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<document id="00954B19A69536C5AB8EEC67CA5B3CE2" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E36EE21DFE6DA34B5E88F973" docLanguage="en" docName="JNATHIST.53.2526.15331584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Cristispira caballasensis DeVries 2019, sp. nov." docType="treatment" docVersion="22" lastPageNumber="1570" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1570" updateTime="1698783606523" updateUser="ExternalLinkService">
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<mods:title id="F2A98B006D78F3C7417EB6591C9D1879">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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<mods:namePart id="F41CA69D5668C9D28DF9EC347996E6E0">DeVries, Thomas J.</mods:namePart>
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<mods:title id="8483487A5BE3DD2979B8452959E2CE9A">Journal of Natural History</mods:title>
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<mods:part id="288B70A2AD4ED9DB34C09FA1BCA86F96">
<mods:date id="065D714A2606C9C7E61F649853048B9A">2019</mods:date>
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<mods:number id="5F969893D9D3460944C74B2AFEC24DCB">2018-12-17</mods:number>
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<mods:number id="FBC09FEB3457B1424D5697D91F5BF01A">53</mods:number>
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<treatment id="03BB2338E36EE21DFE6DA34B5E88F973" ID-DOI="http://doi.org/10.5281/zenodo.3671276" ID-GBIF-Taxon="162146551" ID-Zenodo-Dep="3671276" LSID="urn:lsid:plazi:treatment:03BB2338E36EE21DFE6DA34B5E88F973" httpUri="http://treatment.plazi.org/id/03BB2338E36EE21DFE6DA34B5E88F973" lastPageNumber="1570" pageId="38" pageNumber="1570">
<subSubSection id="C308C1A5E36EE21DFE6DA34B5954FDED" box="[460,856,520,545]" pageId="38" pageNumber="1570" type="nomenclature">
<paragraph id="8BAD922EE36EE21DFE6DA34B5954FDED" blockId="38.[460,856,520,580]" box="[460,856,520,545]" pageId="38" pageNumber="1570">
<heading id="D0E52542E36EE21DFE6DA34B5954FDED" box="[460,856,520,545]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="1570" reason="2">
<taxonomicName id="4C12E9ADE36EE21DFE6DA34B58FEFDED" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[460,754,520,545]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="caballasensis" status="sp. nov.">
<emphasis id="B9664E3CE36EE21DFE6DA34B58FEFDED" bold="true" box="[460,754,520,545]" italics="true" pageId="38" pageNumber="1570">Cristispira caballasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36EE21DFD5AA34B5954FDED" box="[763,856,520,545]" pageId="38" pageNumber="1570" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36EE21DFDECA36858DAFD88" box="[589,726,555,580]" pageId="38" pageNumber="1570" type="description">
<paragraph id="8BAD922EE36EE21DFDECA36858DAFD88" blockId="38.[460,856,520,580]" box="[589,726,555,580]" pageId="38" pageNumber="1570">
<heading id="D0E52542E36EE21DFDECA36858DAFD88" box="[589,726,555,580]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="1570" reason="5">
(
<figureCitation id="13298EABE36EE21DFDF4A36858B7FD88" box="[597,699,555,580]" captionStart="Figure 5" captionStartId="14.[160,225,1068,1091]" captionTargetBox="[164,1150,144,1035]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 5. Gastropods from the Caballas Formation. (ae) Caballasea segmentata gen. nov., sp. nov. B8769. (a) UWBM 107607, paratype, L 12.4 mm, apertural view. (b) UWBM 107607, abapertural view. (c) UWBM 107606, holotype, L 12.6 mm, abapertural view. (d) MUSM INV 262, paratype, L 8.3 mm, oblique view of parietal callus and columella. (e) UWBM 107608, paratype, L 15.3 mm, abapertural view. (f, g) Turritella capistrata sp. nov. (f) MUSM INV 259, paratype, B8772, L 16.3 mm, lateral view. (g) UWBM 107601, holotype, B8770, L 14.4 mm, apertural view. (h) Turritella saposa Olsson, 1944, UWBM 107602, B8770, L 15.5 mm. (i) Cristispira caballasensis sp. nov., UWBM 107605, holotype, B8769, L 5.1 mm, fragment. (j, k, m, n) Rhombopsis longinquus sp. nov. (j) UWBM 107630, holotype, B8772, L 18.3 mm, apertural view, outer lip missing. (k) MUSM INV 269, paratype, B8769, L 19.9 mm, abapertural view. (m) UWBM 107631, paratype, B8772, L 22.5 mm, abapertural view. (n) UWBM 107632, paratype, B8769, L 16.4 mm, lateral view of spire and columella. (l, oq) Tristeirostoma saxaquadrata sp. nov. B8771. (l) UWBM 107636, paratype, L 10.0 mm, spire. (o) UWBM 107,635, holotype, L 7.6 mm, abapertural view. (p) UWBM 107635, lateral view of columella. (q) UWBM 107635, apertural/lateral view." figureDoi="http://doi.org/10.5281/zenodo.3670237" httpUri="https://zenodo.org/record/3670237/files/figure.png" pageId="38" pageNumber="1570">Figure 5</figureCitation>
(i))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36EE21DFF01A33D5886FD11" pageId="38" pageNumber="1570" type="diagnosis">
<paragraph id="8BAD922EE36EE21DFF01A33D5B18FD54" blockId="38.[160,1155,638,733]" box="[160,276,638,664]" pageId="38" pageNumber="1570">
<emphasis id="B9664E3CE36EE21DFF01A33D5B18FD54" bold="true" box="[160,276,638,664]" italics="true" pageId="38" pageNumber="1570">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD922EE36EE21DFF01A3E15886FD11" blockId="38.[160,1155,638,733]" pageId="38" pageNumber="1570">Hypercampanulate; three strong keeled spiral cords on anterior two-thirds of spire whorls; finely spirally threaded interspaces.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36EE21DFF01A25459D1FBCD" pageId="38" pageNumber="1570" type="description">
<paragraph id="8BAD922EE36EE21DFF01A2545B2AFCFD" blockId="38.[160,1156,791,1025]" box="[160,294,791,817]" pageId="38" pageNumber="1570">
<emphasis id="B9664E3CE36EE21DFF01A2545B2AFCFD" bold="true" box="[160,294,791,817]" italics="true" pageId="38" pageNumber="1570">Description</emphasis>
</paragraph>
<paragraph id="8BAD922EE36EE21DFF01A27859D1FBCD" blockId="38.[160,1156,791,1025]" pageId="38" pageNumber="1570">Shell small; pleural angle about 12°. Whorls hypercampanulate, sutural area deeply concave. Protoconch unknown, number of whorls in spire cannot be estimated. Spiral cords keeled; spiral formula dCBAr; primary cords about equally strong. Spacing formula C14B45A67; n = 1). Interspaces with microscopic spiral threads. Lateral growth line orthocline; posterior inflexion absent, anterior inflexion barely discernible. Lateral sinus moderately deep, apex close to spiral cord B. Base and aperture not preserved. Allmon formula?-1-M-B-OR.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36EE21DFF01A57858BFFA1E" pageId="38" pageNumber="1570" type="discussion">
<paragraph id="8BAD922EE36EE21DFF01A5785B0AFB99" blockId="38.[160,1156,1083,1490]" box="[160,262,1083,1109]" pageId="38" pageNumber="1570">
<emphasis id="B9664E3CE36EE21DFF01A5785B0AFB99" bold="true" box="[160,262,1083,1109]" italics="true" pageId="38" pageNumber="1570">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE36EE21DFF01A51C58BFFA1E" blockId="38.[160,1156,1083,1490]" pageId="38" pageNumber="1570">
The elements of the spiral sculpture of
<taxonomicName id="4C12E9ADE36EE21DFDD5A51C5974FBBB" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[628,888,1119,1143]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="caballasensis" status="sp. nov.">
<emphasis id="B9664E3CE36EE21DFDD5A51C5974FBBB" box="[628,888,1119,1143]" italics="true" pageId="38" pageNumber="1570">Cristispira caballasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36EE21DFC23A51C59ECFBBB" box="[898,992,1119,1143]" pageId="38" pageNumber="1570" rank="species">sp. nov.</taxonomicNameLabel>
, evident even from the partial single specimen available, are nearly identical with those of two Peruvian species, the Late Cretaceous
<taxonomicName id="4C12E9ADE36EE21DFDDCA5E659BFFB71" authority="(Olssoni 1944)" baseAuthorityName="Olssoni" baseAuthorityYear="1944" box="[637,947,1189,1213]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="prechira">
<emphasis id="B9664E3CE36EE21DFDDCA5E658F7FB71" box="[637,763,1189,1213]" italics="true" pageId="38" pageNumber="1570">C. prechira</emphasis>
(Olsson, 1944)
</taxonomicName>
and the middle Eocene
<taxonomicName id="4C12E9ADE36EE21DFF5AA5845845FB13" authority="(Riverai 1957)" baseAuthorityName="Riverai" baseAuthorityYear="1957" box="[251,585,1223,1247]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="paracasensis">
<emphasis id="B9664E3CE36EE21DFF5AA5845BA8FB13" box="[251,420,1223,1247]" italics="true" pageId="38" pageNumber="1570">C. paracasensis</emphasis>
(Rivera, 1957)
</taxonomicName>
: three keeled primary spiral cords, a deeply con- cave sutural area, wide concave interspaces with microscopic spiral threads, and in some specimens a secondary spiral cord or thread near the posterior suture. The spacing formula for
<taxonomicName id="4C12E9ADE36EE21DFE8AA46C5BD4FA8B" authorityName="DeVries" authorityYear="2019" box="[299,472,1327,1351]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="caballasensis">
<emphasis id="B9664E3CE36EE21DFE8AA46C5BD4FA8B" box="[299,472,1327,1351]" italics="true" pageId="38" pageNumber="1570">C. caballasensis</emphasis>
</taxonomicName>
(C14B45A67; n = 1) does differ from that of the younger
<taxonomicName id="4C12E9ADE36EE21DFBCEA46C5B26FAA6" baseAuthorityName="Riverai" baseAuthorityYear="1957" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="paracasensis">
<emphasis id="B9664E3CE36EE21DFBCEA46C5B26FAA6" italics="true" pageId="38" pageNumber="1570">C. paracasensis</emphasis>
</taxonomicName>
(C35B58A72; n = 8;
<bibRefCitation id="EF83EFDFE36EE21DFDABA41158ADFAA6" author="DeVries TJ" box="[522,673,1362,1386]" pageId="38" pageNumber="1570" pagination="331 - 351" refId="ref22853" refString="DeVries TJ. 2007. Cenozoic Turritellidae (Gastropoda) from southern Peru. J Paleontol. 81 (2): 331 - 351." type="journal article" year="2007">DeVries 2007</bibRefCitation>
) but is the same as that for the older
<taxonomicName id="4C12E9ADE36EE21DFBCEA4115AF5FA41" baseAuthorityName="Olssoni" baseAuthorityYear="1944" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="prechira">
<emphasis id="B9664E3CE36EE21DFBCEA4115AF5FA41" italics="true" pageId="38" pageNumber="1570">C. prechira</emphasis>
</taxonomicName>
(C14B40A69; n = 1). The strength of the anteriormost primary spiral cord of
<taxonomicName id="4C12E9ADE36EE21DFBCEA4365B21FA63" authorityName="DeVries" authorityYear="2019" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="caballasensis">
<emphasis id="B9664E3CE36EE21DFBCEA4365B21FA63" italics="true" pageId="38" pageNumber="1570">C. caballasensis</emphasis>
</taxonomicName>
, however, is like that of
<taxonomicName id="4C12E9ADE36EE21DFDD1A4D45911FA63" baseAuthorityName="Riverai" baseAuthorityYear="1957" box="[624,797,1431,1455]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="paracasensis">
<emphasis id="B9664E3CE36EE21DFDD1A4D45911FA63" box="[624,797,1431,1455]" italics="true" pageId="38" pageNumber="1570">C. paracasensis</emphasis>
</taxonomicName>
, i.e. much stronger than the anteriormost primary spiral cord of
<taxonomicName id="4C12E9ADE36EE21DFD99A4F958A2FA1E" baseAuthorityName="Olssoni" baseAuthorityYear="1944" box="[568,686,1466,1490]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="38" pageNumber="1570" phylum="Spirochaetes" rank="species" species="prechira">
<emphasis id="B9664E3CE36EE21DFD99A4F958A2FA1E" box="[568,686,1466,1490]" italics="true" pageId="38" pageNumber="1570">C. prechira</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36EE21DFF01A74F5E88F973" pageId="38" pageNumber="1570" type="materials_examined">
<paragraph id="8BAD922EE36EE21DFF01A74F5B0FF9EA" blockId="38.[160,871,1548,1609]" box="[160,259,1548,1574]" pageId="38" pageNumber="1570">
<emphasis id="B9664E3CE36EE21DFF01A74F5B0FF9EA" bold="true" box="[160,259,1548,1574]" italics="true" pageId="38" pageNumber="1570">Material</emphasis>
</paragraph>
<paragraph id="8BAD922EE36EE21DFF01A772596BF985" blockId="38.[160,871,1548,1609]" box="[160,871,1585,1609]" pageId="38" pageNumber="1570">
<materialsCitation id="3B7A9873E36EE21DFF01A772596BF985" ID-GBIF-Occurrence="2565890643" box="[160,871,1585,1609]" collectionCode="UWBM" pageId="38" pageNumber="1570" specimenCode="UWBM 107605" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB43A55E36EE21DFF01A7725B45F985" box="[160,329,1585,1609]" pageId="38" pageNumber="1570">UWBM 107605</specimenCode>
,
<typeStatus id="54A92C8CE36EE21DFEF4A7725BB0F985" box="[341,444,1585,1609]" pageId="38" pageNumber="1570" type="holotype">holotype</typeStatus>
, B8769 (type locality), L (5.1), W (4.4).
</materialsCitation>
</paragraph>
<paragraph id="8BAD922EE36EE21DFF01A7C05B28F951" blockId="38.[160,1156,1667,1727]" box="[160,292,1667,1693]" pageId="38" pageNumber="1570">
<emphasis id="B9664E3CE36EE21DFF01A7C05B28F951" bold="true" box="[160,292,1667,1693]" italics="true" pageId="38" pageNumber="1570">Occurrence</emphasis>
</paragraph>
<paragraph id="8BAD922EE36EE21DFF01A7E45E88F973" blockId="38.[160,1156,1667,1727]" box="[160,1156,1703,1727]" pageId="38" pageNumber="1570">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry id="F305D2BEE36EE21DFBEAA7E45E8DF973" box="[1099,1153,1703,1727]" name="Peru" pageId="38" pageNumber="1570">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>