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<document id="4C5A0F8904E842FDB645B8164FA9236E" ID-DOI="10.11646/zootaxa.2288.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10093582" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629654284239" checkinUser="felipe" docAuthor="HENDRICH, LARS, HAWLITSCHEK, OLIVER &amp; BALKE, MICHAEL" docDate="2009" docId="03AF802609680316FF2653C13482FA3E" docLanguage="en" docName="zt02288p041.pdf" docOrigin="Zootaxa 2288 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.2288.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Neobidessodes samkrisi Hendrich &amp; Balke, sp.n." docType="treatment" docVersion="3" lastPageNumber="25" masterDocId="FF96F85E097F030EFFB155633620FFEB" masterDocTitle="The epigean Australasian species of Neobidessodes gen. n. diving beetles- a revision integrating morphology, cybertaxonomy, DNA taxonomy and phylogeny (Coleoptera: Dytiscidae, Bidessini)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="24" updateTime="1699563497581" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0B6B9B856A7765C618CFD820AC1A92ED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C30A198E77E0D4937BB03FA1F9E2C77">
<mods:title id="80B3490605FAB6AAB2D45DCD374E13E1">The epigean Australasian species of Neobidessodes gen. n. diving beetles- a revision integrating morphology, cybertaxonomy, DNA taxonomy and phylogeny (Coleoptera: Dytiscidae, Bidessini)</mods:title>
</mods:titleInfo>
<mods:name id="C1456CFF05A84531E7B008A8EF825E6F" type="personal">
<mods:role id="08A15DB739AFE5C7BBF6014DD1CE7270">
<mods:roleTerm id="FF79855700E53D16B6428C1425E118EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="205A0283CE78B6D5ECBE4D45F65AD445">HENDRICH, LARS</mods:namePart>
</mods:name>
<mods:name id="9B2BEBA9E1136FCB52D296C516BAA64B" type="personal">
<mods:role id="C5DF775D36D7FF8CF99552F70BDCAF05">
<mods:roleTerm id="05F6B76D04A65A0BE8D63A11B4B4F53E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14FCED8277AC1C7050E9AC6BD53D38BA">HAWLITSCHEK, OLIVER</mods:namePart>
</mods:name>
<mods:name id="001AC15ED023DBE1F793A16D31CEE564" type="personal">
<mods:role id="4963CA20F4B07818ECB9982ABD3F1E03">
<mods:roleTerm id="1F85ED2375E50706C5B65BD4789D5C3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B6E80E4699BDA9DFD87CBB8C075552EA">BALKE, MICHAEL</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D836ECE6F207F3431D50A41A708972B">text</mods:typeOfResource>
<mods:relatedItem id="88DDE12719CCF1174C7A589D2E84137A" type="host">
<mods:titleInfo id="0C0CA92B7F24D166E0CFE03CF3886FDC">
<mods:title id="599D79BD3CC711A47F780498FB4968FF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="13C081E459F467180B16A37DA5F087AA">
<mods:date id="F32DE4AE7516D70E8FDB2BEB197CC900">2009</mods:date>
<mods:detail id="1B7C487C470CFA6F69D817A20CB78F60" type="pubDate">
<mods:number id="CA0C639735F0C30E251DF0E0373EA2EE">2009-11-12</mods:number>
</mods:detail>
<mods:detail id="1A9A8D1B597EB05F051102F797C12014" type="volume">
<mods:number id="0325629FC370AC81AAB031CAF5EAF17D">2288</mods:number>
</mods:detail>
<mods:detail id="96D3842D16453DBBE379395681B5DA4D" type="issue">
<mods:number id="15DD4163757353E14546C3D1360624C0">1</mods:number>
</mods:detail>
<mods:extent id="4EC5D3DAA1E4BC44DD605A15B0D3C049" unit="page">
<mods:start id="5B516C32A638321FE3792F4A873BF9ED">1</mods:start>
<mods:end id="D21CE84CCE4614A013682E11B84A0080">41</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1BAE82A4C5BB07000AD47AAAC55215DD">
<mods:url id="B8642BCDA1E22251CA21D2C52317A560">http://dx.doi.org/10.11646/zootaxa.2288.1.1</mods:url>
</mods:location>
<mods:classification id="EAB2887F4AE5EBE8424BD1092B337735">journal article</mods:classification>
<mods:identifier id="6774BFB93125F698AB901F787169DF31" type="DOI">10.11646/zootaxa.2288.1.1</mods:identifier>
<mods:identifier id="64545DF07B6EF0183B09113A64E61555" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9A8C9341E31EDF4EAF94F17F7183DD37" type="Zenodo-Dep">10093582</mods:identifier>
</mods:mods>
<treatment id="03AF802609680316FF2653C13482FA3E" LSID="urn:lsid:plazi:treatment:03AF802609680316FF2653C13482FA3E" httpUri="http://treatment.plazi.org/id/03AF802609680316FF2653C13482FA3E" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="C31C62BB09680319FF2653C134C3F957" box="[151,739,1698,1724]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BB9313009680319FF2653C134C3F957" blockId="23.[151,739,1698,1759]" box="[151,739,1698,1724]" pageId="23" pageNumber="24">
<heading id="D0F1865C09680319FF2653C134C3F957" bold="true" box="[151,739,1698,1724]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B972ED2209680319FF2653C134C3F957" bold="true" box="[151,739,1698,1724]" pageId="23" pageNumber="24">
<taxonomicName id="4C064AB309680319FF2653C13481F957" ID-CoL="6RV3W" authority="Hendrich &amp; Balke" authorityName="Hendrich &amp; Balke" box="[151,673,1698,1724]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="samkrisi" status="sp. nov.">
<emphasis id="B972ED2209680319FF2653C13795F957" bold="true" box="[151,437,1698,1724]" italics="true" pageId="23" pageNumber="24">Neobidessodes samkrisi</emphasis>
Hendrich &amp; Balke
</taxonomicName>
<taxonomicNameLabel id="A241505909680319FD1853C134C3F957" box="[681,739,1698,1724]" pageId="23" pageNumber="24" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09680319FF2653A63782F934" box="[151,418,1733,1759]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BB9313009680319FF2653A63782F934" blockId="23.[151,739,1698,1759]" box="[151,418,1733,1759]" pageId="23" pageNumber="24">
(
<figureCitation id="133D2DB509680319FF2E53A636CAF934" box="[159,234,1733,1759]" captionStart="FIGURES 5–8" captionStartId="8.[151,275,1973,1997]" captionTargetBox="[155,1427,235,1928]" captionTargetId="figure-48@8.[151,1436,194,1950]" captionTargetPageId="8" captionText="FIGURES 5–8. Habitus of 5) Neobidessodes samkrisi sp.n. (holotype, male); 6) N. grossus; 7) N. mjobergi; 8) Neobidessodes bilita (female) (scale bar = 1 mm) (Photos: A. Riedel)." figureDoi="http://doi.org/10.5281/zenodo.10093586" httpUri="https://zenodo.org/record/10093586/files/figure.png" pageId="23" pageNumber="24">Figs 5</figureCitation>
,
<figureCitation id="133D2DB509680319FF4953A63737F934" box="[248,279,1733,1759]" captionStart="FIGURES 13–17" captionStartId="11.[151,269,1942,1966]" captionTargetBox="[211,1388,187,1914]" captionTargetId="figure-67@11.[197,1403,164,1924]" captionTargetPageId="11" captionText="FIGURES 13–17. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 13) Neobidessodes thoracicus sp.n. (paratype); 14) N. thoracicus sp.n. (“light form”); 15) N. flavosignatus; 16) N. mjobergi and 17) N. samkrisi sp.n. (scale bar = 0.5 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.10093590" httpUri="https://zenodo.org/record/10093590/files/figure.png" pageId="23" pageNumber="24">17</figureCitation>
,
<figureCitation id="133D2DB509680319FE9253A63763F934" box="[291,323,1733,1759]" captionStart="FIGURES 23–24" captionStartId="29.[151,271,1972,1996]" captionTargetBox="[151,1436,455,1949]" captionTargetId="figure-124@29.[151,1436,455,1949]" captionTargetPageId="29" captionText="FIGURES 23–24. Distribution of Neobidessodes: 23) N. flavosignatus; 24) N. samkrisi sp.n. (square) and N. grossus (dots)." figureDoi="http://doi.org/10.5281/zenodo.10093598" httpUri="https://zenodo.org/record/10093598/files/figure.png" pageId="23" pageNumber="24">24</figureCitation>
,
<figureCitation id="133D2DB509680319FEE153A63750F934" box="[336,368,1733,1759]" captionStart="FIGURE 27" captionStartId="32.[151,260,1876,1900]" captionTargetBox="[151,1436,194,1852]" captionTargetId="figure-78@32.[151,1436,194,1852]" captionTargetPageId="32" captionText="FIGURE 27. Phylogram of the tree obtained using GARLI and cox1 and 16S data for Australasian Bidessini, Neotropical Bidessodes and outgroups. Node support, when above 50%: bold (GARLI bootstrap), italics (MrBayes posterior propablities&gt;0.5, x100), normal font (TNT jackknife values). Note: “Clypeodytes migrator” will be transferred to Leiodytes in a forthcoming revision (Hendrich et al. in prep.)." figureDoi="http://doi.org/10.5281/zenodo.10093602" httpUri="https://zenodo.org/record/10093602/files/figure.png" pageId="23" pageNumber="24">27</figureCitation>
,
<figureCitation id="133D2DB509680319FECD53A637BAF934" box="[380,410,1733,1759]" captionStart="FIGURE 28" captionStartId="33.[151,255,1645,1669]" captionTargetBox="[151,1436,197,1621]" captionTargetId="figure-148@33.[151,1436,194,1621]" captionTargetPageId="33" captionText="FIGURE 28. Phylogram of the tree obtained using GARLI and with cox1 data for Neobidessodes alone. Node support, when above 50%: bold (GARLI bootstrap), normal font (TNT parsimony jackknife values). Abbreviations behind N. thoracicus sp.n.: “b.s” = black specimens; “l.f.” = “light form”." figureDoi="http://doi.org/10.5281/zenodo.10093604" httpUri="https://zenodo.org/record/10093604/files/figure.png" pageId="23" pageNumber="24">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09680319FF265271350CF805" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BB9313009680319FF26527135AAF8C7" blockId="23.[151,1436,1810,2030]" box="[151,906,1810,1836]" pageId="23" pageNumber="24">
<materialsCitation id="3B6E3B6D09680319FF26527135A6F8C7" box="[151,902,1810,1836]" country="Indonesia" location="Merauke" municipality="Wasur Reserve" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<emphasis id="B972ED2209680319FF265271371CF8C7" bold="true" box="[151,316,1810,1836]" pageId="23" pageNumber="24">
<typeStatus id="54BD8F9209680319FF26527136F4F8C7" box="[151,212,1810,1836]" pageId="23" pageNumber="24">Type</typeStatus>
locality.
</emphasis>
<location id="8ED967EB09680319FEF55271378FF8C7" LSID="urn:lsid:plazi:treatment:03AF802609680316FF2653C13482FA3E:8ED967EB09680319FEF55271378FF8C7" box="[324,431,1810,1836]" country="Indonesia" municipality="Wasur Reserve" name="Merauke" pageId="23" pageNumber="24" stateProvince="Papua Barat">Merauke</location>
,
<collectingMunicipality id="6BDDAB4A09680319FE0B5271344DF8C7" box="[442,621,1810,1836]" pageId="23" pageNumber="24">Wasur Reserve</collectingMunicipality>
,
<collectingRegion id="49C2FFD209680319FDC852713525F8C7" box="[633,773,1810,1836]" country="Indonesia" name="Papua Barat" pageId="23" pageNumber="24">West Papua</collectingRegion>
,
<collectingCountry id="F31171A009680319FCA1527135A6F8C7" box="[784,902,1810,1836]" name="Indonesia" pageId="23" pageNumber="24">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9313009680319FF74525B350CF805" blockId="23.[151,1436,1810,2030]" pageId="23" pageNumber="24">
<emphasis id="B972ED2209680319FF74525B3758F8B9" bold="true" box="[197,376,1848,1874]" pageId="23" pageNumber="24">Type material</emphasis>
.
<materialsCitation id="3B6E3B6D09680319FE36525B356DF84B" collectingDate="2006-11" collectionCode="ZSM" collectorName="M. Balke &amp; K. Tindige" country="Indonesia" location="Male" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Papua Barat" typeStatus="holotype">
<emphasis id="B972ED2209680319FE36525B3423F8B9" bold="true" box="[391,515,1848,1874]" pageId="23" pageNumber="24">
<typeStatus id="54BD8F9209680319FE36525B37DDF8B9" box="[391,509,1848,1874]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8ED967EB09680319FDBA525A346CF8B8" LSID="urn:lsid:plazi:treatment:03AF802609680316FF2653C13482FA3E:8ED967EB09680319FDBA525A346CF8B8" box="[523,588,1849,1875]" country="Indonesia" name="Male" pageId="23" pageNumber="24" stateProvince="Papua Barat">Male</location>
: “DNA
<collectorName id="26F354E609680319FD1E525A3536F8B8" box="[687,790,1849,1875]" pageId="23" pageNumber="24">M.Balke</collectorName>
2843” [green printed label], “
<collectingCountry id="F31171A009680319FB30525A32DBF8B8" box="[1153,1275,1849,1875]" name="Indonesia" pageId="23" pageNumber="24">Indonesia</collectingCountry>
,
<collectingRegion id="49C2FFD209680319FAB9525A33B7F8B8" box="[1288,1431,1849,1875]" country="Indonesia" name="Papua Barat" pageId="23" pageNumber="24">West Papua</collectingRegion>
, Merauke, Wasur Reserve,
<date id="FFB817F009680319FE535203348EF891" box="[482,686,1888,1914]" pageId="23" pageNumber="24" value="2006-11">
<collectingDate id="EFFCEE1809680319FE535203348EF891" box="[482,686,1888,1914]" pageId="23" pageNumber="24" value="2006-11">November 2006</collectingDate>
</date>
,
<collectorName id="26F354E609680319FD0D52033562F891" box="[700,834,1888,1914]" pageId="23" pageNumber="24">K. Tindige</collectorName>
leg.”, “
<typeStatus id="54BD8F9209680319FC155203323AF891" box="[932,1050,1888,1914]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C064AB309680319FB955203336FF892" box="[1060,1359,1888,1913]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="samkrisi" status="sp. nov.">
<emphasis id="B972ED2209680319FB955203336FF892" box="[1060,1359,1888,1913]" italics="true" pageId="23" pageNumber="24">Neobidessodes samkrisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241505909680319FAE9520333BCF891" box="[1368,1436,1888,1914]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
Hendrich &amp; Balke des. 2009” [red printed label] (
<collectionCode id="ED17A9F509680319FD4E52E53564F84B" box="[767,836,1926,1952]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="23" pageNumber="24" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<emphasis id="B972ED2209680319FCEB52E535C3F84B" bold="true" box="[858,995,1926,1952]" pageId="23" pageNumber="24">
<typeStatus id="54BD8F9209680319FCEB52E535FFF84B" box="[858,991,1926,1952]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D00FAB909680319FC5C52E5321FF84B" box="[1005,1087,1926,1952]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
with “DNA M.Balke 2844” [green printed label] and
<materialsCitation id="3B6E3B6D09680319FE7E52CE3509F805" collectingDate="2006-11" collectionCode="ZSM, CLH" collectorName="M. Balke &amp; K. Tindige" country="Indonesia" location="Male" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="9D00FAB909680319FE7E52CE3465F82C" box="[463,581,1965,1991]" pageId="23" pageNumber="24" type="female">2 females</specimenCount>
with same data as holotype but without DNA label (
<collectionCode id="ED17A9F509680319FB7E52CE3331F82C" box="[1231,1297,1965,1991]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="23" pageNumber="24" type="Museum">ZSM</collectionCode>
,
<collectionCode id="ED17A9F509680319FAAE52CE3340F82C" box="[1311,1376,1965,1991]" pageId="23" pageNumber="24">CLH</collectionCode>
). All
<typeStatus id="54BD8F9209680319FF2652B73727F805" box="[151,263,2004,2030]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
are provided with a red printed
<typeStatus id="54BD8F9209680319FD3152B734C5F805" box="[640,741,2004,2030]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09670316FF7455F437E5FD1D" pageId="24" pageNumber="25" type="description">
<paragraph id="8BB9313009670316FF7455F4355BFF33" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF7455F4377DFF5A" bold="true" box="[197,349,151,177]" pageId="24" pageNumber="25">Description.</emphasis>
Measurements.
<emphasis id="B972ED2209670316FDAC55F434AFFF5A" bold="true" box="[541,655,151,177]" pageId="24" pageNumber="25">
<typeStatus id="54BD8F9209670316FDAC55F434AFFF5A" box="[541,655,151,177]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
: TL = 2.00 mm, TL-H =
<quantity id="4CFE9CD509670316FC0955FB3203FF59" box="[952,1059,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="25" unit="mm" value="1.8">1.80 mm</quantity>
; MW =
<quantity id="4CFE9CD509670316FB3255FB32C0FF59" box="[1155,1248,152,178]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="mm" value="0.9">0.9 mm</quantity>
.
<emphasis id="B972ED2209670316FB5D55F43349FF5A" bold="true" box="[1260,1385,151,177]" pageId="24" pageNumber="25">
<typeStatus id="54BD8F9209670316FB5D55F43349FF5A" box="[1260,1385,151,177]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
</emphasis>
: TL = 2.00–
<quantity id="4CFE9CD509670316FF4055DD377BFF33" box="[241,347,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="24" pageNumber="25" unit="mm" value="2.05">2.05 mm</quantity>
, TL-H =
<quantity id="4CFE9CD509670316FE7855DD3456FF33" box="[457,630,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="1.825" metricValueMax="1.85" metricValueMin="1.8" pageId="24" pageNumber="25" unit="mm" value="1.825" valueMax="1.85" valueMin="1.8">1.80–1.85 mm</quantity>
; MW = 0.9–1.00 mm.
</paragraph>
<paragraph id="8BB9313009670316FF7455863541FE98" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF7455863702FF14" bold="true" box="[197,290,229,255]" pageId="24" pageNumber="25">Colour.</emphasis>
Antennae, palpi and head pale reddish-brown, posterior angles of head and near eyes darkened anteriorly. Pronotum pale reddish-brown with thin blackish band basally. Elytron dark brown with some well marked elongate yellow markings (
<figureCitation id="133D2DB509670316FDE554513483FEA7" box="[596,675,306,332]" captionStart="FIGURES 5–8" captionStartId="8.[151,275,1973,1997]" captionTargetBox="[155,1427,235,1928]" captionTargetId="figure-48@8.[151,1436,194,1950]" captionTargetPageId="8" captionText="FIGURES 5–8. Habitus of 5) Neobidessodes samkrisi sp.n. (holotype, male); 6) N. grossus; 7) N. mjobergi; 8) Neobidessodes bilita (female) (scale bar = 1 mm) (Photos: A. Riedel)." figureDoi="http://doi.org/10.5281/zenodo.10093586" httpUri="https://zenodo.org/record/10093586/files/figure.png" pageId="24" pageNumber="25">Fig. 5</figureCitation>
). Ventral side, including legs and epipleura, reddish-yellow, prosternal process and metacoxal plates somewhat darkened.
</paragraph>
<paragraph id="8BB9313009670316FF74541C372FFDB0" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF74541C37DCFE72" bold="true" box="[197,508,383,409]" pageId="24" pageNumber="25">Sculpture and structure.</emphasis>
Elongate oval. Head with scattered fine punctures and a few larger ones near base. Pronotum and elytron weakly microreticulate and with moderately dense but rather small, and weakly impressed punctures, particularly weak on disc of pronotum. Pronotal striae short but well marked, 1/4 of length of pronotum, weakly incurved. Elytra lacking basal and sutural striae. Underside laterally with scattered weak to moderate punctures, denser in midline. Metacoxal lines raised, very weakly diverging anteriorly.
</paragraph>
<paragraph id="8BB9313009670316FF74570435C9FD24" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF745704372BFD6A" bold="true" box="[197,267,615,641]" pageId="24" pageNumber="25">Male.</emphasis>
Median lobe (
<figureCitation id="133D2DB509670316FE09570B341CFD69" box="[440,572,616,642]" captionStart="FIGURES 13–17" captionStartId="11.[151,269,1942,1966]" captionTargetBox="[211,1388,187,1914]" captionTargetId="figure-67@11.[197,1403,164,1924]" captionTargetPageId="11" captionText="FIGURES 13–17. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 13) Neobidessodes thoracicus sp.n. (paratype); 14) N. thoracicus sp.n. (“light form”); 15) N. flavosignatus; 16) N. mjobergi and 17) N. samkrisi sp.n. (scale bar = 0.5 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.10093590" httpUri="https://zenodo.org/record/10093590/files/figure.png" pageId="24" pageNumber="25">Fig. 17a, b</figureCitation>
). Parameres thin, slender and very elongate, only slightly bifid anteriorly (17c). Much smaller than the parameres of
<taxonomicName id="4C064AB309670316FD7057EC354CFD43" box="[705,876,655,680]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="thoracicus" status="sp. nov.">
<emphasis id="B972ED2209670316FD7057EC354CFD43" box="[705,876,655,680]" italics="true" pageId="24" pageNumber="25">N. thoracicus</emphasis>
</taxonomicName>
<emphasis id="B972ED2209670316FCC757ED3595FD43" bold="true" box="[886,949,654,680]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A241505909670316FCC757ED3595FD43" box="[886,949,654,680]" pageId="24" pageNumber="25" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C064AB309670316FC4757EC32ACFD43" authorityName="Hendrich &amp; Balke" baseAuthorityName="Zimmermann" baseAuthorityYear="1922" box="[1014,1164,655,680]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B972ED2209670316FC4757EC32ACFD43" box="[1014,1164,655,680]" italics="true" pageId="24" pageNumber="25">N. mjobergi</emphasis>
</taxonomicName>
. Protarsus somewhat expanded. Mesotarsus less so. Inner edge of mesotibia slightly incurved.
</paragraph>
<paragraph id="8BB9313009670316FF7457B837E5FD1D" blockId="24.[151,1436,151,1493]" box="[197,453,731,758]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF7457B83707FD1E" bold="true" box="[197,295,731,757]" pageId="24" pageNumber="25">Female.</emphasis>
Tarsi simple.
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09670316FF7456613365FBC0" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BB9313009670316FF74566132B1FCA8" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF745661377AFCF7" bold="true" box="[197,346,770,796]" pageId="24" pageNumber="25">Etymology.</emphasis>
The species is dedicated in memory to our late colleague and friend Samkris Tindige (†
<date id="FFB817F009670316FF07564A3710FCA8" box="[182,304,809,835]" pageId="24" pageNumber="25" value="2007-08-20">20.8.2007</date>
), Indonesian naturalist and conservationist from Manokwari,
<collectingRegion id="49C2FFD209670316FBB1564A32ADFCA8" box="[1024,1165,809,835]" country="Indonesia" name="Papua Barat" pageId="24" pageNumber="25">West Papua</collectingRegion>
.
</paragraph>
<paragraph id="8BB9313009670316FF74562C3365FBC0" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF74562C3766FC82" bold="true" box="[197,326,847,873]" pageId="24" pageNumber="25">Affinities.</emphasis>
<taxonomicName id="4C064AB309670316FEFE56333456FC82" box="[335,630,848,873]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="samkrisi" status="sp. nov.">
<emphasis id="B972ED2209670316FEFE56333456FC82" box="[335,630,848,873]" italics="true" pageId="24" pageNumber="25">Neobidessodes samkrisi</emphasis>
</taxonomicName>
<emphasis id="B972ED2209670316FDCE562C349CFC82" bold="true" box="[639,700,847,873]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A241505909670316FDCE562C349CFC82" box="[639,700,847,873]" pageId="24" pageNumber="25" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is much smaller than almost all other species of the genus, except
<taxonomicName id="4C064AB309670316FF5B5614376AFC7B" authorityName="S QLD, NSW, VIC" baseAuthorityName="Watts" baseAuthorityYear="1978" box="[234,330,887,912]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bilita">
<emphasis id="B972ED2209670316FF5B5614376AFC7B" box="[234,330,887,912]" italics="true" pageId="24" pageNumber="25">N. bilita</emphasis>
</taxonomicName>
from
<collectingRegion id="49C2FFD209670316FE2056153409FC7B" box="[401,553,886,912]" country="Australia" name="Western Australia" pageId="24" pageNumber="25">SE Australia</collectingRegion>
, but can be very easily distinguished from the latter by the form of the male genitalia and the colour of the dorsal surface (
<figureCitation id="133D2DB509670316FD0E56FE350CFC5C" box="[703,812,925,951]" captionStart="FIGURES 5–8" captionStartId="8.[151,275,1973,1997]" captionTargetBox="[155,1427,235,1928]" captionTargetId="figure-48@8.[151,1436,194,1950]" captionTargetPageId="8" captionText="FIGURES 5–8. Habitus of 5) Neobidessodes samkrisi sp.n. (holotype, male); 6) N. grossus; 7) N. mjobergi; 8) Neobidessodes bilita (female) (scale bar = 1 mm) (Photos: A. Riedel)." figureDoi="http://doi.org/10.5281/zenodo.10093586" httpUri="https://zenodo.org/record/10093586/files/figure.png" pageId="24" pageNumber="25">Figs 5, 8</figureCitation>
). From its colouration it is close to
<taxonomicName id="4C064AB309670316FB6B56FE33BBFC5D" authorityName="Hendrich &amp; Balke" baseAuthorityName="Zimmermann" baseAuthorityYear="1922" box="[1242,1435,925,950]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B972ED2209670316FB6B56FE33BBFC5D" box="[1242,1435,925,950]" italics="true" pageId="24" pageNumber="25">N. flavosignatus</emphasis>
</taxonomicName>
(
<figureCitation id="133D2DB509670316FF2E56A73728FC35" box="[159,264,964,990]" captionStart="FIGURES 1–4" captionStartId="7.[151,270,1900,1924]" captionTargetBox="[173,1401,199,1857]" captionTargetId="figure-77@7.[167,1411,180,1869]" captionTargetPageId="7" captionText="FIGURES 1–4. Habitus of 1) N. denticulatus (WA, Pilbara, Millstream Chichester National Park, Fortescue River side branch); 2) N. denticulatus (NT, Kakadu N.P., Jim Jim District, Gungurul Lookout); 3) N. flavosignatus (WA, East Kimberley, Gibb Range, Gibb River Road, Russ Creek Crossing); 4) N. flavosignatus (NT, Manton Dam Recreation Area, 46 km S Darwin) (scale bar = 1 mm), (Photos: A. Riedel)." figureDoi="http://doi.org/10.5281/zenodo.10093584" httpUri="https://zenodo.org/record/10093584/files/figure.png" pageId="24" pageNumber="25">Figs 3, 4</figureCitation>
) and the “light form” of
<taxonomicName id="4C064AB309670316FD9D56A734EAFC36" box="[556,714,964,990]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="thoracicus" status="sp. nov.">
<emphasis id="B972ED2209670316FD9D56A7341FFC36" box="[556,575,964,989]" italics="true" pageId="24" pageNumber="25">N</emphasis>
.
<emphasis id="B972ED2209670316FDFF56A734EAFC36" box="[590,714,964,989]" italics="true" pageId="24" pageNumber="25">thoracicus</emphasis>
</taxonomicName>
<emphasis id="B972ED2209670316FD6056A0352CFC36" bold="true" box="[721,780,963,989]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A241505909670316FD6056A0352CFC36" box="[721,780,963,989]" pageId="24" pageNumber="25" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133D2DB509670316FCAD56A73580FC35" box="[796,928,964,990]" captionStart="FIGURES 9–12" captionStartId="9.[151,274,1833,1857]" captionTargetBox="[158,1433,206,1780]" captionTargetId="figure-68@9.[148,1433,194,1809]" captionTargetPageId="9" captionText="FIGURES 9–12. 9) N. thoracicus sp.n. (paratype); 10) N. thoracicus sp.n. (paratype, “black specimen”); 11) N. thoracicus sp.n. (“light form”, WA, Kimberley Region, 50 km S Wyndham, Black Flag Creek); 12) N. thoracicus sp.n. (“light form”, NT, Kakadu N.P., Jim Jim District, Gungurul Lookout) (scale bar = 1 mm) (Photos: A. Riedel)." figureDoi="http://doi.org/10.5281/zenodo.10093588" httpUri="https://zenodo.org/record/10093588/files/figure.png" pageId="24" pageNumber="25">Figs 11, 12</figureCitation>
) but can be separated from both species by its smaller size and the transverse basal marking on the elytra which is very narrow compared to the other species. The form of the median lobe (
<figureCitation id="133D2DB509670316FDEB5172349EFBC0" box="[602,702,1041,1067]" captionStart="FIGURES 13–17" captionStartId="11.[151,269,1942,1966]" captionTargetBox="[211,1388,187,1914]" captionTargetId="figure-67@11.[197,1403,164,1924]" captionTargetPageId="11" captionText="FIGURES 13–17. Median lobe of aedeagus in ventral (a) and lateral view (b), and right paramere in lateral view (c): 13) Neobidessodes thoracicus sp.n. (paratype); 14) N. thoracicus sp.n. (“light form”); 15) N. flavosignatus; 16) N. mjobergi and 17) N. samkrisi sp.n. (scale bar = 0.5 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.10093590" httpUri="https://zenodo.org/record/10093590/files/figure.png" pageId="24" pageNumber="25">Fig. 17b</figureCitation>
) distinguished
<taxonomicName id="4C064AB309670316FCC3517235D7FBC1" box="[882,1015,1041,1066]" class="Insecta" family="Dytiscidae" genus="Neobidessodes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="samkrisi" status="sp. nov.">
<emphasis id="B972ED2209670316FCC3517235D7FBC1" box="[882,1015,1041,1066]" italics="true" pageId="24" pageNumber="25">N. samkrisi</emphasis>
</taxonomicName>
<emphasis id="B972ED2209670316FC4F51733219FBC1" bold="true" box="[1022,1081,1040,1066]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A241505909670316FC4F51733219FBC1" box="[1022,1081,1040,1066]" pageId="24" pageNumber="25" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
from all other species.
</paragraph>
</subSubSection>
<subSubSection id="C31C62BB09670316FF7451543482FA3E" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BB9313009670316FF7451543532FB74" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF7451543746FBBA" bold="true" box="[197,358,1079,1105]" pageId="24" pageNumber="25">Distribution.</emphasis>
Only known from the few
<typeStatus id="54BD8F9209670316FD1D515B34FEFBB9" box="[684,734,1080,1106]" pageId="24" pageNumber="25">type</typeStatus>
specimens collected near Merauke, southern coast of West Papua (New
<collectingCountry id="F31171A009670316FE88513D37B8FB93" box="[313,408,1118,1144]" name="Guinea-Bissau" pageId="24" pageNumber="25">Guinea</collectingCountry>
),
<collectingCountry id="F31171A009670316FE1E513D3408FB93" box="[431,552,1118,1144]" name="Indonesia" pageId="24" pageNumber="25">Indonesia</collectingCountry>
(
<figureCitation id="133D2DB509670316FD8B513D34BAFB93" box="[570,666,1118,1144]" captionStart="FIGURES 23–24" captionStartId="29.[151,271,1972,1996]" captionTargetBox="[151,1436,455,1949]" captionTargetId="figure-124@29.[151,1436,455,1949]" captionTargetPageId="29" captionText="FIGURES 23–24. Distribution of Neobidessodes: 23) N. flavosignatus; 24) N. samkrisi sp.n. (square) and N. grossus (dots)." figureDoi="http://doi.org/10.5281/zenodo.10093598" httpUri="https://zenodo.org/record/10093598/files/figure.png" pageId="24" pageNumber="25">Fig. 24</figureCitation>
). Probably more widespread in wet savannah areas along the southern coast of
<collectingRegion id="49C2FFD209670316FED751E637CFFB74" box="[358,495,1157,1183]" country="Indonesia" name="Papua Barat" pageId="24" pageNumber="25">West Papua</collectingRegion>
and
<collectingCountry id="F31171A009670316FD9951E6352EFB74" box="[552,782,1157,1183]" name="Papua New Guinea" pageId="24" pageNumber="25">Papua New Guinea</collectingCountry>
.
</paragraph>
<paragraph id="8BB9313009670316FF7451C83482FA3E" blockId="24.[151,1436,151,1493]" pageId="24" pageNumber="25">
<emphasis id="B972ED2209670316FF7451C8370CFB2E" bold="true" box="[197,300,1195,1221]" pageId="24" pageNumber="25">Habitat.</emphasis>
Nothing is known about the life habitat. The specimens were obtained from the Wasur Reserve, a low-lying wetland in the monsoon climate zone of southern New
<collectingCountry id="F31171A009670316FC1451B035DAFB06" box="[933,1018,1235,1261]" name="Guinea-Bissau" pageId="24" pageNumber="25">Guinea</collectingCountry>
with intertidal mudflats and coastal mangroves with extensive seasonally inundated grasslands, savannas, reed swamps, and swamp and monsoon forests. The specimens were collected with the dytiscids
<taxonomicName id="4C064AB309670316FCE450433351FAD1" authority="(W. J. Macleay, 1871)" baseAuthorityName="W. J. Macleay" baseAuthorityYear="1871" box="[853,1393,1312,1338]" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B972ED2209670316FCE45043327FFAD2" box="[853,1119,1312,1337]" italics="true" pageId="24" pageNumber="25">Hydroglyphus basalis</emphasis>
(W.J. Macleay, 1871)
</taxonomicName>
,
<taxonomicName id="4C064AB309670316FACE504337E2FA8B" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="godeffroyi">
<emphasis id="B972ED2209670316FACE50433731FA8B" italics="true" pageId="24" pageNumber="25">H. godeffroyi</emphasis>
(
<bibRefCitation id="EF974CC109670316FE935025379AFA8B" author="Sharp, D." box="[290,442,1350,1376]" pageId="24" pageNumber="25" pagination="179 - 1003" refId="ref29626" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. S cientific Transactions of the Royal Dublin Society, (2) 2, 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C064AB309670316FE61502435A4FA8B" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[464,900,1350,1376]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cingulatus">
<emphasis id="B972ED2209670316FE61502434C8FA8B" box="[464,744,1351,1376]" italics="true" pageId="24" pageNumber="25">Laccophilus cingulatus</emphasis>
<bibRefCitation id="EF974CC109670316FD40502535A4FA8B" author="Sharp, D." box="[753,900,1350,1376]" pageId="24" pageNumber="25" pagination="179 - 1003" refId="ref29626" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. S cientific Transactions of the Royal Dublin Society, (2) 2, 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C064AB309670316FC23502432B6FA8B" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[914,1174,1350,1376]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B972ED2209670316FC23502435D9FA8B" box="[914,1017,1351,1376]" italics="true" pageId="24" pageNumber="25">L. clarki</emphasis>
<bibRefCitation id="EF974CC109670316FBB3502532B6FA8B" author="Sharp, D." box="[1026,1174,1350,1376]" pageId="24" pageNumber="25" pagination="179 - 1003" refId="ref29626" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. S cientific Transactions of the Royal Dublin Society, (2) 2, 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C064AB309670316FB15502436F2FA6C" authority="Fauvel, 1883" authorityName="Fauvel" authorityYear="1883" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="seminiger">
<emphasis id="B972ED2209670316FB155024331AFA8B" box="[1188,1338,1351,1376]" italics="true" pageId="24" pageNumber="25">L. seminiger</emphasis>
Fauvel, 1883
</taxonomicName>
and
<taxonomicName id="4C064AB309670316FEBC500E351BFA6C" authority="transversalis Regimbart, 1877" authorityName="Regimbart" authorityYear="1877" box="[269,827,1389,1415]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="transversalis" subSpecies="transversalis">
<emphasis id="B972ED2209670316FEBC500E3448FA6D" box="[269,616,1389,1414]" italics="true" pageId="24" pageNumber="25">L. transversalis transversalis</emphasis>
Régimbart, 1877
</taxonomicName>
, and the noterid
<taxonomicName id="4C064AB309670316FBB5500E33BBFA6C" authority="(Clark, 1863)" baseAuthorityName="Clark" baseAuthorityYear="1863" box="[1028,1435,1389,1415]" class="Insecta" family="Noteridae" genus="Notomicrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="B972ED2209670316FBB5500E32D3FA6D" box="[1028,1267,1389,1414]" italics="true" pageId="24" pageNumber="25">Notomicrus tenellus</emphasis>
(Clark, 1863)
</taxonomicName>
which indicate a more lotic environment such as a slow flowing and well vegetated stream or an oxbow lake (billabong) in the floodzone of a larger river.
</paragraph>
</subSubSection>
</treatment>
</document>
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