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<document id="6529CB30C63E71FF2BDFB8BDA59682B8" ID-CLB-Dataset="25319" ID-DOI="10.11646/zootaxa.4679.3.12" ID-GBIF-Dataset="44e80db5-8f5d-4bed-a189-baed3b1020c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099173655" checkinUser="plazi" docAuthor="Montreuil, Olivier &amp; Frolov, Andrey V." docDate="2019" docId="03B6CB587A44B968FF18FD66A4F85DF8" docLanguage="en" docName="zootaxa.4679.3.12.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trigonochilus macrotarsus Montreuil &amp; Frolov 2019, new species" docType="treatment" docVersion="6" lastPageNumber="598" masterDocId="FF8FB3207A46B96BFF8FFFD3A7795C0D" masterDocTitle="A review of the Malagasy species of the genus Trigonochilus Brenske (Coleoptera Scarabaeidae: Rutelinae: Adoretini) with description of a new species" masterLastPageNumber="600" masterPageNumber="596" pageNumber="598" updateTime="1698757505706" updateUser="ExternalLinkService">
<mods:mods id="A2737DA921F7EC3316B7E04A73331ECB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF227263F9C3C8F0B9ECB87CEE6D90E9">
<mods:title id="EDE923FD16046E491416799F4EDC1628">A review of the Malagasy species of the genus Trigonochilus Brenske (Coleoptera Scarabaeidae: Rutelinae: Adoretini) with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="1B38EF6102470446CF437A354D87F924" type="personal">
<mods:role id="DD855C22DDA654595249BBC3402BF0BD">
<mods:roleTerm id="C5E39E7FC30D284A432E0329CF3D5D8D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9AF93F60ED9F2A41707B88138AB7058">Montreuil, Olivier</mods:namePart>
</mods:name>
<mods:name id="BF7603554A6DED24A11596B188A689F2" type="personal">
<mods:role id="526339C731B843AF501347235106F364">
<mods:roleTerm id="30DF364819C354C4EDF562502F096353">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B42D7568FAC38A834391F220DE79A3E1">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="602C1766834CD1B0F6E67F3AB0881296">text</mods:typeOfResource>
<mods:relatedItem id="23848B00E5973E4D5803E6CBFBEBA20B" type="host">
<mods:titleInfo id="31F21F7982FEFFB995C0B26882F57B5A">
<mods:title id="0B72E0B03522606300CD7004DBC95B9C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="047DE865B7542F8F98C6942C9DE44FEC">
<mods:date id="4C96DFE69E439DAD1C50C4F4778A80B9">2019</mods:date>
<mods:detail id="01DEC5CA5F0446413A6F084F54EA5367" type="pubDate">
<mods:number id="59732B8721E4DC71509B34C471ED4780">2019-10-03</mods:number>
</mods:detail>
<mods:detail id="45C206E980B8B19756402FA987179AF4" type="volume">
<mods:number id="4D1BE3D4DB83178733B0F86040E9EF38">4679</mods:number>
</mods:detail>
<mods:detail id="7429BAC9A1AF7259C7B83C45D89FDF2B" type="issue">
<mods:number id="3927F113C933A7EC4C593339F450B427">3</mods:number>
</mods:detail>
<mods:extent id="9DE50FA5B0587D74A054D107EBB52E9C" unit="page">
<mods:start id="28058DFED20D5DBC3895EC0C2E29A511">596</mods:start>
<mods:end id="8FF7A6A107DEF50321A3D5D5CDA3927E">600</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="909F8AF99019098C869B06AF21BD0B4F">journal article</mods:classification>
<mods:identifier id="EAB874B7B1A9FA45A871E9450D0B767C" type="CLB-Dataset">25319</mods:identifier>
<mods:identifier id="384C0301A2AB57300D6787DFAE545695" type="DOI">10.11646/zootaxa.4679.3.12</mods:identifier>
<mods:identifier id="9DDF09F749D71966F603D8CE2E67E4A8" type="GBIF-Dataset">44e80db5-8f5d-4bed-a189-baed3b1020c8</mods:identifier>
<mods:identifier id="C0227B708E2F1ADCD1EAAC17EAADBD58" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A65157FC2B9F9042846EC87E73B0E698" type="Zenodo-Dep">3471116</mods:identifier>
<mods:identifier id="5A8CA7736E6E71B0C5BE9A63B62402D8" type="ZooBank">F1C41402-60E4-4245-9144-8EF62FAA919C</mods:identifier>
</mods:mods>
<treatment id="03B6CB587A44B968FF18FD66A4F85DF8" ID-GBIF-Taxon="159628563" LSID="urn:lsid:plazi:treatment:03B6CB587A44B968FF18FD66A4F85DF8" httpUri="http://treatment.plazi.org/id/03B6CB587A44B968FF18FD66A4F85DF8" lastPageId="3" lastPageNumber="598" pageId="2" pageNumber="598">
<subSubSection id="C30529C57A44B969FF18FD66A4495EC0" box="[151,816,693,718]" pageId="2" pageNumber="598" type="nomenclature">
<paragraph id="8BA07A4E7A44B969FF18FD66A4495EC0" blockId="2.[151,1437,693,2014]" box="[151,816,693,718]" pageId="2" pageNumber="598">
<heading id="D0E8CD227A44B969FF18FD66A4495EC0" bold="true" box="[151,816,693,718]" fontSize="10" level="2" pageId="2" pageNumber="598" reason="2">
<emphasis id="B96BA65C7A44B969FF18FD66A4495EC0" bold="true" box="[151,816,693,718]" pageId="2" pageNumber="598">
<taxonomicName id="4C1F01CD7A44B969FF18FD66A5D95EC3" ID-CoL="7CRYY" authority="Montreuil &amp; Frolov" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[151,672,693,718]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A44B969FF18FD66A6C15EC0" bold="true" box="[151,440,693,717]" italics="true" pageId="2" pageNumber="598">Trigonochilus macrotarsus</emphasis>
Montreuil &amp; Frolov
</taxonomicName>
,
<taxonomicNameLabel id="A2581B277A44B969FD23FD65A4495EC0" box="[684,816,693,718]" pageId="2" pageNumber="598" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A44B969FF18FD09A6765EFF" box="[151,271,730,754]" pageId="2" pageNumber="598" type="description">
<paragraph id="8BA07A4E7A44B969FF18FD09A6765EFF" blockId="2.[151,1437,693,2014]" box="[151,271,730,754]" pageId="2" pageNumber="598">
<figureCitation id="132466CB7A44B969FF18FD09A7A35EFF" box="[151,218,730,754]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Figs. 1</figureCitation>
D–F
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A44B969FF18FD2DA57E5FAB" pageId="2" pageNumber="598" type="materials_examined">
<paragraph id="8BA07A4E7A44B969FF18FD2DA57E5FAB" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF18FD2DA64C5F18" bold="true" box="[151,309,765,790]" pageId="2" pageNumber="598">Type material</emphasis>
.
<materialsCitation id="3B7770137A44B969FECDFD2DA6435F37" ID-GBIF-Occurrence="2423812412" collectionCode="MNHN" country="Madagascar" elevation="1000" location="Ile de Masoala" pageId="2" pageNumber="598" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A4C4EC7A44B969FECDFD2DA6DC5F1B" box="[322,421,766,790]" pageId="2" pageNumber="598" type="holotype">Holotype</typeStatus>
<specimenCount id="9D19B1C77A44B969FE24FD2DA6B85F18" box="[427,449,766,789]" pageId="2" pageNumber="598" type="male">♂</specimenCount>
, “
<collectingCountry id="F3083ADE7A44B969FE5AFD2DA5215F1B" box="[469,600,766,790]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
Est, pr.
<location id="8EC02C957A44B969FD21FD2FA4345F1B" LSID="urn:lsid:plazi:treatment:03B6CB587A44B968FF18FD66A4F85DF8:8EC02C957A44B969FD21FD2FA4345F1B" box="[686,845,764,790]" country="Madagascar" name="Ile de Masoala" pageId="2" pageNumber="598">Île de Masoala</location>
, m. if, Ambohitsitondrona,
<quantity id="4CE7D7AB7A44B969FBFFFD2DA3C55F1B" box="[1136,1212,766,790]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="598" unit="m" value="1000.0">
<elevation id="00329D7D7A44B969FBFFFD2DA3C55F1B" box="[1136,1212,766,790]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="598" unit="m" value="1000.0">1000m</elevation>
</quantity>
, XII, Vadon et Peyri- eras” (
<collectionCode id="ED0EE28B7A44B969FF54FCF1A64A5F37" box="[219,307,802,826]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7770137A44B969FECBFCF1A4E55F37" ID-GBIF-Occurrence="2423812418" box="[324,924,802,826]" collectionCode="MNHN" country="Madagascar" elevation="1000" location="Ile de Masoala" pageId="2" pageNumber="598" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54A4C4EC7A44B969FECBFCF1A6C95F37" box="[324,432,802,826]" pageId="2" pageNumber="598" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D19B1C77A44B969FE36FCF1A69B5F34" box="[441,482,802,826]" pageId="2" pageNumber="598" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D19B1C77A44B969FE62FCF1A56D5F34" box="[493,532,802,826]" pageId="2" pageNumber="598" type="female">1 ♀</specimenCount>
, same data as the holotype (
<collectionCode id="ED0EE28B7A44B969FCB2FCF1A4EC5F37" box="[829,917,802,826]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7770137A44B969FC29FCF1A59E5F53" ID-GBIF-Occurrence="2423812415" collectingDate="1959-11" collectionCode="MNHN" collectorName="P. Soga" country="Madagascar" elevation="1600" location="Est" municipality="Marojejy-Ouest" pageId="2" pageNumber="598" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D19B1C77A44B969FC29FCF1A4B65F34" box="[934,975,802,826]" pageId="2" pageNumber="598" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D19B1C77A44B969FC55FCF1A3785F34" box="[986,1025,802,826]" pageId="2" pageNumber="598" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F3083ADE7A44B969FB98FCF1A3E35F37" box="[1047,1178,802,826]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
<location id="8EC02C957A44B969FB2FFCF1A3BC5F34" LSID="urn:lsid:plazi:treatment:03B6CB587A44B968FF18FD66A4F85DF8:8EC02C957A44B969FB2FFCF1A3BC5F34" box="[1184,1221,802,825]" country="Madagascar" municipality="Marojejy-Ouest" name="Est" pageId="2" pageNumber="598">Est</location>
, dct. Sambava, R.N. XII,
<collectingMunicipality id="6BC4E0347A44B969FF46FC95A60C5F53" box="[201,373,838,862]" pageId="2" pageNumber="598">Marojejy-Ouest</collectingMunicipality>
,
<quantity id="4CE7D7AB7A44B969FEF1FC95A6B25F53" box="[382,459,838,862]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="598" unit="m" value="1600.0">
<elevation id="00329D7D7A44B969FEF1FC95A6B25F53" box="[382,459,838,862]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="598" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FFA15C8E7A44B969FE5AFC96A56E5F50" box="[469,535,837,861]" pageId="2" pageNumber="598" value="1959-11">
<collectingDate id="EFE5A5667A44B969FE5AFC96A56E5F50" box="[469,535,837,861]" pageId="2" pageNumber="598" value="1959-11">XI-59</collectingDate>
</date>
,
<collectorName id="26EA1F987A44B969FDADFC95A5085F53" box="[546,625,838,862]" pageId="2" pageNumber="598">P. Soga</collectorName>
” (
<collectionCode id="ED0EE28B7A44B969FD06FC95A5985F53" box="[649,737,838,862]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7770137A44B969FD7DFC95A5585F8F" ID-GBIF-Occurrence="2423812419" collectionCode="MNHN" collectorName="P. Soga" country="Madagascar" elevation="1300" location="Est" municipality="Marojejy-Ouest" pageId="2" pageNumber="598" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D19B1C77A44B969FD7DFC95A4625F50" box="[754,795,838,862]" pageId="2" pageNumber="598" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="F3083ADE7A44B969FCA0FC95A4CB5F53" box="[815,946,838,862]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
<location id="8EC02C957A44B969FC37FC95A4A45F50" LSID="urn:lsid:plazi:treatment:03B6CB587A44B968FF18FD66A4F85DF8:8EC02C957A44B969FC37FC95A4A45F50" box="[952,989,838,861]" country="Madagascar" municipality="Marojejy-Ouest" name="Est" pageId="2" pageNumber="598">Est</location>
, dct. Sambava, R.N. XII,
<collectingMunicipality id="6BC4E0347A44B969FB62FC95A2E05F53" box="[1261,1433,838,862]" pageId="2" pageNumber="598">Marojejy-Ouest</collectingMunicipality>
,
<quantity id="4CE7D7AB7A44B969FF18FCB9A79D5F8F" box="[151,228,874,898]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="598" unit="m" value="1300.0">
<elevation id="00329D7D7A44B969FF18FCB9A79D5F8F" box="[151,228,874,898]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="598" unit="m" value="1300.0">1300m</elevation>
</quantity>
, IX-X-59,
<collectorName id="26EA1F987A44B969FED6FCB9A6D35F8F" box="[345,426,874,898]" pageId="2" pageNumber="598">P. Soga</collectorName>
” (
<collectionCode id="ED0EE28B7A44B969FE4CFCB9A5625F8F" box="[451,539,874,898]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7770137A44B969FDA2FCB9A57A5FAB" ID-GBIF-Occurrence="2423812414" collectingDate="1972-12-08" collectionCode="MNHN" collectorName="A. Peyrieras" country="Madagascar" elevation="1300" location="Est" pageId="2" pageNumber="598" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D19B1C77A44B969FDA2FCB9A5215F8C" box="[557,600,874,898]" pageId="2" pageNumber="598" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D19B1C77A44B969FDEBFCB9A5F55F8C" box="[612,652,874,898]" pageId="2" pageNumber="598" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F3083ADE7A44B969FD2BFCB9A45E5F8F" box="[676,807,874,898]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
<location id="8EC02C957A44B969FCA1FCB9A42A5F8C" LSID="urn:lsid:plazi:treatment:03B6CB587A44B968FF18FD66A4F85DF8:8EC02C957A44B969FCA1FCB9A42A5F8C" box="[814,851,874,897]" country="Madagascar" name="Est" pageId="2" pageNumber="598">Est</location>
, Massif du Marojejy (rés. nat. intégr. 12),
<quantity id="4CE7D7AB7A44B969FA96FCB9A21F5F8F" box="[1305,1382,874,898]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="598" unit="m" value="1300.0">
<elevation id="00329D7D7A44B969FA96FCB9A21F5F8F" box="[1305,1382,874,898]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="598" unit="m" value="1300.0">1300m</elevation>
</quantity>
, 2/
<date id="FFA15C8E7A44B969FA07FCB9A7845FAB" pageId="2" pageNumber="598" value="1972-12-08">
<collectingDate id="EFE5A5667A44B969FA07FCB9A7845FAB" pageId="2" pageNumber="598" value="1972-12-08">8- XII-1972</collectingDate>
</date>
,
<collectorName id="26EA1F987A44B969FE89FC5DA6F45FAB" box="[262,397,910,934]" pageId="2" pageNumber="598">A. Peyrieras</collectorName>
” (
<collectionCode id="ED0EE28B7A44B969FE2BFC5DA6855FAB" box="[420,508,910,934]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="598">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A44B969FF48FC62A52D5AEF" pageId="2" pageNumber="598" type="description">
<paragraph id="8BA07A4E7A44B969FF48FC62A5EF5FC7" blockId="2.[151,1437,693,2014]" box="[199,662,945,970]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FC62A6325FC4" bold="true" box="[199,331,945,969]" pageId="2" pageNumber="598">Description</emphasis>
.
<typeStatus id="54A4C4EC7A44B969FED7FC61A6C75FC7" box="[344,446,946,970]" pageId="2" pageNumber="598" type="holotype">Holotype</typeStatus>
, male (
<figureCitation id="132466CB7A44B969FD84FC61A5F35FC7" box="[523,650,946,970]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Figs. 1D, E</figureCitation>
).
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FC05A5C8583B" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
Length 14.0 mm, maximum width
<quantity id="4CE7D7AB7A44B969FDCDFC06A5E15FE0" box="[578,664,981,1005]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="598" unit="mm" value="6.5">6.5 mm</quantity>
. Head uniformly dark brown, pronotum light brown with 2 dark brown longitudinal stripes; elytra light brown with darker humoral and apical umbones and irregular dark brown stripes; anterior and middle legs light brown, posterior legs darker.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FB91A69858CB" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FB91A6785857" box="[199,257,1090,1114]" italics="true" pageId="2" pageNumber="598">Head</emphasis>
. Clypeus broadly rounded, its anterior margin somewhat upturned. Punctures large and rather dense, separated by less than a puncture diameter on disc, each puncture bearing short, laying setae. Frons flat, punctate as clypeus. Frontoclypeal suture distinct. Labrum with a long, dilated medial process. Antennal club elongated, straight, shorter than funiculus and scape combined.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FB01A628593C" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FB01A66D58E7" box="[199,276,1234,1258]" italics="true" pageId="2" pageNumber="598">Thorax</emphasis>
. Pronotal bordered all around, border narrow. Lateral margins regularly curved, anterolateral angles narrowly rounded, posterolateral angles obtusely rounded. Punctuation somewhat sparser than that on head, punctures bear short and laying setae.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FAEDA2715958" blockId="2.[151,1437,693,2014]" box="[199,1288,1342,1366]" pageId="2" pageNumber="598">
<taxonomicName id="4C1F01CD7A44B969FF48FAEDA649595B" box="[199,304,1342,1366]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="B96BA65C7A44B969FF48FAEDA649595B" box="[199,304,1342,1366]" italics="true" pageId="2" pageNumber="598">Scutellum</emphasis>
</taxonomicName>
. Triangular, punctate, bearing short and laying setae similar to those on head and pronotum.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FAB1A39A59CF" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FAB1A6735977" box="[199,266,1378,1402]" italics="true" pageId="2" pageNumber="598">Elytra</emphasis>
. Striae poorly developed, noticeable as rows of punctures, odd interstriae much narrower than even and slightly raised. Punctuation slightly denser on disc, punctures bearing short, laying setae, similar to those on head and pronotum. Lateral margins somewhat crenulate (in dorsal view), with a bit longer and thicker setae in apical third.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FA1DA39359EB" blockId="2.[151,1437,693,2014]" box="[199,1258,1486,1510]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FA1DA63B59E8" box="[199,322,1486,1509]" italics="true" pageId="2" pageNumber="598">Tergite VIII</emphasis>
. Moderately convex, prominent, with denser and somewhat longer setae than on elytra.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48FA21A3025A07" blockId="2.[151,1437,693,2014]" box="[199,1147,1522,1546]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48FA21A6345A07" box="[199,333,1522,1546]" italics="true" pageId="2" pageNumber="598">Ventral side.</emphasis>
Metasternum and abdominal sternites with setation similar to that on elytra.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48F9C5A3D75A23" blockId="2.[151,1437,693,2014]" box="[199,1198,1558,1582]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F9C5A7805A20" box="[199,249,1558,1581]" italics="true" pageId="2" pageNumber="598">Legs</emphasis>
. Protibiae tridentate, basal tooth smaller than 2 basal. Metatibiae and metatarsi enlarged.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48F9E9A4005A5F" blockId="2.[151,1437,693,2014]" box="[199,889,1594,1618]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F9E9A6565A5F" box="[199,303,1594,1618]" italics="true" pageId="2" pageNumber="598">Aedeagus</emphasis>
. Aedeagus with completely fused parameres (
<figureCitation id="132466CB7A44B969FC95F9E9A4155A5F" box="[794,876,1594,1618]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48F98DA6185A97" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F98DA6635A7B" bold="true" box="[199,282,1630,1654]" pageId="2" pageNumber="598">Female</emphasis>
(
<figureCitation id="132466CB7A44B969FEA7F98DA6015A7B" box="[296,376,1630,1654]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Fig. 1F</figureCitation>
). Similar to the male in colour and setation, but have shorter antennae and relatively slender metatibiae and metatarsi.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48F975A52D5AEF" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F975A6395AB3" bold="true" box="[199,320,1702,1726]" pageId="2" pageNumber="598">
<typeStatus id="54A4C4EC7A44B969FF48F975A6425AB3" box="[199,315,1702,1726]" pageId="2" pageNumber="598" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Body length of the
<typeStatus id="54A4C4EC7A44B969FD94F975A5F85AB3" box="[539,641,1702,1726]" pageId="2" pageNumber="598" type="paratype">paratypes</typeStatus>
varies from
<quantity id="4CE7D7AB7A44B969FC83F976A4D25AB0" box="[780,939,1701,1725]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" metricValueMax="1.45" metricValueMin="1.28" pageId="2" pageNumber="598" unit="mm" value="13.65" valueMax="14.5" valueMin="12.8">12.8–14.5 mm</quantity>
. Coloration of some specimens is lighter with darker stripes on elytra poorly noticeable.
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A44B969FF48F93DA3D85B98" pageId="2" pageNumber="598" type="etymology">
<paragraph id="8BA07A4E7A44B969FF48F93DA3C15B0B" blockId="2.[151,1437,693,2014]" box="[199,1208,1774,1798]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F93DA6315B0B" bold="true" box="[199,328,1774,1798]" pageId="2" pageNumber="598">Etymology.</emphasis>
The species is named in reference to the highly enlarged metatibiae and metatarsi.
</paragraph>
<paragraph id="8BA07A4E7A44B969FF48F8C2A3D85B98" blockId="2.[151,1437,693,2014]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F8C2A6BA5B24" bold="true" box="[199,451,1809,1833]" pageId="2" pageNumber="598">Differential diagnosis.</emphasis>
<taxonomicName id="4C1F01CD7A44B969FE43F8C1A5945B24" authority="Montreuil &amp; Frolov, 2019" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[460,749,1810,1834]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A44B969FE43F8C1A5945B24" box="[460,749,1810,1834]" italics="true" pageId="2" pageNumber="598">Trigonochilus macrotarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2581B277A44B969FD79F8C1A4015B27" box="[758,888,1810,1834]" pageId="2" pageNumber="598" rank="species">new species</taxonomicNameLabel>
differs from
<taxonomicName id="4C1F01CD7A44B969FB8EF8C1A3135B27" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[1025,1130,1810,1834]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A44B969FB8EF8C1A3135B27" box="[1025,1130,1810,1834]" italics="true" pageId="2" pageNumber="598">T. oedipus</emphasis>
</taxonomicName>
in the following characters: pronotum and head light brown with a pattern of longitudinal darker stripes; anterior and middle legs are light brown, posterior legs are uniformly dark brown; the elytra are more or less parallel-sided; apical margin of labrum with a long, broadened process; metatarsi with two claws in both sexes; aedeagus with parameres split at the apex.
</paragraph>
</subSubSection>
<subSubSection id="C30529C57A44B968FF48F872A4F85DF8" lastPageId="3" lastPageNumber="599" pageId="2" pageNumber="598" type="distribution">
<paragraph id="8BA07A4E7A44B969FF48F872A4F45BB7" blockId="2.[151,1437,693,2014]" box="[199,909,1953,1978]" pageId="2" pageNumber="598">
<emphasis id="B96BA65C7A44B969FF48F872A6235BB4" bold="true" box="[199,346,1953,1977]" pageId="2" pageNumber="598">Distribution.</emphasis>
Northern
<collectingCountry id="F3083ADE7A44B969FE48F871A5335BB7" box="[455,586,1954,1978]" name="Madagascar" pageId="2" pageNumber="598">Madagascar</collectingCountry>
, Masoala Peninsula (
<figureCitation id="132466CB7A44B969FCA4F871A4F95BB7" box="[811,896,1954,1978]" captionStart="FIGURES 1" captionStartId="1.[151,264,1895,1920]" captionTargetBox="[187,1410,347,1860]" captionTargetId="figure@1.[178,1410,336,1868]" captionTargetPageId="1" captionText="FIGURES 1A–F. Trigonochilus oedipus (Fairmaire, 1903) (A–C), T. macrotarsus Montreuil &amp; Frolov, new species (D–F) and locality map (G). A, male, dorsal view; B, aedeagus, lateral and dorsal view; C, female, dorsal view; D, male, holotype, dorsal view; E, aedeagus, lateral and dosal view; F, female, paratype, dorsal view; G, locality map (black circle – T. oedipus, white circle – T. macrotarsus). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471118" httpUri="https://zenodo.org/record/3471118/files/figure.png" pageId="2" pageNumber="598">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8BA07A4E7A44B968FF48F815A4F85DF8" blockId="2.[151,1437,693,2014]" lastBlockId="3.[151,1437,154,502]" lastPageId="3" lastPageNumber="599" pageId="2" pageNumber="598">
The taxonomic position of
<taxonomicName id="4C1F01CD7A44B969FE61F815A5F45BD0" authority="Montreuil &amp; Frolov, 2019" authorityName="Montreuil &amp; Frolov" authorityYear="2019" box="[494,653,1990,2013]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="macrotarsus" status="sp. nov.">
<emphasis id="B96BA65C7A44B969FE61F815A5F45BD0" box="[494,653,1990,2013]" italics="true" pageId="2" pageNumber="598">T. macrotarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2581B277A44B969FD19F815A46E5BD3" box="[662,791,1990,2014]" pageId="2" pageNumber="598" rank="species">new species</taxonomicNameLabel>
within
<taxonomicName id="4C1F01CD7A44B969FCE3F815A3795BD3" box="[876,1024,1990,2014]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="B96BA65C7A44B969FCE3F815A3795BD3" box="[876,1024,1990,2014]" italics="true" pageId="2" pageNumber="598">Trigonochilus</emphasis>
</taxonomicName>
is provisional. We placed this species in this genus based on the enlarged metatibiae and metatarsi in males similar to those of
<taxonomicName id="4C1F01CD7A45B968FBDCFF49A3C35CBF" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[1107,1210,154,178]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A45B968FBDCFF49A3C35CBF" box="[1107,1210,154,178]" italics="true" pageId="3" pageNumber="599">T. oedipus</emphasis>
</taxonomicName>
. However, a number of other characters listed in the diagnoses suggest that the two species may not be closely related or even congeneric. Also close relations of the Malagasy taxa with African ones are not well supported by data. Both African species have the metatibia of males with very long, acute processes on posterior margin (
<bibRefCitation id="EF8E07BF7A45B968FC17FED5A3675D13" author="Arrow, G. J." box="[920,1054,262,286]" pageId="3" pageNumber="599" pagination="393 - 401" refId="ref2207" refString="Arrow, G. J. (1901) Remarks on secondary sexual differences in rutelid Coleoptera, with descriptions of some new forms. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 7 (7), 393 - 401. https: // doi. org / 10.1080 / 00222930108678492" type="journal article" year="1901">Arrow 1901</bibRefCitation>
;
<bibRefCitation id="EF8E07BF7A45B968FBA6FED5A3B95D13" author="Krikken J." box="[1065,1216,262,286]" pageId="3" pageNumber="599" pagination="41 - 45" refId="ref2595" refString="Krikken J. (1979) Suralcis Machatschke, a remarkable genus of African ruteline chafers. Entomologische Berichten, 39 (3), 41 - 45." type="journal article" year="1979">Krikken 1979</bibRefCitation>
). Such processes are absent in Malagasy species. Also, the tarsi and claws of the African species are rather slender while those in Malagasy species are much more robust with greatly enlarged apical tarsomeres and claws. Within Malagasy taxa, only
<taxonomicName id="4C1F01CD7A45B968FAA0FE9DA2E55D6B" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" box="[1327,1436,334,358]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="species" species="oedipus">
<emphasis id="B96BA65C7A45B968FAA0FE9DA2E55D6B" box="[1327,1436,334,358]" italics="true" pageId="3" pageNumber="599">T. oedipus</emphasis>
</taxonomicName>
shows pointed medial process on labrum, while the new species has a broadened process. Phylogenetic analysis of African and Malagasy
<taxonomicName id="4C1F01CD7A45B968FEBEFE45A6EE5DA3" authorityName="Burmeister" authorityYear="1844" box="[305,407,406,430]" class="Insecta" family="Rutelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="tribe" tribe="Adoretini">Adoretini</taxonomicName>
may suggest establishing another taxon for the Malagasy species or placing them in other nominal taxa. However, until such an analysis is available we follow Ohaus’ classification and consider both Malagasy species with enlarged metatibia and metatarsi within the genus
<taxonomicName id="4C1F01CD7A45B968FD69FE0DA4035DFB" box="[742,890,478,502]" class="Insecta" family="Scarabaeidae" genus="Trigonochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="genus">
<emphasis id="B96BA65C7A45B968FD69FE0DA4035DFB" box="[742,890,478,502]" italics="true" pageId="3" pageNumber="599">Trigonochilus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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