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<document id="9D670158A9B8C534B30701DD2C396E4E" ID-DOI="10.1206/3961.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5370399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268866948" checkinUser="carolina" docAuthor="Sinclair, Bradley J. &amp; Grimaldi, David A." docDate="2020" docId="03B94C5BFFD13C6767CE34B2FEB47027" docLanguage="en" docName="N3961.pdf" docOrigin="American Museum Novitates 2020 (3961)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Alavesia spinosa Sinclair &amp; Grimaldi 2020, new species" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="FF803423FFCC3C46662D3721FC617321" masterDocTitle="Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="30" updateTime="1699205599378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7239702CE323B6483A80F21B0966AE3B">Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)</mods:title>
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<mods:namePart id="970267DE36CFD86BAF7B7C3C89F6F3B7">Sinclair, Bradley J.</mods:namePart>
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<mods:namePart id="CB1E38D01E57EABBC1583B339FEFB565">Grimaldi, David A.</mods:namePart>
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<mods:title id="079A2CB4F142383E0D9C247D4EC56544">American Museum Novitates</mods:title>
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<mods:date id="B1A5C66F5FE541331D76B555D7A104D2">2020</mods:date>
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<mods:number id="656EC1D7B5DC25E226F39B14F9AEA5BC">2020-10-06</mods:number>
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<treatment id="03B94C5BFFD13C6767CE34B2FEB47027" ID-DOI="http://doi.org/10.5281/zenodo.5462049" ID-Zenodo-Dep="5462049" LSID="urn:lsid:plazi:treatment:03B94C5BFFD13C6767CE34B2FEB47027" httpUri="http://treatment.plazi.org/id/03B94C5BFFD13C6767CE34B2FEB47027" lastPageId="33" lastPageNumber="34" pageId="29" pageNumber="30">
<subSubSection id="C30AAEC6FFD13C5B67CE34B2FF24708F" box="[483,837,915,943]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BAFFD4DFFD13C5B67CE34B2FF24708F" blockId="29.[483,837,915,943]" box="[483,837,915,943]" pageId="29" pageNumber="30">
<heading id="D0E74A21FFD13C5B67CE34B2FF24708F" box="[483,837,915,943]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<emphasis id="B964215FFFD13C5B67CE34B2FED3708F" bold="true" box="[483,690,915,942]" pageId="29" pageNumber="30">
<taxonomicName id="4C1086CEFFD13C5B67CE34B2FECA708F" ID-CoL="89WYN" authority="Sinclair &amp; Grimaldi, 2020" authorityName="Sinclair &amp; Grimaldi" authorityYear="2020" box="[483,683,915,942]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="B964215FFFD13C5B67CE34B2FECA708F" bold="true" box="[483,683,915,942]" italics="true" pageId="29" pageNumber="30">Alavesia spinosa</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2579C24FFD13C5B649734B4FF24708F" box="[698,837,916,943]" pageId="29" pageNumber="30" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFD13C5B67C334EDFF5A70C7" box="[494,827,972,999]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BAFFD4DFFD13C5B67C334EDFF5A70C7" blockId="29.[494,827,972,999]" box="[494,827,972,999]" pageId="29" pageNumber="30">
<heading id="D0E74A21FFD13C5B67C334EDFF5A70C7" box="[494,827,972,999]" centered="true" fontSize="36" level="3" pageId="29" pageNumber="30" reason="7">
<figureCitation id="132BE1C8FFD13C5B67C334EDFE0A70C7" box="[494,619,972,998]" captionStart="FIGURE" captionStartId="6.[108,197,1285,1306]" captionTargetBox="[200,1125,232,1256]" captionTargetId="figure-108@6.[108,1212,224,1262]" captionTargetPageId="6" captionText="FIGURE. 1. Antennae of Burmese amber Alavesia species, lateral views. All are males except N, O, P. Some antennae were reversed in direction for optimal comparison; longicornuta and longistylata are slightly reconstructed to account for curvature of the stylus. All are to the same scale. Specimen numbers: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-KL2-7. F. AMNH Bu-1476. G. JZC Bu-556 (AMNH). H. USNM PAL 726872. K. USNM PAL 726869. L. AMNH Bu-KL7-18. M. JZC Bu-282. N. Undetermined species. O. A. zigrasi. P. A. zigrasi." figureDoi="http://doi.org/10.5281/zenodo.5370401" httpUri="https://zenodo.org/record/5370401/files/figure.png" pageId="29" pageNumber="30">Figures 1L</figureCitation>
,
<figureCitation id="132BE1C8FFD13C5B645534EDFEFC70C7" box="[632,669,972,998]" captionStart="FIGURE 4" captionStartId="9.[108,194,1276,1297]" captionTargetBox="[116,1200,232,1241]" captionTargetId="figure-39@9.[108,1212,224,1244]" captionTargetPageId="9" captionText="FIGURE 4. Wings of Alavesia spp. in Burmese amber. A. A. spinosa, AMNH Bu-KL7-18. BD. A. zigrasi, showing some differences in venation among three specimens: B. male; C. male; D. female." figureDoi="http://doi.org/10.5281/zenodo.5370407" httpUri="https://zenodo.org/record/5370407/files/figure.png" pageId="29" pageNumber="30">4A</figureCitation>
,
<figureCitation id="132BE1C8FFD13C5B648434EDFEAC70C6" box="[681,717,972,999]" captionStart="FIGURE 5" captionStartId="10.[108,194,1523,1544]" captionTargetBox="[253,1069,228,1487]" captionTargetId="figure-69@10.[108,1212,224,1493]" captionTargetPageId="10" captionText="FIGURE 5. Thoraces of three species of Alavesia in Burmese amber, showing some of the differences in notal chaetotaxy. A. A. longicornuta (JZC Bu-556). B. A. magna (AMNH Bu-KL2-7). C. A. spinosa (AMNH Bu-KL7-18). All are to the same scale. Setal abbreviations: acr, acrostichals; dcs, dorsocentrals; ntpl, notopleurals; palr, postalar; pscdc, prescutellar dorsocentrals; salr, supraalar, sct, scutellars." figureDoi="http://doi.org/10.5281/zenodo.5370409" httpUri="https://zenodo.org/record/5370409/files/figure.png" pageId="29" pageNumber="30">5C</figureCitation>
,
<figureCitation id="132BE1C8FFD13C5B64F734EDFE9C70C7" box="[730,765,972,998]" captionStart="FIGURE 7" captionStartId="12.[108,194,1180,1201]" captionTargetBox="[125,1209,228,1145]" captionTargetId="figure-74@12.[108,1212,224,1152]" captionTargetPageId="12" captionText="FIGURE 7. Male terminalia of three species of Alavesia in Burmese amber, shown in various views, to the same scale. A. A. pankowskiorum, AMNH Bu-KL7-19, lateral view. B. A. pankowskiorum, USNM PAL 726869, anterolateral view. C. A. spinosa, AMNH KL7-18, ventrolateral view, with detail of surstylus tip showing clavate setula. D. A. zigrasi, JZC Bu-281. For abbreviations see figure 6." figureDoi="http://doi.org/10.5281/zenodo.5370413" httpUri="https://zenodo.org/record/5370413/files/figure.png" pageId="29" pageNumber="30">7C</figureCitation>
,
<figureCitation id="132BE1C8FFD13C5B652734EDFF5A70C7" box="[778,827,972,998]" captionStart="FIGURE 17" captionStartId="31.[108,194,1672,1693]" captionTargetBox="[108,1135,224,1652]" captionTargetId="figure-54@31.[108,1135,224,1652]" captionTargetPageId="31" captionText="FIGURE 17. Alavesia spp. in Burmese amber. BF are all females unidentified to species. A. A. spinosa holotype, AMNH Bu-KL7-18. B. JZC Bu-1729. C. USNM PAL 726870. D. USNM PAL 726874. E. USNM PAL 726870. F. USNM PAL 726875." figureDoi="http://doi.org/10.5281/zenodo.5370431" httpUri="https://zenodo.org/record/5370431/files/figure.png" pageId="29" pageNumber="30">17A</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFD13C5B668E3325FD8977AF" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8BAFFD4DFFD13C5B668E3325FD8977AF" blockId="29.[108,1214,1028,1727]" pageId="29" pageNumber="30">
<materialsCitation id="3B78F710FFD13C5B668E3325FD8577AF" collectionCode="R" longitude="-4.88" pageId="29" pageNumber="30" specimenCode="R4" specimenCount="1">
DIAGNOSIS: Wing moderately slender (fig. 17A), L/
<geoCoordinate id="EE249B8AFFD13C5B64D53325FF2C773F" box="[760,845,1028,1054]" degrees="2.88" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-2.88">W 2.88</geoCoordinate>
, with distinctly narrowed apex;
<specimenCode id="DBB65536FFD13C5B6641330BFCE97769" box="[108,136,1066,1096]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="30">
R
<subScript id="1794FF08FFD13C5B66533316FCE97769" attach="left" box="[126,136,1079,1096]" fontSize="7" pageId="29" pageNumber="30">4</subScript>
</specimenCode>
+5 and M
<subScript id="1794FF08FFD13C5B66DE3316FD707769" attach="left" box="[243,273,1079,1096]" fontSize="7" pageId="29" pageNumber="30">1+2</subScript>
slightly curved; bm-cu and CuA nearly symmetrical; cell dm long and slender. Postpedicel very large, L/
<geoCoordinate id="EE249B8AFFD13C5B67B7336EFD937748" box="[410,498,1103,1129]" degrees="4.88" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-4.88">W 4.88</geoCoordinate>
, lanceolate. Male genitalia with pair of thick, spinelike setae just anterior to bases of surstyli
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFD13C67668933B8FDBF7150" lastPageId="33" lastPageNumber="34" pageId="29" pageNumber="30" type="description">
<paragraph id="8BAFFD4DFFD13C5B668933B8F871764F" blockId="29.[108,1214,1028,1727]" pageId="29" pageNumber="30">
DESCRIPTION:
<emphasis id="B964215FFFD13C5B677F33B8FDED7792" box="[338,396,1177,1203]" italics="true" pageId="29" pageNumber="30">Male</emphasis>
.
<emphasis id="B964215FFFD13C5B67B733B8FE407792" bold="true" box="[410,545,1177,1203]" pageId="29" pageNumber="30">Coloration</emphasis>
partially visible, much of cuticle with layer of air. Mesoscutum and scutellum dark tan/light brown; postpedicel brown; pleura brown at least in center, perhaps more; halter knob light brown; tarsi entirely dark brown, tibiae and femora lighter; tergites 14 dark brown, others yellowish; wing completely hyaline.
<emphasis id="B964215FFFD13C5B65573228FFA37602" bold="true" box="[890,962,1289,1315]" pageId="29" pageNumber="30">Head:</emphasis>
Scape slender, longer than pedicel; pedicel small, quadrate, setose; postpedicel long, lanceolate (fig. 1L), L/
<geoCoordinate id="EE249B8AFFD13C5B624D320EF8D67668" box="[1120,1207,1327,1353]" degrees="4.88" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-4.88">W 4.88</geoCoordinate>
; stylus relatively short, 0.39
<emphasis id="B964215FFFD13C5B67883275FDD7764F" bold="true" box="[421,438,1364,1390]" pageId="29" pageNumber="30">×</emphasis>
length of postpedicel, with 2 minute basal articles.
</paragraph>
<paragraph id="8BAFFD4DFFD13C5B668E3258FF837508" blockId="29.[108,1214,1028,1727]" pageId="29" pageNumber="30">
<emphasis id="B964215FFFD13C5B668E3258FD6376B2" bold="true" box="[163,258,1401,1427]" pageId="29" pageNumber="30">Thorax:</emphasis>
Macrosetae thickened (fig. 5C); acrostichals small and fine, in biserial rows; dorsocentrals similar to acrostichals, ending in pair of long prescutellar setae plus 3 smaller macrosetae per side. Postpronotal setae only slightly larger than acrostichals; 5 notopleural setae of variable lengths; 1 postsutural supraalar seta; 1 postalar seta; 3 pairs scutellar setae, apical pair upright, crossed at tips, only slightly larger than lateralmost scutellars.
</paragraph>
<paragraph id="8BAFFD4DFFD13C5B668B3115F8C5759F" blockId="29.[108,1214,1028,1727]" pageId="29" pageNumber="30">
<emphasis id="B964215FFFD13C5B668B3115FC84756F" bold="true" box="[166,229,1588,1614]" pageId="29" pageNumber="30">Legs:</emphasis>
Forecoxa setose on anterior surface, longer at apex; forefemur with anteroventral row of ~30 short, spinulelike setae along length of femur; mid and hind femora with such setae only near apex. Tibia with long, apical ventral seta(e): 1-2-2. Hind tibia with short dorsal row of 34 thick setae, lengths ~0.7
<emphasis id="B964215FFFD13C5B67F63185FD8D759F" bold="true" box="[475,492,1700,1726]" pageId="29" pageNumber="30">×</emphasis>
width of tibia, posterior surface with brush of fine setulae.
</paragraph>
<caption id="DF6FADC5FFD23C5866413183F8F075F4" ID-DOI="http://doi.org/10.5281/zenodo.5370429" ID-Zenodo-Dep="5370429" httpUri="https://zenodo.org/record/5370429/files/figure.png" pageId="30" pageNumber="31" startId="30.[108,194,1698,1719]" targetBox="[121,1140,224,1675]" targetPageId="30">
<paragraph id="8BAFFD4DFFD23C5866413183F8F075F4" blockId="30.[108,1212,1695,1750]" pageId="30" pageNumber="31">
FIGURE 16.
<taxonomicName id="4C1086CEFFD23C5866C33181FDE97596" box="[238,392,1696,1720]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="zigrasi">
<emphasis id="B964215FFFD23C5866C33181FDE97596" box="[238,392,1696,1720]" italics="true" pageId="30" pageNumber="31">Alavesia zigrasi</emphasis>
</taxonomicName>
in Burmese amber.
<emphasis id="B964215FFFD23C58647A31BEFE117596" bold="true" box="[599,624,1695,1719]" pageId="30" pageNumber="31">A.</emphasis>
Holotype, JZC Bu-281.
<emphasis id="B964215FFFD23C58654B3181FF147599" bold="true" box="[870,885,1696,1720]" pageId="30" pageNumber="31">B</emphasis>
. Male paratype, JZC Bu-282.
<emphasis id="B964215FFFD23C58628B31BEF8D77596" bold="true" box="[1190,1206,1695,1719]" pageId="30" pageNumber="31">C</emphasis>
. Female paratype, AMNH Bu-KL9-21.
<emphasis id="B964215FFFD23C5867D8319CFE6675F4" bold="true" box="[501,519,1725,1749]" pageId="30" pageNumber="31">D</emphasis>
. Female paratype, JZC-165a.
<emphasis id="B964215FFFD23C58651A319CFF2475F4" bold="true" box="[823,837,1725,1749]" pageId="30" pageNumber="31">E</emphasis>
. Female paratype, JZC Bu-165b.
</paragraph>
</caption>
<caption id="DF6FADC5FFD33C59664131A9FDCC75F9" ID-DOI="http://doi.org/10.5281/zenodo.5370431" ID-Zenodo-Dep="5370431" httpUri="https://zenodo.org/record/5370431/files/figure.png" pageId="31" pageNumber="32" startId="31.[108,194,1672,1693]" targetBox="[108,1135,224,1652]" targetPageId="31">
<paragraph id="8BAFFD4DFFD33C59664131A9FDCC75F9" blockId="31.[108,1212,1669,1752]" pageId="31" pageNumber="32">
FIGURE 17.
<taxonomicName id="4C1086CEFFD33C5966C231A7FD1275BC" box="[239,371,1670,1694]" class="Insecta" family="Atelestidae" genus="Alavesia" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B964215FFFD33C5966C231A7FD2375BF" box="[239,322,1670,1694]" italics="true" pageId="31" pageNumber="32">Alavesia</emphasis>
spp.
</taxonomicName>
in Burmese amber. BF are all females unidentified to species.
<emphasis id="B964215FFFD33C5965D631A4F86D75BC" bold="true" box="[1019,1036,1669,1693]" pageId="31" pageNumber="32">A</emphasis>
.
<taxonomicName id="4C1086CEFFD33C59623531A7F81E75BC" authorityName="Sinclair &amp; Grimaldi" authorityYear="2020" box="[1048,1151,1670,1693]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="B964215FFFD33C59623531A7F81E75BC" box="[1048,1151,1670,1693]" italics="true" pageId="31" pageNumber="32">A. spinosa</emphasis>
</taxonomicName>
holotype, AMNH Bu-KL7-18.
<emphasis id="B964215FFFD33C5967513182FDED759A" bold="true" box="[380,396,1699,1723]" pageId="31" pageNumber="32">B</emphasis>
. JZC Bu-1729.
<emphasis id="B964215FFFD33C5964033182FE5E759A" bold="true" box="[558,575,1699,1723]" pageId="31" pageNumber="32">C</emphasis>
. USNM PAL 726870.
<emphasis id="B964215FFFD33C5965053182FF5A759A" bold="true" box="[808,827,1699,1723]" pageId="31" pageNumber="32">D</emphasis>
. USNM PAL 726874.
<emphasis id="B964215FFFD33C5962093182F852759A" bold="true" box="[1060,1075,1699,1723]" pageId="31" pageNumber="32">E</emphasis>
. USNM PAL 726870.
<emphasis id="B964215FFFD33C5966EF31E1FCB175F9" bold="true" box="[194,208,1728,1752]" pageId="31" pageNumber="32">F</emphasis>
. USNM PAL 726875.
</paragraph>
</caption>
<caption id="DF6FADC5FFEC3C6666413188F84C75FD" ID-DOI="http://doi.org/10.5281/zenodo.5370435" ID-Zenodo-Dep="5370435" httpUri="https://zenodo.org/record/5370435/files/figure.png" pageId="32" pageNumber="33" startId="32.[108,194,1705,1726]" targetBox="[123,1158,227,1675]" targetPageId="32">
<paragraph id="8BAFFD4DFFEC3C6666413188F84C75FD" blockId="32.[108,1212,1703,1757]" pageId="32" pageNumber="33">
FIGURE 18. Two pieces of Burmese amber with small swarms of
<taxonomicName id="4C1086CEFFEC3C66653E3186FF05759E" authorityName="Sinclair &amp; Grimaldi" authorityYear="2020" box="[787,868,1703,1727]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B964215FFFEC3C66653E3186FF05759E" box="[787,868,1703,1727]" italics="true" pageId="32" pageNumber="33">Alavesia</emphasis>
</taxonomicName>
. Numbers indicate
<taxonomicName id="4C1086CEFFEC3C66621C3186F8E5759E" authorityName="Sinclair &amp; Grimaldi" authorityYear="2020" box="[1073,1156,1703,1727]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B964215FFFEC3C66621C3186F8E5759E" box="[1073,1156,1703,1727]" italics="true" pageId="32" pageNumber="33">Alavesia</emphasis>
</taxonomicName>
individuals. See text for descriptions and interpretations.
<emphasis id="B964215FFFEC3C6664A331E5FEC775FD" bold="true" box="[654,678,1732,1756]" pageId="32" pageNumber="33">A.</emphasis>
AMNH Bu-177.
<emphasis id="B964215FFFEC3C66657531E4FF0E75FC" bold="true" box="[856,879,1733,1757]" pageId="32" pageNumber="33">B.</emphasis>
AMNH Bu-KL15.
</paragraph>
</caption>
<paragraph id="8BAFFD4DFFED3C67668E37C0FFF572B0" blockId="33.[108,1213,225,775]" pageId="33" pageNumber="34">
<emphasis id="B964215FFFED3C67668E37C0FC8C73DA" bold="true" box="[163,237,225,251]" pageId="33" pageNumber="34">Wing:</emphasis>
Length
<quantity id="4CE850A8FFED3C67676337C0FDC673DD" box="[334,423,225,252]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="33" pageNumber="34" unit="mm" value="1.7">1.7 mm</quantity>
(fig. 4A), vein C terminating slightly beyond M
<subScript id="1794FF08FFED3C6765F837CEFF927221" attach="left" box="[981,1011,239,256]" fontSize="7" pageId="33" pageNumber="34">1+2</subScript>
; Sc nearly reaching costal margin; R
<subScript id="1794FF08FFED3C6767773635FD197204" attach="left" box="[346,376,276,293]" fontSize="7" pageId="33" pageNumber="34">4+5</subScript>
parallel to M
<subScript id="1794FF08FFED3C67643A3635FE547204" attach="left" box="[535,565,276,293]" fontSize="7" pageId="33" pageNumber="34">1+2</subScript>
but diverging at apices, both veins slightly curved (not straight); crossvein r-m at distal third of cell dm, cell long and slender; apex of M
<subScript id="1794FF08FFED3C67627B361BF83E726A" attach="left" box="[1110,1119,314,331]" fontSize="7" pageId="33" pageNumber="34">4</subScript>
slightly closer to M
<subScript id="1794FF08FFED3C6766DF367EFD717251" attach="left" box="[242,272,351,368]" fontSize="7" pageId="33" pageNumber="34">1+2</subScript>
than to CuA+CuP; CuA not aligned with bm-m; apex of cell cua acute, cell cua almost symmetrical with cell bm; CuA+CuP bent slightly in middle.
</paragraph>
<paragraph id="8BAFFD4DFFED3C67668E36BDFDBF7150" blockId="33.[108,1213,225,775]" pageId="33" pageNumber="34">
<emphasis id="B964215FFFED3C67668E36BDFD427297" bold="true" box="[163,291,412,438]" pageId="33" pageNumber="34">Abdomen:</emphasis>
Slightly longer than thorax. Setae short, with little differentiation; sternites 24 with slightly thicker, short setulae on posterior and lateral margins. Terminalia (fig. 7C): Epandrium short, broad, dorsally complete, with some setulae and fringe of 56 long, fine setae on ventral margins. Cercus small. Surstylus long, slender, digitate, with medial row of 5 fine setulae, apical setula clavate. Phallus short, bifid. Genitalia uniquely with pair of stout, large, spinelike setae near bases of surstyli.
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFED3C676688355CFFE8719A" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BAFFD4DFFED3C676688355CFFE8719A" blockId="33.[108,1213,225,775]" pageId="33" pageNumber="34">
<materialsCitation id="3B78F710FFED3C676688355CFFE8719A" collectionCode="TYPE, MATERIAL, AMNH" pageId="33" pageNumber="34" specimenCode="KL7-18" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED016588FFED3C676688355CFD0471B7" box="[165,357,636,663]" pageId="33" pageNumber="34">TYPE MATERIAL</collectionCode>
:
<typeStatus id="54AB43EFFFED3C676759355DFD8271B7" box="[372,483,636,662]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
, male:
<collectionCode id="ED016588FFED3C676414355DFEF571B7" box="[569,660,636,662]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="34">AMNH</collectionCode>
<specimenCode id="DBB65536FFED3C6764B0355DFE9971B7" box="[669,760,636,662]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="34">KL7-18</specimenCode>
. The fly is in excellent condition, preserved in clear yellow amber with many microscopic details visible.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFED3C67668835E6FEB47027" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BAFFD4DFFED3C67668835E6FEB47027" blockId="33.[108,1213,225,775]" pageId="33" pageNumber="34">
ETYMOLOGY: The specific epithet is from the Latin
<emphasis id="B964215FFFED3C67652435E6FF0971C1" box="[777,872,711,736]" italics="true" pageId="33" pageNumber="34">spinosus</emphasis>
(thorny), in reference to the pair of thick, spinelike setae on the male terminalia.
</paragraph>
</subSubSection>
</treatment>
</document>