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<document ID-DOI="10.1590/S1679-62252008000300003" ID-ISSN="1982-0224" ID-Zenodo-Dep="4647417" approvalRequired="12" approvalRequired_for_textStreams="11" approvalRequired_for_treatments="1" checkinTime="1617111262820" checkinUser="carolina" docAuthor="Castellanos-Morales, Cesar A." docDate="2008" docId="03AFBC062034DF7BFC63FE9AFE0CF9B4" docLanguage="en" docName="NeotropIchthyol.6.3.307-314.pdf" docOrigin="Neotropical Ichthyology 6 (3)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252008000300003&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.3:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="3" docTitle="Trichomycterus uisae Castellanos-Morales 2008, new species" docType="treatment" docVersion="2" lastPageId="4" lastPageNumber="311" masterDocId="FF96C47E2035DF7FFFA0FF97FFA2FFAD" masterDocTitle="Trichomycterus uisae: a new species of hypogean catfish (Siluriformes: Trichomycteridae) from the northeastern Andean Cordillera of Colombia" masterLastPageNumber="314" masterPageNumber="307" pageId="1" pageNumber="308" updateTime="1617112864615" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Trichomycterus uisae: a new species of hypogean catfish (Siluriformes: Trichomycteridae) from the northeastern Andean Cordillera of Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Castellanos-Morales, Cesar A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
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<mods:start>307</mods:start>
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</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252008000300003</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4647417</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03AFBC062034DF7BFC63FE9AFE0CF9B4" httpUri="http://treatment.plazi.org/id/03AFBC062034DF7BFC63FE9AFE0CF9B4" lastPageId="4" lastPageNumber="311" pageId="1" pageNumber="308">
<subSubSection box="[963,1343,269,292]" pageId="1" pageNumber="308" type="nomenclature">
<paragraph blockId="1.[963,1343,269,324]" box="[963,1343,269,292]" pageId="1" pageNumber="308">
<heading box="[963,1343,269,292]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="308" reason="2">
<emphasis bold="true" box="[963,1343,269,292]" pageId="1" pageNumber="308">
<emphasis bold="true" box="[963,1208,269,292]" italics="true" pageId="1" pageNumber="308">
<taxonomicName authority="Castellanos-Morales, 2008" authorityName="Castellanos-Morales" authorityYear="2008" box="[963,1201,269,292]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="uisae" status="new species">Trichomycterus uisae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[1213,1343,269,292]" pageId="1" pageNumber="308" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1101,1204,301,324]" pageId="1" pageNumber="308" type="description">
<paragraph blockId="1.[963,1343,269,324]" box="[1101,1204,301,324]" pageId="1" pageNumber="308">
<figureCitation box="[1101,1204,301,324]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[153,192,1920,1943]" captionStartId-1="2.[158,197,620,643]" captionTargetBox-0="[151,1474,1572,1895]" captionTargetBox-1="[158,801,213,602]" captionTargetId-0="figure-557@1.[151,1474,1572,1895]" captionTargetId-1="figure-478@2.[158,801,213,602]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 2. Trichomycterus uisae, holotype, CAC-CDMB 72.5 mm Colombia. Lateral view of the left side." captionText-1="Fig. 3. Trichomycterus uisae, holotype, CAC-CDMB 72, 54 mm SL. Cueva El Misterio, upper Sogamoso River basin, Santander, Colombia. Dorsal view of head." figureDoi-0="http://doi.org/10.5281/zenodo.4647421" figureDoi-1="http://doi.org/10.5281/zenodo.4647423" httpUri-0="https://zenodo.org/record/4647421/files/figure.png" httpUri-1="https://zenodo.org/record/4647423/files/figure.png" pageId="1" pageNumber="308">Figs. 2 - 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="308" type="materials_examined">
<paragraph blockId="1.[831,1475,365,904]" pageId="1" pageNumber="308">
<materialsCitation collectingDate="2006-11-01" collectorName="Cueva El Misterio &amp; L. L. Marino-Zamudio &amp; M. Pardo-Penaloza" country="Colombia" county="Municipio de Los Santos" elevation="1600" latitude="6.8391666" location="Acuarela" longLatPrecision="21" longitude="-73.08833" municipality="Vereda Mesa de Los Santos" pageId="1" pageNumber="308" specimenCount="1" stateProvince="Santander" typeStatus="holotype">
<emphasis bold="true" box="[831,932,365,386]" pageId="1" pageNumber="308">
<typeStatus box="[831,928,365,386]" pageId="1" pageNumber="308">Holotype</typeStatus>
.
</emphasis>
CAC-CDMB
<quantity box="[1071,1157,365,386]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" pageId="1" pageNumber="308" unit="mm" value="72.5">72.5 mm</quantity>
SL,
<collectingCountry box="[1201,1300,365,386]" name="Colombia" pageId="1" pageNumber="308">Colombia</collectingCountry>
,
<collectingRegion country="Colombia" name="Santander" pageId="1" pageNumber="308">Departamento de Santander</collectingRegion>
,
<collectingCounty box="[940,1187,394,415]" pageId="1" pageNumber="308">Municipio de Los Santos</collectingCounty>
,
<collectingMunicipality box="[1197,1470,394,415]" pageId="1" pageNumber="308">Vereda Mesa de Los Santos</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03AFBC062034DF7BFC63FE9AFE0CF9B4:8ED95BCB2034DF7EFC9FFE31FC37FE16" box="[831,917,422,443]" country="Colombia" county="Municipio de Los Santos" latitude="6.8391666" longLatPrecision="21" longitude="-73.08833" municipality="Vereda Mesa de Los Santos" name="Acuarela" pageId="1" pageNumber="308" stateProvince="Santander">Acuarela</location>
road
<quantity box="[968,1035,422,443]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="308" unit="km" value="3.5">3.5 km</quantity>
,
<collectorName box="[1043,1214,422,443]" pageId="1" pageNumber="308">
<location LSID="urn:lsid:plazi:treatment:03AFBC062034DF7BFC63FE9AFE0CF9B4:8ED95BCB2034DF7EFBB3FE31FB1CFE16" box="[1043,1214,422,443]" country="Colombia" county="Municipio de Los Santos" latitude="6.8391666" longLatPrecision="21" longitude="-73.08833" municipality="Vereda Mesa de Los Santos" name="Cueva El Misterio" pageId="1" pageNumber="308" stateProvince="Santander">Cueva El Misterio</location>
</collectorName>
(
<geoCoordinate box="[1225,1342,422,443]" degrees="06" direction="north" minutes="50" orientation="latitude" pageId="1" pageNumber="308" precision="15" seconds="21" value="6.8391666">06º5021”N</geoCoordinate>
,
<geoCoordinate box="[1350,1470,422,443]" degrees="73" direction="west" minutes="05" orientation="longitude" pageId="1" pageNumber="308" precision="15" seconds="18" value="-73.08833">73º0518”W</geoCoordinate>
,
<elevation box="[831,999,451,472]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="308" unit="m" value="1600.0">
elevation
<quantity box="[924,999,451,472]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="308" unit="m" value="1600.0">1600 m</quantity>
</elevation>
),
<location LSID="urn:lsid:plazi:treatment:03AFBC062034DF7BFC63FE9AFE0CF9B4:8ED95BCB2034DF7EFC57FE54FB70FE75" box="[1015,1234,451,472]" country="Colombia" county="Municipio de Los Santos" latitude="6.8391666" longLatPrecision="21" longitude="-73.08833" municipality="Vereda Mesa de Los Santos" name="upper Sogamoso River" pageId="1" pageNumber="308" stateProvince="Santander">upper Sogamoso River</location>
basin; C. Castellanos-M,
<collectorName box="[831,1054,480,501]" pageId="1" pageNumber="308">L. L. Marino-Zamudio</collectorName>
&amp;
<collectorName box="[1086,1276,480,501]" pageId="1" pageNumber="308">M. Pardo-Peñaloza</collectorName>
,
<date box="[1286,1404,480,501]" pageId="1" pageNumber="308" value="2006-11-01">
<collectingDate box="[1286,1404,480,501]" pageId="1" pageNumber="308" value="2006-11-01">1 Nov 2006</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="1.[831,1475,365,904]" pageId="1" pageNumber="308">
<materialsCitation box="[831,1270,509,530]" location="All" pageId="1" pageNumber="308" specimenCount="2" typeStatus="paratype">
<emphasis bold="true" box="[831,941,509,530]" pageId="1" pageNumber="308">
<typeStatus box="[831,937,509,530]" pageId="1" pageNumber="308">Paratypes</typeStatus>
.
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03AFBC062034DF7BFC63FE9AFE0CF9B4:8ED95BCB2034DF7EFC13FE6AFC70FDBF" box="[947,978,509,530]" name="All" pageId="1" pageNumber="308">All</location>
collected at
<typeStatus box="[1103,1188,509,530]" pageId="1" pageNumber="308">holotype</typeStatus>
locality
</materialsCitation>
.
<materialsCitation collectingDate="2007-02-28" collectionCode="ANSP" collectorName="Marino-Zamudio" pageId="1" pageNumber="308" specimenCode="ANSP 187498" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1278,1421,509,530]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="1" pageNumber="308">ANSP 187498</specimenCode>
,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.55" pageId="1" pageNumber="308" unit="mm" value="45.5">45.5 mm</quantity>
SL, C. Castellanos-M &amp; L. L.
<collectorName box="[1168,1338,538,559]" pageId="1" pageNumber="308">Marino-Zamudio</collectorName>
,
<date box="[1347,1471,538,559]" pageId="1" pageNumber="308" value="2007-02-28">
<collectingDate box="[1347,1471,538,559]" pageId="1" pageNumber="308" value="2007-02-28">28 Feb 2007</collectingDate>
</date>
</materialsCitation>
. CAC-CDMB 88,
<quantity box="[1004,1091,566,587]" metricMagnitude="-2" metricUnit="m" metricValue="5.59" pageId="1" pageNumber="308" unit="mm" value="55.9">55.9 mm</quantity>
SL, C. Castellanos-M &amp; L. L. Marino- Zamudio,
<date box="[934,1064,595,616]" pageId="1" pageNumber="308" value="2007-02-28">28 Feb 2007</date>
.
<materialsCitation collectingDate="2007-03-01" collectorName="L. L. Marino-Zamudio &amp; Pardo-Penaloza" pageId="1" pageNumber="308" specimenCode="IAvH-P 10806, 2, 52.2" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1075,1221,595,616]" collectionCode="IAvH-P" pageId="1" pageNumber="308">IAvH-P 10806</specimenCode>
,
<specimenCode box="[1234,1248,595,616]" collectionCode="IAvH-P" pageId="1" pageNumber="308">2</specimenCode>
,
<specimenCode box="[1260,1304,595,616]" collectionCode="IAvH-P" pageId="1" pageNumber="308">52.2</specimenCode>
-57.0 mm SL, C. Castellanos-M,
<collectorName box="[989,1218,624,645]" pageId="1" pageNumber="308">L. L. Marino-Zamudio</collectorName>
&amp; M.
<collectorName box="[1293,1449,624,645]" pageId="1" pageNumber="308">Pardo-Peñaloza</collectorName>
,
<date pageId="1" pageNumber="308" value="2007-03-01">
<collectingDate pageId="1" pageNumber="308" value="2007-03-01">1 Mar 2007</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectingDate="2007-03-01" collectorName="Zamudio &amp; M. Pardo-Penaloza" pageId="1" pageNumber="308" specimenCode="UIS-T 1698, 49" specimenCount="1" typeStatus="paratype">
<specimenCode box="[934,1050,653,674]" collectionCode="UIS-T" pageId="1" pageNumber="308">UIS-T 1698</specimenCode>
,
<specimenCode box="[1059,1083,653,674]" collectionCode="UIS-T" pageId="1" pageNumber="308">49</specimenCode>
mm SL, C. Castellanos-M, L. L. Marino-
<collectorName box="[831,919,682,703]" pageId="1" pageNumber="308">Zamudio</collectorName>
&amp;
<collectorName box="[956,1149,682,703]" pageId="1" pageNumber="308">M. Pardo-Peñaloza</collectorName>
,
<date box="[1161,1284,682,703]" pageId="1" pageNumber="308" value="2007-03-01">
<collectingDate box="[1161,1284,682,703]" pageId="1" pageNumber="308" value="2007-03-01">1 Mar 2007</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectingDate="2007-03-02" collectorName="L. L. Marino-Zamudio &amp; Penaloza" pageId="1" pageNumber="308" specimenCode="UIS-T 1699" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1296,1418,682,703]" collectionCode="UIS-T" pageId="1" pageNumber="308">UIS-T 1699</specimenCode>
,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.35" pageId="1" pageNumber="308" unit="mm" value="43.5">43.5 mm</quantity>
SL, C. Castellanos-M,
<collectorName box="[1109,1336,710,731]" pageId="1" pageNumber="308">L. L. Marino-Zamudio</collectorName>
&amp; M. Pardo-
<collectorName box="[831,922,739,760]" pageId="1" pageNumber="308">Peñaloza</collectorName>
,
<date box="[932,1050,739,760]" pageId="1" pageNumber="308" value="2007-03-02">
<collectingDate box="[932,1050,739,760]" pageId="1" pageNumber="308" value="2007-03-02">2 Mar 2007</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[831,1475,365,904]" pageId="1" pageNumber="308">
<emphasis bold="true" box="[831,1023,768,789]" pageId="1" pageNumber="308">
<typeStatus box="[831,1020,768,789]" isOtherTypeStatus="true" pageId="1" pageNumber="308">Non-type material</typeStatus>
.
</emphasis>
All collected at
<typeStatus box="[1176,1260,768,789]" pageId="1" pageNumber="308">holotype</typeStatus>
locality. CAC-CDMB 90, 2 C&amp;S,
<quantity box="[947,1085,797,818]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" metricValueMax="4.11" metricValueMin="3.21" pageId="1" pageNumber="308" unit="mm" value="36.6" valueMax="41.1" valueMin="32.1">32.1-41.1 mm</quantity>
SL,
<date box="[1132,1249,797,818]" pageId="1" pageNumber="308" value="2007-03-01">1 Mar 2007</date>
. CAC-CDMB 91,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.55" pageId="1" pageNumber="308" unit="mm" value="45.5">45.5 mm</quantity>
SL,
<date box="[917,1035,826,847]" pageId="1" pageNumber="308" value="2007-03-02">2 Mar 2007</date>
. CAC-CDMB 92, C&amp;S,
<quantity box="[1279,1367,826,847]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="1" pageNumber="308" unit="mm" value="28.3">28.3 mm</quantity>
SL,
<date pageId="1" pageNumber="308" value="2007-03-02">2 Mar 2007</date>
. UIS-T 1700,
<quantity box="[1015,1099,854,875]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" pageId="1" pageNumber="308" unit="mm" value="39.8">39.8 mm</quantity>
SL, C. Castellanos-M. &amp; L. L. Marino- Zamudio,
<date box="[932,1050,883,904]" pageId="1" pageNumber="308" value="2007-03-02">2 Mar 2007</date>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="309" pageId="1" pageNumber="308" type="diagnosis">
<paragraph blockId="1.[831,1474,938,1537]" pageId="1" pageNumber="308">
<emphasis bold="true" box="[831,949,938,961]" pageId="1" pageNumber="308">Diagnosis.</emphasis>
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[954,1184,938,961]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[954,1184,938,961]" italics="true" pageId="1" pageNumber="308">Trichomycterus uisae</emphasis>
</taxonomicName>
(
<figureCitation box="[1197,1293,938,961]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[153,192,1920,1943]" captionStartId-1="2.[158,197,620,643]" captionTargetBox-0="[151,1474,1572,1895]" captionTargetBox-1="[158,801,213,602]" captionTargetId-0="figure-557@1.[151,1474,1572,1895]" captionTargetId-1="figure-478@2.[158,801,213,602]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 2. Trichomycterus uisae, holotype, CAC-CDMB 72.5 mm Colombia. Lateral view of the left side." captionText-1="Fig. 3. Trichomycterus uisae, holotype, CAC-CDMB 72, 54 mm SL. Cueva El Misterio, upper Sogamoso River basin, Santander, Colombia. Dorsal view of head." figureDoi-0="http://doi.org/10.5281/zenodo.4647421" figureDoi-1="http://doi.org/10.5281/zenodo.4647423" httpUri-0="https://zenodo.org/record/4647421/files/figure.png" httpUri-1="https://zenodo.org/record/4647423/files/figure.png" pageId="1" pageNumber="308">Figs. 2-3</figureCitation>
) is distinguished from both epigean and troglomorphic species of the genus by the following combination of characters: anterior and posterior fontanels separated but connected by a narrow channel (
<emphasis box="[881,903,1066,1089]" italics="true" pageId="1" pageNumber="308">vs</emphasis>
. fontanels separated by epiphyseal bar); five to nine opercular odontodes (
<emphasis box="[1059,1081,1098,1121]" italics="true" pageId="1" pageNumber="308">vs</emphasis>
. 8 to 16); reduced mouth width (32.6- 41.2% of HL); relatively deep head (48.6-66.3% of HL); reduced pigmentation (except for epigean
<taxonomicName authorityName="Fernandez &amp; Schaefer" authorityYear="2005" box="[1254,1368,1162,1185]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="gorgona">
<emphasis box="[1254,1368,1162,1185]" italics="true" pageId="1" pageNumber="308">T. gorgona</emphasis>
</taxonomicName>
and caverestricted congeners
<emphasis italics="true" pageId="1" pageNumber="308">
<taxonomicName box="[1075,1205,1194,1217]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="chaberti">T. chaberti</taxonomicName>
,
<taxonomicName box="[1221,1438,1194,1217]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="itacarambiensis">T. itacarambiensis</taxonomicName>
,
<taxonomicName class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="sandovali">T. sandovali</taxonomicName>
,
<taxonomicName box="[949,1135,1226,1249]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="santanderensis">T. santanderensis</taxonomicName>
</emphasis>
and
<taxonomicName box="[1185,1304,1226,1249]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="spelaeus">
<emphasis box="[1185,1304,1226,1249]" italics="true" pageId="1" pageNumber="308">T. spelaeus</emphasis>
</taxonomicName>
) consisting of a narrow predorsal dark bluish-gray stripe from nape to origin of dorsal fin and ground coloration light-brown without spots (
<emphasis box="[840,862,1322,1345]" italics="true" pageId="1" pageNumber="308">vs</emphasis>
. pigmented body with dark spots); variable reduction of eyes from 7.4 to 11.1% of HL (except for epigean
<taxonomicName authorityName="Fernandez &amp; Schaefer" authorityYear="2005" box="[1358,1473,1354,1377]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="gorgona">
<emphasis box="[1358,1473,1354,1377]" italics="true" pageId="1" pageNumber="308">T. gorgona</emphasis>
</taxonomicName>
and hypogean
<emphasis box="[1013,1385,1386,1409]" italics="true" pageId="1" pageNumber="308">
<taxonomicName box="[1013,1150,1386,1409]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="chaberti">T. chaberti</taxonomicName>
,
<taxonomicName box="[1168,1385,1386,1409]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="itacarambensis">T. itacarambensis</taxonomicName>
</emphasis>
and
<taxonomicName class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="santanderensis">
<emphasis italics="true" pageId="1" pageNumber="308">T. santanderensis</emphasis>
</taxonomicName>
); extended nasal, maxillary and rictal barbels (85.2-108%, 93.6-111.8% and 55.1-70.2% of HL, respectively); caudal fin with slightly convex margin, dorsal lobe of caudalfin longer than ventral lobe. Other characters shared with
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647421" ID-Zenodo-Dep="4647421" httpUri="https://zenodo.org/record/4647421/files/figure.png" pageId="1" pageNumber="308" startId="1.[153,192,1920,1943]" targetBox="[151,1474,1572,1895]" targetPageId="1">
<paragraph blockId="1.[153,805,1920,1975]" pageId="1" pageNumber="308">
<emphasis bold="true" box="[153,221,1920,1943]" pageId="1" pageNumber="308">Fig. 2.</emphasis>
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[226,451,1920,1943]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="308" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[226,451,1920,1943]" italics="true" pageId="1" pageNumber="308">Trichomycterus uisae</emphasis>
</taxonomicName>
, holotype, CAC-CDMB 72.5 mm Colombia. Lateral view of the left side.
</paragraph>
</caption>
<paragraph blockId="1.[810,1476,1920,1943]" lastBlockId="2.[158,801,760,943]" lastPageId="2" lastPageNumber="309" pageId="1" pageNumber="308">
SL. Cueva El Misterio, upper Sogamoso River basin,
<collectingRegion box="[1365,1472,1920,1943]" country="Colombia" name="Santander" pageId="1" pageNumber="308">Santander</collectingRegion>
, various other species of
<taxonomicName authorityName="Valenciennes in Humboldt" authorityYear="1832" box="[420,588,760,783]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis box="[420,588,760,783]" italics="true" pageId="2" pageNumber="309">Trichomycterus</emphasis>
</taxonomicName>
yet useful for identification include origin of pelvic fin anterior to vertical through dorsal-fin origin; short caudal peduncle (19-21.9% of SL); tip of first pectoral-fin ray prolonged as a long filament (52.1- 89% of pectoral-fin length) and pelvic fins not widely separated.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647423" ID-Zenodo-Dep="4647423" httpUri="https://zenodo.org/record/4647423/files/figure.png" pageId="2" pageNumber="309" startId="2.[158,197,620,643]" targetBox="[158,801,213,602]" targetPageId="2">
<paragraph blockId="2.[158,801,620,707]" pageId="2" pageNumber="309">
<emphasis bold="true" box="[158,228,620,643]" pageId="2" pageNumber="309">Fig. 3.</emphasis>
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[235,465,620,643]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="309" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[235,465,620,643]" italics="true" pageId="2" pageNumber="309">Trichomycterus uisae</emphasis>
</taxonomicName>
, holotype, CAC-CDMB 72, 54 mm SL. Cueva El Misterio, upper Sogamoso River basin, Santander, Colombia. Dorsal view of head.
</paragraph>
</caption>
<subSubSection lastPageId="4" lastPageNumber="311" pageId="2" pageNumber="309" type="description">
<paragraph blockId="2.[158,802,984,1935]" pageId="2" pageNumber="309">
<emphasis bold="true" box="[158,299,984,1007]" pageId="2" pageNumber="309">Description.</emphasis>
Morphometric data presented in
<tableCitation box="[652,731,984,1007]" captionStart="Table 1" captionStartId="5.[831,894,1280,1303]" captionText="Table 1. Morphometric data of Trichomycterus uisae. Standard length is expressed in millimeters. Measurements 2 to 11 are expressed as percents of standard length, measurements 12 to 19 are expressed as percents of head length." pageId="2" pageNumber="309">Table 1</tableCitation>
. Body elongated, deeper than wide, gradually deeper from trunk toward caudal peduncle; dorsal profile of trunk convex. Ventral profile of trunk straight. Dorsal and ventral profiles of caudal peduncle slightly convex. Integument thick with lateral cutaneous folds forming vertical rings between pectoral and anal fins in specimens preserved in alcohol. Anal and urogenital openings closer to pelvic-fin base than to anal-fin origin, totally covered when pelvic fin extended. Free vertebrae 35. Pleural ribs 11-12.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647425" ID-Zenodo-Dep="4647425" httpUri="https://zenodo.org/record/4647425/files/figure.png" pageId="2" pageNumber="309" startId="2.[839,878,991,1014]" targetBox="[846,1470,227,946]" targetPageId="2">
<paragraph blockId="2.[839,1481,991,1206]" pageId="2" pageNumber="309">
<emphasis bold="true" box="[839,910,991,1014]" pageId="2" pageNumber="309">Fig. 4.</emphasis>
Dorsal view of the neurocranium of
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="309" phylum="Chordata" rank="species" species="uisae">
<emphasis italics="true" pageId="2" pageNumber="309">Trichomycterus uisae</emphasis>
</taxonomicName>
. CAC-CDMB 90, 32.1 mm SL. af: anterior fontanel; f: frontal; i1-3: infraorbital sensory pores 1-3; i10-11: infraorbital sensory pores 10-11; le: lateral ethmoid; me: mesethmoid; pf: posterior fontanel; psc: postotic sensory canal; s+pro+ptr: sphenotic+ prootic+pterosphenoid; su: supraoccipital; ssc: supraorbital sensory canal; w: Weberian capsule. Scale bar =1 mm.
</paragraph>
</caption>
<paragraph blockId="2.[158,802,984,1935]" lastBlockId="2.[839,1482,1304,1935]" pageId="2" pageNumber="309">
Head wide, trapezoidal, depressed in dorsal view. Dorsal profile of head straight, ventral and lateral profiles convex. Eye small, black, rounded, well defined, with variable diameter, positioned dorsally on anterior half of head. Mouth subterminal, with corners oriented backwards. Lower lip with conspicuously fleshy lateral lobes. Teeth conical, curved, arranged in 3-4 irregular rows on upper jaw and three rows on lower jaw. Jaw muscles not particularly developed and not bulging from surface of head. Neurocranium with elongate mesethmoid T-shape. Anterior fontanel small, oval in shape, located between frontals at level of infraorbital laterosensory canal exit 10-11. Posterior fontanel long and broad. Anterior one-third of posterior fontanel situated between frontal bones, with remainder in anterior portion of supraoccipital bone (
<figureCitation box="[223,288,1752,1775]" captionStart="Fig" captionStartId="2.[839,878,991,1014]" captionTargetBox="[846,1470,227,946]" captionTargetId="figure-504@2.[836,1479,215,961]" captionTargetPageId="2" captionText="Fig. 4. Dorsal view of the neurocranium of Trichomycterus uisae. CAC-CDMB90,32.1 mmSL.af:anteriorfontanel;f:frontal; i1-3: infraorbital sensory pores 1-3; i10-11: infraorbital sensory pores 10-11; le: lateral ethmoid; me: mesethmoid; pf: posterior fontanel; psc: postotic sensory canal; s+pro+ptr: sphenotic+ prootic+pterosphenoid; su: supraoccipital; ssc: supraorbital sensory canal; w: Weberian capsule. Scale bar =1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647425" httpUri="https://zenodo.org/record/4647425/files/figure.png" pageId="2" pageNumber="309">Fig. 4</figureCitation>
). Both fontanels connected by a narrow channel. Branchial membranes thick, united to isthmus anteromedially and forming free fold across isthmus. Gill opening wide. Seven branchiostegal rays (
<figureCitation box="[381,459,1848,1871]" captionStart="Fig" captionStartId="3.[151,190,1344,1367]" captionTargetBox="[198,774,220,1316]" captionTargetId="figure-296@3.[151,794,198,1321]" captionTargetPageId="3" captionText="Fig. 5. a. Left hyoid arch of Trichomycterus uisae in ventral view.CAC-CDMB 90, 32.1 mm SL. ac:anterior ceratohyal; br: branchiostegal rays; pc: posterior ceratohyal; vh: hypohyal. b. Opercular series of Trichomycterus uisae. op: opercle; io: interopercle. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647429" httpUri="https://zenodo.org/record/4647429/files/figure.png" pageId="2" pageNumber="309">Fig. 5a</figureCitation>
), six externally visible from below. Nasal and maxillary barbels surpassing base of pectoral fin. Maxillary barbel longer than nasal barbel. Anterior nostril surrounded by slightly raised thick integument, continuous with nasal barbel, both forming a tubular-shaped structure around nostril. Posterior nostril oriented transversally, its anterior edge delimited by thin and long flap of integument. Interopecular patch of odontodes well developed, with 27-32 conical and elongated odontodes arranged in four irregular rows, with interopercular odontodes on posterior edge larger. Opercular patch of odontodes small, with 5-9 conical odontodes arranged in three irregular rows (
<figureCitation box="[1306,1386,1560,1583]" captionStart="Fig" captionStartId="3.[151,190,1344,1367]" captionTargetBox="[198,774,220,1316]" captionTargetId="figure-296@3.[151,794,198,1321]" captionTargetPageId="3" captionText="Fig. 5. a. Left hyoid arch of Trichomycterus uisae in ventral view.CAC-CDMB 90, 32.1 mm SL. ac:anterior ceratohyal; br: branchiostegal rays; pc: posterior ceratohyal; vh: hypohyal. b. Opercular series of Trichomycterus uisae. op: opercle; io: interopercle. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647429" httpUri="https://zenodo.org/record/4647429/files/figure.png" pageId="2" pageNumber="309">Fig. 5b</figureCitation>
).
</paragraph>
<paragraph blockId="2.[839,1482,1304,1935]" lastBlockId="3.[151,794,1541,1948]" lastPageId="3" lastPageNumber="310" pageId="2" pageNumber="309">
Pectoral fin rounded with i,8 rays. First ray thin and fragile, prolonged as a long filament. Scapulocoracoid with long anteriorly directed process, located close to first pectoral-fin ray base (
<figureCitation box="[948,1018,1688,1711]" captionStart="Fig" captionStartId="3.[831,870,1201,1224]" captionTargetBox="[850,1443,207,1150]" captionTargetId="figure-341@3.[831,1475,192,1158]" captionTargetPageId="3" captionText="Fig. 6. Ventral view of the right pectoral girdle and fin of Trichomycterus uisae. CAC-CDMB 90, 41.1 mm SL. ap: anterior process; pr: pectoral radial; r: rays. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647427" httpUri="https://zenodo.org/record/4647427/files/figure.png" pageId="2" pageNumber="309">Fig. 6</figureCitation>
). Dorsal-fin rounded, located posterior to vertical through midbody, with iv,7 rays (only two unbranched rays externally visible). First pterygiophore inserted between neural spines of 15-16
<superScript attach="left" box="[1072,1084,1782,1796]" fontSize="6" pageId="2" pageNumber="309">th</superScript>
free vertebra. Pelvic-fin rays i,4, with a lateral splint. Pelvic-fin origin anterior to vertical through dorsal-fin origin and its posterior edge slightly surpasses urogenital opening. Pelvic-fin bases not widely separated. Basipterygium with two long anterior processes narrowing from base to distal tip, one or two medial processes and one short posterior process (
<figureCitation box="[404,468,1573,1596]" captionStart="Fig" captionStartId="4.[158,197,862,885]" captionTargetBox="[206,784,204,830]" captionTargetId="figure-343@4.[158,801,191,847]" captionTargetPageId="4" captionText="Fig. 7. Ventral view of the basipterygium of Trichomycterus uisae. CAC-CDMB 90, 41.1 mm SL. ap: anterior process; mp: medial process; pp: posterior process. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647431" httpUri="https://zenodo.org/record/4647431/files/figure.png" pageId="3" pageNumber="310">Fig. 7</figureCitation>
). Anal-fin similar to dorsal fin, but smaller, with ii, 5 rays, its origin at level of last dorsal-fin ray. First pterygiophore inserted between hemal spines of free vertebrae 20-21. Caudal-fin rays i, 5+6, i. Parhypural and hypurals 1+2 associated with eight rays, hypural 3 with 3 rays, and hypurals 4+5 with 3 rays. 17-18 dorsal and 12 ventral procurrent rays. Caudal-fin edge slightly convex, uppermost rays larger. Caudal skeleton with neural spine of preural centrum 2 well developed. Hypurals 1 and 2 fused to parhypural; hypural 3 partially fused to hypurals 4+
<quantity box="[685,723,1861,1884]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="310" unit="in" value="5.0">5 in</quantity>
young specimens (
<figureCitation box="[278,357,1893,1916]" captionStart="Fig" captionStartId="4.[839,878,1304,1327]" captionTargetBox="[854,1480,204,1296]" captionTargetId="figure-342@4.[840,1483,191,1304]" captionTargetPageId="4" captionText="Fig. 8. Caudal skeleton of Trichomycterus uisae. a: CAC- CDMB 92, 28.3 mm SL, young specimen. b: CAC-CDMB 90, 41.1 mm SL, adult specimen. a: apophysis; hy 1-5: hypurals 1- 5; un: uroneural; ph: parhypural; pu1+u1: preural centrum 1+ural centrum 1; pu2: preural centrum 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647433" httpUri="https://zenodo.org/record/4647433/files/figure.png" pageId="3" pageNumber="310">Fig. 8a</figureCitation>
), and separated in adult specimens (
<figureCitation captionStart="Fig" captionStartId="4.[839,878,1304,1327]" captionTargetBox="[854,1480,204,1296]" captionTargetId="figure-342@4.[840,1483,191,1304]" captionTargetPageId="4" captionText="Fig. 8. Caudal skeleton of Trichomycterus uisae. a: CAC- CDMB 92, 28.3 mm SL, young specimen. b: CAC-CDMB 90, 41.1 mm SL, adult specimen. a: apophysis; hy 1-5: hypurals 1- 5; un: uroneural; ph: parhypural; pu1+u1: preural centrum 1+ural centrum 1; pu2: preural centrum 2. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4647433" httpUri="https://zenodo.org/record/4647433/files/figure.png" pageId="3" pageNumber="310">Fig. 8b</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647427" ID-Zenodo-Dep="4647427" httpUri="https://zenodo.org/record/4647427/files/figure.png" pageId="3" pageNumber="310" startId="3.[831,870,1201,1224]" targetBox="[850,1443,207,1150]" targetPageId="3">
<paragraph blockId="3.[831,1473,1201,1288]" pageId="3" pageNumber="310">
<emphasis bold="true" box="[831,905,1201,1224]" pageId="3" pageNumber="310">Fig. 6.</emphasis>
Ventral view of the right pectoral girdle and fin of
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[831,1054,1233,1256]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="310" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[831,1054,1233,1256]" italics="true" pageId="3" pageNumber="310">Trichomycterus uisae</emphasis>
</taxonomicName>
. CAC-CDMB 90, 41.1 mm SL. ap: anterior process; pr: pectoral radial; r: rays. Scale bar = 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647429" ID-Zenodo-Dep="4647429" httpUri="https://zenodo.org/record/4647429/files/figure.png" pageId="3" pageNumber="310" startId="3.[151,190,1344,1367]" targetBox="[198,774,220,1316]" targetPageId="3">
<paragraph blockId="3.[151,794,1344,1495]" pageId="3" pageNumber="310">
<emphasis bold="true" box="[151,247,1344,1367]" pageId="3" pageNumber="310">Fig. 5. a.</emphasis>
Left hyoid arch of
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[454,685,1344,1367]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="310" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[454,685,1344,1367]" italics="true" pageId="3" pageNumber="310">Trichomycterus uisae</emphasis>
</taxonomicName>
in ventral view. CAC-CDMB 90, 32.1 mm SL. ac: anterior ceratohyal; br: branchiostegal rays; pc: posterior ceratohyal; vh: hypohyal.
<emphasis bold="true" box="[151,172,1440,1463]" pageId="3" pageNumber="310">b.</emphasis>
Opercular series of
<emphasis box="[388,626,1440,1463]" italics="true" pageId="3" pageNumber="310">
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[388,622,1440,1463]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="310" phylum="Chordata" rank="species" species="uisae">Trichomycterus uisae</taxonomicName>
.
</emphasis>
op: opercle; io: interopercle. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph blockId="3.[831,1474,1383,1502]" pageId="3" pageNumber="310">
<emphasis bold="true" box="[831,1132,1383,1406]" pageId="3" pageNumber="310">Coloration in live specimens.</emphasis>
Body color light-brown-(
<emphasis bold="true" box="[1384,1409,1383,1406]" pageId="3" pageNumber="310">M</emphasis>
10YR - 5/6). Base of all fins with yellow tones from base to edge (
<emphasis bold="true" box="[1449,1474,1415,1438]" pageId="3" pageNumber="310">M</emphasis>
5YR 6/8 to
<emphasis bold="true" box="[968,993,1447,1470]" pageId="3" pageNumber="310">M</emphasis>
2.5Y 7/6). One predorsal narrow dark bluish gray band (
<emphasis bold="true" box="[946,971,1479,1502]" pageId="3" pageNumber="310">M</emphasis>
G-5PB).
</paragraph>
<paragraph blockId="3.[831,1474,1543,1662]" pageId="3" pageNumber="310">
<emphasis bold="true" box="[831,1083,1543,1566]" pageId="3" pageNumber="310">Coloration in alcohol.</emphasis>
All specimens preserved in alcohol with ground color yellowish (
<emphasis bold="true" box="[1146,1171,1575,1598]" pageId="3" pageNumber="310">M</emphasis>
10YR - 5/6). Base of all fins with yellow tones (
<emphasis bold="true" box="[1031,1056,1607,1630]" pageId="3" pageNumber="310">M</emphasis>
2.5Y 7/6), with exception of two nontype specimens with ground color brown-yellowish.
</paragraph>
<paragraph blockId="3.[831,1474,1703,1950]" lastBlockId="4.[158,802,994,1337]" lastPageId="4" lastPageNumber="311" pageId="3" pageNumber="310">
<emphasis bold="true" box="[831,1019,1703,1726]" pageId="3" pageNumber="310">Ecological notes.</emphasis>
The Cueva El Misterio is located at the east side of a plateau known as Mesa de los Santos, in the Municipio de los Santos. The plateau has Cretaceous sedimentary rocks and limestones of the Rosablanca Formation (
<bibRefCitation author="Williams, J" box="[839,1007,1831,1854]" pageId="3" pageNumber="310" refId="ref5417" refString="Williams, J. 1990. Barrita y fluorita en la Mesa de Los Santos y" type="book" year="1990">Williams, 1990</bibRefCitation>
); reaches 1800 meters above sea level and is located on the oriental versant of the Chicamocha Canyon, in the Colombian Andes, of
<collectingRegion box="[1111,1358,1895,1918]" country="Colombia" name="Santander" pageId="3" pageNumber="310">Santander Department</collectingRegion>
. The cave, with a total of
<quantity box="[978,1044,1927,1950]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="310" unit="m" value="110.0">110 m</quantity>
of explored passages, is isolated from the epigean stream and is oriented longitudinally with galleries formed by gentle slope tunnels and narrow passageways. The cave has small wells interconnected by reduced descending channels where water infiltration was observed. In dry months, an isolated sump pool was observed at each gallery. The bottom of each well is rocky and contains much sediment composed chiefly of bat excrement. Cydnid bugs (Hemiptera: Heteroptera) were found inside the wells. Diptera, crayfish and bats were observed in the interior of the galleries (
<figureCitation box="[166,230,1282,1305]" captionStart="Fig" captionStartId="5.[151,190,1117,1140]" captionTargetBox="[151,794,192,1097]" captionTargetId="figure-518@5.[151,794,192,1097]" captionTargetPageId="5" captionText="Fig. 9. Inside Cueva El Misterio, upper Sogamoso River basin, Santander, Colombia. Type-locality of Trichomycterus uisae." figureDoi="http://doi.org/10.5281/zenodo.4647435" httpUri="https://zenodo.org/record/4647435/files/figure.png" pageId="4" pageNumber="311">Fig. 9</figureCitation>
). Water temperature, when the
<typeStatus box="[554,647,1282,1305]" pageId="4" pageNumber="311">holotype</typeStatus>
was collected, was 20°C and cave temperature was 18.5°C.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647431" ID-Zenodo-Dep="4647431" httpUri="https://zenodo.org/record/4647431/files/figure.png" pageId="4" pageNumber="311" startId="4.[158,197,862,885]" targetBox="[206,784,204,830]" targetPageId="4">
<paragraph blockId="4.[158,801,862,949]" pageId="4" pageNumber="311">
<emphasis bold="true" box="[158,228,862,885]" pageId="4" pageNumber="311">Fig. 7.</emphasis>
Ventral view of the basipterygium of
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="311" phylum="Chordata" rank="species" species="uisae">
<emphasis italics="true" pageId="4" pageNumber="311">Trichomycterus uisae</emphasis>
</taxonomicName>
. CAC-CDMB 90, 41.1 mm SL. ap: anterior process; mp: medial process; pp: posterior process. Scale bar = 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647433" ID-Zenodo-Dep="4647433" httpUri="https://zenodo.org/record/4647433/files/figure.png" pageId="4" pageNumber="311" startId="4.[839,878,1304,1327]" targetBox="[854,1480,204,1296]" targetPageId="4">
<paragraph blockId="4.[839,1481,1304,1455]" pageId="4" pageNumber="311">
<emphasis bold="true" box="[839,913,1304,1327]" pageId="4" pageNumber="311">Fig. 8.</emphasis>
Caudal skeleton of
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[1138,1372,1304,1327]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="311" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[1138,1372,1304,1327]" italics="true" pageId="4" pageNumber="311">Trichomycterus uisae</emphasis>
</taxonomicName>
. a: CAC- CDMB 92, 28.3 mm SL, young specimen. b: CAC-CDMB 90, 41.1 mm SL, adult specimen. a: apophysis; hy 1-5: hypurals 1- 5; un: uroneural; ph: parhypural; pu1+u1: preural centrum 1+ural centrum 1; pu2: preural centrum 2. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection pageId="4" pageNumber="311" type="etymology">
<paragraph blockId="4.[158,801,1378,1497]" pageId="4" pageNumber="311">
<emphasis bold="true" box="[158,288,1378,1401]" pageId="4" pageNumber="311">Etymology.</emphasis>
The specific epithet “
<taxonomicName authorityName="Castellanos-Morales" authorityYear="2008" box="[533,590,1378,1401]" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="311" phylum="Chordata" rank="species" species="uisae">
<emphasis box="[533,590,1378,1401]" italics="true" pageId="4" pageNumber="311">uisae</emphasis>
</taxonomicName>
” refers to the acronym UIS corresponding to the Universidad Industrial de
<collectingRegion box="[158,267,1442,1465]" country="Colombia" name="Santander" pageId="4" pageNumber="311">Santander</collectingRegion>
, in the
<collectingRegion box="[339,634,1442,1465]" country="Colombia" name="Santander" pageId="4" pageNumber="311">Departamento de Santander</collectingRegion>
,
<collectingCountry box="[643,752,1442,1465]" name="Colombia" pageId="4" pageNumber="311">Colombia</collectingCountry>
. The name is used as a noun in apposition.
</paragraph>
<paragraph blockId="4.[158,430,1538,1561]" box="[158,430,1538,1561]" pageId="4" pageNumber="311">
<emphasis bold="true" box="[158,332,1538,1561]" pageId="4" pageNumber="311">Common name.</emphasis>
Trepador
</paragraph>
</subSubSection>
</treatment>
</document>