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<document id="CED98A93C7EE750075057FBEB60C16E4" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E356E224FE60A5DE5900FE70" docLanguage="en" docName="JNATHIST.53.2526.15331584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Terebralia Swainsoni 1840" docType="treatment" docVersion="21" lastPageNumber="1563" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1562" updateTime="1698783606523" updateUser="ExternalLinkService">
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<mods:title id="3E015B2526DD5057E1EA37BE4E885E98">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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<mods:namePart id="CFB3461378738701C2F54D2F5B0A525A">DeVries, Thomas J.</mods:namePart>
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<treatment id="03BB2338E356E224FE60A5DE5900FE70" ID-DOI="http://doi.org/10.5281/zenodo.3671212" ID-GBIF-Taxon="162146505" ID-Zenodo-Dep="3671212" LSID="urn:lsid:plazi:treatment:03BB2338E356E224FE60A5DE5900FE70" httpUri="http://treatment.plazi.org/id/03BB2338E356E224FE60A5DE5900FE70" lastPageId="31" lastPageNumber="1563" pageId="30" pageNumber="1562">
<subSubSection id="C308C1A5E356E225FE60A5DE596EFB7A" box="[449,866,1181,1206]" pageId="30" pageNumber="1562" type="nomenclature">
<paragraph id="8BAD922EE356E225FE60A5DE596EFB7A" blockId="30.[449,866,1181,1206]" box="[449,866,1181,1206]" pageId="30" pageNumber="1562">
<heading id="D0E52542E356E225FE60A5DE596EFB7A" box="[449,866,1181,1206]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="1562" reason="2">
Genus
<taxonomicName id="4C12E9ADE356E225FDB4A5DE596EFB7A" ID-CoL="7PJ78" authority="Swainsoni 1840" authorityName="Swainsoni" authorityYear="1840" box="[533,866,1181,1206]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FDB4A5DE596EFB7A" bold="true" box="[533,866,1181,1206]" pageId="30" pageNumber="1562">
<emphasis id="B9664E3CE356E225FDB4A5DE589EFB7A" bold="true" box="[533,658,1181,1206]" italics="true" pageId="30" pageNumber="1562">Terebralia</emphasis>
Swainsonı 1840
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E356E225FF01A5A35AE8FAD7" pageId="30" pageNumber="1562" type="materials_examined">
<paragraph id="8BAD922EE356E225FF01A5A35AE8FAD7" blockId="30.[160,1155,1248,1307]" pageId="30" pageNumber="1562">
<typeStatus id="54A92C8CE356E225FF01A5A35ADAFB34" box="[160,214,1248,1272]" pageId="30" pageNumber="1562">Type</typeStatus>
species:
<emphasis id="B9664E3CE356E225FEEBA5A35817FB34" box="[330,539,1248,1272]" italics="true" pageId="30" pageNumber="1562">strombus palustris</emphasis>
Linnaeus,
<date id="FFACB4EEE356E225FD00A5A358D2FB34" box="[673,734,1248,1272]" pageId="30" pageNumber="1562" value="1767-01">1767,</date>
by subsequent designation (
<bibRefCitation id="EF83EFDFE356E225FBE1A5A35ADBFAD7" author="Sacco F" pageId="30" pageNumber="1562" refId="ref26201" refString="Sacco F. 1895. I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte XVII. Cerithiidaei Triphoridaei Cerithiopsidaei e Diastomidae. Torino: Carlo Clausen." type="book" year="1895">Sacco 1895</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E356E224FF01A4085900FE70" lastPageId="31" lastPageNumber="1563" pageId="30" pageNumber="1562" type="discussion">
<paragraph id="8BAD922EE356E225FF01A4085B0AFAA9" blockId="30.[160,1155,1355,1727]" box="[160,262,1355,1381]" pageId="30" pageNumber="1562">
<emphasis id="B9664E3CE356E225FF01A4085B0AFAA9" bold="true" box="[160,262,1355,1381]" italics="true" pageId="30" pageNumber="1562">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE356E224FF01A42C580CFEC2" blockId="30.[160,1155,1355,1727]" lastBlockId="31.[160,1155,142,444]" lastPageId="31" lastPageNumber="1563" pageId="30" pageNumber="1562">
Two small specimens from the Caballas Formation strongly resemble the early teleoconch whorls of species of
<taxonomicName id="4C12E9ADE356E225FE76A4D158F7FA66" authority="Swainsoni 1840" authorityName="Swainsoni" authorityYear="1840" box="[471,763,1426,1450]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FE76A4D1584FFA66" box="[471,579,1426,1450]" italics="true" pageId="30" pageNumber="1562">Terebralia</emphasis>
Swainson, 1840
</taxonomicName>
, except that both specimens from the East Pisco Basin lack varices and a columellar fold. The absence of varices alone is insufficient to rule out
<taxonomicName id="4C12E9ADE356E225FE15A494582CFA23" authorityName="Swainsoni" authorityYear="1840" box="[436,544,1495,1519]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FE15A494582CFA23" box="[436,544,1495,1519]" italics="true" pageId="30" pageNumber="1562">Terebralia</emphasis>
</taxonomicName>
as a correct generic placement, inasmuch as some individuals of
<taxonomicName id="4C12E9ADE356E225FE9EA4B95BA7F9DE" authorityName="Swainsoni" authorityYear="1840" box="[319,427,1530,1554]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FE9EA4B95BA7F9DE" box="[319,427,1530,1554]" italics="true" pageId="30" pageNumber="1562">Terebralia</emphasis>
</taxonomicName>
lack varices on early teleoconch whorls. Also, the absence alone of a columellar fold (or columellar callus, an extension of the anteriormost of two internal columellar plaits; see
<bibRefCitation id="EF83EFDFE356E225FD83A77C58E5F99B" author="Houbrick RS" box="[546,745,1599,1623]" pageId="30" pageNumber="1562" pagination="289 - 338" refId="ref24191" refString="Houbrick RS. 1991. Systematic review and fuctlional morphology of the mangrove snails Terebralia and Telescopium (Potamididae; Prosobranchia). Malacologia. 33: 289 - 338." type="journal article" year="1991">Houbrick (1991))</bibRefCitation>
may be insufficient to exclude
<taxonomicName id="4C12E9ADE356E225FF01A7215B00F9B6" authorityName="Swainsoni" authorityYear="1840" box="[160,268,1634,1658]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FF01A7215B00F9B6" box="[160,268,1634,1658]" italics="true" pageId="30" pageNumber="1562">Terebralia</emphasis>
</taxonomicName>
; the Miocene
<taxonomicName id="4C12E9ADE356E225FE0DA7215969F9B6" authority="Pilsbry and Harbisoni 1933" authorityName="Pilsbry and Harbisoni" authorityYear="1933" box="[428,869,1634,1658]" class="Gastropoda" family="Potamididae" genus="Cerithideops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FE0DA721583EF9B6" box="[428,562,1634,1658]" italics="true" pageId="30" pageNumber="1562">Cerithideops</emphasis>
Pilsbry and Harbison, 1933
</taxonomicName>
, which lacks a columellar fold, is included among fossil
<taxonomicName id="4C12E9ADE356E225FDAFA7C65876F951" authorityName="Swainsoni" authorityYear="1840" box="[526,634,1669,1693]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE356E225FDAFA7C65876F951" box="[526,634,1669,1693]" italics="true" pageId="30" pageNumber="1562">Terebralia</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF83EFDFE356E225FD11A7C65973F951" author="Reid DGi Dyal Pi Lozouet Pi Glaubrecht Mi Williams ST" box="[688,895,1669,1693]" pageId="30" pageNumber="1562" pagination="680 - 699" refId="ref26089" refString="Reid DGi Dyal Pi Lozouet Pi Glaubrecht Mi Williams ST. 2008. Mudwhelks and mangroves: the evolutionary history of an ecological association (Gastropoda: Potamididae). Molecular Phylogenetics Evol. 47: 680 - 699." type="journal article" year="2008">Reid et al. (2008)</bibRefCitation>
. Assignment to other potamidid genera, however, seems unwarranted; other genera do not have flat-sided whorls that exhibit the pattern of sculpture seen on the whorls of Caballas Formation specimens and early teleoconch whorls of specimens of
<taxonomicName id="4C12E9ADE357E224FCE6A1F259BFFF05" authorityName="Swainsoni" authorityYear="1840" box="[839,947,177,201]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="31" pageNumber="1563" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE357E224FCE6A1F259BFFF05" box="[839,947,177,201]" italics="true" pageId="31" pageNumber="1563">Terebralia</emphasis>
</taxonomicName>
: few and equally strong spiral cords that are equal in strength to the axial ribs and do not produce nodes or spines at their intersections.
</paragraph>
<paragraph id="8BAD922EE357E224FF61A05A5900FE70" blockId="31.[160,1155,142,444]" pageId="31" pageNumber="1563">
<bibRefCitation id="EF83EFDFE357E224FF61A05A5B85FEFD" author="Reid DGi Dyal Pi Lozouet Pi Glaubrecht Mi Williams ST" box="[192,393,281,305]" pageId="31" pageNumber="1563" pagination="680 - 699" refId="ref26089" refString="Reid DGi Dyal Pi Lozouet Pi Glaubrecht Mi Williams ST. 2008. Mudwhelks and mangroves: the evolutionary history of an ecological association (Gastropoda: Potamididae). Molecular Phylogenetics Evol. 47: 680 - 699." type="journal article" year="2008">Reid et al. (2008)</bibRefCitation>
propose that the oldest examples of
<taxonomicName id="4C12E9ADE357E224FCEEA05A59B7FEFD" authorityName="Swainsoni" authorityYear="1840" box="[847,955,281,305]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="31" pageNumber="1563" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE357E224FCEEA05A59B7FEFD" box="[847,955,281,305]" italics="true" pageId="31" pageNumber="1563">Terebralia</emphasis>
</taxonomicName>
lived during the middle Eocene in Europe and made their appearance in the Americas during the Miocene. The presence of
<taxonomicName id="4C12E9ADE357E224FE67A01D583EFEBA" authorityName="Swainsoni" authorityYear="1840" box="[454,562,350,374]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="31" pageNumber="1563" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE357E224FE67A01D583EFEBA" box="[454,562,350,374]" italics="true" pageId="31" pageNumber="1563">Terebralia</emphasis>
</taxonomicName>
in southern
<collectingCountry id="F305D2BEE357E224FD61A01D58FFFEBA" box="[704,755,350,374]" name="Peru" pageId="31" pageNumber="1563">Peru</collectingCountry>
well before the Miocene, therefore, constitutes a biogeographical puzzle that can only be resolved by finding larger specimens of the two species from the Caballas Formation.
</paragraph>
</subSubSection>
</treatment>
</document>