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<document id="82B5BD84FE38A56EF6DA1B8A98899B60" ID-DOI="10.11646/zootaxa.2288.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10093582" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629654284239" checkinUser="felipe" docAuthor="HENDRICH, LARS, HAWLITSCHEK, OLIVER &amp; BALKE, MICHAEL" docDate="2009" docId="03AF802609790308FF2654C13576F802" docLanguage="en" docName="zt02288p041.pdf" docOrigin="Zootaxa 2288 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.2288.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Neobidessodes Hendrich &amp; Balke, gen.n." docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FF96F85E097F030EFFB155633620FFEB" masterDocTitle="The epigean Australasian species of Neobidessodes gen. n. diving beetles- a revision integrating morphology, cybertaxonomy, DNA taxonomy and phylogeny (Coleoptera: Dytiscidae, Bidessini)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="7" updateTime="1699563497581" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6969AAAC64B94BE8542E4769C80F2DB8">The epigean Australasian species of Neobidessodes gen. n. diving beetles- a revision integrating morphology, cybertaxonomy, DNA taxonomy and phylogeny (Coleoptera: Dytiscidae, Bidessini)</mods:title>
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<mods:namePart id="8A53DA302D33FB16C0B695E4BF76523E">HENDRICH, LARS</mods:namePart>
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<mods:namePart id="5D8525D88010AF9E1DCA38F06BA97ABF">HAWLITSCHEK, OLIVER</mods:namePart>
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<mods:namePart id="4523689B137CFCA3B02056306840E9CD">BALKE, MICHAEL</mods:namePart>
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<mods:title id="8636D42A28D7CC634D4F530AFE2A5405">Zootaxa</mods:title>
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<mods:date id="C71E6D0A3E30E8E201943C75B7108400">2009</mods:date>
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<mods:number id="F74690D20863943DD168C12608024B82">2288</mods:number>
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<mods:identifier id="F950CEE0221ADE8C87017DB3D3FC6C3A" type="DOI">10.11646/zootaxa.2288.1.1</mods:identifier>
<mods:identifier id="B9AF7565F8E5E934EA7C6FA04BA6B033" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C31C62BB09790308FF2654C134A5FE57" box="[151,645,418,444]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BB9313009790308FF2654C134A5FE57" blockId="6.[151,645,418,479]" box="[151,645,418,444]" pageId="6" pageNumber="7">
<heading id="D0F1865C09790308FF2654C134A5FE57" bold="true" box="[151,645,418,444]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B972ED2209790308FF2654C134A5FE57" bold="true" box="[151,645,418,444]" pageId="6" pageNumber="7">
<taxonomicName id="4C064AB309790308FF2654C13413FE57" ID-CoL="63DPZ" authority="Hendrich &amp; Balke" authorityName="Hendrich &amp; Balke" box="[151,563,418,444]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FF2654C13767FE57" bold="true" box="[151,327,418,444]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
Hendrich &amp; Balke
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<taxonomicNameLabel id="A241505909790308FD8B54C134A5FE57" box="[570,645,418,444]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C31C62BB09790308FF2654A6373EFE34" box="[151,286,453,479]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BB9313009790308FF2654A6373EFE34" blockId="6.[151,645,418,479]" box="[151,286,453,479]" pageId="6" pageNumber="7">
(
<figureCitation id="133D2DB509790308FF2E54A63735FE34" box="[159,277,453,479]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–8" captionStart-2="FIGURES 9–12" captionStart-3="FIGURES 13–17" captionStart-4="FIGURES 18–20" captionStart-5="FIGURES 21" captionStartId-0="7.[151,270,1900,1924]" captionStartId-1="8.[151,275,1973,1997]" captionStartId-2="9.[151,274,1833,1857]" captionStartId-3="11.[151,269,1942,1966]" captionStartId-4="12.[151,269,1750,1774]" captionStartId-5="13.[151,269,1906,1930]" captionTargetBox-0="[173,1401,199,1857]" captionTargetBox-1="[155,1427,235,1928]" captionTargetBox-2="[158,1433,206,1780]" captionTargetBox-3="[211,1388,187,1914]" captionTargetBox-4="[171,1417,199,1703]" captionTargetBox-5="[155,1431,1125,1855]" captionTargetId-0="figure-77@7.[167,1411,180,1869]" captionTargetId-1="figure-48@8.[151,1436,194,1950]" captionTargetId-2="figure-68@9.[148,1433,194,1809]" captionTargetId-3="figure-67@11.[197,1403,164,1924]" captionTargetId-4="figure-116@12.[151,1436,187,1726]" captionTargetId-5="figure-361@13.[151,1436,1125,1882]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionText-0="FIGURES 1–4. Habitus of 1) N. denticulatus (WA, Pilbara, Millstream Chichester National Park, Fortescue River side branch); 2) N. denticulatus (NT, Kakadu N.P., Jim Jim District, Gungurul Lookout); 3) N. flavosignatus (WA, East Kimberley, Gibb Range, Gibb River Road, Russ Creek Crossing); 4) N. flavosignatus (NT, Manton Dam Recreation Area, 46 km S Darwin) (scale bar = 1 mm), (Photos: A. Riedel)." captionText-1="FIGURES 5–8. Habitus of 5) Neobidessodes samkrisi sp.n. (holotype, male); 6) N. grossus; 7) N. mjobergi; 8) Neobidessodes bilita (female) (scale bar = 1 mm) (Photos: A. Riedel)." captionText-2="FIGURES 9–12. 9) N. thoracicus sp.n. (paratype); 10) N. thoracicus sp.n. (paratype, “black specimen”); 11) N. thoracicus sp.n. (“light form”, WA, Kimberley Region, 50 km S Wyndham, Black Flag Creek); 12) N. thoracicus sp.n. (“light form”, NT, Kakadu N.P., Jim Jim District, Gungurul Lookout) (scale bar = 1 mm) (Photos: A. Riedel)." captionText-3="FIGURES 13–17. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 13) Neobidessodes thoracicus sp.n. (paratype); 14) N. thoracicus sp.n. (“light form”); 15) N. flavosignatus; 16) N. mjobergi and 17) N. samkrisi sp.n. (scale bar = 0.5 mm) (Photos: L. Hendrich)." captionText-4="FIGURES 18–20. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 18) Neobidessodes bilita; 19) N. denticulatus and 20) N. grossus (scale bar = 0.5 mm) (Photos: L. Hendrich)." captionText-5="FIGURES 21. Denticulate tip of elytra in Neobidessodes denticulatus (Photo: L. Hendrich)." figureDoi-0="http://doi.org/10.5281/zenodo.10093584" figureDoi-1="http://doi.org/10.5281/zenodo.10093586" figureDoi-2="http://doi.org/10.5281/zenodo.10093588" figureDoi-3="http://doi.org/10.5281/zenodo.10093590" figureDoi-4="http://doi.org/10.5281/zenodo.10093592" figureDoi-5="http://doi.org/10.5281/zenodo.10093594" httpUri-0="https://zenodo.org/record/10093584/files/figure.png" httpUri-1="https://zenodo.org/record/10093586/files/figure.png" httpUri-2="https://zenodo.org/record/10093588/files/figure.png" httpUri-3="https://zenodo.org/record/10093590/files/figure.png" httpUri-4="https://zenodo.org/record/10093592/files/figure.png" httpUri-5="https://zenodo.org/record/10093594/files/figure.png" pageId="6" pageNumber="7">Figs 1–21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09790308FF26577135C9FDC7" box="[151,1001,530,556]" pageId="6" pageNumber="7" type="type_taxon">
<paragraph id="8BB9313009790308FF26577135C9FDC7" blockId="6.[151,1436,530,2026]" box="[151,1001,530,556]" pageId="6" pageNumber="7">
<emphasis id="B972ED2209790308FF2657713712FDC7" bold="true" box="[151,306,530,556]" pageId="6" pageNumber="7">
<typeStatus id="54BD8F9209790308FF26577136F4FDC7" box="[151,212,530,556]" pageId="6" pageNumber="7">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C064AB309790308FEF1577134F2FDC7" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[320,722,530,556]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B972ED2209790308FEF15771341DFDC0" box="[320,573,530,555]" italics="true" pageId="6" pageNumber="7">Bidessus denticulatus</emphasis>
<bibRefCitation id="EF974CC109790308FDF4577134F2FDC7" author="Sharp, D." box="[581,722,530,556]" pageId="6" pageNumber="7" pagination="179 - 1003" refId="ref29626" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. S cientific Transactions of the Royal Dublin Society, (2) 2, 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09790308FF77575B36D3FD2C" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BB9313009790308FF77575B36D3FD2C" blockId="6.[151,1436,530,2026]" pageId="6" pageNumber="7">
<emphasis id="B972ED2209790308FF77575B3766FDB9" bold="true" box="[198,326,568,594]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
<taxonomicName id="4C064AB309790308FEE5575A3522FDB9" authority="Hendrich &amp; Balke" authorityName="Hendrich &amp; Balke" box="[340,770,568,594]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FEE5575A3422FDB9" box="[340,514,569,594]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
<emphasis id="B972ED2209790308FDA1575B3522FDB9" bold="true" box="[528,770,568,594]" pageId="6" pageNumber="7">Hendrich &amp; Balke</emphasis>
</taxonomicName>
<emphasis id="B972ED2209790308FCA1575B357BFDB9" bold="true" box="[784,859,568,594]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A241505909790308FCA1575B357BFDB9" box="[784,859,568,594]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
is assigned to the Bidessini based on the bisegmented parameres of the aedeagus (
<figureCitation id="133D2DB509790308FDDF570334EBFD91" box="[622,715,608,634]" captionStart="FIGURES 13–17" captionStartId="11.[151,269,1942,1966]" captionTargetBox="[211,1388,187,1914]" captionTargetId="figure-67@11.[197,1403,164,1924]" captionTargetPageId="11" captionText="FIGURES 13–17. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 13) Neobidessodes thoracicus sp.n. (paratype); 14) N. thoracicus sp.n. (“light form”); 15) N. flavosignatus; 16) N. mjobergi and 17) N. samkrisi sp.n. (scale bar = 0.5 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.10093590" httpUri="https://zenodo.org/record/10093590/files/figure.png" pageId="6" pageNumber="7">Figs 17</figureCitation>
,
<figureCitation id="133D2DB509790308FD68570334D7FD91" box="[729,759,608,634]" captionStart="FIGURES 18–20" captionStartId="12.[151,269,1750,1774]" captionTargetBox="[171,1417,199,1703]" captionTargetId="figure-116@12.[151,1436,187,1726]" captionTargetPageId="12" captionText="FIGURES 18–20. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 18) Neobidessodes bilita; 19) N. denticulatus and 20) N. grossus (scale bar = 0.5 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.10093592" httpUri="https://zenodo.org/record/10093592/files/figure.png" pageId="6" pageNumber="7">18</figureCitation>
) (
<bibRefCitation id="EF974CC109790308FCA05703359BFD91" author="Bistrom, O." box="[785,955,608,634]" pageId="6" pageNumber="7" pagination="1 - 41" refId="ref28436" refString="Bistrom, O. (1988) Generic review of the Bidessini (Coleoptera, Dytiscidae). Acta Zoologica Fennica, 184, 1 - 41." type="journal article" year="1988">Biström 1988</bibRefCitation>
). It is a genus of small to medium sized Bidessini (
<quantity id="4CFE9CD509790308FEA757E5378CFD4B" box="[278,428,646,672]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.2" metricValueMin="2.3" pageId="6" pageNumber="7" unit="mm" value="3.25" valueMax="4.2" valueMin="2.3">2.3–4.2 mm</quantity>
), in
<collectingCountry id="F31171A009790308FE5057E5346EFD4B" box="[481,590,646,672]" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
now represented by eight species and by one species in southern New
<collectingCountry id="F31171A009790308FF2657CE36D0FD2C" box="[151,240,685,711]" name="Guinea-Bissau" pageId="6" pageNumber="7">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09790308FF7457B0344EF95E" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BB9313009790308FF7457B0328DFCFF" blockId="6.[151,1436,530,2026]" pageId="6" pageNumber="7">
<emphasis id="B972ED2209790308FF7457B03772FD06" bold="true" box="[197,338,723,749]" pageId="6" pageNumber="7">Etymology.</emphasis>
The name
<taxonomicName id="4C064AB309790308FE6657B734A5FD06" box="[471,645,724,749]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FE6657B734A5FD06" box="[471,645,724,749]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
is derived from the Greek word
<emphasis id="B972ED2209790308FBBB57B53215FD06" box="[1034,1077,726,749]" italics="true" pageId="6" pageNumber="7">Neo</emphasis>
[new] and
<taxonomicName id="4C064AB309790308FB0A57B7331DFD06" authorityName="Regimbart" authorityYear="1900" box="[1211,1341,724,749]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FB0A57B7331DFD06" box="[1211,1341,724,749]" italics="true" pageId="6" pageNumber="7">Bidessodes</emphasis>
</taxonomicName>
, a valid genus name for a group of Neotropical predaceous water beetles. Its gender is masculine.
</paragraph>
<paragraph id="8BB9313009790308FF745642356EFB7C" blockId="6.[151,1436,530,2026]" pageId="6" pageNumber="7">
The new genus is well separated from all other Bidessini by the combination of the following combination of characters: 1) body elongate oval; 2) basal pronotal striae in some species absent or fine, in others sharply incised, oblique, not connected by a transverse groove; 3) elytra lacking basal striae and sutural striae; 4) epipleura lacking transverse carina; 5) head lacking cervical line and its foremargin not bordered; 6) prosternal process broad, distinctly excavated and marginated; 7) inner margin of both metacoxal wings strongly ridged; 8) hind margin of abdominal ventrites 3–5 without row of minor irregular dentate processes; 9) well-developed finger-like apical lobe on distal segment of paramere. All but two (
<taxonomicName id="4C064AB309790308FB36516A3367FBC9" authorityName="WA (Yilgarn" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2003" box="[1159,1351,1033,1058]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="limestonensis">
<emphasis id="B972ED2209790308FB36516A3367FBC9" box="[1159,1351,1033,1058]" italics="true" pageId="6" pageNumber="7">N. limestonensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064AB309790308FA30516A3738FBA2" authorityName="WA (Yilgarn" baseAuthorityName="Watts &amp; Humphreys" baseAuthorityYear="2003" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gutteridgei">
<emphasis id="B972ED2209790308FA30516A3738FBA2" italics="true" pageId="6" pageNumber="7">N. gutteridgei</emphasis>
</taxonomicName>
) of the species studied have a more or less contrasting black/yellow surface. The ground pattern of these species includes various yellow or reddish spots (
<figureCitation id="133D2DB509790308FC8651353595FB9B" box="[823,949,1110,1136]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–8" captionStart-2="FIGURES 9–12" captionStartId-0="7.[151,270,1900,1924]" captionStartId-1="8.[151,275,1973,1997]" captionStartId-2="9.[151,274,1833,1857]" captionTargetBox-0="[173,1401,199,1857]" captionTargetBox-1="[155,1427,235,1928]" captionTargetBox-2="[158,1433,206,1780]" captionTargetId-0="figure-77@7.[167,1411,180,1869]" captionTargetId-1="figure-48@8.[151,1436,194,1950]" captionTargetId-2="figure-68@9.[148,1433,194,1809]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 1–4. Habitus of 1) N. denticulatus (WA, Pilbara, Millstream Chichester National Park, Fortescue River side branch); 2) N. denticulatus (NT, Kakadu N.P., Jim Jim District, Gungurul Lookout); 3) N. flavosignatus (WA, East Kimberley, Gibb Range, Gibb River Road, Russ Creek Crossing); 4) N. flavosignatus (NT, Manton Dam Recreation Area, 46 km S Darwin) (scale bar = 1 mm), (Photos: A. Riedel)." captionText-1="FIGURES 5–8. Habitus of 5) Neobidessodes samkrisi sp.n. (holotype, male); 6) N. grossus; 7) N. mjobergi; 8) Neobidessodes bilita (female) (scale bar = 1 mm) (Photos: A. Riedel)." captionText-2="FIGURES 9–12. 9) N. thoracicus sp.n. (paratype); 10) N. thoracicus sp.n. (paratype, “black specimen”); 11) N. thoracicus sp.n. (“light form”, WA, Kimberley Region, 50 km S Wyndham, Black Flag Creek); 12) N. thoracicus sp.n. (“light form”, NT, Kakadu N.P., Jim Jim District, Gungurul Lookout) (scale bar = 1 mm) (Photos: A. Riedel)." figureDoi-0="http://doi.org/10.5281/zenodo.10093584" figureDoi-1="http://doi.org/10.5281/zenodo.10093586" figureDoi-2="http://doi.org/10.5281/zenodo.10093588" httpUri-0="https://zenodo.org/record/10093584/files/figure.png" httpUri-1="https://zenodo.org/record/10093586/files/figure.png" httpUri-2="https://zenodo.org/record/10093588/files/figure.png" pageId="6" pageNumber="7">Figs 1–12</figureCitation>
). The median lobes are simple and very elongate, in ventral view strongly tapered or rounded at tip.
</paragraph>
<paragraph id="8BB9313009790308FF7451C73540FAE0" blockId="6.[151,1436,530,2026]" pageId="6" pageNumber="7">
The Neotropical genus
<taxonomicName id="4C064AB309790308FE6651C73479FB56" authorityName="Regimbart" authorityYear="1900" box="[471,601,1188,1213]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FE6651C73479FB56" box="[471,601,1188,1213]" italics="true" pageId="6" pageNumber="7">Bidessodes</emphasis>
</taxonomicName>
can be separated from Australian
<taxonomicName id="4C064AB309790308FC5D51C732BAFB56" box="[1004,1178,1188,1213]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FC5D51C732BAFB56" box="[1004,1178,1188,1213]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
<emphasis id="B972ED2209790308FB1051C032CBFB56" bold="true" box="[1185,1259,1187,1213]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A241505909790308FB1051C032CBFB56" box="[1185,1259,1187,1213]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
by having 1) a non excavated and marginated prosternal process, and 2) a visible distinct row of minor irregular dentate processes on the hind margin of the abdominal ventrites 3–5.
</paragraph>
<paragraph id="8BB9313009790308FF745074344EF95E" blockId="6.[151,1436,530,2026]" pageId="6" pageNumber="7">
<emphasis id="B972ED2209790308FF74507437C4FADA" bold="true" box="[197,484,1303,1329]" pageId="6" pageNumber="7">Molecular systematics.</emphasis>
Our phylogenetic analyses show (
<figureCitation id="133D2DB509790308FCCF507B35F5FAD9" box="[894,981,1304,1330]" captionStart="FIGURE 27" captionStartId="32.[151,260,1876,1900]" captionTargetBox="[151,1436,194,1852]" captionTargetId="figure-78@32.[151,1436,194,1852]" captionTargetPageId="32" captionText="FIGURE 27. Phylogram of the tree obtained using GARLI and cox1 and 16S data for Australasian Bidessini, Neotropical Bidessodes and outgroups. Node support, when above 50%: bold (GARLI bootstrap), italics (MrBayes posterior propablities&gt;0.5, x100), normal font (TNT jackknife values). Note: “Clypeodytes migrator” will be transferred to Leiodytes in a forthcoming revision (Hendrich et al. in prep.)." figureDoi="http://doi.org/10.5281/zenodo.10093602" httpUri="https://zenodo.org/record/10093602/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
) that
<taxonomicName id="4C064AB309790308FBAB507B32E9FADA" box="[1050,1225,1304,1329]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FBAB507B32E9FADA" box="[1050,1225,1304,1329]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
<emphasis id="B972ED2209790308FB605074333DFADA" bold="true" box="[1233,1309,1303,1329]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A241505909790308FB605074333DFADA" box="[1233,1309,1303,1329]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
is not part of the
<taxonomicName id="4C064AB309790308FF54505C3745FAB3" authorityName="Guignot" authorityYear="1953" box="[229,357,1343,1368]" class="Insecta" family="Dytiscidae" genus="Allodessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FF54505C3745FAB3" box="[229,357,1343,1368]" italics="true" pageId="6" pageNumber="7">Allodessus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C064AB309790308FECE505C3439FAB3" authorityName="Guignot" authorityYear="1939" box="[383,537,1343,1368]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FECE505C3439FAB3" box="[383,537,1343,1368]" italics="true" pageId="6" pageNumber="7">Limbodessus</emphasis>
</taxonomicName>
, the
<taxonomicName id="4C064AB309790308FDE7505C348DFAB3" authorityName="Guignot" authorityYear="1939" box="[598,685,1343,1368]" class="Insecta" family="Dytiscidae" genus="Uvarus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FDE7505C348DFAB3" box="[598,685,1343,1368]" italics="true" pageId="6" pageNumber="7">Uvarus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C064AB309790308FD76505C3579FAB3" authorityName="Watts" authorityYear="1978" box="[711,857,1343,1368]" class="Insecta" family="Dytiscidae" genus="Gibbidessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FD76505C3579FAB3" box="[711,857,1343,1368]" italics="true" pageId="6" pageNumber="7">Gibbidessus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C064AB309790308FCC2505C323EFAB3" authorityName="Hendrich &amp; Balke" authorityYear="2009" box="[883,1054,1343,1368]" class="Insecta" family="Dytiscidae" genus="Kakadudessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FCC2505C323EFAB3" box="[883,1054,1343,1368]" italics="true" pageId="6" pageNumber="7">Kakadudessus</emphasis>
</taxonomicName>
or the
<taxonomicName id="4C064AB309790308FBCC505C32D0FAB3" authorityName="Guignot" authorityYear="1936" box="[1149,1264,1343,1368]" class="Insecta" family="Dytiscidae" genus="Leiodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FBCC505C32D0FAB3" box="[1149,1264,1343,1368]" italics="true" pageId="6" pageNumber="7">Leiodytes</emphasis>
</taxonomicName>
/
<taxonomicName id="4C064AB309790308FABB505C33BCFAB3" baseAuthorityName="Hendrich &amp; Wang" baseAuthorityYear="2006" box="[1290,1436,1343,1368]" class="Insecta" family="Dytiscidae" genus="Clypeodytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FABB505C33BCFAB3" box="[1290,1436,1343,1368]" italics="true" pageId="6" pageNumber="7">Clypeodytes</emphasis>
</taxonomicName>
clades. It is rather part of a separate lineage sister to the first two clades. The DNA sequence data from the mitochondrial cytochrome
<emphasis id="B972ED2209790308FE6D50EF37C9FA4E" box="[476,489,1420,1445]" italics="true" pageId="6" pageNumber="7">c</emphasis>
oxidase 1 and 16S rRNA genes also show that
<taxonomicName id="4C064AB309790308FB9950EF32F9FA4E" box="[1064,1241,1420,1445]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FB9950EF32F9FA4E" box="[1064,1241,1420,1445]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
<emphasis id="B972ED2209790308FB5350E8330FFA4E" bold="true" box="[1250,1327,1419,1445]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A241505909790308FB5350E8330FFA4E" box="[1250,1327,1419,1445]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
does not belong to any of the known Oriental and Australasian Bidessini genera (the morphologically strongly deviating
<taxonomicName id="4C064AB309790308FEBD50BA35E9FA18" authority="Balke, Hendrich, Mazzoldi &amp; Bistrom, 2002" authorityName="Balke, Hendrich, Mazzoldi &amp; Bistrom" authorityYear="2002" box="[268,969,1497,1523]" class="Insecta" family="Dytiscidae" genus="Borneodessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FEBD50BA378FFA19" box="[268,431,1497,1522]" italics="true" pageId="6" pageNumber="7">Borneodessus</emphasis>
Balke, Hendrich, Mazzoldi &amp; Biström, 2002
</taxonomicName>
was not available for sequencing). Our data show that Australian
<taxonomicName id="4C064AB309790308FE65536334A6F9F2" box="[468,646,1536,1561]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FE65536334A6F9F2" box="[468,646,1536,1561]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
<emphasis id="B972ED2209790308FD3F509C34FBF9F2" bold="true" box="[654,731,1535,1561]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A241505909790308FD3F509C34FBF9F2" box="[654,731,1535,1561]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
are not closely related to South American
<taxonomicName id="4C064AB309790308FB5253633348F9F2" authorityName="Regimbart" authorityYear="1900" box="[1251,1384,1536,1561]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FB5253633348F9F2" box="[1251,1384,1536,1561]" italics="true" pageId="6" pageNumber="7">Bidessodes</emphasis>
</taxonomicName>
and recognition of
<taxonomicName id="4C064AB309790308FEF4534437D4F9AB" box="[325,500,1575,1600]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972ED2209790308FEF4534437D4F9AB" box="[325,500,1575,1600]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
<emphasis id="B972ED2209790308FE4C53453467F9AB" bold="true" box="[509,583,1574,1600]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A241505909790308FE4C53453467F9AB" box="[509,583,1574,1600]" pageId="6" pageNumber="7" rank="genus">gen.n.</taxonomicNameLabel>
</emphasis>
does not create paraphyly among other Australasian genera. This result is well supported in all analyses (
<figureCitation id="133D2DB509790308FD9B532E34A5F98C" box="[554,645,1613,1639]" captionStart="FIGURE 27" captionStartId="32.[151,260,1876,1900]" captionTargetBox="[151,1436,194,1852]" captionTargetId="figure-78@32.[151,1436,194,1852]" captionTargetPageId="32" captionText="FIGURE 27. Phylogram of the tree obtained using GARLI and cox1 and 16S data for Australasian Bidessini, Neotropical Bidessodes and outgroups. Node support, when above 50%: bold (GARLI bootstrap), italics (MrBayes posterior propablities&gt;0.5, x100), normal font (TNT jackknife values). Note: “Clypeodytes migrator” will be transferred to Leiodytes in a forthcoming revision (Hendrich et al. in prep.)." figureDoi="http://doi.org/10.5281/zenodo.10093602" httpUri="https://zenodo.org/record/10093602/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
). Essentially the same tree topology was recovered with different analytical approaches (maximum likelihood, parsimony and Bayesian probabilities as implemented in MrBayes,
<figureCitation id="133D2DB509790308FEA053F83747F95E" box="[273,359,1691,1717]" captionStart="FIGURE 27" captionStartId="32.[151,260,1876,1900]" captionTargetBox="[151,1436,194,1852]" captionTargetId="figure-78@32.[151,1436,194,1852]" captionTargetPageId="32" captionText="FIGURE 27. Phylogram of the tree obtained using GARLI and cox1 and 16S data for Australasian Bidessini, Neotropical Bidessodes and outgroups. Node support, when above 50%: bold (GARLI bootstrap), italics (MrBayes posterior propablities&gt;0.5, x100), normal font (TNT jackknife values). Note: “Clypeodytes migrator” will be transferred to Leiodytes in a forthcoming revision (Hendrich et al. in prep.)." figureDoi="http://doi.org/10.5281/zenodo.10093602" httpUri="https://zenodo.org/record/10093602/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
, node support values).
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09790308FF7453A33576F802" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BB9313009790308FF7453A33576F802" blockId="6.[151,1436,530,2026]" pageId="6" pageNumber="7">
<emphasis id="B972ED2209790308FF7453A33761F931" bold="true" box="[197,321,1728,1754]" pageId="6" pageNumber="7">Remarks.</emphasis>
The placement of the two stygobitic species in this genus was primarily based on evidence from
<emphasis id="B972ED2209790308FF6A538B3733F8EA" box="[219,275,1768,1793]" italics="true" pageId="6" pageNumber="7">cox1</emphasis>
and 16S sequence data (
<bibRefCitation id="EF974CC109790308FDF4538B35A9F8E9" author="Watts, C. H. S. &amp; Humphreys, W. F." box="[581,905,1768,1794]" pageId="6" pageNumber="7" pagination="135 - 187" refId="ref29904" refString="Watts, C. H. S. &amp; Humphreys, W. F. (2003) Twenty-five new Dytiscidae (Coleoptera) of the genera Tjirtudessus Watts &amp; Humphreys, Nirripirti Watts &amp; Humphreys and Bidessodes Regimbart from underground waters in Australia. Records of the South Australian Museum, 36 (2), 135 - 187." type="journal article" year="2003">Watts &amp; Humphreys 2003</bibRefCitation>
) which suggested a relationship of the two species with
<taxonomicName id="4C064AB309790308FE9C526C3415F8C3" authority="(Watts, 1978)" authorityName="S QLD, NSW, VIC" baseAuthorityName="Watts" baseAuthorityYear="1978" box="[301,565,1806,1832]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bilita">
<emphasis id="B972ED2209790308FE9C526C37ADF8C3" box="[301,397,1807,1832]" italics="true" pageId="6" pageNumber="7">N. bilita</emphasis>
(
<bibRefCitation id="EF974CC109790308FE2D526D340CF8C3" author="Watts, C. H. S." box="[412,556,1806,1832]" pageId="6" pageNumber="7" pagination="1 - 166" refId="ref29702" refString="Watts, C. H. S. (1978) A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology, Supplement Series, 57, 1 - 166." type="journal article" year="1978">Watts, 1978</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C064AB309790308FDF1526C35EDF8C3" authority="(Zimmermann, 1922)" authorityName="N WA, NT, N QLD" baseAuthorityName="Zimmermann" baseAuthorityYear="1922" box="[576,973,1806,1832]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B972ED2209790308FDF1526C34EAF8C3" box="[576,714,1807,1832]" italics="true" pageId="6" pageNumber="7">N. mjobergi</emphasis>
(
<bibRefCitation id="EF974CC109790308FD6B526D35E5F8C3" author="Zimmermann, A." box="[730,965,1806,1832]" pageId="6" pageNumber="7" pagination="1 - 4" refId="ref30407" refString="Zimmermann, A. (1922) Results of Dr. E. Mjoberg's Swedish Expeditions to Australia 1910 - 1913, 28. Dytiscidae. Arkiv for Zoologie, 14 (16), 1 - 4." type="journal article" year="1922">Zimmermann, 1922</bibRefCitation>
)
</taxonomicName>
and/or
<taxonomicName id="4C064AB309790308FB97526C36C0F8A4" authority="(Zimmermann, 1922)" authorityName="N WA, NT, N QLD" baseAuthorityName="Zimmermann" baseAuthorityYear="1922" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B972ED2209790308FB97526C32C4F8C3" box="[1062,1252,1807,1832]" italics="true" pageId="6" pageNumber="7">N. flavosignatus</emphasis>
(
<bibRefCitation id="EF974CC109790308FB42526D36F7F8A4" author="Zimmermann, A." pageId="6" pageNumber="7" pagination="1 - 4" refId="ref30407" refString="Zimmermann, A. (1922) Results of Dr. E. Mjoberg's Swedish Expeditions to Australia 1910 - 1913, 28. Dytiscidae. Arkiv for Zoologie, 14 (16), 1 - 4." type="journal article" year="1922">Zimmermann, 1922</bibRefCitation>
)
</taxonomicName>
. A subsequent, comprehensive analysis of Australian stygobitic Bidessini and their epigean relatives (
<bibRefCitation id="EF974CC109790308FF2E523F3741F89D" author="Leys, R. &amp; Watts, C. H. S. &amp; Cooper, S. J. B. &amp; Humphrey, W. F." box="[159,353,1884,1910]" pageId="6" pageNumber="7" pagination="2819 - 2834" refId="ref29230" refString="Leys, R., Watts, C. H. S., Cooper, S. J. B. &amp; Humphrey, W. F. (2003) Evolution of subterranean diving beetles (Coleoptera: Dytiscidae) in the arid zone of Australia. Evolution, 57 (12), 2819 - 2834." type="journal article" year="2003">
Leys
<emphasis id="B972ED2209790308FF50523F3735F89E" box="[225,277,1884,1909]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2003
</bibRefCitation>
) suggests a sister-group relation of the stygobitic and the epigean
<taxonomicName id="4C064AB309790308FBDF523F333CF89E" box="[1134,1308,1884,1909]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FBDF523F333CF89E" box="[1134,1308,1884,1909]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
(under the name
<taxonomicName id="4C064AB309790308FF5152E03745F877" authorityName="Regimbart" authorityYear="1900" box="[224,357,1923,1948]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FF5152E03745F877" box="[224,357,1923,1948]" italics="true" pageId="6" pageNumber="7">Bidessodes</emphasis>
</taxonomicName>
at the time). There are no morphological characters that would negate their placement in
<taxonomicName id="4C064AB309790308FF2652CA3767F829" box="[151,327,1961,1986]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972ED2209790308FF2652CA3767F829" box="[151,327,1961,1986]" italics="true" pageId="6" pageNumber="7">Neobidessodes</emphasis>
</taxonomicName>
as currently defined. Our species level
<emphasis id="B972ED2209790308FC9152CA3577F829" box="[800,855,1961,1986]" italics="true" pageId="6" pageNumber="7">cox1</emphasis>
data analysis (
<figureCitation id="133D2DB509790308FBBB52CA3241F828" box="[1034,1121,1961,1987]" captionStart="FIGURE 28" captionStartId="33.[151,255,1645,1669]" captionTargetBox="[151,1436,197,1621]" captionTargetId="figure-148@33.[151,1436,194,1621]" captionTargetPageId="33" captionText="FIGURE 28. Phylogram of the tree obtained using GARLI and with cox1 data for Neobidessodes alone. Node support, when above 50%: bold (GARLI bootstrap), normal font (TNT parsimony jackknife values). Abbreviations behind N. thoracicus sp.n.: “b.s” = black specimens; “l.f.” = “light form”." figureDoi="http://doi.org/10.5281/zenodo.10093604" httpUri="https://zenodo.org/record/10093604/files/figure.png" pageId="6" pageNumber="7">Fig. 28</figureCitation>
) confirms the sister-group relationship between stygobitic and epigean
<emphasis id="B972ED2209790308FD1152B33576F802" box="[672,854,2000,2025]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C064AB309790308FD1152B33572F802" box="[672,850,2000,2025]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Neobidessodes</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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