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<document id="302C4B71FA80EB1988AFB3E0D0071BAA" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E35CE22EFE5EA2265E88FC59" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Nerita squatina DeVries 2019, sp. nov." docType="treatment" docVersion="23" lastPageNumber="1553" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1552" updateTime="1698783606523" updateUser="ExternalLinkService">
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<mods:title id="243C25CA10B065020A2C69CAC3B97CCA">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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<mods:namePart id="28329B2FE6FE56A46AA2B42696FFA643">DeVries, Thomas J.</mods:namePart>
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<mods:title id="8C0B26764DF844D565C759F5039B752C">Journal of Natural History</mods:title>
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<mods:part id="842799530CCCCD31E2471A0A055625E4">
<mods:date id="B2CE192086448F1C355DDD1FDAE631A4">2019</mods:date>
<mods:detail id="AFA33EDBABE617D6DD180B49789DA1F7" type="pubDate">
<mods:number id="67D7542EC94837CB151B1D351983D8AE">2018-12-17</mods:number>
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<mods:detail id="F147B647C4F2D0D377D4BFF8E7804B98" type="volume">
<mods:number id="0BC90B0B8233124A0342C53E1CD74278">53</mods:number>
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<mods:number id="9A2EB7A8300216AC6EB811C9CC02DB43">25</mods:number>
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<mods:classification id="5ACBB28CD1CF367227880C2EFF8CFDB2">journal article</mods:classification>
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<mods:identifier id="33706A8F682BA3BCAC2F0FB7F1A281AA" type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
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<mods:identifier id="222A6D65791801C83A81512B447D3B17" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03BB2338E35CE22EFE5EA2265E88FC59" ID-DOI="http://doi.org/10.5281/zenodo.3671230" ID-GBIF-Taxon="162146501" ID-Zenodo-Dep="3671230" LSID="urn:lsid:plazi:treatment:03BB2338E35CE22EFE5EA2265E88FC59" httpUri="http://treatment.plazi.org/id/03BB2338E35CE22EFE5EA2265E88FC59" lastPageId="21" lastPageNumber="1553" pageId="20" pageNumber="1552">
<subSubSection id="C308C1A5E35CE22FFE5EA2265929FCB2" box="[511,805,869,894]" pageId="20" pageNumber="1552" type="nomenclature">
<paragraph id="8BAD922EE35CE22FFE5EA2265929FCB2" blockId="20.[511,805,869,928]" box="[511,805,869,894]" pageId="20" pageNumber="1552">
<heading id="D0E52542E35CE22FFE5EA2265929FCB2" box="[511,805,869,894]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="1552" reason="2">
<taxonomicName id="4C12E9ADE35CE22FFE5EA22658B3FCB2" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[511,703,869,894]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="20" pageNumber="1552" phylum="Mollusca" rank="species" species="squatina" status="sp. nov.">
<emphasis id="B9664E3CE35CE22FFE5EA22658B3FCB2" bold="true" box="[511,703,869,894]" italics="true" pageId="20" pageNumber="1552">Nerita squatina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E35CE22FFD69A2265929FCB2" box="[712,805,869,894]" pageId="20" pageNumber="1552" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22FFD9DA2C458EBFC6C" box="[572,743,903,928]" pageId="20" pageNumber="1552" type="description">
<paragraph id="8BAD922EE35CE22FFD9DA2C458EBFC6C" blockId="20.[511,805,869,928]" box="[572,743,903,928]" pageId="20" pageNumber="1552">
<heading id="D0E52542E35CE22FFD9DA2C458EBFC6C" box="[572,743,903,928]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="1552" reason="5">
(
<figureCitation id="13298EABE35CE22FFDE5A2C458ABFC6C" box="[580,679,903,928]" captionStart="Figure 4" captionStartId="13.[160,225,936,959]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 4. Gastropods from the Caballas Formation. (a–e) Nerita squatina sp. nov., B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649,paratype, W 10.1 mm, anterior of outer lip.(d) UWBM 107647,paratype,W 7.8 mm,inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f) Calyptraea cf. C. aperta Solander, 1766, UWBM 107578, B8772,W (diameter) 9.9 mm, external mould. (g) Ampullina ortoni Gabb, 1870, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l) Melanella indicaformis sp. nov. (h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769,L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q) Potamides henryi sp. nov. (m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t) Potamides janeae sp. nov., B8771.(r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x) Papposilenus utriculus gen.nov., sp. nov. (u) UWBM 107592,paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view.(w, y,z,jj) Rhinotamides everriculum sp.nov. (w) UWBM 107598,paratype, B8772,L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view.(aa,bb) Terebralia pauli gen. nov.sp.nov., UWBM 107599,holotype,B8771, L 9.2 mm.(aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc) Terebralia marki gen. nov. sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii). Nodifaunus gainesi sp. nov. (dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing." figureDoi="http://doi.org/10.5281/zenodo.3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="20" pageNumber="1552">Figure 4</figureCitation>
(a
<emphasis id="B9664E3CE35CE22FFD1AA2C458C5FC6C" box="[699,713,903,928]" italics="true" pageId="20" pageNumber="1552">–</emphasis>
e))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22FFF01A28F581DFBE7" pageId="20" pageNumber="1552" type="diagnosis">
<paragraph id="8BAD922EE35CE22FFF01A28F5B18FC2A" blockId="20.[160,1156,972,1067]" box="[160,276,972,998]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A28F5B18FC2A" bold="true" box="[160,276,972,998]" italics="true" pageId="20" pageNumber="1552">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD922EE35CE22FFF01A2B2581DFBE7" blockId="20.[160,1156,972,1067]" pageId="20" pageNumber="1552">Spire flattened. Whorls with strong spiral cords. Columella with strong teeth, without papillae. Outer lip deeply fluted.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22FFF01A5145998FA23" pageId="20" pageNumber="1552" type="description">
<paragraph id="8BAD922EE35CE22FFF01A5145B2AFBBD" blockId="20.[160,1156,1111,1519]" box="[160,294,1111,1137]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A5145B2AFBBD" bold="true" box="[160,294,1111,1137]" italics="true" pageId="20" pageNumber="1552">Description</emphasis>
</paragraph>
<paragraph id="8BAD922EE35CE22FFF01A53F5998FA23" blockId="20.[160,1156,1111,1519]" pageId="20" pageNumber="1552">
(Based on fragments of multiple specimens): shell thick; length and width less than
<quantity id="4CEA3FCBE35CE22FFB91A53F5E8CFB58" box="[1072,1152,1148,1172]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="1552" unit="mm" value="20.0">20 mm</quantity>
. Spire flat to slightly sunken. Four whorls, angulate posteriorly and medially. Sculpture of about 15 steeply rounded primary spiral cords, subequal and evenly spaced anteriorly and posteriorly, alternatingly larger medially; rarely with secondary spiral cords. Interspaces flat. Axial sculpture absent except growth lines. Colour maculated black and cream. Apertural margin of columellar callus with strong anterior tooth, weaker medial tooth, and strong posterior bifid tooth; other teeth possibly present posteriorward. Abapertural portion of columellar callus with six well-spaced transversely elongate teeth, not aligned with apertural teeth; no papillae present. Outer lip with posterior inner edge bevelled and fluted; inflexions of bevel with small, spirally elongate spirally denticles along entire length. Base of columella separated from basal portion of bevelled outer lip by shallow sulcus.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22EFF01A7585E3AFE70" lastPageId="21" lastPageNumber="1553" pageId="20" pageNumber="1552" type="discussion">
<paragraph id="8BAD922EE35CE22FFF01A7585B0AF9F9" blockId="20.[160,1155,1563,1727]" box="[160,262,1563,1589]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A7585B0AF9F9" bold="true" box="[160,262,1563,1589]" italics="true" pageId="20" pageNumber="1552">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE35CE22EFF01A77C5940FF20" blockId="20.[160,1155,1563,1727]" lastBlockId="21.[160,1156,142,444]" lastPageId="21" lastPageNumber="1553" pageId="20" pageNumber="1552">
<taxonomicName id="4C12E9ADE35CE22FFF01A77C5B45F99B" authorityName="DeVries" authorityYear="2019" box="[160,329,1599,1623]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="20" pageNumber="1552" phylum="Mollusca" rank="species" species="squatina">
<emphasis id="B9664E3CE35CE22FFF01A77C5B45F99B" box="[160,329,1599,1623]" italics="true" pageId="20" pageNumber="1552">Nerita squatina</emphasis>
</taxonomicName>
does not resemble the smooth-shelled
<taxonomicName id="4C12E9ADE35CE22FFCAEA77C5E4AF99B" authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[783,1094,1599,1623]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="20" pageNumber="1552" phylum="Mollusca" rank="species" species="jayanca">
<emphasis id="B9664E3CE35CE22FFCAEA77C59A2F99B" box="[783,942,1599,1623]" italics="true" pageId="20" pageNumber="1552">Nerita jayanca</emphasis>
Olsson, 1944
</taxonomicName>
from the upper Campanian
<collectingCountry id="F305D2BEE35CE22FFE01A7215BF4F9B6" box="[416,504,1634,1658]" name="Haiti" pageId="20" pageNumber="1552">Tortuga</collectingCountry>
Formation in the Sechura Basin of northern
<collectingCountry id="F305D2BEE35CE22FFC55A7215E26F9B6" box="[1012,1066,1634,1658]" name="Peru" pageId="20" pageNumber="1552">Peru</collectingCountry>
. Rather, the flattened shell form and arrangement of spiral cords of
<taxonomicName id="4C12E9ADE35CE22FFCEDA7C659C6F951" authorityName="DeVries" authorityYear="2019" box="[844,970,1669,1693]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="20" pageNumber="1552" phylum="Mollusca" rank="species" species="squatina">
<emphasis id="B9664E3CE35CE22FFCEDA7C659C6F951" box="[844,970,1669,1693]" italics="true" pageId="20" pageNumber="1552">N. squatina</emphasis>
</taxonomicName>
resemble specimens of the late Eocene
<taxonomicName id="4C12E9ADE35CE22FFE1BA7E45907F973" authority="Woodringi 1973" authorityName="Woodringi" authorityYear="1973" box="[442,779,1703,1727]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="20" pageNumber="1552" phylum="Mollusca" rank="species" species="listrota">
<emphasis id="B9664E3CE35CE22FFE1BA7E45843F973" box="[442,591,1703,1727]" italics="true" pageId="20" pageNumber="1552">Nerita listrota</emphasis>
Woodring, 1973
</taxonomicName>
from
<collectingCountry id="F305D2BEE35CE22FFCEEA7E459A5F973" box="[847,937,1703,1727]" name="Panama" pageId="20" pageNumber="1552">Panama</collectingCountry>
and the Paleocene to middle Eocene
<taxonomicName id="4C12E9ADE35DE22EFED1A1CD58B1FF6A" authority="Gabbi 1869" authorityName="Gabbi" authorityYear="1869" box="[368,701,142,166]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="species" species="triangulata">
<emphasis id="B9664E3CE35DE22EFED1A1CD583FFF6A" box="[368,563,142,166]" italics="true" pageId="21" pageNumber="1553">Nerita triangulata</emphasis>
Gabb, 1869
</taxonomicName>
from western North America, but some spiral cords on the latter two species are beaded and the columellar callus of those species is papillate (
<bibRefCitation id="EF83EFDFE35DE22EFE22A1975838FF20" author="Woodring WP" box="[387,564,212,236]" pageId="21" pageNumber="1553" pagination="453 - 539" refId="ref27334" refString="Woodring WP. 1973. Geology and paleontology of Canal Zone and adjoining parts of Panama. Descriptions of Tertiary mollusks (Additions to gastropodsi scaphopodsi pelecypods: Nuculidae to Malleidae). U S Geol Survey Prof Pap. 306 - E: 453 - 539." type="book chapter" year="1973">Woodring 1973</bibRefCitation>
;
<bibRefCitation id="EF83EFDFE35DE22EFDE3A1975933FF20" author="Squires RLi Saul LR" box="[578,831,212,236]" pageId="21" pageNumber="1553" pagination="177 - 192" refId="ref26560" refString="Squires RLi Saul LR. 2002. New information on Late Cretaceousi Paleocenei and Eocene neritid gastropods from the North Pacific slope. Veliger. 45: 177 - 192." type="journal article" year="2002">Squires and Saul 2002</bibRefCitation>
).
</paragraph>
<paragraph id="8BAD922EE35DE22EFF61A1B55E3AFE70" blockId="21.[160,1156,142,444]" pageId="21" pageNumber="1553">
The coarse adapertural and abapertural teeth on the columellar callus and the prominence of the spiral cords on
<taxonomicName id="4C12E9ADE35DE22EFE6EA05A584BFEFD" authorityName="DeVries" authorityYear="2019" box="[463,583,281,305]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="species" species="squatina">
<emphasis id="B9664E3CE35DE22EFE6EA05A584BFEFD" box="[463,583,281,305]" italics="true" pageId="21" pageNumber="1553">N. squatina</emphasis>
</taxonomicName>
appear on modern species of the subgenera
<taxonomicName id="4C12E9ADE35DE22EFB8FA05A5E63FEFD" authorityName="Linnaeusi" authorityYear="1758" box="[1070,1135,281,305]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="B9664E3CE35DE22EFB8FA05A5E63FEFD" box="[1070,1135,281,305]" italics="true" pageId="21" pageNumber="1553">Nerita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E35DE22EFBD2A05A5ABFFE98" pageId="21" pageNumber="1553" sensu="stricto">s. s.</taxonomicNameLabel>
,
<emphasis id="B9664E3CE35DE22EFF1FA07F5AC3FE98" box="[190,207,316,340]" italics="true" pageId="21" pageNumber="1553">N</emphasis>
. (
<taxonomicName id="4C12E9ADE35DE22EFF44A07F5B24FE98" authorityName="Gray" authorityYear="1858" box="[229,296,316,340]" class="Gastropoda" family="Neritidae" genus="Ritena" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE35DE22EFF44A07F5B24FE98" box="[229,296,316,340]" italics="true" pageId="21" pageNumber="1553">Ritena</emphasis>
</taxonomicName>
) Gray,
<date id="FFACB4EEE35DE22EFEDAA07F5BBAFE98" box="[379,438,316,340]" pageId="21" pageNumber="1553" value="1858-01">1858,</date>
and
<emphasis id="B9664E3CE35DE22EFE52A07F5808FE98" box="[499,516,316,340]" italics="true" pageId="21" pageNumber="1553">N</emphasis>
. (
<taxonomicName id="4C12E9ADE35DE22EFDB8A07F588BFE98" authorityName="Martens" authorityYear="1887" box="[537,647,316,340]" class="Gastropoda" family="Neritidae" genus="Cymostyla" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE35DE22EFDB8A07F588BFE98" box="[537,647,316,340]" italics="true" pageId="21" pageNumber="1553">Cymostyla</emphasis>
</taxonomicName>
) Martens,
<date id="FFACB4EEE35DE22EFCA1A07F5930FE98" box="[768,828,316,340]" pageId="21" pageNumber="1553" value="1887-01">1887,</date>
closely related clades with a common ancestor inferred to have lived during the Oligocene or earlier (
<bibRefCitation id="EF83EFDFE35DE22EFC67A01D5ADAFE55" author="Frey MAi Vermeij GJ" pageId="21" pageNumber="1553" pagination="1067 - 1086" refId="ref23600" refString="Frey MAi Vermeij GJ. 2008. Molecular phylogenies and historical biogeography of a circumtropical group of gastropods (Genus: Nerita): implications for regional diversity patterns in the marine tropics. Molecular Phylogenetics Evol. 48: 1067 - 1086." type="journal article" year="2008">Frey and Vermeij 2008</bibRefCitation>
;
<bibRefCitation id="EF83EFDFE35DE22EFF40A0C25B46FE55" author="Frey MA" box="[225,330,385,409]" pageId="21" pageNumber="1553" pagination="1 - 7" refId="ref23575" refString="Frey MA. 2010. A revised classification of the gastropod genus Nerita. Veliger. 51 (1): 1 - 7." type="journal article" year="2010">Frey 2010</bibRefCitation>
). Absent from the Caballas Formation species of
<taxonomicName id="4C12E9ADE35DE22EFCFDA0C25990FE55" authorityName="Linnaeusi" authorityYear="1758" box="[860,924,385,409]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE35DE22EFCFDA0C25990FE55" box="[860,924,385,409]" italics="true" pageId="21" pageNumber="1553">Nerita</emphasis>
</taxonomicName>
, however, is the high spire characteristic of the
<taxonomicName id="4C12E9ADE35DE22EFE1DA0E75BF0FE70" authorityName="Linnaeusi" authorityYear="1758" box="[444,508,420,444]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="21" pageNumber="1553" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="B9664E3CE35DE22EFE1DA0E75BF0FE70" box="[444,508,420,444]" italics="true" pageId="21" pageNumber="1553">Nerita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E35DE22EFDA2A0E75828FE70" box="[515,548,420,444]" pageId="21" pageNumber="1553" sensu="stricto">s.s.</taxonomicNameLabel>
-
<emphasis id="B9664E3CE35DE22EFD8BA0E758EFFE70" box="[554,739,420,444]" italics="true" pageId="21" pageNumber="1553">Ritena-Cymostyla</emphasis>
clade (
<bibRefCitation id="EF83EFDFE35DE22EFC90A0E75E26FE70" author="Frey MAi Vermeij GJ" box="[817,1066,420,444]" pageId="21" pageNumber="1553" pagination="1067 - 1086" refId="ref23600" refString="Frey MAi Vermeij GJ. 2008. Molecular phylogenies and historical biogeography of a circumtropical group of gastropods (Genus: Nerita): implications for regional diversity patterns in the marine tropics. Molecular Phylogenetics Evol. 48: 1067 - 1086." type="journal article" year="2008">Frey and Vermeij 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35DE22EFF01A0AC5905FD82" pageId="21" pageNumber="1553" type="etymology">
<paragraph id="8BAD922EE35DE22EFF01A0AC5B11FDC5" blockId="21.[160,1155,495,590]" box="[160,285,495,521]" pageId="21" pageNumber="1553">
<emphasis id="B9664E3CE35DE22EFF01A0AC5B11FDC5" bold="true" box="[160,285,495,521]" italics="true" pageId="21" pageNumber="1553">Etymology</emphasis>
</paragraph>
<paragraph id="8BAD922EE35DE22EFF01A3505905FD82" blockId="21.[160,1155,495,590]" pageId="21" pageNumber="1553">
<emphasis id="B9664E3CE35DE22EFF01A3505AAAFDE7" box="[160,166,531,555]" italics="true" pageId="21" pageNumber="1553">‘</emphasis>
Squatina
<emphasis id="B9664E3CE35DE22EFEA9A3505B01FDE7" box="[264,269,531,555]" italics="true" pageId="21" pageNumber="1553">’</emphasis>
, Latin noun meaning
<emphasis id="B9664E3CE35DE22EFDA4A3505807FDE7" box="[517,523,531,555]" italics="true" pageId="21" pageNumber="1553">‘</emphasis>
shark
<emphasis id="B9664E3CE35DE22EFDE7A3505847FDE7" box="[582,587,531,555]" italics="true" pageId="21" pageNumber="1553">’</emphasis>
, referring to the row of adapertural teeth and row of parallel abapertural ridges on the columellar callus.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35DE22EFF01A3C2586DFCE9" pageId="21" pageNumber="1553" type="materials_examined">
<paragraph id="8BAD922EE35DE22EFF01A3C25B0FFD57" blockId="21.[160,1157,641,805]" box="[160,259,641,667]" pageId="21" pageNumber="1553">
<emphasis id="B9664E3CE35DE22EFF01A3C25B0FFD57" bold="true" box="[160,259,641,667]" italics="true" pageId="21" pageNumber="1553">Material</emphasis>
</paragraph>
<paragraph id="8BAD922EE35DE22EFF01A3E6586DFCE9" blockId="21.[160,1157,641,805]" pageId="21" pageNumber="1553">
All localities B8771 (
<typeStatus id="54A92C8CE35DE22EFE2FA3E65BCDFD71" box="[398,449,677,701]" pageId="21" pageNumber="1553">type</typeStatus>
locality); all material fragmentary.
<materialsCitation id="3B7A9873E35DE22EFCFFA3E65B3AFD2C" ID-GBIF-Occurrence="2571180958" collectionCode="UWBM" pageId="21" pageNumber="1553" specimenCode="UWBM 107577" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB43A55E35DE22EFCFFA3E65E07FD71" box="[862,1035,677,701]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="21" pageNumber="1553" type="Museum">UWBM 107577</specimenCode>
,
<typeStatus id="54A92C8CE35DE22EFBB8A3E65E8CFD71" box="[1049,1152,677,701]" pageId="21" pageNumber="1553" type="holotype">holotype</typeStatus>
, spire, W (5.7)
</materialsCitation>
;
<materialsCitation id="3B7A9873E35DE22EFEE3A38B59C6FD2C" ID-GBIF-Occurrence="2571180960" box="[322,970,712,736]" collectionCode="UWBM" pageId="21" pageNumber="1553" specimenCode="UWBM 107646" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB43A55E35DE22EFEE3A38B5BE0FD2C" box="[322,492,712,736]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="21" pageNumber="1553" type="Museum">UWBM 107646</specimenCode>
,
<typeStatus id="54A92C8CE35DE22EFE59A38B586DFD2C" box="[504,609,712,736]" pageId="21" pageNumber="1553" type="paratype">paratype,</typeStatus>
columellar callus teeth, W (6.2)
</materialsCitation>
;
<materialsCitation id="3B7A9873E35DE22EFC76A38B5BDFFCCF" ID-GBIF-Occurrence="2571180959" collectionCode="UWBM" pageId="21" pageNumber="1553" specimenCode="UWBM 107647" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB43A55E35DE22EFC76A38B5E8DFD2C" box="[983,1153,712,736]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="21" pageNumber="1553" type="Museum">UWBM 107647</specimenCode>
,
<typeStatus id="54A92C8CE35DE22EFF01A3A85B05FCCF" box="[160,265,747,771]" pageId="21" pageNumber="1553" type="paratype">paratype,</typeStatus>
outer lip, W (7.8)
</materialsCitation>
;
<materialsCitation id="3B7A9873E35DE22EFE41A3A859C6FCCF" ID-GBIF-Occurrence="2571180956" box="[480,970,747,771]" collectionCode="UWBM" pageId="21" pageNumber="1553" specimenCode="UWBM 107648" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB43A55E35DE22EFE41A3A85886FCCF" box="[480,650,747,771]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="21" pageNumber="1553" type="Museum">UWBM 107648</specimenCode>
,
<typeStatus id="54A92C8CE35DE22EFD37A3A858F3FCCF" box="[662,767,747,771]" pageId="21" pageNumber="1553" type="paratype">paratype,</typeStatus>
shoulder, W (8.5)
</materialsCitation>
;
<materialsCitation id="3B7A9873E35DE22EFC76A3A8586DFCE9" ID-GBIF-Occurrence="2571180955" collectionCode="UWBM" pageId="21" pageNumber="1553" specimenCode="UWBM 107649" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB43A55E35DE22EFC76A3A85E8CFCCF" box="[983,1152,747,771]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="21" pageNumber="1553" type="Museum">UWBM 107649</specimenCode>
,
<typeStatus id="54A92C8CE35DE22EFF01A24E5B05FCE9" box="[160,265,781,805]" pageId="21" pageNumber="1553" type="paratype">paratype,</typeStatus>
anterior of outer lip, W (10.1)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35DE22EFDC1A24E5E88FC59" pageId="21" pageNumber="1553" type="distribution">
<paragraph id="8BAD922EE35DE22EFDC1A24E5868FCE9" blockId="21.[160,1157,641,805]" box="[608,612,781,805]" pageId="21" pageNumber="1553">.</paragraph>
<paragraph id="8BAD922EE35DE22EFF01A21A5B28FCBF" blockId="21.[160,1156,857,917]" box="[160,292,857,883]" pageId="21" pageNumber="1553">
<emphasis id="B9664E3CE35DE22EFF01A21A5B28FCBF" bold="true" box="[160,292,857,883]" italics="true" pageId="21" pageNumber="1553">Occurrence</emphasis>
</paragraph>
<paragraph id="8BAD922EE35DE22EFF01A23E5E88FC59" blockId="21.[160,1156,857,917]" box="[160,1156,893,917]" pageId="21" pageNumber="1553">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry id="F305D2BEE35DE22EFBEAA23E5E8DFC59" box="[1099,1153,893,917]" name="Peru" pageId="21" pageNumber="1553">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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