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<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E355E225FE1FA2265E88FB93" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Rhinotamides everriculum DeVries 2019, sp. nov." docType="treatment" docVersion="21" lastPageNumber="1562" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1561" updateTime="1636522412784" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>DeVries, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>25</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1533</mods:start>
<mods:end>1584</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671206" ID-GBIF-Taxon="162146470" ID-Zenodo-Dep="3671206" LSID="urn:lsid:plazi:treatment:03BB2338E355E225FE1FA2265E88FB93" httpUri="http://treatment.plazi.org/id/03BB2338E355E225FE1FA2265E88FB93" lastPageId="30" lastPageNumber="1562" pageId="29" pageNumber="1561">
<subSubSection box="[446,870,869,894]" pageId="29" pageNumber="1561" type="nomenclature">
<paragraph blockId="29.[446,870,869,929]" box="[446,870,869,894]" pageId="29" pageNumber="1561">
<heading box="[446,870,869,894]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="1561" reason="2">
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[446,768,869,894]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="29" pageNumber="1561" phylum="Mollusca" rank="species" species="everriculum" status="sp. nov.">
<emphasis bold="true" box="[446,768,869,894]" italics="true" pageId="29" pageNumber="1561">Rhinotamides everriculum</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[777,870,869,894]" pageId="29" pageNumber="1561" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[495,820,904,929]" pageId="29" pageNumber="1561" type="description">
<paragraph blockId="29.[446,870,869,929]" box="[495,820,904,929]" pageId="29" pageNumber="1561">
<heading box="[495,820,904,929]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="1561" reason="5">
(
<figureCitation box="[503,602,904,929]" captionStart="Figure 4" captionStartId="13.[160,225,936,959]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 4. Gastropods from the Caballas Formation. (a–e) Nerita squatina sp. nov., B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649,paratype, W 10.1 mm, anterior of outer lip.(d) UWBM 107647,paratype,W 7.8 mm,inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f) Calyptraea cf. C. aperta Solander, 1766, UWBM 107578, B8772,W (diameter) 9.9 mm, external mould. (g) Ampullina ortoni Gabb, 1870, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l) Melanella indicaformis sp. nov. (h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769,L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q) Potamides henryi sp. nov. (m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t) Potamides janeae sp. nov., B8771.(r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x) Papposilenus utriculus gen.nov., sp. nov. (u) UWBM 107592,paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view.(w, y,z,jj) Rhinotamides everriculum sp.nov. (w) UWBM 107598,paratype, B8772,L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view.(aa,bb) Terebralia pauli gen. nov.sp.nov., UWBM 107599,holotype,B8771, L 9.2 mm.(aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc) Terebralia marki gen. nov. sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii). Nodifaunus gainesi sp. nov. (dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing." figureDoi="http://doi.org/10.5281/zenodo.3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="29" pageNumber="1561">Figure 4</figureCitation>
(w), 4(y), 4(z), 4(jj))
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="1561" type="diagnosis">
<paragraph blockId="29.[160,1155,973,1068]" box="[160,276,973,999]" pageId="29" pageNumber="1561">
<emphasis bold="true" box="[160,276,973,999]" italics="true" pageId="29" pageNumber="1561">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="29.[160,1155,973,1068]" pageId="29" pageNumber="1561">Reticulate sculpture of three thin primary spiral cords and bluntly spinose axial ribs on later teleoconch whorls.</paragraph>
</subSubSection>
<subSubSection lastPageId="30" lastPageNumber="1562" pageId="29" pageNumber="1561" type="description">
<paragraph blockId="29.[160,1156,1112,1727]" box="[160,294,1112,1138]" pageId="29" pageNumber="1561">
<emphasis bold="true" box="[160,294,1112,1138]" italics="true" pageId="29" pageNumber="1561">Description</emphasis>
</paragraph>
<paragraph blockId="29.[160,1156,1112,1727]" lastBlockId="30.[160,1155,142,201]" lastPageId="30" lastPageNumber="1562" pageId="29" pageNumber="1561">Shell length estimated about 35 mm, cerithiform, spire angle about 25° for early whorls, 15° for later whorls. Protoconch unknown. Spire about 11 whorls, flat-sided to weakly convex. Sutures impressed. Earliest teleoconch whorls with reticulated sculpture produced by three strong spiral cords and about eight stronger axial ribs that extend suture to suture. Subsequent whorls with stronger, more nodular, and more numerous axial ribs (11), three distinct primary spiral cords and on some specimens, an additional basal peripheral cord; the three to four cords cross axial ribs to produce raised nodes; a secondary spiral cord inserted between each pair of primary spiral cords and between anterior and posterior primary cords and the respective sutures. Mid-spire whorls with more numerous axial ribs (14), insertion of intercalated tertiary spiral threads between three primary spiral cords; and posteriormost spiral cord with nodes becoming larger and bluntly spinose. Last whorl with smooth anterior primary spiral cord, reduced middle spiral cord, varying losses of tertiary spiral threads, and bluntly spinose posterior primary spiral cord; axial ribs diminished anteriorly. Varices, ventrolateral varix absent. Base with peripheral primary spiral cord, flattened adaxially with at least one additional strong spiral cord. Outer lip flaring, extended posteriorly adjacent to suture, producing subsutural blunt spine, but without posterior canal. Parietal area with callus; columella short, without fold. Anterior canal twisted abaxially, with keeled posterior margin; poorly preserved.</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="1562" type="discussion">
<paragraph blockId="30.[160,1156,248,621]" box="[160,262,248,274]" pageId="30" pageNumber="1562">
<emphasis bold="true" box="[160,262,248,274]" italics="true" pageId="30" pageNumber="1562">Remarks</emphasis>
</paragraph>
<paragraph blockId="30.[160,1156,248,621]" pageId="30" pageNumber="1562">
The Caballas Formation specimens attributed to
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[752,900,285,309]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">
<emphasis box="[752,900,285,309]" italics="true" pageId="30" pageNumber="1562">Rhinotamides</emphasis>
</taxonomicName>
differ from the Late Cretaceous species from the
<collectingCountry box="[483,571,320,344]" name="Haiti" pageId="30" pageNumber="1562">Tortuga</collectingCountry>
Formation,
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[706,784,320,344]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="species" species="rudis">
<emphasis box="[706,784,320,344]" italics="true" pageId="30" pageNumber="1562">R. rudis</emphasis>
</taxonomicName>
, in having axial ribs that become increasingly numerous with each spire whorl, thus preventing axial ribs on successive whorls from remaining axially aligned, as they do on the spire of
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[888,967,389,413]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="species" species="rudis">
<emphasis box="[888,967,389,413]" italics="true" pageId="30" pageNumber="1562">R. rudis</emphasis>
</taxonomicName>
. Specimens of
<taxonomicName authorityName="DeVries" authorityYear="2019" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="species" species="everriculum">
<emphasis italics="true" pageId="30" pageNumber="1562">R. everriculum</emphasis>
</taxonomicName>
share more characters with early Eocene specimens of a species herein transferred to
<emphasis box="[325,479,458,482]" italics="true" pageId="30" pageNumber="1562">
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[325,475,458,482]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="genus">Rhinotamides</taxonomicName>
,
</emphasis>
<taxonomicName authority="Woodsi 1922" authorityName="Woodsi" authorityYear="1922" box="[489,896,458,482]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="species" species="occidentalis">
<emphasis box="[489,739,458,482]" italics="true" pageId="30" pageNumber="1562">Potamides occidentalis</emphasis>
Woods, 1922
</taxonomicName>
, from the Talara Basin of northern
<collectingCountry box="[297,351,493,517]" name="Peru" pageId="30" pageNumber="1562">Peru</collectingCountry>
. Those shared characters include a short smooth columella adjacent to an abaxially twisted anterior canal with a keeled posterior margin. Specimens of
<taxonomicName box="[160,450,562,586]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="1562" phylum="Mollusca" rank="species" species="occidentalis">
<emphasis box="[160,450,562,586]" italics="true" pageId="30" pageNumber="1562">Rhinotamides occidentalis</emphasis>
</taxonomicName>
, however, have less sharply defined spiral cords on later whorls.
</paragraph>
</subSubSection>
<subSubSection box="[160,1147,669,729]" pageId="30" pageNumber="1562" type="etymology">
<paragraph blockId="30.[160,1147,669,729]" box="[160,285,669,695]" pageId="30" pageNumber="1562">
<emphasis bold="true" box="[160,285,669,695]" italics="true" pageId="30" pageNumber="1562">Etymology</emphasis>
</paragraph>
<paragraph blockId="30.[160,1147,669,729]" box="[160,1147,705,729]" pageId="30" pageNumber="1562">
<emphasis box="[160,166,705,729]" italics="true" pageId="30" pageNumber="1562">‘</emphasis>
Everriculum
<emphasis box="[298,303,705,729]" italics="true" pageId="30" pageNumber="1562">’</emphasis>
, Latin for
<emphasis box="[422,428,705,729]" italics="true" pageId="30" pageNumber="1562">‘</emphasis>
fishing net
<emphasis box="[549,555,705,729]" italics="true" pageId="30" pageNumber="1562">’</emphasis>
, referencing the reticulate sculpture of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="1562" type="materials_examined">
<paragraph blockId="30.[160,1157,777,1011]" box="[160,259,777,803]" pageId="30" pageNumber="1562">
<emphasis bold="true" box="[160,259,777,803]" italics="true" pageId="30" pageNumber="1562">Material</emphasis>
</paragraph>
<paragraph blockId="30.[160,1157,777,1011]" pageId="30" pageNumber="1562">
<materialsCitation ID-GBIF-Occurrence="2572160120" box="[160,911,814,838]" collectionCode="UWBM" pageId="30" pageNumber="1562" specimenCode="UWBM 107593" specimenCount="1" typeStatus="holotype">
<specimenCode box="[160,331,814,838]" pageId="30" pageNumber="1562">UWBM 107593</specimenCode>
,
<typeStatus box="[345,448,814,838]" pageId="30" pageNumber="1562">holotype</typeStatus>
, B8769 (type locality), L (21.1), W (14.7)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572160119" pageId="30" pageNumber="1562" specimenCount="1" typeStatus="paratype">
remainder are
<typeStatus pageId="30" pageNumber="1562">paratypes</typeStatus>
:
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890708" box="[234,691,848,872]" pageId="30" pageNumber="1562">
<collectionCode box="[234,310,848,872]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="30" pageNumber="1562" type="Museum">UWBM</collectionCode>
107594, B8772, L (19.4), W (17.0);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890687" box="[699,1156,848,872]" pageId="30" pageNumber="1562">
<collectionCode box="[699,775,848,872]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="30" pageNumber="1562" type="Museum">UWBM</collectionCode>
107595, B8769, L (29.7), W (17.6);
</materialsCitation>
</paragraph>
<paragraph blockId="30.[160,1157,777,1011]" pageId="30" pageNumber="1562">
<materialsCitation ID-GBIF-Occurrence="2565890653" box="[160,619,883,907]" pageId="30" pageNumber="1562">
<collectionCode box="[160,236,883,907]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="30" pageNumber="1562" type="Museum">UWBM</collectionCode>
107596, B8772, L (25.4), W (15.4);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890714" box="[627,1072,883,907]" pageId="30" pageNumber="1562">
<collectionCode box="[627,703,883,907]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="30" pageNumber="1562" type="Museum">UWBM</collectionCode>
107597, B8769, L (12.2), W (8.9);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890736" pageId="30" pageNumber="1562">
<collectionCode box="[1079,1155,883,907]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="30" pageNumber="1562" type="Museum">UWBM</collectionCode>
107598, B8772, L (23.5), W (11.0);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890669" box="[551,1024,918,942]" pageId="30" pageNumber="1562">
<collectionCode box="[551,624,918,942]" pageId="30" pageNumber="1562">MUSM</collectionCode>
INV 255, B8769, L (24.8), W (15.2);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890745" pageId="30" pageNumber="1562">
<collectionCode box="[1033,1106,918,942]" pageId="30" pageNumber="1562">MUSM</collectionCode>
INV 256, B8769, L (28.4), W (15.0);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890674" box="[509,968,952,976]" pageId="30" pageNumber="1562">
<collectionCode box="[509,582,952,976]" pageId="30" pageNumber="1562">MUSM</collectionCode>
INV 257, B8769, L (13.6), W (9.0);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2565890748" pageId="30" pageNumber="1562">
<collectionCode box="[977,1050,952,976]" pageId="30" pageNumber="1562">MUSM</collectionCode>
INV 258, B8772, L (14.4), W (9.8).
</materialsCitation>
</paragraph>
<paragraph blockId="30.[160,1156,1058,1119]" box="[160,292,1058,1084]" pageId="30" pageNumber="1562">
<emphasis bold="true" box="[160,292,1058,1084]" italics="true" pageId="30" pageNumber="1562">Occurrence</emphasis>
</paragraph>
<paragraph blockId="30.[160,1156,1058,1119]" box="[160,1156,1095,1119]" pageId="30" pageNumber="1562">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry box="[1099,1153,1095,1119]" name="Peru" pageId="30" pageNumber="1562">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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