumber="258" masterDocId="FFF3FFFBFFB0D65A5666FF84FC13FFFC" masterDocTitle="Two New Species of the Genus Ataenius Schmidt, 1810 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini)" masterLastPageNumber="264" masterPageNumber="255" pageNumber="256" updateTime="1722973450802" updateUser="ExternalLinkService">
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<mods:title id="EB6512CA565D0DFB482183AFB0B1A415">Two New Species of the Genus Ataenius Schmidt, 1810 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini)</mods:title>
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<mods:namePart id="B298E78163C33B7965446C88A900F00E">Minkina, Łukasz</mods:namePart>
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<mods:namePart id="835463A8C8B61CA28D1B3280457C85D5">Gama, Emanuel R. R.</mods:namePart>
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<mods:namePart id="5FFC61C7A98C94EDB232E01183244C84">Skelley, Paul E.</mods:namePart>
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<mods:title id="800774BC3D2AEC669A71EC9706053707">The Coleopterists Bulletin</mods:title>
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<mods:part id="3E3A722ECD6E6F01830BF73F63492EE9">
<mods:date id="4BA5A4E22B5D63CB78197145A734ED5B">2024</mods:date>
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<mods:number id="A893C9E0D4EA36E1EEC7D71C7D41A4F3">2024-06-21</mods:number>
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<mods:detail id="287764D5B1B55C65E42685FC1608BE48" type="volume">
<mods:number id="B3C82602CE23181B01F3241A808EECF2">78</mods:number>
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<mods:number id="35F768377D7DE0380BE4EE23300C3868">2</mods:number>
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<mods:identifier id="F528E2CDD0D90361C101D2CACA96DB22" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="C379651EFFB2D65856FBFAE6FE1BFA1D" pageId="2" pageNumber="256" type="nomenclature">
<paragraph id="8BDC3695FFB2D65856FBFAE6FE1BFA1D" blockId="2.[157,633,1377,1506]" pageId="2" pageNumber="256">
<heading id="D09481F9FFB2D65856FBFAE6FDD5FA6E" centered="true" fontSize="9" level="2" pageId="2" pageNumber="256" reason="2">
<emphasis id="B917EA87FFB2D65856FBFAE6FDD5FA6E" bold="true" pageId="2" pageNumber="256">
<taxonomicName id="4C634D16FFB2D65856FBFAE6FE67FA8B" ID-CoL="CGP9K" authority="Minkina, Gama, and Skelley" authorityName="Minkina, Gama, and SkelleY" authorityYear="2024" box="[157,628,1377,1399]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="species" species="kudrnai" status="sp. nov.">
<emphasis id="B917EA87FFB2D65856FBFAE6FD43FA8B" bold="true" box="[157,336,1377,1399]" italics="true" pageId="2" pageNumber="256">Ataenius kudrnai</emphasis>
Minkina, Gama, and Skelley
</taxonomicName>
,
<taxonomicNameLabel id="A22457FCFFB2D6585736FAF8FDD5FA6E" box="[336,454,1404,1426]" pageId="2" pageNumber="256" rank="species">new species</taxonomicNameLabel>
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zoobank.org/
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="13582A10FFB2D6585773FA48FD63FA1D" box="[277,368,1484,1506]" captionStart="Figs" captionStartId="3.[197,237,1451,1470]" captionTargetBox="[185,1181,196,1434]" captionTargetId="figure-114@3.[173,1193,193,1434]" captionTargetPageId="3" captionText="Figs. 1–3. Ataenius kudrnai, new species, ♂, holotYpe. 1) Dorsal view; 2) Ventral view; 3) Lateral view. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243972" httpUri="https://zenodo.org/record/13243972/files/figure.png" pageId="2" pageNumber="256">Figs. 1–3</figureCitation>
,
<figureCitation id="13582A10FFB2D658571DFA48FD86FA1D" box="[379,405,1484,1505]" captionStart="Figs" captionStartId="6.[176,215,836,855]" captionTargetBox="[155,1160,203,819]" captionTargetId="figure-414@6.[151,1171,193,819]" captionTargetPageId="6" captionText="Figs. 10–13. Heads, dorso-anterior view. 10) Ataenius kudrnai, new species, ♂, holotYpe; 11) A. guriensis, ♂, para- tYpe; 12) A. punctipennis, ♂; 13) A. vazdemelloi, new species, ♂, holotYpe. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243978" httpUri="https://zenodo.org/record/13243978/files/figure.png" pageId="2" pageNumber="256">10</figureCitation>
,
<figureCitation id="13582A10FFB2D65857F9FA48FE13FA1D" box="[415,512,1484,1506]" captionStart="Figs" captionStartId="7.[197,236,1305,1324]" captionTargetBox="[193,1167,204,1273]" captionTargetId="figure-131@7.[173,1193,193,1287]" captionTargetPageId="7" captionText="Figs. 14–23. Aedeagi and epipharynges. Aedeagus: 14) Ataenius kudrnai, new species, ♂, holotYpe, dorsal view; 15) Same, lateral view; 16) A. guriensis, ♂, paratYpe, dorsal view; 17) Same, lateral view; 18) A. punctipennis, ♂, lateral view; 19) A. vazdemelloi, new species, ♂, holotYpe, lateral view. EpipharYnx: 20) A. kudrnai, new species, ♂, holotYpe; 21) A. guriensis, ♂, paratYpe; 22) A. punctipennis, ♂; 23) A. vazdemelloi, new species, ♂, holotYpe. Scale lines: Figs. 14–19 = 0.5 mm; Figs. 20–23 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13243980" httpUri="https://zenodo.org/record/13243980/files/figure.png" pageId="2" pageNumber="256">14, 15, 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C379651EFFB2D65856D6FA73FDD5F96E" pageId="2" pageNumber="256" type="materials_examined">
<paragraph id="8BDC3695FFB2D65856D6FA73FD1CF9D4" blockId="2.[152,638,1527,1736]" pageId="2" pageNumber="256">
<materialsCitation id="3B0B3CC8FFB2D65856D6FA73FD1CF9D4" country="Paraguay" location="Mariscal Estigarribia." pageId="2" pageNumber="256" specimenCount="1" typeStatus="holotype">
<emphasis id="B917EA87FFB2D65856D6FA73FD2FF9F1" bold="true" box="[176,316,1527,1549]" pageId="2" pageNumber="256">
<typeStatus id="54D88837FFB2D65856D6FA73FCF2F9F0" box="[176,225,1527,1548]" pageId="2" pageNumber="256">Type</typeStatus>
Locality.
</emphasis>
<collectingCountry id="F3747605FFB2D6585726FA73FD8BF9F0" box="[320,408,1527,1548]" name="Paraguay" pageId="2" pageNumber="256">ParaguaY</collectingCountry>
,
<quantity id="4C9B9B70FFB2D65857C6FA73FDEFF9F1" box="[416,508,1527,1549]" metricMagnitude="4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="2" pageNumber="256" unit="km" value="82.5" valueMax="85.0" valueMin="80.0">80–85 km</quantity>
NW
<location id="8EBC604EFFB2D658544AFA73FD1CF9D4" LSID="urn:lsid:plazi:treatment:03CA8783FFB2D65E56FBFAE6F82AF94E:8EBC604EFFB2D658544AFA73FD1CF9D4" country="Paraguay" name="Mariscal Estigarribia." pageId="2" pageNumber="256">Mariscal Estigarribia.</location>
</materialsCitation>
</paragraph>
<paragraph id="8BDC3695FFB2D65856D6F9A8FDD5F96E" blockId="2.[152,638,1527,1736]" pageId="2" pageNumber="256">
<emphasis id="B917EA87FFB2D65856D6F9A8FD52F9BE" bold="true" box="[176,321,1580,1602]" pageId="2" pageNumber="256">Type Material.</emphasis>
<materialsCitation id="3B0B3CC8FFB2D6585723F9A8FDD5F96E" collectingDate="2008-01-30" collectingDateMax="2008-01-31" collectingDateMin="2008-01-30" collectorName="Arnost Kudrna" country="Paraguay" determinerName="Det. Ing. Jiri Chromy" location="Estigarribia" pageId="2" pageNumber="256" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D88837FFB2D6585723F9A8FD88F9BE" box="[325,411,1580,1602]" pageId="2" pageNumber="256" type="holotype">HolotYpe</typeStatus>
(
<specimenCount id="9D65FD1CFFB2D65857C1F9A8FDA9F9BD" box="[423,442,1580,1601]" count="1" pageId="2" pageNumber="256" type="male">♂</specimenCount>
, MMBC): “
<collectingCountry id="F3747605FFB2D6585443F9A8FE6DF9BD" box="[549,638,1580,1601]" name="Paraguay" pageId="2" pageNumber="256">ParaguaY</collectingCountry>
-06, ABCJCH |
<quantity id="4C9B9B70FFB2D6585759F9C3FD89F9A1" box="[319,410,1607,1629]" metricMagnitude="4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="2" pageNumber="256" unit="km" value="82.5" valueMax="85.0" valueMin="80.0">80–85km</quantity>
NW Mariscal- |
<location id="8EBC604EFFB2D6585436F9C3FCF5F984" LSID="urn:lsid:plazi:treatment:03CA8783FFB2D65E56FBFAE6F82AF94E:8EBC604EFFB2D6585436F9C3FCF5F984" country="Paraguay" name="Estigarribia" pageId="2" pageNumber="256">Estigarribia</location>
<date id="FFDD1055FFB2D658568BF9E6FD92F984" box="[237,385,1634,1656]" pageId="2" pageNumber="256" value="2008-01-30" valueMax="2008-01-31" valueMin="2008-01-30">
<collectingDate id="EF99E9BDFFB2D658568BF9E6FD92F984" box="[237,385,1634,1656]" pageId="2" pageNumber="256" value="2008-01-30" valueMax="2008-01-31" valueMin="2008-01-30">30.i.-31.i.2008</collectingDate>
</date>
| leg.
<collectorName id="26965343FFB2D65857A6F9E6FE46F984" box="[448,597,1634,1656]" pageId="2" pageNumber="256">Arnošt Kudrna</collectorName>
jr. | Coll. &amp;
<determinerName id="7E00DA51FFB2D6585680F9F9FDAAF96E" box="[230,441,1660,1682]" pageId="2" pageNumber="256">Det. Ing. Jiří Chromý</determinerName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C379651EFFB2D65E56D6F913FDFEF9BB" lastPageId="4" lastPageNumber="258" pageId="2" pageNumber="256" type="description">
<paragraph id="8BDC3695FFB2D65856D6F913FE6DF934" blockId="2.[152,638,1527,1736]" pageId="2" pageNumber="256">
<emphasis id="B917EA87FFB2D65856D6F913FDC0F950" bold="true" box="[176,467,1687,1709]" pageId="2" pageNumber="256">
Description of the
<typeStatus id="54D88837FFB2D6585716F913FDDCF950" box="[368,463,1687,1708]" pageId="2" pageNumber="256" type="holotype">Holotype</typeStatus>
.
</emphasis>
Dorsum (
<figureCitation id="13582A10FFB2D6585453F913FE7DF950" box="[565,622,1687,1709]" captionStart="Figs" captionStartId="3.[197,237,1451,1470]" captionTargetBox="[185,1181,196,1434]" captionTargetId="figure-114@3.[173,1193,193,1434]" captionTargetPageId="3" captionText="Figs. 1–3. Ataenius kudrnai, new species, ♂, holotYpe. 1) Dorsal view; 2) Ventral view; 3) Lateral view. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243972" httpUri="https://zenodo.org/record/13243972/files/figure.png" pageId="2" pageNumber="256">Fig. 1</figureCitation>
). Length:
<quantity id="4C9B9B70FFB2D6585688F936FD2DF93B" box="[238,318,1714,1735]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="256" unit="mm" value="3.4">3.4 mm</quantity>
. Dorsal surface reddish brown;
</paragraph>
<paragraph id="8BDC3695FFB2D65854CBFF46F86DFEBE" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">
tarsomeres, antennae and mouthparts pale brown. Body small-sized for
<taxonomicName id="4C634D16FFB2D65855EFFF59FFCEFF0E" baseAuthorityName="Stebnicka" baseAuthorityYear="2007" box="[905,989,221,242]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA87FFB2D65855EFFF59FFCEFF0E" box="[905,989,221,242]" italics="true" pageId="2" pageNumber="256">Ataenius</emphasis>
</taxonomicName>
, elongate, moderatelY convex; indistinctlY macrosetate on sides of pronotum and distinctly macrosetate on sides and before apex of elytra, macrosetae short and thin.
</paragraph>
<paragraph id="8BDC3695FFB2D65854A3FEC3F85BFDCE" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">
Head (
<figureCitation id="13582A10FFB2D6585561FEC3FF43FEA0" box="[775,848,327,349]" captionStart="Figs" captionStartId="6.[176,215,836,855]" captionTargetBox="[155,1160,203,819]" captionTargetId="figure-414@6.[151,1171,193,819]" captionTargetPageId="6" captionText="Figs. 10–13. Heads, dorso-anterior view. 10) Ataenius kudrnai, new species, ♂, holotYpe; 11) A. guriensis, ♂, para- tYpe; 12) A. punctipennis, ♂; 13) A. vazdemelloi, new species, ♂, holotYpe. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243978" httpUri="https://zenodo.org/record/13243978/files/figure.png" pageId="2" pageNumber="256">Fig. 10</figureCitation>
) moderatelY convex, moderatelY shinY, with distinct microreticulation; transverselY trapezoidal; clYpeus anteriorlY distinctlY sinuate, sides widely rounded, latter truncate, not notched before genae, which are obtuse and distinctly exceeding eYes. FrontoclYpeal suture absent. Punctures on clypeus irregular in size and shape (basally and on the disc regularlY rounded, anteriorlY transverse, not wrinkled), dense, regularly distributed.
</paragraph>
<paragraph id="8BDC3695FFB2D65854A3FDB3FF4DFCF4" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">
EpipharYnx (
<figureCitation id="13582A10FFB2D658552AFDB3FF88FDB0" box="[844,923,567,589]" captionStart="Figs" captionStartId="7.[197,236,1305,1324]" captionTargetBox="[193,1167,204,1273]" captionTargetId="figure-131@7.[173,1193,193,1287]" captionTargetPageId="7" captionText="Figs. 14–23. Aedeagi and epipharynges. Aedeagus: 14) Ataenius kudrnai, new species, ♂, holotYpe, dorsal view; 15) Same, lateral view; 16) A. guriensis, ♂, paratYpe, dorsal view; 17) Same, lateral view; 18) A. punctipennis, ♂, lateral view; 19) A. vazdemelloi, new species, ♂, holotYpe, lateral view. EpipharYnx: 20) A. kudrnai, new species, ♂, holotYpe; 21) A. guriensis, ♂, paratYpe; 22) A. punctipennis, ♂; 23) A. vazdemelloi, new species, ♂, holotYpe. Scale lines: Figs. 14–19 = 0.5 mm; Figs. 20–23 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13243980" httpUri="https://zenodo.org/record/13243980/files/figure.png" pageId="2" pageNumber="256">Fig. 20</figureCitation>
) transverse, with sides rounded, anterior margin truncate, with weakly produced corypha bearing two short, thick celtes at apex.Acanthopariae, acropariae, prophobae without chaetae. Chaetopariae, adelochaetae, chaetopedia with dense belts of thick, long chaetae. Epitorma cup shaped, shortened, not reaching base of epipharynx. Tormae long.
</paragraph>
<paragraph id="8BDC3695FFB2D65854A3FC89F887FB9E" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">Pronotum moderatelY convex, distinctlY transverse, as wide as base of elYtra, widest near base, moderately shiny, with distinct microreticulation. Anterior angles widelY rounded, sides verY widelY rounded; truncate before hind angles.Anterior margin not bordered. Sides and basal margin distinctly bordered. Punctures on sides bear indistinct, verY short macrosetation. Punctation double: smaller punctures dense, not fine, quite regularlY distributed, regular in size; larger punctures dense, quite regularly distributed, irregular in size, punctures near base with diameter about twice as large as smaller punctures, anteriorly all punctures smaller.</paragraph>
<paragraph id="8BDC3695FFB2D65854A3FBE3FFFCFB4E" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">Scutellar shield small, triangular, with ogival sides, matte, with verY distinct microreticulation, with few small punctures basally.</paragraph>
<paragraph id="8BDC3695FFB2D65854A3FB33FF36FA2B" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">Base of elytra thinly bordered, with small but distinct humeral denticles. ElYtra elongate, convex, nearly parallel sided, widest at midlength. Elytron with ten striae and ten interstriae. Interstriae distinctlY convex, weaklY shinY with distinct microreticulation and with irregular row of moderately large, quite dense punctures near each border; bor- ders smooth. Striae shiny, with distinct, moderately coarse, dense punctation; punctures of striae not encroaching on interstriae; striae variablY connected before apex.</paragraph>
<paragraph id="8BDC3695FFB2D65854A3FA58FF92F9F1" blockId="2.[684,1172,194,1736]" pageId="2" pageNumber="256">Pygidium with similar structure as abdominal ventrites, disc eroded.</paragraph>
<paragraph id="8BDC3695FFB2D65954A3F996FE1BF92E" blockId="2.[684,1172,194,1736]" lastBlockId="3.[173,660,1538,1747]" lastPageId="3" lastPageNumber="257" pageId="2" pageNumber="256">Legs. Profemur shiny, with trace of microreticulation, distinctly bordered anteriorly and basally, surface with coarse punctures; meso- and metafemora shinY, with trace of microreticulation, verY thinly bordered only basally, surface with fine punctures; all femora denselY punctate, punctures weaklY variable in size and shape, some with macrosetae. Protibia distinctly tridentate laterally, proximally not serrulate, with dorsal side smooth; apical spur inwardlY curved before apex, somewhat downwardlY bent. Metatibial superior apical spur somewhat longer than metatarsomere 1, latter as long as following three tarsomeres combined. Apex of meso- and metatibiae with weak accessory spine. Claws short, thin, distinctly arcuate.</paragraph>
<caption id="DF1C661DFFB3D65956A3FA2FFD24FA2B" ID-DOI="http://doi.org/10.5281/zenodo.13243972" ID-Zenodo-Dep="13243972" httpUri="https://zenodo.org/record/13243972/files/figure.png" pageId="3" pageNumber="257" startId="3.[197,237,1451,1470]" targetBox="[185,1181,196,1434]" targetPageId="3" targetType="figure">
<paragraph id="8BDC3695FFB3D65956A3FA2FFD24FA2B" blockId="3.[173,1192,1451,1495]" pageId="3" pageNumber="257">
<emphasis id="B917EA87FFB3D65956A3FA2FFD0DFA42" bold="true" box="[197,286,1451,1470]" pageId="3" pageNumber="257">Figs. 1–3.</emphasis>
<taxonomicName id="4C634D16FFB3D6595752FA28FDD8FA42" authorityName="Minkina &amp; Gama &amp; Skelley" authorityYear="2024" box="[308,459,1451,1471]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="257" phylum="Arthropoda" rank="species" species="kudrnai" status="sp. nov.">
<emphasis id="B917EA87FFB3D6595752FA28FDD8FA42" box="[308,459,1451,1471]" italics="true" pageId="3" pageNumber="257">Ataenius kudrnai</emphasis>
</taxonomicName>
,
<emphasis id="B917EA87FFB3D65957B0FA28FE51FA42" bold="true" box="[470,578,1451,1471]" pageId="3" pageNumber="257">
<taxonomicNameLabel id="A22457FCFFB3D65957B0FA28FE51FA42" box="[470,578,1451,1471]" pageId="3" pageNumber="257" rank="species">new species</taxonomicNameLabel>
</emphasis>
, ♂, holotYpe.
<emphasis id="B917EA87FFB3D65954A5FA2FFEC7FA42" bold="true" box="[707,724,1451,1470]" pageId="3" pageNumber="257">1)</emphasis>
Dorsal view;
<emphasis id="B917EA87FFB3D6595535FA2FFF77FA42" bold="true" box="[851,868,1451,1470]" pageId="3" pageNumber="257">2)</emphasis>
Ventral view;
<emphasis id="B917EA87FFB3D6595581FA2FFFE9FA42" bold="true" box="[999,1018,1451,1470]" pageId="3" pageNumber="257">3)</emphasis>
Lateral view. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BDC3695FFB3D65954BDF987FF74F9E4" blockId="3.[706,1192,1539,1747]" box="[731,871,1539,1560]" pageId="3" pageNumber="257">Macropterous.</paragraph>
<paragraph id="8BDC3695FFB3D65E54BDF999FE6CFA5B" blockId="3.[706,1192,1539,1747]" lastBlockId="4.[151,639,1372,1714]" lastPageId="4" lastPageNumber="258" pageId="3" pageNumber="257">
Venter (
<figureCitation id="13582A10FFB3D659554EF999FF76F9CE" box="[808,869,1565,1587]" captionStart="Figs" captionStartId="3.[197,237,1451,1470]" captionTargetBox="[185,1181,196,1434]" captionTargetId="figure-114@3.[173,1193,193,1434]" captionTargetPageId="3" captionText="Figs. 1–3. Ataenius kudrnai, new species, ♂, holotYpe. 1) Dorsal view; 2) Ventral view; 3) Lateral view. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243972" httpUri="https://zenodo.org/record/13243972/files/figure.png" pageId="3" pageNumber="257">Fig. 2</figureCitation>
). Metaventral plate shinY, with a trace of microreticulation, verY weaklY convex, with distinct, quite wide, deep longitudinal line in the middle; medial surface with moderatelY coarse punctation, regular in size, moderately dense, some punctures bearing verY short macrosetae.Abdominal ventrites shinY, without microreticulation; basal margins narrowly fluted, except the last one which is distinctlY fluted; with distinct, dense punctation variable in size; some punctures bearing short macrosetae.
</paragraph>
<caption id="DF1C661DFFB4D65E56D6FA89F882FADC" ID-DOI="http://doi.org/10.5281/zenodo.13243974" ID-Zenodo-Dep="13243974" box="[176,1169,1293,1312]" httpUri="https://zenodo.org/record/13243974/files/figure.png" pageId="4" pageNumber="258" startId="4.[176,215,1293,1312]" targetBox="[163,1159,198,1275]" targetPageId="4" targetType="figure">
<paragraph id="8BDC3695FFB4D65E56D6FA89F882FADC" blockId="4.[176,1169,1293,1312]" box="[176,1169,1293,1312]" pageId="4" pageNumber="258">
<emphasis id="B917EA87FFB4D65E56D6FA89FD15FADC" bold="true" box="[176,262,1293,1312]" pageId="4" pageNumber="258">Figs. 4–6.</emphasis>
<taxonomicName id="4C634D16FFB4D65E577DFA89FDAAFAE3" authorityName="Stebnicka" authorityYear="2005" box="[283,441,1293,1312]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="guriensis">
<emphasis id="B917EA87FFB4D65E577DFA89FDAAFAE3" box="[283,441,1293,1312]" italics="true" pageId="4" pageNumber="258">Ataenius guriensis</emphasis>
</taxonomicName>
, ♂, paratYpe.
<emphasis id="B917EA87FFB4D65E5455FA89FE57FADC" bold="true" box="[563,580,1293,1312]" pageId="4" pageNumber="258">4)</emphasis>
Dorsal view;
<emphasis id="B917EA87FFB4D65E54DBFA89FEDCFADC" bold="true" box="[701,719,1293,1312]" pageId="4" pageNumber="258">5)</emphasis>
Ventral view;
<emphasis id="B917EA87FFB4D65E552DFA89FF4DFADC" bold="true" box="[843,862,1293,1312]" pageId="4" pageNumber="258">6)</emphasis>
Lateral view. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BDC3695FFB4D65E56D6FA28FE61F9ED" blockId="4.[151,639,1372,1714]" pageId="4" pageNumber="258">
Aedeagus (
<figureCitation id="13582A10FFB4D65E5779FA28FD89FA3E" box="[287,410,1452,1474]" captionStart="Figs" captionStartId="7.[197,236,1305,1324]" captionTargetBox="[193,1167,204,1273]" captionTargetId="figure-131@7.[173,1193,193,1287]" captionTargetPageId="7" captionText="Figs. 14–23. Aedeagi and epipharynges. Aedeagus: 14) Ataenius kudrnai, new species, ♂, holotYpe, dorsal view; 15) Same, lateral view; 16) A. guriensis, ♂, paratYpe, dorsal view; 17) Same, lateral view; 18) A. punctipennis, ♂, lateral view; 19) A. vazdemelloi, new species, ♂, holotYpe, lateral view. EpipharYnx: 20) A. kudrnai, new species, ♂, holotYpe; 21) A. guriensis, ♂, paratYpe; 22) A. punctipennis, ♂; 23) A. vazdemelloi, new species, ♂, holotYpe. Scale lines: Figs. 14–19 = 0.5 mm; Figs. 20–23 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13243980" httpUri="https://zenodo.org/record/13243980/files/figure.png" pageId="4" pageNumber="258">Figs. 14, 15</figureCitation>
). Parameres somewhat shorter than phallobase, distinctly elongate, weakly downturned toward rounded apex; apex with small, membranous process not extending beyond apex.
</paragraph>
<paragraph id="8BDC3695FFB4D65E56D6F992FD91F9D0" blockId="4.[151,639,1372,1714]" box="[176,386,1558,1580]" pageId="4" pageNumber="258">
<emphasis id="B917EA87FFB4D65E56D6F992FD05F9D0" bold="true" box="[176,278,1558,1580]" pageId="4" pageNumber="258">Variation.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BDC3695FFB4D65E56D6F9B5FDFEF9BB" blockId="4.[151,639,1372,1714]" box="[176,493,1585,1607]" pageId="4" pageNumber="258">
<emphasis id="B917EA87FFB4D65E56D6F9B5FD92F9BB" bold="true" box="[176,385,1585,1607]" pageId="4" pageNumber="258">Sexual Dimorphism.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C379651EFFB4D65E56D6F9C8F82AF94E" pageId="4" pageNumber="258" type="etymology">
<paragraph id="8BDC3695FFB4D65E56D6F9C8FDCDF980" blockId="4.[151,639,1372,1714]" pageId="4" pageNumber="258">
<emphasis id="B917EA87FFB4D65E56D6F9C8FD30F99D" bold="true" box="[176,291,1612,1633]" pageId="4" pageNumber="258">Etymology.</emphasis>
PatronYmic. Dedicated to Arnošt Kudrna, collector of the
<typeStatus id="54D88837FFB4D65E57E5F9E2FDC9F980" box="[387,474,1638,1660]" pageId="4" pageNumber="258" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BDC3695FFB4D65E56D6F905F82AF94E" blockId="4.[151,639,1372,1714]" lastBlockId="4.[685,1172,1372,1714]" pageId="4" pageNumber="258">
<emphasis id="B917EA87FFB4D65E56D6F905FDB8F96B" bold="true" box="[176,427,1665,1687]" pageId="4" pageNumber="258">Affinity and Discussion.</emphasis>
Based on the transverselY punctate clYpeus lacking wrinkles, elongate aedeagus with parameres rounded at apex and somewhat shorter than phallobase, body reddish brown, weak accessory spine on metatibia, and apical spur of protibia in male with inwardly hooked apex, we can place
<taxonomicName id="4C634D16FFB4D65E556BFA43FF62FA27" authorityName="Minkina &amp; Gama &amp; Skelley" authorityYear="2024" box="[781,881,1478,1500]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="kudrnai">
<emphasis id="B917EA87FFB4D65E556BFA43FF62FA27" box="[781,881,1478,1500]" italics="true" pageId="4" pageNumber="258">A. kudrnai</emphasis>
</taxonomicName>
in the
<emphasis id="B917EA87FFB4D65E55D5FA43FFD2FA20" box="[947,961,1479,1500]" italics="true" pageId="4" pageNumber="258">A</emphasis>
.
<emphasis id="B917EA87FFB4D65E55A8FA42F880FA27" box="[974,1171,1478,1499]" italics="true" pageId="4" pageNumber="258">
aequalis-A.
<taxonomicName id="4C634D16FFB4D65E5226FA42F880FA27" baseAuthorityName="Blanchard" baseAuthorityYear="1846" box="[1088,1171,1478,1499]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="platensis">platensis</taxonomicName>
</emphasis>
species group. In
<bibRefCitation id="EFF24B64FFB4D65E5533FA65F814FA0B" author="Stebnicka, Z." box="[853,1031,1505,1527]" pageId="4" pageNumber="258" pagination="1 - 2" refId="ref4598" refString="Stebnicka, Z. 2005. The New World species of Ataenius Harold, 1867. VI. Revision of the A. aequalisplatensis - group. Acta Zoologica Cracoviensia 48 B (1 - 2): 99 - 138." type="journal article" year="2005">Stebnicka’s (2005)</bibRefCitation>
keY to species of the
<emphasis id="B917EA87FFB4D65E5492FA78FF11F9ED" box="[756,770,1532,1553]" italics="true" pageId="4" pageNumber="258">A</emphasis>
.
<emphasis id="B917EA87FFB4D65E5572FA78FF6AF9ED" box="[788,889,1532,1553]" italics="true" pageId="4" pageNumber="258">aequalis-A</emphasis>
.
<taxonomicName id="4C634D16FFB4D65E55ECFA78FFCEF9ED" baseAuthorityName="Blanchard" baseAuthorityYear="1846" box="[906,989,1532,1553]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="platensis">
<emphasis id="B917EA87FFB4D65E55ECFA78FFCEF9ED" box="[906,989,1532,1553]" italics="true" pageId="4" pageNumber="258">platensis</emphasis>
</taxonomicName>
species group,
<taxonomicName id="4C634D16FFB4D65E5218FA78FEE5F9D7" authorityName="Minkina &amp; Gama &amp; Skelley" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="kudrnai">
<emphasis id="B917EA87FFB4D65E5218FA78FEE5F9D7" italics="true" pageId="4" pageNumber="258">A. kudrnai</emphasis>
</taxonomicName>
keys to
<taxonomicName id="4C634D16FFB4D65E5524F993FFC0F9D0" authorityName="Harold" authorityYear="1868" box="[834,979,1559,1580]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B917EA87FFB4D65E5524F993FFC0F9D0" box="[834,979,1559,1580]" italics="true" pageId="4" pageNumber="258">A. punctipennis</emphasis>
</taxonomicName>
, but it is also similar to
<taxonomicName id="4C634D16FFB4D65E54A0F9B5FF25F9BB" authorityName="Stebnicka" authorityYear="2005" box="[710,822,1585,1607]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="guriensis">
<emphasis id="B917EA87FFB4D65E54A0F9B5FF25F9BB" box="[710,822,1585,1607]" italics="true" pageId="4" pageNumber="258">A. guriensis</emphasis>
</taxonomicName>
, having a similar aedeagus. Because of their similarities, we present important distinguishing characters in
<tableCitation id="C6E1032EFFB4D65E552DF9E2FF83F980" box="[843,912,1638,1660]" captionStart="Table 1" captionStartId="8.[176,226,194,213]" captionText="Table 1. Differential characters of Ataenius kudrnai, new species and the most similar species." pageId="4" pageNumber="258">Table 1</tableCitation>
and fully illustrate
<taxonomicName id="4C634D16FFB4D65E5222F9E3FEEFF96B" authorityName="Harold" authorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B917EA87FFB4D65E5222F9E3F841F980" box="[1092,1106,1639,1660]" italics="true" pageId="4" pageNumber="258">A</emphasis>
.
<emphasis id="B917EA87FFB4D65E523BF9E3FEEFF96B" italics="true" pageId="4" pageNumber="258">punctipennis</emphasis>
</taxonomicName>
(
<figureCitation id="13582A10FFB4D65E556AF905FF7EF96A" box="[780,877,1665,1687]" captionStart="Figs" captionStartId="5.[197,236,1378,1397]" captionTargetBox="[190,1183,211,1346]" captionTargetId="figure-145@5.[173,1193,193,1360]" captionTargetPageId="5" captionText="Figs. 7–9. Ataenius punctipennis, ♂. 7) Dorsal view; 8) Ventral view; 9) Lateral view. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243976" httpUri="https://zenodo.org/record/13243976/files/figure.png" pageId="4" pageNumber="258">Figs. 7–9</figureCitation>
,
<figureCitation id="13582A10FFB4D65E551FF905FF87F96A" box="[889,916,1665,1686]" captionStart="Figs" captionStartId="6.[176,215,836,855]" captionTargetBox="[155,1160,203,819]" captionTargetId="figure-414@6.[151,1171,193,819]" captionTargetPageId="6" captionText="Figs. 10–13. Heads, dorso-anterior view. 10) Ataenius kudrnai, new species, ♂, holotYpe; 11) A. guriensis, ♂, para- tYpe; 12) A. punctipennis, ♂; 13) A. vazdemelloi, new species, ♂, holotYpe. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243978" httpUri="https://zenodo.org/record/13243978/files/figure.png" pageId="4" pageNumber="258">12</figureCitation>
,
<figureCitation id="13582A10FFB4D65E55C6F905FFF3F96A" box="[928,992,1665,1686]" captionStart="Figs" captionStartId="7.[197,236,1305,1324]" captionTargetBox="[193,1167,204,1273]" captionTargetId="figure-131@7.[173,1193,193,1287]" captionTargetPageId="7" captionText="Figs. 14–23. Aedeagi and epipharynges. Aedeagus: 14) Ataenius kudrnai, new species, ♂, holotYpe, dorsal view; 15) Same, lateral view; 16) A. guriensis, ♂, paratYpe, dorsal view; 17) Same, lateral view; 18) A. punctipennis, ♂, lateral view; 19) A. vazdemelloi, new species, ♂, holotYpe, lateral view. EpipharYnx: 20) A. kudrnai, new species, ♂, holotYpe; 21) A. guriensis, ♂, paratYpe; 22) A. punctipennis, ♂; 23) A. vazdemelloi, new species, ♂, holotYpe. Scale lines: Figs. 14–19 = 0.5 mm; Figs. 20–23 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13243980" httpUri="https://zenodo.org/record/13243980/files/figure.png" pageId="4" pageNumber="258">18, 22</figureCitation>
) and
<taxonomicName id="4C634D16FFB4D65E527CF905F880F96B" authorityName="Stebnicka" authorityYear="2005" box="[1050,1171,1665,1687]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="258" phylum="Arthropoda" rank="species" species="guriensis">
<emphasis id="B917EA87FFB4D65E527CF905F83AF96A" box="[1050,1065,1665,1686]" italics="true" pageId="4" pageNumber="258">A</emphasis>
.
<emphasis id="B917EA87FFB4D65E5251F906F880F96B" box="[1079,1171,1666,1687]" italics="true" pageId="4" pageNumber="258">guriensis</emphasis>
</taxonomicName>
(
<figureCitation id="13582A10FFB4D65E54D2F918FF1DF94E" box="[692,782,1692,1714]" captionStart="Figs" captionStartId="4.[176,215,1293,1312]" captionTargetBox="[163,1159,198,1275]" captionTargetId="figure-205@4.[151,1171,193,1275]" captionTargetPageId="4" captionText="Figs. 4–6. Ataenius guriensis, ♂, paratYpe. 4) Dorsal view; 5) Ventral view; 6) Lateral view. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243974" httpUri="https://zenodo.org/record/13243974/files/figure.png" pageId="4" pageNumber="258">Figs. 4–6</figureCitation>
,
<figureCitation id="13582A10FFB4D65E5571F918FF3CF94D" box="[791,815,1692,1713]" captionStart="Figs" captionStartId="6.[176,215,836,855]" captionTargetBox="[155,1160,203,819]" captionTargetId="figure-414@6.[151,1171,193,819]" captionTargetPageId="6" captionText="Figs. 10–13. Heads, dorso-anterior view. 10) Ataenius kudrnai, new species, ♂, holotYpe; 11) A. guriensis, ♂, para- tYpe; 12) A. punctipennis, ♂; 13) A. vazdemelloi, new species, ♂, holotYpe. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13243978" httpUri="https://zenodo.org/record/13243978/files/figure.png" pageId="4" pageNumber="258">11</figureCitation>
,
<figureCitation id="13582A10FFB4D65E555FF918FF84F94D" box="[825,919,1692,1714]" captionStart="Figs" captionStartId="7.[197,236,1305,1324]" captionTargetBox="[193,1167,204,1273]" captionTargetId="figure-131@7.[173,1193,193,1287]" captionTargetPageId="7" captionText="Figs. 14–23. Aedeagi and epipharynges. Aedeagus: 14) Ataenius kudrnai, new species, ♂, holotYpe, dorsal view; 15) Same, lateral view; 16) A. guriensis, ♂, paratYpe, dorsal view; 17) Same, lateral view; 18) A. punctipennis, ♂, lateral view; 19) A. vazdemelloi, new species, ♂, holotYpe, lateral view. EpipharYnx: 20) A. kudrnai, new species, ♂, holotYpe; 21) A. guriensis, ♂, paratYpe; 22) A. punctipennis, ♂; 23) A. vazdemelloi, new species, ♂, holotYpe. Scale lines: Figs. 14–19 = 0.5 mm; Figs. 20–23 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13243980" httpUri="https://zenodo.org/record/13243980/files/figure.png" pageId="4" pageNumber="258">16, 17, 21</figureCitation>
) for comparison.
</paragraph>
</subSubSection>
</treatment>
</document>
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