533 lines
68 KiB
XML
533 lines
68 KiB
XML
<document id="861B15DFF93E1A82B4DCE193AEA665B6" ID-CLB-Dataset="198080" ID-DOI="10.11646/palaeoentomology.5.5.12" ID-GBIF-Dataset="e5693547-f032-4d8d-b943-ff8e946c4525" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333651" ID-ZooBank="9C4266AB-52BA-477E-9D55-506D244892FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668766416390" checkinUser="plazi" docAuthor="De SOUZA AMORIM, DALTON & BROWN, BRIAN V." docDate="2022" docId="03C58784361FFFDDB10B2A8AFB4AF99A" docLanguage="en" docName="palaeoentomology.5.5.12.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.12" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Docidiadia grimaldii De SOUZA AMORIM & BROWN 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="510" masterDocId="FFFCFFFC361EFFD8B2212C06FF9FFFF1" masterDocTitle="Docidiadia grimaldii sp. nov. from Myanmar amber (Diptera: Diadocidiidae), with comments on the genus Docidiadia Blagoderov & Grimaldi" masterLastPageNumber="512" masterPageNumber="505" pageNumber="506" updateTime="1699411746723" updateUser="ExternalLinkService">
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<mods:mods id="BEE6677C8A7DDDD600C4B3D383EC540F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3243986054602CBE4269B251FC2C7486">
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<mods:title id="7603E23676116623CC3482944E101B3A">Docidiadia grimaldii sp. nov. from Myanmar amber (Diptera: Diadocidiidae), with comments on the genus Docidiadia Blagoderov & Grimaldi</mods:title>
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<mods:namePart id="D0B82E7C59421DEE35725A472ACB8A25">De SOUZA AMORIM, DALTON</mods:namePart>
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<mods:roleTerm id="A8425720B8FEEE41735E8960B39597EE">Author</mods:roleTerm>
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<mods:namePart id="174D9EA66AACBE8A025FAD8C5088275D">BROWN, BRIAN V.</mods:namePart>
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<mods:typeOfResource id="217DB43A4331C256022D58746C666020">text</mods:typeOfResource>
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<mods:title id="758173AFE45A8996E844BF93F6EFA077">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="CA5E6656E8DFA4FDAE8A08E9E4A1D7D8">
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<mods:date id="31462F7B732A5F678AD4D86EA4C12417">2022</mods:date>
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<mods:number id="47E2C77F9A189186D0C563D7C9A14D99">2022-09-22</mods:number>
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<mods:number id="FD44B7BB31601AB6B6AFF2C79F19FB36">5</mods:number>
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<mods:number id="C10E360955891951513DDEDBB366A1CE">5</mods:number>
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<mods:url id="0F973BA947DBB558480A04E30A95D2ED">http://dx.doi.org/10.11646/palaeoentomology.5.5.12</mods:url>
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<mods:classification id="41F99DE1A99F4C663479D70193CC29F9">journal article</mods:classification>
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<mods:identifier id="41C9D9FB45FB58BE0AD2830DFE3E691F" type="CLB-Dataset">198080</mods:identifier>
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<mods:identifier id="7F27E6DB90A32FD1857DA7A27D532EB2" type="DOI">10.11646/palaeoentomology.5.5.12</mods:identifier>
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<mods:identifier id="5DCD9A7118733460B2F284494D1A60C0" type="ISSN">2624-2834</mods:identifier>
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<mods:identifier id="B4101A9BCB587C152D75AF2950778D59" type="Zenodo-Dep">7333651</mods:identifier>
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<treatment id="03C58784361FFFDDB10B2A8AFB4AF99A" ID-DOI="http://doi.org/10.5281/zenodo.7383741" ID-GBIF-Taxon="205243674" ID-Zenodo-Dep="7383741" LSID="urn:lsid:plazi:treatment:03C58784361FFFDDB10B2A8AFB4AF99A" httpUri="http://treatment.plazi.org/id/03C58784361FFFDDB10B2A8AFB4AF99A" lastPageId="5" lastPageNumber="510" pageId="1" pageNumber="506">
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<subSubSection id="C3766519361FFFD9B10B2A8AFBE6F956" box="[810,1145,1676,1703]" pageId="1" pageNumber="506" type="nomenclature">
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<paragraph id="8BD33692361FFFD9B10B2A8AFBE6F956" blockId="1.[810,1145,1676,1738]" box="[810,1145,1676,1703]" pageId="1" pageNumber="506">
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<heading id="D09B81FE361FFFD9B10B2A8AFBE6F956" bold="true" box="[810,1145,1676,1703]" fontSize="10" level="2" pageId="1" pageNumber="506" reason="3">
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<emphasis id="B918EA80361FFFD9B10B2A8AFBE6F956" bold="true" box="[810,1145,1676,1703]" pageId="1" pageNumber="506">
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<taxonomicName id="4C6C4D11361FFFD9B10B2A8AFB86F956" ID-CoL="9J73Z" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[810,1049,1676,1703]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
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<emphasis id="B918EA80361FFFD9B10B2A8AFB86F956" bold="true" box="[810,1049,1676,1703]" italics="true" pageId="1" pageNumber="506">Docidiadia grimaldii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22B57FB361FFFD9B63E2A8BFBE6F956" box="[1055,1145,1677,1703]" pageId="1" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3766519361FFFD9B10B2AB6FC02F93B" box="[810,925,1712,1738]" pageId="1" pageNumber="506" type="description">
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<paragraph id="8BD33692361FFFD9B10B2AB6FC02F93B" blockId="1.[810,1145,1676,1738]" box="[810,925,1712,1738]" pageId="1" pageNumber="506">
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||
(
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<figureCitation id="13572A17361FFFD9B1132AB6FC0BF93B" box="[818,916,1712,1738]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[136,235,1516,1540]" captionStartId-1="3.[136,235,1516,1540]" captionStartId-2="4.[136,235,861,885]" captionTargetBox-1="[362,1225,189,1493]" captionTargetId-0="figure-269@2.[365,1222,189,1493]" captionTargetId-1="figure-244@3.[362,1225,189,1493]" captionTargetId-2="figure-634@4.[280,1308,189,837]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Docidiadia grimaldii sp. nov., female holotype, habitus." captionText-1="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." captionText-2="FIGURE 3. Docidiadia grimaldii sp. nov., female paratype, habitus." figureDoi-0="http://doi.org/10.5281/zenodo.7382648" figureDoi-1="http://doi.org/10.5281/zenodo.7333655" figureDoi-2="http://doi.org/10.5281/zenodo.7382650" httpUri-0="https://zenodo.org/record/7382648/files/figure.png" httpUri-1="https://zenodo.org/record/7333655/files/figure.png" httpUri-2="https://zenodo.org/record/7382650/files/figure.png" pageId="1" pageNumber="506">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3766519361FFFD9B10B2AF0FB5EF831" pageId="1" pageNumber="506" type="materials_examined">
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<paragraph id="8BD33692361FFFD9B10B2AF0FB5EF831" blockId="1.[810,1452,1782,2093]" pageId="1" pageNumber="506">
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<emphasis id="B918EA80361FFFD9B10B2AF0FC06F8E1" bold="true" box="[810,921,1782,1808]" pageId="1" pageNumber="506">Material.</emphasis>
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<materialsCitation id="3B043CCF361FFFD9B1BF2AF0FA37F8A6" ID-GBIF-Occurrence="3978642302" collectionCode="LACM" pageId="1" pageNumber="506" specimenCode="ENT 479021" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D78830361FFFD9B1BF2AF0FB96F8E1" box="[926,1033,1782,1808]" pageId="1" pageNumber="506" type="holotype">Holotype</typeStatus>
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, male,
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<collectionCode id="ED7DAE57361FFFD9B6772AF0FB38F8E1" box="[1110,1191,1782,1808]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="506" type="Museum">LACM</collectionCode>
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<specimenCode id="DBCA9EE9361FFFD9B68D2AF1FAA4F8E1" box="[1196,1339,1782,1809]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="506" type="Museum">ENT 479021</specimenCode>
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(
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<collectionCode id="ED7DAE57361FFFD9B7682AF0FA3FF8E1" box="[1353,1440,1782,1808]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="506" type="Museum">LACM</collectionCode>
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), virtually complete specimen in a greenish yellow trapezoid piece of amber trimmed to 25 × 13 ×
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<quantity id="4C949B77361FFFD9B69D2B3AFB64F8A6" box="[1212,1275,1852,1879]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="506" unit="mm" value="5.0">5 mm</quantity>
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(
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<figureCitation id="13572A17361FFFD9B7262B3BFAD3F8A6" box="[1287,1356,1853,1879]" captionStart="FIGURE 1" captionStartId="2.[136,235,1516,1540]" captionTargetId="figure-269@2.[365,1222,189,1493]" captionTargetPageId="2" captionText="FIGURE 1. Docidiadia grimaldii sp. nov., female holotype, habitus." figureDoi="http://doi.org/10.5281/zenodo.7382648" httpUri="https://zenodo.org/record/7382648/files/figure.png" pageId="1" pageNumber="506">Figs 1</figureCitation>
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,
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<figureCitation id="13572A17361FFFD9B7792B3BFA00F8A6" box="[1368,1439,1853,1879]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="1" pageNumber="506">2A–C</figureCitation>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B043CCF361FFFD9B10B2B66FB22F831" ID-GBIF-Occurrence="3978642301" collectionCode="LACM" pageId="1" pageNumber="506" specimenCode="ENT 479022" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54D78830361FFFD9B10B2B66FC10F88B" box="[810,911,1888,1914]" pageId="1" pageNumber="506" type="paratype">Paratype</typeStatus>
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, male,
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<collectionCode id="ED7DAE57361FFFD9B1C02B66FBADF88B" box="[993,1074,1888,1914]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="506" type="Museum">LACM</collectionCode>
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<specimenCode id="DBCA9EE9361FFFD9B61A2B66FB4EF88B" box="[1083,1233,1888,1914]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="506" type="Museum">ENT 479022</specimenCode>
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, virtually complete specimen in a greenish yellow ellipsoid piece of amber trimmed to 17 × 11 ×
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<quantity id="4C949B77361FFFD9B63E2BA0FBFFF830" box="[1055,1120,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="506" unit="mm" value="4.0">4 mm</quantity>
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(
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<figureCitation id="13572A17361FFFD9B64E2BA0FB2CF831" box="[1135,1203,1958,1984]" captionStart="FIGURE 3" captionStartId="4.[136,235,861,885]" captionTargetId="figure-634@4.[280,1308,189,837]" captionTargetPageId="4" captionText="FIGURE 3. Docidiadia grimaldii sp. nov., female paratype, habitus." figureDoi="http://doi.org/10.5281/zenodo.7382650" httpUri="https://zenodo.org/record/7382650/files/figure.png" pageId="1" pageNumber="506">Fig. 3</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3766519361FFFDAB17E2BCCFD18F94F" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506" type="etymology">
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<paragraph id="8BD33692361FFFDAB17E2BCCFD75F985" blockId="1.[810,1452,1782,2093]" lastBlockId="2.[136,778,1589,2093]" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506">
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<emphasis id="B918EA80361FFFD9B17E2BCCFC79F815" bold="true" box="[863,998,1994,2020]" pageId="1" pageNumber="506">Etymology.</emphasis>
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The species name honors our good friend David Grimaldi, who has brought an extraordinary contribution to the understanding of insect evolution, dealing with large groups of extant and fossil species of different fly families and of different orders of insects.
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</paragraph>
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<caption id="DF13661A361CFFDAB2A929EAFCD9F9F5" ID-DOI="http://doi.org/10.5281/zenodo.7382648" ID-Zenodo-Dep="7382648" box="[136,838,1516,1541]" httpUri="https://zenodo.org/record/7382648/files/figure.png" pageId="2" pageNumber="507" startId="2.[136,235,1516,1540]" targetBox="[365,1222,189,1493]" targetPageId="2">
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<paragraph id="8BD33692361CFFDAB2A929EAFCD9F9F5" blockId="2.[136,838,1516,1541]" box="[136,838,1516,1541]" pageId="2" pageNumber="507">
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<emphasis id="B918EA80361CFFDAB2A929EAFE9BF9F5" bold="true" box="[136,260,1516,1540]" pageId="2" pageNumber="507">FIGURE 1.</emphasis>
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<taxonomicName id="4C6C4D11361CFFDAB32B29EAFE7EF9F5" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[266,481,1516,1540]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
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<emphasis id="B918EA80361CFFDAB32B29EAFE7EF9F5" box="[266,481,1516,1540]" italics="true" pageId="2" pageNumber="507">Docidiadia grimaldii</emphasis>
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</taxonomicName>
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<emphasis id="B918EA80361CFFDAB3C629EBFDA7F9F4" bold="true" box="[487,568,1517,1541]" pageId="2" pageNumber="507">
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<taxonomicNameLabel id="A22B57FB361CFFDAB3C629EBFDA7F9F4" box="[487,568,1517,1541]" pageId="2" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, female holotype, habitus.
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</paragraph>
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</caption>
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<paragraph id="8BD33692361CFFDAB29C2A78FD18F94F" blockId="2.[136,778,1589,2093]" pageId="2" pageNumber="507">
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<emphasis id="B918EA80361CFFDAB29C2A78FE52F969" bold="true" box="[189,461,1662,1689]" pageId="2" pageNumber="507">Locality and horizon.</emphasis>
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Known only from amber fossils from the mid-Cretaceous of
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<collectingCountry id="F37B7602361CFFDAB0352AA2FD1CF94F" box="[532,643,1700,1726]" name="Myanmar" pageId="2" pageNumber="507">Myanmar</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3766519361CFFDAB29C2ACEFD5BF7DD" pageId="2" pageNumber="507" type="diagnosis">
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<paragraph id="8BD33692361CFFDAB29C2ACEFD5BF7DD" blockId="2.[136,778,1589,2093]" pageId="2" pageNumber="507">
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<emphasis id="B918EA80361CFFDAB29C2ACEFEA7F913" bold="true" box="[189,312,1736,1762]" pageId="2" pageNumber="507">Diagnosis.</emphasis>
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Flagellum (
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<figureCitation id="13572A17361CFFDAB3EE2ACEFDB1F913" box="[463,558,1736,1762]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="2" pageNumber="507">Fig. 2A</figureCitation>
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) slightly wider at base, gradually slenderer towards apex, 15 flagellomeres, last flagellomere 8× longer than wide, first seven flagellomeres at least slightly wider than long. First sector of Rs long (
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<figureCitation id="13572A17361CFFDAB32D2B5DFEFCF884" box="[268,355,1883,1909]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="2" pageNumber="507">Fig. 2B</figureCitation>
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), oblique, originating at basal third of wing; cell r1 wide. Veins r-m, base of M
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<subScript id="17E834D7361CFFDAB0712B94FDC7F851" attach="left" box="[592,600,1938,1952]" fontSize="6" pageId="2" pageNumber="507">4</subScript>
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and m-cu more or less aligned and pretty much transverse. Distal fifth of M
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<subScript id="17E834D7361CFFDAB2802BDDFF36F818" attach="left" box="[161,169,2011,2025]" fontSize="6" pageId="2" pageNumber="507">4</subScript>
|
||
, but especially distally fourth of CuA strongly bent posteriorly. Female terminalia (
|
||
<figureCitation id="13572A17361CFFDAB3CF2BE8FDD9F7F9" box="[494,582,2030,2056]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="2" pageNumber="507">Fig. 2C</figureCitation>
|
||
) with long distal lobes of sternite 8; cercus with tip directed distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3766519361CFFDCB17E2A34FEE4F815" lastPageId="4" lastPageNumber="509" pageId="2" pageNumber="507" type="description">
|
||
<paragraph id="8BD33692361CFFDAB17E2A34FB54F9BC" blockId="2.[810,1452,1586,2081]" box="[863,1227,1586,1613]" pageId="2" pageNumber="507">
|
||
<emphasis id="B918EA80361CFFDAB17E2A34FC6EF9BD" bold="true" box="[863,1009,1586,1612]" pageId="2" pageNumber="507">Description.</emphasis>
|
||
Female (
|
||
<figureCitation id="13572A17361CFFDAB6782A34FB00F9BC" box="[1113,1183,1586,1613]" captionStart="FIGURE 1" captionStartId="2.[136,235,1516,1540]" captionTargetId="figure-269@2.[365,1222,189,1493]" captionTargetPageId="2" captionText="FIGURE 1. Docidiadia grimaldii sp. nov., female holotype, habitus." figureDoi="http://doi.org/10.5281/zenodo.7382648" httpUri="https://zenodo.org/record/7382648/files/figure.png" pageId="2" pageNumber="507">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13572A17361CFFDAB68C2A35FB22F9BC" box="[1197,1213,1587,1613]" captionStart="FIGURE 3" captionStartId="4.[136,235,861,885]" captionTargetId="figure-634@4.[280,1308,189,837]" captionTargetPageId="4" captionText="FIGURE 3. Docidiadia grimaldii sp. nov., female paratype, habitus." figureDoi="http://doi.org/10.5281/zenodo.7382650" httpUri="https://zenodo.org/record/7382650/files/figure.png" pageId="2" pageNumber="507">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD33692361CFFDBB17E2A50FDCBF916" blockId="2.[810,1452,1586,2081]" lastBlockId="3.[136,778,1597,2091]" lastPageId="3" lastPageNumber="508" pageId="2" pageNumber="507">
|
||
<emphasis id="B918EA80361CFFDAB17E2A50FC03F981" box="[863,924,1622,1648]" italics="true" pageId="2" pageNumber="507">Head</emphasis>
|
||
(
|
||
<figureCitation id="13572A17361CFFDAB18F2A50FB95F980" box="[942,1034,1622,1649]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="2" pageNumber="507">Fig. 2A</figureCitation>
|
||
). Eyes not large, occupying most of lateral surface of head, no gena; eyes largely separated from each other above antennae, no dorso-ventral differentiation of ommatidia, ommatricha not visible. Three ocelli, placed on a prominent mound; no strong ocellar seta between ocelli. Labellum and palpus well developed, three palpomeres, second palpomere pyriform, about as long as distal palpomere, distal palpomere elongate, with apical setulae. Antenna with short scape, about half of pedicel length, with some few setulae distally, pedicel longer than flagellomere 1, with a whorl of setulae around distal margin. Flagellum with 15 flagellomeres, no scales, first flagellomere about twice as wide as long, gradually slenderer and elongate towards apex; flagellomeres 2–8 slightly wider than long, flagellomeres 9–14 as wide as long or slightly longer than wide, last flagellomere long, slender, style-like, 8× longer than wide, apparently a subdivision of the previous flagellomere.
|
||
</paragraph>
|
||
<caption id="DF13661A361DFFDBB2A929EAFAD5F9F5" ID-DOI="http://doi.org/10.5281/zenodo.7333655" ID-Zenodo-Dep="7333655" box="[136,1354,1516,1541]" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="3" pageNumber="508" startId="3.[136,235,1516,1540]" targetBox="[362,1225,189,1493]" targetPageId="3">
|
||
<paragraph id="8BD33692361DFFDBB2A929EAFAD5F9F5" blockId="3.[136,1354,1516,1541]" box="[136,1354,1516,1541]" pageId="3" pageNumber="508">
|
||
<emphasis id="B918EA80361DFFDBB2A929EAFE9BF9F5" bold="true" box="[136,260,1516,1540]" pageId="3" pageNumber="508">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6C4D11361DFFDBB32B29EAFE7EF9F5" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[266,481,1516,1540]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361DFFDBB32B29EAFE7EF9F5" box="[266,481,1516,1540]" italics="true" pageId="3" pageNumber="508">Docidiadia grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361DFFDBB3C629EBFDA7F9F4" bold="true" box="[487,568,1517,1541]" pageId="3" pageNumber="508">
|
||
<taxonomicNameLabel id="A22B57FB361DFFDBB3C629EBFDA7F9F4" box="[487,568,1517,1541]" pageId="3" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype.
|
||
<emphasis id="B918EA80361DFFDBB0D429EAFC98F9F5" bold="true" box="[757,775,1516,1540]" pageId="3" pageNumber="508">A</emphasis>
|
||
, Head and thorax.
|
||
<emphasis id="B918EA80361DFFDBB1E929EBFC46F9F4" bold="true" box="[968,985,1517,1541]" pageId="3" pageNumber="508">B</emphasis>
|
||
, Wing.
|
||
<emphasis id="B918EA80361DFFDBB60B29EAFBA3F9F5" bold="true" box="[1066,1084,1516,1540]" pageId="3" pageNumber="508">C</emphasis>
|
||
, Abdomen and terminalia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD33692361DFFDBB29C2AF7FB76F8DF" blockId="3.[136,778,1597,2091]" lastBlockId="3.[810,1452,1595,2093]" pageId="3" pageNumber="508">
|
||
<emphasis id="B918EA80361DFFDBB29C2AF7FE91F8FA" box="[189,270,1777,1803]" italics="true" pageId="3" pageNumber="508">Thorax</emphasis>
|
||
. Scutum short, strongly arched, high at anterior end, curved downwards at posterior half, scutellum small. Scattered longer setae over scutum in addition to a pair of rows of slightly longer dorso-central setae, scutellum with a couple of slightly divergent long setae at posterior margin, no setulae on dorsal surface of scutellar disc. Halteres long, reaching almost posterior margin of second abdominal segment. Antepronotum short, proepisternum long, propepimeron short, triangular, clearly separated from katepisternum. Anepisternum with a deep dorsal membranous notch; katepisternum elongate dorsoventrally. Mesepimeron wide, reaching ventral margin of thoracic pleura. Laterotergite not bulging, mediotergite slightly curved. Metepisternum relatively large. Apparently only antepronotum and proepisternum with elongate setae, other sclerites bare.
|
||
</paragraph>
|
||
<paragraph id="8BD33692361DFFDCB17E2B3CFF59FBC3" blockId="3.[810,1452,1595,2093]" lastBlockId="4.[136,778,939,2093]" lastPageId="4" lastPageNumber="509" pageId="3" pageNumber="508">
|
||
<emphasis id="B918EA80361DFFDBB17E2B3CFC0BF8A2" box="[863,916,1850,1875]" italics="true" pageId="3" pageNumber="508">Legs</emphasis>
|
||
. Coxae elongate, all of same length. Femora and tibiae thin, long, hind tibia 1.4 longer than femur. Femora with scattered fine setae besides a short row of ventral setae at distal end. Tibiae thin, no distal differentiated area of regular comb of setae at distal inner end of front tibia; mid tibia with a long row of longer thin setae; hind tibia with a row of five stronger dorsal setae. First tarsomeres more than twice length of second tarsomere. Tibial spurs 1:2:2; spurs finely pilose, slender, about twice as long as width of tibiae at apex. Tarsal claws small, apparently no teeth.
|
||
</paragraph>
|
||
<caption id="DF13661A361AFFDCB2A92F5BFCDAFC84" ID-DOI="http://doi.org/10.5281/zenodo.7382650" ID-Zenodo-Dep="7382650" box="[136,837,861,885]" httpUri="https://zenodo.org/record/7382650/files/figure.png" pageId="4" pageNumber="509" startId="4.[136,235,861,885]" targetBox="[280,1308,189,837]" targetPageId="4">
|
||
<paragraph id="8BD33692361AFFDCB2A92F5BFCDAFC84" blockId="4.[136,837,861,885]" box="[136,837,861,885]" pageId="4" pageNumber="509">
|
||
<emphasis id="B918EA80361AFFDCB2A92F5BFE9BFC84" bold="true" box="[136,260,861,885]" pageId="4" pageNumber="509">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6C4D11361AFFDCB32B2F5BFE7EFC84" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[266,481,861,885]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361AFFDCB32B2F5BFE7EFC84" box="[266,481,861,885]" italics="true" pageId="4" pageNumber="509">Docidiadia grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361AFFDCB3C62F5BFDA7FC84" bold="true" box="[487,568,861,885]" pageId="4" pageNumber="509">
|
||
<taxonomicNameLabel id="A22B57FB361AFFDCB3C62F5BFDA7FC84" box="[487,568,861,885]" pageId="4" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female paratype, habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD33692361AFFDCB29C2838FF59F96D" blockId="4.[136,778,939,2093]" pageId="4" pageNumber="509">
|
||
<emphasis id="B918EA80361AFFDCB29C2838FF68FBA6" box="[189,247,1086,1111]" italics="true" pageId="4" pageNumber="509">Wing</emphasis>
|
||
(
|
||
<figureCitation id="13572A17361AFFDCB329283BFEFCFBA6" box="[264,355,1085,1111]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="4" pageNumber="509">Fig. 2B</figureCitation>
|
||
). Elliptical,
|
||
<quantity id="4C949B77361AFFDCB3D1283BFDD7FBA6" box="[496,584,1085,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="509" unit="mm" value="1.8">1.8 mm</quantity>
|
||
long. C, Sc, and radial veins more sclerotized, posterior veins fainter. Macrotrichia on bR, R
|
||
<subScript id="17E834D7361AFFDCB3A9289EFE0FFB57" attach="left" box="[392,400,1176,1190]" fontSize="6" pageId="4" pageNumber="509">1</subScript>
|
||
, second sector of Rs, r-m and on distal end of veins M
|
||
<subScript id="17E834D7361AFFDCB35628BAFEE0FB3B" attach="left" box="[375,383,1212,1226]" fontSize="6" pageId="4" pageNumber="509">1</subScript>
|
||
, M
|
||
<subScript id="17E834D7361AFFDCB38728BAFE31FB3B" attach="left" box="[422,430,1212,1226]" fontSize="6" pageId="4" pageNumber="509">2</subScript>
|
||
, M
|
||
<subScript id="17E834D7361AFFDCB3F728BAFE41FB3B" attach="left" box="[470,478,1212,1226]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
and CuA, no macrotrichia on wing membrane. Sc incomplete, ending before level of origin of Rs. C extending beyond apex of R
|
||
<subScript id="17E834D7361AFFDCB05C2903FD1AFAE2" attach="left" box="[637,645,1285,1299]" fontSize="6" pageId="4" pageNumber="509">5</subScript>
|
||
for about a third of distance to apex of M
|
||
<subScript id="17E834D7361AFFDCB3F6292FFE40FAC6" attach="left" box="[471,479,1321,1335]" fontSize="6" pageId="4" pageNumber="509">1</subScript>
|
||
. bR forking at about basal third of wing, cell r1 wide, first sector of Rs oblique, long, more weakly sclerotized; R
|
||
<subScript id="17E834D7361AFFDCB39E2974FE58FA71" attach="left" box="[447,455,1394,1408]" fontSize="6" pageId="4" pageNumber="509">1</subScript>
|
||
long, reaching almost distal 2/3 of wing length; R
|
||
<subScript id="17E834D7361AFFDCB3542990FEE2FA55" attach="left" box="[373,381,1430,1444]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
absent; r-m almost transverse. M
|
||
<subScript id="17E834D7361AFFDCB0D12990FC96FA55" attach="left" box="[752,777,1430,1444]" fontSize="6" pageId="4" pageNumber="509">1+2</subScript>
|
||
and M
|
||
<subScript id="17E834D7361AFFDCB2EE29BDFF48FA38" attach="left" box="[207,215,1467,1481]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
running almost parallel, no sign of bM; connection of base of M
|
||
<subScript id="17E834D7361AFFDCB30329D9FEB5FA1C" attach="left" box="[290,298,1503,1517]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
and m-cu aligned with r-m. Medial fork slightly longer than M
|
||
<subScript id="17E834D7361AFFDCB3BD2A05FE2AF9E0" attach="left" box="[412,437,1539,1553]" fontSize="6" pageId="4" pageNumber="509">1+2</subScript>
|
||
. Distal fifth of M
|
||
<subScript id="17E834D7361AFFDCB0BF2A05FD39F9E0" attach="left" box="[670,678,1539,1553]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
slightly arched posteriorly, distal fourth of CuA strongly arched posteriorly. CuP present, incomplete, ending at about two thirds of distance to margin. Anal lobe rounded; no visible alula.
|
||
</paragraph>
|
||
<paragraph id="8BD33692361AFFDCB29C2AA1FD98F8FB" blockId="4.[136,778,939,2093]" pageId="4" pageNumber="509">
|
||
<emphasis id="B918EA80361AFFDCB29C2AA1FEB8F930" box="[189,295,1703,1729]" italics="true" pageId="4" pageNumber="509">Abdomen</emphasis>
|
||
. Tergites and sternites well sclerotized, with scattered setae, segments 1 to 8 visible, segment 8 less than half length of segment 7.
|
||
</paragraph>
|
||
<paragraph id="8BD33692361AFFDCB29C2B12FD58F84E" blockId="4.[136,778,939,2093]" pageId="4" pageNumber="509">
|
||
<emphasis id="B918EA80361AFFDCB29C2B12FEA8F8DF" box="[189,311,1812,1838]" italics="true" pageId="4" pageNumber="509">Terminalia</emphasis>
|
||
(
|
||
<figureCitation id="13572A17361AFFDCB3662B12FE3EF8DF" box="[327,417,1812,1838]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="4" pageNumber="509">Fig. 2C</figureCitation>
|
||
). Sternite 8 with a pair of large, more or less pointed setose lobes with a deep medial incision. Tergite 9 wide, about half the length of segment 7, cercus apparently 1-segmented, large, setose, projected more distally than level of tip of lobes of sternite 8.
|
||
</paragraph>
|
||
<paragraph id="8BD33692361AFFDCB29C2BCCFEE4F815" blockId="4.[136,778,939,2093]" box="[189,379,1994,2020]" pageId="4" pageNumber="509">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3766519361AFFDDB29C2BE8FB4AF99A" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509" type="discussion">
|
||
<paragraph id="8BD33692361AFFDCB29C2BE8FC0DFBFF" blockId="4.[136,778,939,2093]" lastBlockId="4.[810,1452,939,2099]" pageId="4" pageNumber="509">
|
||
<emphasis id="B918EA80361AFFDCB29C2BE8FEAEF7F9" bold="true" box="[189,305,2030,2056]" pageId="4" pageNumber="509">Remarks.</emphasis>
|
||
Both specimens are clearly conspecific. Not only the wing venation of both specimens is identical in details, but the shape of the unusual structure of the female terminalia in both specimens is basically identical.
|
||
</paragraph>
|
||
<paragraph id="8BD33692361AFFDCB17E281EFC72F86A" blockId="4.[810,1452,939,2099]" pageId="4" pageNumber="509">
|
||
<emphasis id="B918EA80361AFFDCB17E281EFC7BFBC3" bold="true" box="[863,996,1048,1074]" pageId="4" pageNumber="509">Discussion.</emphasis>
|
||
In general terms,
|
||
<taxonomicName id="4C6C4D11361AFFDCB69A281EFA34FBC3" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[1211,1451,1048,1074]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361AFFDCB69A281EFA34FBC3" box="[1211,1451,1048,1074]" italics="true" pageId="4" pageNumber="509">Docidiadia grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361AFFDCB10B283BFC1AFBA6" bold="true" box="[810,901,1085,1111]" pageId="4" pageNumber="509">
|
||
<taxonomicNameLabel id="A22B57FB361AFFDCB10B283BFC1AFBA6" box="[810,901,1085,1111]" pageId="4" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is very similar to the type-species of the genus,
|
||
<taxonomicName id="4C6C4D11361AFFDCB10B2867FAA7FB8A" authority="Blagoderov & Grimaldi" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[810,1336,1121,1147]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB10B2867FB81FB8A" box="[810,1054,1121,1147]" italics="true" pageId="4" pageNumber="509">Docidiadia burmitica</emphasis>
|
||
Blagoderov & Grimaldi
|
||
</taxonomicName>
|
||
and there is no question about the species being congeneric. The differences indicating that they are separate species are evident in different aspects of the morphology. In the antenna (
|
||
<figureCitation id="13572A17361AFFDCB1AF28F4FC7AFAFC" box="[910,997,1266,1293]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="4" pageNumber="509">Fig. 2A</figureCitation>
|
||
), the flagellomeres of the distal half of the antenna are more slender in
|
||
<taxonomicName id="4C6C4D11361AFFDCB643291EFB74FAC0" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[1122,1259,1303,1329]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB643291EFBE9FAC0" box="[1122,1142,1304,1329]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB6A22911FB74FAC0" box="[1155,1259,1303,1329]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C6C4D11361AFFDCB737291EFA3BFAC0" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1302,1444,1303,1329]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB737291EFAB5FAC0" box="[1302,1322,1304,1329]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB7162911FA3BFAC0" box="[1335,1444,1303,1329]" italics="true" pageId="4" pageNumber="509">burmitica</emphasis>
|
||
</taxonomicName>
|
||
, the distal palpomere seems to be slightly longer than in
|
||
<taxonomicName id="4C6C4D11361AFFDCB10B2966FC2AFA88" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[810,949,1375,1402]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB10B2966FCA1FA88" box="[810,830,1376,1401]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB16C2959FC2AFA88" box="[845,949,1375,1401]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
. R
|
||
<quantity id="4C949B77361AFFDCB1F62974FC62FA88" box="[983,1021,1375,1408]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="509" unit="in" value="1.0">
|
||
<subScript id="17E834D7361AFFDCB1F62974FC40FA71" attach="left" box="[983,991,1394,1408]" fontSize="6" pageId="4" pageNumber="509">1</subScript>
|
||
in
|
||
</quantity>
|
||
<taxonomicName id="4C6C4D11361AFFDCB6252966FB10FA88" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[1028,1167,1375,1402]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB6252966FB87FA88" box="[1028,1048,1376,1401]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB6062959FB10FA88" box="[1063,1167,1375,1401]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
is slightly longer than in
|
||
<taxonomicName id="4C6C4D11361AFFDCB10B2983FC23FA6F" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[810,956,1412,1438]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB10B2983FCA1FA6F" box="[810,830,1413,1438]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB16E2982FC23FA6F" box="[847,956,1412,1438]" italics="true" pageId="4" pageNumber="509">burmitica</emphasis>
|
||
</taxonomicName>
|
||
, while the sequence of veins r-m/M
|
||
<subScript id="17E834D7361AFFDCB7432990FAF5FA55" attach="left" box="[1378,1386,1430,1444]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
/m-cu (
|
||
<figureCitation id="13572A17361AFFDCB11329AEFC16FA33" box="[818,905,1448,1474]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="4" pageNumber="509">Fig. 2B</figureCitation>
|
||
) is more transverse in
|
||
<taxonomicName id="4C6C4D11361AFFDCB6AC29AFFA82FA33" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1165,1309,1448,1475]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB6AC29AFFB3EFA33" box="[1165,1185,1449,1474]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB69129AEFA82FA33" box="[1200,1309,1448,1474]" italics="true" pageId="4" pageNumber="509">burmitica</emphasis>
|
||
</taxonomicName>
|
||
, while in
|
||
<taxonomicName id="4C6C4D11361AFFDCB7B129AFFC0DFA16" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB7B129AFFA3BFA33" box="[1424,1444,1449,1474]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB10B29CBFC0DFA16" box="[810,914,1485,1511]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
it is slightly oblique, especially the first sector of M
|
||
<subScript id="17E834D7361AFFDCB1402A05FCF6F9E0" attach="left" box="[865,873,1539,1553]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
and m-cu. The type series of
|
||
<taxonomicName id="4C6C4D11361AFFDCB69829F7FA34F9FA" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1209,1451,1521,1547]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB69829F7FA34F9FA" box="[1209,1451,1521,1547]" italics="true" pageId="4" pageNumber="509">Docidiadia burmitica</emphasis>
|
||
</taxonomicName>
|
||
has a female
|
||
<typeStatus id="54D78830361AFFDCB19A2A10FB84F9C1" box="[955,1051,1558,1584]" pageId="4" pageNumber="509" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="13572A17361AFFDCB60B2A13FBF1F9DE" box="[1066,1134,1557,1583]" captionStart="FIGURE 4" captionStartId="5.[136,235,927,951]" captionTargetBox="[280,1308,189,904]" captionTargetId="figure-528@5.[280,1308,189,904]" captionTargetPageId="5" captionText="FIGURE 4. Docidiadia burmitica, female paratype, habitus (photo credit, Vladimir Blagoderov and David Grimaldi)." figureDoi="http://doi.org/10.5281/zenodo.7333657" httpUri="https://zenodo.org/record/7333657/files/figure.png" pageId="4" pageNumber="509">Fig. 4</figureCitation>
|
||
). The cercus in
|
||
<taxonomicName id="4C6C4D11361AFFDCB73C2A10FA34F9DE" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1309,1451,1557,1584]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB73C2A10FAAEF9DE" box="[1309,1329,1558,1583]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB71F2A13FA34F9DE" box="[1342,1451,1557,1583]" italics="true" pageId="4" pageNumber="509">burmitica</emphasis>
|
||
</taxonomicName>
|
||
has the distal tip slightly projected ventrally, while in
|
||
<taxonomicName id="4C6C4D11361AFFDCB10B2A59FC29F989" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[810,950,1630,1657]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB10B2A59FCA1F989" box="[810,830,1631,1656]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB16F2A58FC29F989" box="[846,950,1630,1656]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
this tip is directed distally. In
|
||
<taxonomicName id="4C6C4D11361AFFDCB7392A59FA3BF989" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[1304,1444,1630,1657]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB7392A59FAB3F989" box="[1304,1324,1631,1656]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB71D2A58FA3BF989" box="[1340,1444,1630,1656]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
, the distal lobes of the sternite 8 seem longer than in
|
||
<taxonomicName id="4C6C4D11361AFFDCB7B12A85FC08F930" authorityName="Blagoderov & Grimaldi" authorityYear="2004" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB7B12A85FA3BF96D" box="[1424,1444,1667,1692]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB10B2AA1FC08F930" box="[810,919,1703,1729]" italics="true" pageId="4" pageNumber="509">burmitica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13572A17361AFFDCB1862AA1FB9EF930" box="[935,1025,1703,1729]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="4" pageNumber="509">Fig. 2C</figureCitation>
|
||
). The clearest feature separating both species, however, is the distal end of the flagellum: the 14
|
||
<superScript id="7C199BDA361AFFDCB7BE2ACDFA33F928" attach="none" box="[1439,1452,1739,1753]" fontSize="6" pageId="4" pageNumber="509">th</superScript>
|
||
flagellomere in
|
||
<taxonomicName id="4C6C4D11361AFFDCB1C32AF6FBEBF8F8" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[994,1140,1775,1802]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361AFFDCB1C32AF6FC69F8F8" box="[994,1014,1776,1801]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB6262AE9FBEBF8F8" box="[1031,1140,1775,1801]" italics="true" pageId="4" pageNumber="509">burmitica</emphasis>
|
||
</taxonomicName>
|
||
is “secondarily segmented in two parts” (
|
||
<bibRefCitation id="EFFD4B63361AFFDCB1F12B12FAD3F8DF" author="Blagoderov, V. A. & Grimaldi, D." box="[976,1356,1812,1838]" pageId="4" pageNumber="509" pagination="1 - 76" refId="ref4104" refString="Blagoderov, V. A. & Grimaldi, D. (2004) Fossil Sciaroidea (Diptera) in Cretaceous ambers, exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae. American Museum Novitates, 3433, 1 - 76. https: // doi. org / 10.1206 / 0003 - 0082 (2004) 433 <0001:" type="journal article" year="2004">Blagoderov & Grimaldi, 2004: 7</bibRefCitation>
|
||
), with a short distal segment, while in
|
||
<taxonomicName id="4C6C4D11361AFFDCB6A32B3FFA91F8A3" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[1154,1294,1848,1875]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B918EA80361AFFDCB6A32B3FFB09F8A3" box="[1154,1174,1849,1874]" italics="true" pageId="4" pageNumber="509">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361AFFDCB6872B3EFA91F8A3" box="[1190,1294,1848,1874]" italics="true" pageId="4" pageNumber="509">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13572A17361AFFDCB73E2B3EFAE6F8A3" box="[1311,1401,1848,1874]" captionStart="FIGURE 2" captionStartId="3.[136,235,1516,1540]" captionTargetBox="[362,1225,189,1493]" captionTargetId="figure-244@3.[362,1225,189,1493]" captionTargetPageId="3" captionText="FIGURE 2. Docidiadia grimaldii sp. nov., female holotype. A, Head and thorax. B, Wing. C, Abdomen and terminalia." figureDoi="http://doi.org/10.5281/zenodo.7333655" httpUri="https://zenodo.org/record/7333655/files/figure.png" pageId="4" pageNumber="509">Fig. 2A</figureCitation>
|
||
) the subdivided distal segment is very long, about eight times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BD33692361AFFDDB17E2BA3FE9DFA6F" blockId="4.[810,1452,939,2099]" lastBlockId="5.[136,778,1005,2068]" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509">
|
||
This second species of
|
||
<taxonomicName id="4C6C4D11361AFFDCB6562BA3FB6AF84E" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1143,1269,1957,1983]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361AFFDCB6562BA3FB6AF84E" box="[1143,1269,1957,1983]" italics="true" pageId="4" pageNumber="509">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
shows that the association of the genus to the
|
||
<taxonomicName id="4C6C4D11361AFFDCB69F2BCCFAC4F815" authorityName="Winnertz" authorityYear="1863" box="[1214,1371,1994,2020]" class="Insecta" family="Diadocidiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="family">Diadocidiidae</taxonomicName>
|
||
needs additional corroboration. Among the extant families of Sciaroidea, r-m, M
|
||
<subScript id="17E834D7361AFFDCB6222423FB94F7C2" attach="left" box="[1027,1035,2085,2099]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
|
||
and m-cu are perfectly aligned and transverse in the diadocidiids, and there is a short stump of bM projecting basically from the anterior end of M
|
||
<subScript id="17E834D7361BFFDDB0982822FD4DFBC3" attach="left" box="[697,722,1060,1074]" fontSize="6" pageId="5" pageNumber="510">1+2</subScript>
|
||
. The stump of bM is absent in both species of
|
||
<taxonomicName id="4C6C4D11361BFFDDB0442831FD7CFBA0" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[613,739,1079,1105]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB0442831FD7CFBA0" box="[613,739,1079,1105]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C6C4D11361BFFDDB2A9285AFEE7FB87" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[136,376,1116,1142]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361BFFDDB2A9285AFEE7FB87" box="[136,376,1116,1142]" italics="true" pageId="5" pageNumber="510">Docidiadia burmitica</emphasis>
|
||
</taxonomicName>
|
||
, r-m and the base of M
|
||
<subScript id="17E834D7361BFFDDB0532868FDE5FB8D" attach="left" box="[626,634,1134,1148]" fontSize="6" pageId="5" pageNumber="510">4</subScript>
|
||
are perfectly aligned and transverse, with m-cu slightly more oblique. In
|
||
<taxonomicName id="4C6C4D11361BFFDDB28828A1FEA9FB31" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" box="[169,310,1190,1216]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361BFFDDB28828A1FF22FB31" box="[169,189,1191,1216]" italics="true" pageId="5" pageNumber="510">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361BFFDDB2EF28A0FEA9FB31" box="[206,310,1190,1216]" italics="true" pageId="5" pageNumber="510">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361BFFDDB31E28A0FE03FB31" bold="true" box="[319,412,1190,1216]" pageId="5" pageNumber="510">
|
||
<taxonomicNameLabel id="A22B57FB361BFFDDB31E28A0FE03FB31" box="[319,412,1190,1216]" pageId="5" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, r-m is more or less transverse, but the first section of M
|
||
<subScript id="17E834D7361BFFDDB3BC28DBFE3AFB1A" attach="left" box="[413,421,1245,1259]" fontSize="6" pageId="5" pageNumber="510">4</subScript>
|
||
and m-cu are oblique, in such a way that the transverse condition of the set of these three veins is not seen. The distal half of CuA in both species of
|
||
<taxonomicName id="4C6C4D11361BFFDDB286293CFEBAFAA5" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[167,293,1338,1364]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB286293CFEBAFAA5" box="[167,293,1338,1364]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
is particularly different from the condition of CuA in
|
||
<taxonomicName id="4C6C4D11361BFFDDB3202959FEE0FA88" authorityName="Ruthe" authorityYear="1831" box="[257,383,1375,1401]" class="Insecta" family="Diadocidiidae" genus="Diadocidia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB3202959FEE0FA88" box="[257,383,1375,1401]" italics="true" pageId="5" pageNumber="510">Diadocidia</emphasis>
|
||
</taxonomicName>
|
||
, with M
|
||
<subScript id="17E834D7361BFFDDB3C02977FE76FA8E" attach="left" box="[481,489,1393,1407]" fontSize="6" pageId="5" pageNumber="510">4</subScript>
|
||
rather strongly diverging from CuA.
|
||
</paragraph>
|
||
<caption id="DF13661A361BFFDDB2A92F99FAA8FC46" ID-DOI="http://doi.org/10.5281/zenodo.7333657" ID-Zenodo-Dep="7333657" box="[136,1335,927,952]" httpUri="https://zenodo.org/record/7333657/files/figure.png" pageId="5" pageNumber="510" startId="5.[136,235,927,951]" targetBox="[280,1308,189,904]" targetPageId="5">
|
||
<paragraph id="8BD33692361BFFDDB2A92F99FAA8FC46" blockId="5.[136,1335,927,952]" box="[136,1335,927,952]" pageId="5" pageNumber="510">
|
||
<emphasis id="B918EA80361BFFDDB2A92F99FE9BFC46" bold="true" box="[136,260,927,951]" pageId="5" pageNumber="510">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C6C4D11361BFFDDB32B2F99FE79FC46" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[266,486,927,951]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="burmitica">
|
||
<emphasis id="B918EA80361BFFDDB32B2F99FE79FC46" box="[266,486,927,951]" italics="true" pageId="5" pageNumber="510">Docidiadia burmitica</emphasis>
|
||
</taxonomicName>
|
||
, female paratype, habitus (photo credit, Vladimir Blagoderov and David Grimaldi).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD33692361BFFDDB29C29AFFE7DF81F" blockId="5.[136,778,1005,2068]" pageId="5" pageNumber="510">
|
||
On the other hand, the antenna of
|
||
<taxonomicName id="4C6C4D11361BFFDDB0AD29AFFF6FFA19" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361BFFDDB0AD29AFFF6FFA19" italics="true" pageId="5" pageNumber="510">Docidiadia grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361BFFDDB2DF29C8FEC0FA19" bold="true" box="[254,351,1486,1512]" pageId="5" pageNumber="510">
|
||
<taxonomicNameLabel id="A22B57FB361BFFDDB2DF29C8FEC0FA19" box="[254,351,1486,1512]" pageId="5" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to the condition seen in
|
||
<taxonomicName id="4C6C4D11361BFFDDB2A929F5FE86F9FC" authorityName="Rohdendorf" authorityYear="1962" box="[136,281,1523,1549]" class="Insecta" family="Archizelmiridae" genus="Archizelmira" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB2A929F5FE86F9FC" box="[136,281,1523,1549]" italics="true" pageId="5" pageNumber="510">Archizelmira</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B918EA80361BFFDDB37129F5FE44F9FC" box="[336,475,1523,1549]" italics="true" pageId="5" pageNumber="510">Zelmirarcha</emphasis>
|
||
, in the
|
||
<taxonomicName id="4C6C4D11361BFFDDB00F29F5FD77F9FC" authorityName="Rohdendorf" authorityYear="1962" box="[558,744,1523,1549]" class="Insecta" family="Archizelmiridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="family">Archizelmiridae</taxonomicName>
|
||
. In the species of
|
||
<taxonomicName id="4C6C4D11361BFFDDB3112A1EFE31F9C3" authorityName="Ruthe" authorityYear="1831" box="[304,430,1560,1586]" class="Insecta" family="Diadocidiidae" genus="Diadocidia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB3112A1EFE31F9C3" box="[304,430,1560,1586]" italics="true" pageId="5" pageNumber="510">Diadocidia</emphasis>
|
||
</taxonomicName>
|
||
, the antenna is more elongate and curved forward, typical of fungus gnats.
|
||
<taxonomicName id="4C6C4D11361BFFDDB0AD2A3BFF6FF98D" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361BFFDDB0AD2A3BFF6FF98D" italics="true" pageId="5" pageNumber="510">Docidiadia grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361BFFDDB3232A64FEF9F98D" bold="true" box="[258,358,1634,1660]" pageId="5" pageNumber="510">
|
||
<taxonomicNameLabel id="A22B57FB361BFFDDB3232A64FEF9F98D" box="[258,358,1634,1660]" pageId="5" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a straight flagellum and the flagellomeres are slightly wider than long at the basal half of the flagellum and gradually more slender on the distal half, slightly longer than wide. In
|
||
<taxonomicName id="4C6C4D11361BFFDDB0582AD7FD98F8E1" authority="Rohdendorf, 1962" authorityName="Rohdendorf" authorityYear="1962" class="Insecta" family="Archizelmiridae" genus="Archizelmira" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="kazhakstanica">
|
||
<emphasis id="B918EA80361BFFDDB0582AD7FEB8F8E1" italics="true" pageId="5" pageNumber="510">Archizelmira kazhakstanica</emphasis>
|
||
Rohdendorf, 1962
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFFD4B63361BFFDDB0702AF0FF5BF8C4" author="Grimaldi, D. & Amorim, D. S. & Blagoderov, V." pageId="5" pageNumber="510" pagination="368 - 381" refId="ref5343" refString="Grimaldi, D., Amorim, D. S. & Blagoderov, V. (2003) The Mesozoic family Archizelmiridae (Diptera: Insecta). Journal of Paleontology, 77 (2), 368 - 381. https: // doi. org / 10.1666 / 0022 - 3360 (2003) 077 <0368:" type="journal article" year="2003">
|
||
Grimaldi
|
||
<emphasis id="B918EA80361BFFDDB0E52AF1FD63F8E1" box="[708,764,1782,1808]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
|
||
., 2003
|
||
</bibRefCitation>
|
||
: fig. 2.4), the antenna is much less modified than in most archizelmirids, but it is indeed slightly derived, with a straight, setiform flagellum. The flagellum in
|
||
<taxonomicName id="4C6C4D11361BFFDDB0CF2B60FF6FF855" authority="De SOUZA AMORIM & BROWN, 2022" authorityName="De SOUZA AMORIM & BROWN" authorityYear="2022" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis id="B918EA80361BFFDDB0CF2B60FC9DF88E" box="[750,770,1894,1919]" italics="true" pageId="5" pageNumber="510">D</emphasis>
|
||
.
|
||
<emphasis id="B918EA80361BFFDDB2A92B8CFF6FF855" box="[136,240,1930,1956]" italics="true" pageId="5" pageNumber="510">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B918EA80361BFFDDB2D92B8CFECDF855" bold="true" box="[248,338,1930,1956]" pageId="5" pageNumber="510">
|
||
<taxonomicNameLabel id="A22B57FB361BFFDDB2D92B8CFECDF855" box="[248,338,1930,1956]" pageId="5" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in particular is similar to the flagellum of
|
||
<taxonomicName id="4C6C4D11361BFFDDB28C2BA9FD50F838" authority="Grimaldi et al., 2003" authorityName="Grimaldi" authorityYear="2003" box="[173,719,1967,1994]" class="Insecta" family="Archizelmiridae" genus="Archimelzira" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="americana">
|
||
<emphasis id="B918EA80361BFFDDB28C2BA9FE5BF838" box="[173,452,1967,1993]" italics="true" pageId="5" pageNumber="510">Archimelzira americana</emphasis>
|
||
<bibRefCitation id="EFFD4B63361BFFDDB3F02BA9FD50F838" author="Grimaldi, D. & Amorim, D. S. & Blagoderov, V." box="[465,719,1967,1994]" pageId="5" pageNumber="510" pagination="368 - 381" refId="ref5343" refString="Grimaldi, D., Amorim, D. S. & Blagoderov, V. (2003) The Mesozoic family Archizelmiridae (Diptera: Insecta). Journal of Paleontology, 77 (2), 368 - 381. https: // doi. org / 10.1666 / 0022 - 3360 (2003) 077 <0368:" type="journal article" year="2003">
|
||
Grimaldi
|
||
<emphasis id="B918EA80361BFFDDB0652BB6FDE3F838" box="[580,636,1967,1993]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
|
||
., 2003
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFFD4B63361BFFDDB2A92BD2FEE9F81F" author="Grimaldi, D. & Amorim, D. S. & Blagoderov, V." box="[136,374,2004,2031]" pageId="5" pageNumber="510" pagination="368 - 381" refId="ref5343" refString="Grimaldi, D., Amorim, D. S. & Blagoderov, V. (2003) The Mesozoic family Archizelmiridae (Diptera: Insecta). Journal of Paleontology, 77 (2), 368 - 381. https: // doi. org / 10.1666 / 0022 - 3360 (2003) 077 <0368:" type="journal article" year="2003">
|
||
Grimaldi
|
||
<emphasis id="B918EA80361BFFDDB2D52BD3FEBAF81F" box="[244,293,2004,2030]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
|
||
., 2003
|
||
</bibRefCitation>
|
||
: fig. 9.1).
|
||
</paragraph>
|
||
<paragraph id="8BD33692361BFFDDB29C2BFFFA34FABA" blockId="5.[136,778,1005,2068]" lastBlockId="5.[810,1452,1005,1643]" pageId="5" pageNumber="510">
|
||
This is not a simple and direct indication that
|
||
<taxonomicName id="4C6C4D11361BFFDDB10B2FEBFC37FBF6" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[810,936,1005,1031]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB10B2FEBFC37FBF6" box="[810,936,1005,1031]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C6C4D11361BFFDDB64E2FEBFAB6FBF6" authorityName="Rohdendorf" authorityYear="1962" box="[1135,1321,1005,1031]" class="Insecta" family="Archizelmiridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="family">Archizelmiridae</taxonomicName>
|
||
. The wing venation in both species of
|
||
<taxonomicName id="4C6C4D11361BFFDDB67D2817FB45FBDA" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1116,1242,1041,1067]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB67D2817FB45FBDA" box="[1116,1242,1041,1067]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
is definitely much more conservative than in archizelmirids. The wing in
|
||
<taxonomicName id="4C6C4D11361BFFDDB10B285FFC7FFB82" authorityName="Rohdendorf" authorityYear="1962" box="[810,992,1113,1139]" class="Insecta" family="Archizelmiridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="family">Archizelmiridae</taxonomicName>
|
||
has a slender cell r1, a short and rather transverse first sector of Rs, and there is a displacement of the point of contact of the tip of the first sector of M
|
||
<subScript id="17E834D7361BFFDDB78228B5FA34FB30" attach="left" box="[1443,1451,1203,1217]" fontSize="6" pageId="5" pageNumber="510">4</subScript>
|
||
and the base of m-cu towards the base of the wing—that produces an acute angle between the first and second sectors of M
|
||
<subScript id="17E834D7361BFFDDB1962919FC20FADC" attach="left" box="[951,959,1311,1325]" fontSize="6" pageId="5" pageNumber="510">4</subScript>
|
||
. This possibly derived condition is missing in
|
||
<taxonomicName id="4C6C4D11361BFFDDB16E2937FC52FABA" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[847,973,1329,1355]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB16E2937FC52FABA" box="[847,973,1329,1355]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
, as well as in
|
||
<taxonomicName id="4C6C4D11361BFFDDB6B92937FA89FABA" authorityName="Ruthe" authorityYear="1831" box="[1176,1302,1329,1355]" class="Insecta" family="Diadocidiidae" genus="Diadocidia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB6B92937FA89FABA" box="[1176,1302,1329,1355]" italics="true" pageId="5" pageNumber="510">Diadocidia</emphasis>
|
||
</taxonomicName>
|
||
, ditomyiids,
|
||
</paragraph>
|
||
<paragraph id="8BD33692361BFFDDB10B2953FB4AF99A" blockId="5.[810,1452,1005,1643]" pageId="5" pageNumber="510">
|
||
and some other fungus gnats, that have a more or less transverse r-m/M
|
||
<subScript id="17E834D7361BFFDDB1D2298DFC64FA68" attach="left" box="[1011,1019,1419,1433]" fontSize="6" pageId="5" pageNumber="510">4</subScript>
|
||
/m-cu. The condition in
|
||
<taxonomicName id="4C6C4D11361BFFDDB70C297FFA34FA62" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1325,1451,1401,1427]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB70C297FFA34FA62" box="[1325,1451,1401,1427]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
is, however, comparatively plesiomorphic and does not necessarily join
|
||
<taxonomicName id="4C6C4D11361BFFDDB1CE29C7FBF2FA2A" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[1007,1133,1473,1499]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB1CE29C7FBF2FA2A" box="[1007,1133,1473,1499]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
with other genera with a similar wing vein pattern. A more clear solution for the position of
|
||
<taxonomicName id="4C6C4D11361BFFDDB1952A0FFBADF9D2" authorityName="Blagoderov & Grimaldi" authorityYear="2004" box="[948,1074,1545,1571]" class="Insecta" family="Diadocidiidae" genus="Docidiadia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B918EA80361BFFDDB1952A0FFBADF9D2" box="[948,1074,1545,1571]" italics="true" pageId="5" pageNumber="510">Docidiadia</emphasis>
|
||
</taxonomicName>
|
||
amongst the extant and extinct sciaroids actually depends on an analysis of the Sciaroidea as a whole which is still not available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |