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<document id="B8413B734BCE58267EA572AB4DA9AA1F" ID-CLB-Dataset="8958" ID-DOI="10.11646/zootaxa.4820.3.1" ID-GBIF-Dataset="fd413580-2960-463f-a924-8c27eaeafc6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4397861" ID-ZooBank="44BD1E07-C9F3-4488-936E-819C2FCA18C8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609210733456" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="03AF3A058D7FFFB8FAA9FBB6A18FFC0D" docLanguage="en" docName="zootaxa.4820.3.1.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Oculogaster) niger Kluge 2020, sp. n." docType="treatment" docVersion="8" lastPageNumber="420" masterDocId="FF96427D8D72FFABFA3EFFACA367FFC4" masterDocTitle="Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)" masterLastPageNumber="437" masterPageNumber="401" pageNumber="414" updateTime="1698848343517" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CDDF91F4CBCEB3862390D2D63B237569">Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<mods:namePart id="8A63CC7D48A6BCC17D5587E79D6E9C02">Kluge, Nikita J.</mods:namePart>
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<mods:date id="39C0595707A3A256886CFC6E116662F0">2020</mods:date>
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<treatment id="03AF3A058D7FFFB8FAA9FBB6A18FFC0D" ID-DOI="http://doi.org/10.5281/zenodo.4397971" ID-GBIF-Taxon="176573415" ID-Zenodo-Dep="4397971" LSID="urn:lsid:plazi:treatment:03AF3A058D7FFFB8FAA9FBB6A18FFC0D" httpUri="http://treatment.plazi.org/id/03AF3A058D7FFFB8FAA9FBB6A18FFC0D" lastPageId="19" lastPageNumber="420" pageId="13" pageNumber="414">
<subSubSection id="C31CD8988D7FFFA6FAA9FBB6A136FBF1" box="[151,593,1050,1077]" pageId="13" pageNumber="414" type="nomenclature">
<paragraph id="8BB98B138D7FFFA6FAA9FBB6A136FBF1" blockId="13.[151,593,1050,1113]" box="[151,593,1050,1077]" pageId="13" pageNumber="414">
<heading id="D0F13C7F8D7FFFA6FAA9FBB6A136FBF1" bold="true" box="[151,593,1050,1077]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
<emphasis id="B97257018D7FFFA6FAA9FBB6A136FBF1" bold="true" box="[151,593,1050,1077]" pageId="13" pageNumber="414">
<taxonomicName id="4C06F0908D7FFFA6FAA9FBB6A16FFBF1" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[151,520,1050,1077]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D7FFFA6FAA9FBB6A274FBF1" bold="true" box="[151,275,1050,1077]" italics="true" pageId="13" pageNumber="414">Procloeon</emphasis>
(
<emphasis id="B97257018D7FFFA6FB1AFBB6A2D0FBF1" bold="true" box="[292,439,1050,1077]" italics="true" pageId="13" pageNumber="414">Oculogaster</emphasis>
)
<emphasis id="B97257018D7FFFA6FBF6FBB7A16FFBF1" bold="true" box="[456,520,1051,1077]" italics="true" pageId="13" pageNumber="414">niger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241EA7A8D7FFFA6F831FBB7A136FBF1" box="[527,593,1051,1077]" pageId="13" pageNumber="414" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D7FFFA6FAA9FB93A240FB9D" box="[151,295,1087,1113]" pageId="13" pageNumber="414" type="description">
<paragraph id="8BB98B138D7FFFA6FAA9FB93A240FB9D" blockId="13.[151,593,1050,1113]" box="[151,295,1087,1113]" pageId="13" pageNumber="414">
(
<figureCitation id="133D97968D7FFFA6FAA1FB93A278FB9D" box="[159,287,1087,1113]" captionStart-0="FIGURES 4348" captionStart-1="FIGURES 4956" captionStart-2="FIGURES 5766" captionStart-3="FIGURES 6770" captionStart-4="FIGURES 7172" captionStart-5="FIGURES 7377" captionStartId-0="14.[151,264,1916,1940]" captionStartId-1="15.[151,264,1887,1911]" captionStartId-2="16.[151,264,1939,1963]" captionStartId-3="17.[151,264,1918,1942]" captionStartId-4="18.[151,264,1960,1984]" captionStartId-5="20.[151,264,1907,1931]" captionTargetBox-0="[151,1436,181,1892]" captionTargetBox-1="[151,1436,181,1863]" captionTargetBox-2="[157,1431,818,1915]" captionTargetBox-3="[151,1436,181,1895]" captionTargetBox-4="[170,1428,712,1926]" captionTargetBox-5="[151,1436,273,1884]" captionTargetId-0="figure-27@14.[151,1436,181,1892]" captionTargetId-1="figure-16@15.[151,1436,181,1863]" captionTargetId-2="figure-312@16.[151,1436,813,1916]" captionTargetId-3="figure-16@17.[151,1436,181,1895]" captionTargetId-4="figure-305@18.[151,1436,705,1936]" captionTargetId-5="figure-95@20.[151,1436,273,1884]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="20" captionText-0="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." captionText-1="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." captionText-2="FIGURES 5766. Larvae of Procloeon (Oculogaster) niger sp. n. 5762, tergalii IIII and VVII; 6365, tergalii I, IV and VII of other specimens (6465, holotype); 66, claw." captionText-3="FIGURES 6770. Male Procloeon (Oculogaster) niger sp. n. 6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype)." captionText-4="FIGURES 7172. Male Procloeon (Oculogaster) niger sp. n. 71, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 72, their subimaginal exuviae." captionText-5="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi-0="http://doi.org/10.5281/zenodo.4397885" figureDoi-1="http://doi.org/10.5281/zenodo.4397887" figureDoi-2="http://doi.org/10.5281/zenodo.4397889" figureDoi-3="http://doi.org/10.5281/zenodo.4397893" figureDoi-4="http://doi.org/10.5281/zenodo.4397895" figureDoi-5="http://doi.org/10.5281/zenodo.4397897" httpUri-0="https://zenodo.org/record/4397885/files/figure.png" httpUri-1="https://zenodo.org/record/4397887/files/figure.png" httpUri-2="https://zenodo.org/record/4397889/files/figure.png" httpUri-3="https://zenodo.org/record/4397893/files/figure.png" httpUri-4="https://zenodo.org/record/4397895/files/figure.png" httpUri-5="https://zenodo.org/record/4397897/files/figure.png" pageId="13" pageNumber="414">Figs 4377</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D7FFFA6FAA9FB2BA2A5FB01" pageId="13" pageNumber="414" type="etymology">
<paragraph id="8BB98B138D7FFFA6FAA9FB2BA2A5FB01" blockId="13.[151,1437,1159,2013]" pageId="13" pageNumber="414">
<emphasis id="B97257018D7FFFA6FAA9FB2BA279FB65" bold="true" box="[151,286,1159,1185]" pageId="13" pageNumber="414">Etymology.</emphasis>
Allusion to the uniformly dark coloration of larva and uniform dark color pattern on abdominal terga in male and female imago.
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D7FFFB8FAF9FB62A18FFC0D" lastPageId="19" lastPageNumber="420" pageId="13" pageNumber="414" type="materials_examined">
<paragraph id="8BB98B138D7FFFA6FAF9FB62A013FA91" blockId="13.[151,1437,1159,2013]" pageId="13" pageNumber="414">
<emphasis id="B97257018D7FFFA6FAF9FB62A2D6FB2C" bold="true" box="[199,433,1230,1256]" pageId="13" pageNumber="414">Material examined.</emphasis>
<materialsCitation id="3B6E814E8D7FFFA6FB86FB63A002FAF5" ID-GBIF-Occurrence="3014991301" collectingDate="2019-02-07" collectionCode="AM, NW" collectorName="N. Kluge." country="South Africa" county="Nature's Valley" latitude="-33.95" location="Garden Route district" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" pageId="13" pageNumber="414" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B97257018D7FFFA6FB86FB63A142FB2D" bold="true" box="[440,549,1231,1257]" pageId="13" pageNumber="414">
<typeStatus id="54BD35B18D7FFFA6FB86FB63A142FB2D" box="[440,549,1231,1257]" pageId="13" pageNumber="414" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="ED1713D68D7FFFA6F80AFB63A100FB2D" box="[564,615,1231,1257]" country="Australia" name="Australian Museum" pageId="13" pageNumber="414" type="Museum">AM</collectionCode>
): L-S-I
<specimenCount id="9D00409A8D7FFFA6F884FB63A1AAFB2D" box="[698,717,1231,1257]" pageId="13" pageNumber="414" type="male"></specimenCount>
{specimen [XVIII](3)2019},
<collectingCountry id="F311CB838D7FFFA6FE20FB63A796FB2D" box="[1054,1265,1231,1257]" name="South Africa" pageId="13" pageNumber="414">SOUTH AFRICA</collectingCountry>
,
<location id="8ED9DDC88D7FFFA6FF3EFB63A382FAC9" LSID="urn:lsid:plazi:treatment:03AF3A058D7FFFB8FAA9FBB6A18FFC0D:8ED9DDC88D7FFFA6FF3EFB63A382FAC9" country="South Africa" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Garden Route district" pageId="13" pageNumber="414">Garden Route district</location>
(=
<location id="8ED9DDC88D7FFFA6FB30FB5FA2C6FACA" LSID="urn:lsid:plazi:treatment:03AF3A058D7FFFB8FAA9FBB6A18FFC0D:8ED9DDC88D7FFFA6FB30FB5FA2C6FACA" box="[270,417,1267,1294]" country="South Africa" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Eden district" pageId="13" pageNumber="414">Eden district</location>
), lodge «Wild Spirit», tributary of
<collectingMunicipality id="6BDD11698D7FFFA6F914FB5FA095FAC9" box="[810,1010,1267,1293]" pageId="13" pageNumber="414">Bobbejaans River</collectingMunicipality>
(
<quantity id="4CFE26F68D7FFFA6FE3DFB5EA75AFAC9" box="[1027,1085,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="414" unit="km" value="5.0">5 km</quantity>
<collectionCode id="ED1713D68D7FFFA6FE7BFB5FA714FAC9" box="[1093,1139,1267,1293]" pageId="13" pageNumber="414">NW</collectionCode>
<collectingCounty id="62D8F39F8D7FFFA6FE44FB5FA649FACA" box="[1146,1326,1267,1294]" pageId="13" pageNumber="414">Natures Valley</collectingCounty>
).
<geoCoordinate id="EE32EDD48D7FFFA6FF7EFB5EA6FBFAC9" box="[1344,1436,1266,1293]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="13" pageNumber="414" precision="925" value="-33.95">33°57S</geoCoordinate>
<geoCoordinate id="EE32EDD48D7FFFA6FAA9FABBA27BFAF5" box="[151,284,1303,1331]" degrees="23" direction="east" minutes="31" orientation="longitude" pageId="13" pageNumber="414" precision="15" seconds="30" value="23.525">23°3130E</geoCoordinate>
, 3rd pool from waterfall,
<date id="FFB8ADD38D7FFFA6F87BFABBA1C9FAF5" box="[581,686,1303,1329]" pageId="13" pageNumber="414" value="2019-02-07">
<collectingDate id="EFFC543B8D7FFFA6F87BFABBA1C9FAF5" box="[581,686,1303,1329]" pageId="13" pageNumber="414" value="2019-02-07">7.II.2019</collectingDate>
</date>
, coll.
<collectorName id="26F3EEC58D7FFFA6F8CBFABBA002FAF5" box="[757,869,1303,1329]" pageId="13" pageNumber="414">N. Kluge.</collectorName>
</materialsCitation>
<materialsCitation id="3B6E814E8D7FFFA6F953FABBA017FA91" ID-GBIF-Occurrence="3014991305" collectingDate="2019-02-03" collectingDateMax="2019-02-09" collectingDateMin="2019-02-03" collectionCode="ZIN" collectorName="N. Kluge." country="South Africa" county="Nature's Valley" latitude="-33.95" location="Garden Route district" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" pageId="13" pageNumber="414" specimenCount="24" specimenCount-female="2" specimenCount-larva="19" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B97257018D7FFFA6F953FABBA082FAF5" bold="true" box="[877,997,1303,1329]" pageId="13" pageNumber="414">
<typeStatus id="54BD35B18D7FFFA6F953FABBA082FAF5" box="[877,997,1303,1329]" pageId="13" pageNumber="414" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<collectionCode id="ED1713D68D7FFFA6F9C8FABBA74CFAF5" box="[1014,1067,1303,1329]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="13" pageNumber="414" type="Museum">ZIN</collectionCode>
): same locality and collector,
<date id="FFB8ADD38D7FFFA6FFBEFABBA265FA91" pageId="13" pageNumber="414" value="2019-02-03" valueMax="2019-02-09" valueMin="2019-02-03">
<collectingDate id="EFFC543B8D7FFFA6FFBEFABBA265FA91" pageId="13" pageNumber="414" value="2019-02-03" valueMax="2019-02-09" valueMin="2019-02-03">39.II.2019</collectingDate>
</date>
: 2 L-S-I
<specimenCount id="9D00409A8D7FFFA6FB5DFA97A211FA91" box="[355,374,1339,1365]" pageId="13" pageNumber="414" type="male"></specimenCount>
, 3 L-S
<specimenCount id="9D00409A8D7FFFA6FBFAFA97A2BFFA91" box="[452,472,1339,1365]" pageId="13" pageNumber="414" type="male"></specimenCount>
, 1 L-S-I
<specimenCount id="9D00409A8D7FFFA6F800FA97A12BFA91" box="[574,588,1339,1365]" pageId="13" pageNumber="414" type="female"></specimenCount>
, 1 L-S
<specimenCount id="9D00409A8D7FFFA6F8A1FA97A1C9FA91" box="[671,686,1339,1365]" pageId="13" pageNumber="414" type="female"></specimenCount>
, L/S
<specimenCount id="9D00409A8D7FFFA6F8DBFA97A19EFA91" box="[741,761,1339,1365]" pageId="13" pageNumber="414" type="male"></specimenCount>
,
<specimenCount id="9D00409A8D7FFFA6F93BFA97A017FA91" box="[773,880,1339,1365]" pageId="13" pageNumber="414" type="larva">19 larvae</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB98B138D7FFFA6FAF9FAF2A267FA05" blockId="13.[151,1437,1159,2013]" pageId="13" pageNumber="414">
<emphasis id="B97257018D7FFFA6FAF9FAF2A2D2FABD" bold="true" box="[199,437,1374,1401]" pageId="13" pageNumber="414">
Descriptions.
<emphasis id="B97257018D7FFFA6FB54FAF3A2D2FABD" bold="true" box="[362,437,1375,1401]" italics="true" pageId="13" pageNumber="414">Larva.</emphasis>
</emphasis>
CUTICULAR COLORATION. Head, thorax, legs and abdomen nearly uniformly brown (
<figureCitation id="133D97968D7FFFA6FAA1FA2FA244FA59" box="[159,291,1410,1437]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="13" pageNumber="414">Figs 4546</figureCitation>
). Caudalii lighter, brown proximally and near apex, with posterior margin of each 4th segment brown (
<figureCitation id="133D97968D7FFFA6FA9EFA0BA393FA05" box="[160,244,1447,1473]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="13" pageNumber="414">Fig. 43</figureCitation>
).
</paragraph>
<paragraph id="8BB98B138D7FFFA6FAF9FA67A184F9CD" blockId="13.[151,1437,1159,2013]" pageId="13" pageNumber="414">
HYPODERMAL COLORATION. Mature larvae of both sexes with large lateral dark markings on abdominal terga, as in winged stages (as in
<figureCitation id="133D97968D7FFFA6F83FFA43A1E6F9CD" box="[513,641,1519,1545]" captionStart="FIGURES 6770" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-16@17.[151,1436,181,1895]" captionTargetPageId="17" captionText="FIGURES 6770. Male Procloeon (Oculogaster) niger sp. n. 6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397893" httpUri="https://zenodo.org/record/4397893/files/figure.png" pageId="13" pageNumber="414">Figs 6768</figureCitation>
,
<figureCitation id="133D97968D7FFFA6F8B2FA42A1B0F9CD" box="[652,727,1518,1545]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="13" pageNumber="414">7375</figureCitation>
).
</paragraph>
<paragraph id="8BB98B138D7FFFBBFAF9F9BFA061FE31" blockId="13.[151,1437,1159,2013]" lastBlockId="16.[151,1437,151,789]" lastPageId="16" lastPageNumber="417" pageId="13" pageNumber="414">
<collectionCode id="ED1713D68D7FFFA6FAF9F9BFA293F9E9" box="[199,500,1555,1581]" pageId="13" pageNumber="414">SHAPE AND SETATION</collectionCode>
. Labrum as in other
<taxonomicName id="4C06F0908D7FFFA6F8DAF9BFA009F9E9" authorityName="Kluge" authorityYear="2016" box="[740,878,1555,1581]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D7FFFA6F8DAF9BFA009F9E9" box="[740,878,1555,1581]" italics="true" pageId="13" pageNumber="414">Oculogaster</emphasis>
</taxonomicName>
(
<figureCitation id="133D97968D7FFFA6F9BEF9BFA761F9E9" box="[896,1030,1554,1581]" captionStart="FIGURES 4956" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="13" pageNumber="414">Figs 4950</figureCitation>
). Mandibles with incisor and kinetodontium fused more than to middle, with ventral denticle well-developed; incisor of right mandible with 2nd denticle shortest; right prostheca apically with 26 short parallel processes (
<figureCitation id="133D97968D7FFFA6F9F9F9F7A72CF9B1" box="[967,1099,1626,1653]" captionStart="FIGURES 4956" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="13" pageNumber="414">Figs 5152</figureCitation>
). Maxillary palp 2-segmented (
<figureCitation id="133D97968D7FFFA6FA9EF9D3A394F95D" box="[160,243,1662,1689]" captionStart="FIGURES 4956" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="13" pageNumber="414">Fig. 54</figureCitation>
). Labium with glossae and paraglossae of subequal size, palp with median angle moderately projected (
<figureCitation id="133D97968D7FFFA6FF51F9D3A385F979" captionStart="FIGURES 4956" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="13" pageNumber="414">Fig. 5554</figureCitation>
). Femora with very small spine-like setae on outer and inner sides, with 2 spine-like apical setae located in plane of leg; apical setae pointed or blunt, larger than other spine-like setae or equal to them (as in
<figureCitation id="133D97968D7FFFA6FED1F96BA625F925" box="[1263,1346,1735,1761]" captionStart="FIGURES 1718" captionStartId="7.[151,264,1525,1549]" captionTargetBox="[151,1436,561,1502]" captionTargetId="figure-198@7.[151,1436,561,1502]" captionTargetPageId="7" captionText="FIGURES 1718. Larva of Procloeon (Oculogaster) barnardi sp. n., middle leg. 17, apex of femur and base of tibia, anterior view (patella-tibial suture and bases of fine setae are visible as light markings on darker background); 18, more distal part of tibia, posterior view (with continuation of patella-tibial suture visible as light line)." figureDoi="http://doi.org/10.5281/zenodo.4397869" httpUri="https://zenodo.org/record/4397869/files/figure.png" pageId="13" pageNumber="414">Fig. 17</figureCitation>
). Claws moderately long, with two rows of denticles, minute in proximal part and larger more distally, exceeding mid-length of claw (
<figureCitation id="133D97968D7FFFA6FAC4F8A3A229F8EE" box="[250,334,1807,1834]" captionStart="FIGURES 5766" captionStartId="16.[151,264,1939,1963]" captionTargetBox="[157,1431,818,1915]" captionTargetId="figure-312@16.[151,1436,813,1916]" captionTargetPageId="16" captionText="FIGURES 5766. Larvae of Procloeon (Oculogaster) niger sp. n. 5762, tergalii IIII and VVII; 6365, tergalii I, IV and VII of other specimens (6465, holotype); 66, claw." figureDoi="http://doi.org/10.5281/zenodo.4397889" httpUri="https://zenodo.org/record/4397889/files/figure.png" pageId="13" pageNumber="414">Fig. 66</figureCitation>
). Hind protoptera completely absent. 14 lateral spines on posterior part of abdominal segment IX only (
<figureCitation id="133D97968D7FFFA6FAE7F89FA249F889" box="[217,302,1843,1869]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="13" pageNumber="414">Fig. 47</figureCitation>
). Posterolateral spines present on segments VIIX, being minute on most anterior of these segments. Posterior margin of abdominal tergum I with few sparse small spine-like pointed denticles; posterior margins of terga IIX with spine-like pointed denticles of unequal length, shorter on anterior terga and longer on posterior terga (
<figureCitation id="133D97968D7FFFA6FA9EF833A392F87D" box="[160,245,1951,1977]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="13" pageNumber="414">Fig. 47</figureCitation>
); on tergum IX row of denticles interrupted medially, behind pair of submedian setae (
<figureCitation id="133D97968D7FFFA6FEF1F833A643F87D" box="[1231,1316,1951,1977]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="13" pageNumber="414">Fig. 47</figureCitation>
). Abdominal terga with short scales in wide semilunar or W-shaped non-opercula-bearing sockets, with sparse simple fine setae; one pair of bifurcate setae (
<figureCitation id="133D97968D62FFBBF826FF3BA10CFF75" box="[536,619,151,177]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="16" pageNumber="417">Fig. 48</figureCitation>
) near midlength of each tergum
<collectionCode id="ED1713D68D62FFBBF9E1FF3BA75CFF75" box="[991,1083,151,177]" pageId="16" pageNumber="417">VIIIIX</collectionCode>
and one pair of bifurcate setae near posterior margin of each tergum VII and IX (as in
<figureCitation id="133D97968D62FFBBF8C1FF17A025FF11" box="[767,834,187,213]" captionStart="FIGURES 17" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 17. Procloeon (Oculogaster) barnardi sp. n. 16, male larva ready to moult to subimago: 12, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IVIX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="16" pageNumber="417">Fig. 3</figureCitation>
). Posterior margins of abdominal sterna IIV smooth, posterior margin of sternum
<collectionCode id="ED1713D68D62FFBBFBE3FF73A296FF3D" box="[477,497,223,249]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="417" type="Museum">V</collectionCode>
with few minute denticles, posterior margins of sterna VIIX and paraprocts with spine-like pointed denticles of unequal length. Abdominal sterna with sparse scales and sparse fine setae; in lateral areas of sterna both simple and bifid fine setae vary from small to very long, located sparsely and irregularly, not forming regular rows. Tergalius I with costal rib very short and lacking denticles; costal rib on tergalii IIVII longer, either without denticles or with few small denticles on apex; dorsal lamella present either on tergalii IV, or IIIV (
<figureCitation id="133D97968D62FFBBFAA1FE3FA245FE69" box="[159,290,402,429]" captionStart="FIGURES 5766" captionStartId="16.[151,264,1939,1963]" captionTargetBox="[157,1431,818,1915]" captionTargetId="figure-312@16.[151,1436,813,1916]" captionTargetPageId="16" captionText="FIGURES 5766. Larvae of Procloeon (Oculogaster) niger sp. n. 5762, tergalii IIII and VVII; 6365, tergalii I, IV and VII of other specimens (6465, holotype); 66, claw." figureDoi="http://doi.org/10.5281/zenodo.4397889" httpUri="https://zenodo.org/record/4397889/files/figure.png" pageId="16" pageNumber="417">Figs 5765</figureCitation>
). Cerci with especially long and slender distal portions lacking swimming setae and lateral spines (
<figureCitation id="133D97968D62FFBBFF51FE3FA3D0FE15" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="16" pageNumber="417">Fig. 43</figureCitation>
). In distal part of cercus (proximad of slender setae-less distal portion) spine on lateral side of each segment subequal or slightly longer than next segment (
<figureCitation id="133D97968D62FFBBF89BFE77A19EFE31" box="[677,761,475,501]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="16" pageNumber="417">Fig. 43</figureCitation>
).
</paragraph>
<caption id="DF79DB9B8D7CFFA5FAA9F8D0A2C5F818" ID-DOI="http://doi.org/10.5281/zenodo.4397885" ID-Zenodo-Dep="4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="14" pageNumber="415" startId="14.[151,264,1916,1940]" targetBox="[151,1436,181,1892]" targetPageId="14">
<paragraph id="8BB98B138D7CFFA5FAA9F8D0A2C5F818" blockId="14.[151,1437,1916,2012]" pageId="14" pageNumber="415">
<emphasis id="B97257018D7CFFA5FAA9F8D0A233F850" bold="true" box="[151,340,1916,1940]" pageId="14" pageNumber="415">FIGURES 4348.</emphasis>
Larval exuviae of
<taxonomicName id="4C06F0908D7CFFA5F82DF8D0A02CF850" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[531,843,1916,1941]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D7CFFA5F82DF8D0A11BF850" box="[531,636,1916,1940]" italics="true" pageId="14" pageNumber="415">Procloeon</emphasis>
(
<emphasis id="B97257018D7CFFA5F8B4F8D0A060F850" box="[650,775,1916,1940]" italics="true" pageId="14" pageNumber="415">Oculogaster</emphasis>
)
<emphasis id="B97257018D7CFFA5F928F8D1A02CF850" box="[790,843,1917,1940]" italics="true" pageId="14" pageNumber="415">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D7CFFA5F96CF8D1A0ECF851" bold="true" box="[850,907,1917,1941]" pageId="14" pageNumber="415">
<taxonomicNameLabel id="A241EA7A8D7CFFA5F96CF8D1A0ECF851" box="[850,907,1917,1941]" pageId="14" pageNumber="415" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype).
</paragraph>
</caption>
<caption id="DF79DB9B8D7DFFA4FAA9F8F3A3B6F87B" ID-DOI="http://doi.org/10.5281/zenodo.4397887" ID-Zenodo-Dep="4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="15" pageNumber="416" startId="15.[151,264,1887,1911]" targetBox="[151,1436,181,1863]" targetPageId="15">
<paragraph id="8BB98B138D7DFFA4FAA9F8F3A3B6F87B" blockId="15.[151,1436,1887,1983]" pageId="15" pageNumber="416">
<emphasis id="B97257018D7DFFA4FAA9F8F3A232F8B3" bold="true" box="[151,341,1887,1911]" pageId="15" pageNumber="416">FIGURES 4956.</emphasis>
Larva of
<taxonomicName id="4C06F0908D7DFFA4FB85F8F3A192F8B3" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[443,757,1887,1912]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D7DFFA4FB85F8F3A143F8B3" box="[443,548,1887,1911]" italics="true" pageId="15" pageNumber="416">Procloeon</emphasis>
(
<emphasis id="B97257018D7DFFA4F80DF8F3A1D7F8B3" box="[563,688,1887,1911]" italics="true" pageId="15" pageNumber="416">Oculogaster</emphasis>
)
<emphasis id="B97257018D7DFFA4F8FEF8CCA192F8B3" box="[704,757,1888,1911]" italics="true" pageId="15" pageNumber="416">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D7DFFA4F8C3F8CCA050F8BC" bold="true" box="[765,823,1888,1912]" pageId="15" pageNumber="416">
<taxonomicNameLabel id="A241EA7A8D7DFFA4F8C3F8CCA050F8BC" box="[765,823,1888,1912]" pageId="15" pageNumber="416" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides.
</paragraph>
</caption>
<paragraph id="8BB98B138D62FFBBFAF9FE52A103FD6D" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="417">
<emphasis id="B97257018D62FFBBFAF9FE52A224FDDD" bold="true" box="[199,323,510,537]" italics="true" pageId="16" pageNumber="417">Subimago.</emphasis>
CUTICULAR COLORATION. Head colorless. Pronotum with brown and colorless areas. Mesonotum brown with sutures darker brown, with large colorless areas occupying anterior part of submedioscutum and posterior scutal protuberance (
<figureCitation id="133D97968D62FFBBFBD4FDEBA159FDA5" box="[490,574,583,609]" captionStart="FIGURES 6770" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-16@17.[151,1436,181,1895]" captionTargetPageId="17" captionText="FIGURES 6770. Male Procloeon (Oculogaster) niger sp. n. 6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397893" httpUri="https://zenodo.org/record/4397893/files/figure.png" pageId="16" pageNumber="417">Fig. 70</figureCitation>
). Thoracic pleura mostly colorless, with some sclerites brown (
<figureCitation id="133D97968D62FFBBFEC1FDEBA634FDA6" box="[1279,1363,583,610]" captionStart="FIGURES 6770" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-16@17.[151,1436,181,1895]" captionTargetPageId="17" captionText="FIGURES 6770. Male Procloeon (Oculogaster) niger sp. n. 6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397893" httpUri="https://zenodo.org/record/4397893/files/figure.png" pageId="16" pageNumber="417">Fig. 69</figureCitation>
). Legs nearly colorless, with base of tibia brown, sometimes brownish stripes on femur. Abdomen very light brownish with sigilla colorless. Caudalii light brownish.
</paragraph>
<paragraph id="8BB98B138D62FFBBFAF9FD1FA3A3FCD1" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="417">
TEXTURE. On fore leg of male and female, 1st tarsal segment covered partly with microtrichia (as in tibia), partly with pointed microlepides; all other tarsal segments entirely covered with pointed microlepides (
<figureCitation id="133D97968D62FFBBFF09FD7BA3D0FCD1" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="16" pageNumber="417">Figs 7677</figureCitation>
).
</paragraph>
<caption id="DF79DB9B8D62FFBBFAA9F83FA11CF80B" ID-DOI="http://doi.org/10.5281/zenodo.4397889" ID-Zenodo-Dep="4397889" httpUri="https://zenodo.org/record/4397889/files/figure.png" pageId="16" pageNumber="417" startId="16.[151,264,1939,1963]" targetBox="[157,1431,818,1915]" targetPageId="16">
<paragraph id="8BB98B138D62FFBBFAA9F83FA11CF80B" blockId="16.[151,1436,1939,1999]" pageId="16" pageNumber="417">
<emphasis id="B97257018D62FFBBFAA9F83FA233F86F" bold="true" box="[151,340,1939,1963]" pageId="16" pageNumber="417">FIGURES 5766.</emphasis>
Larvae of
<taxonomicName id="4C06F0908D62FFBBFBFEF83FA190F86F" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[448,759,1939,1963]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D62FFBBFBFEF83FA14EF86F" box="[448,553,1939,1963]" italics="true" pageId="16" pageNumber="417">Procloeon</emphasis>
(
<emphasis id="B97257018D62FFBBF809F83FA1D3F86F" box="[567,692,1939,1963]" italics="true" pageId="16" pageNumber="417">Oculogaster</emphasis>
)
<emphasis id="B97257018D62FFBBF8FCF838A190F86F" box="[706,759,1940,1963]" italics="true" pageId="16" pageNumber="417">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D62FFBBF8C0F83FA051F86F" bold="true" box="[766,822,1939,1963]" pageId="16" pageNumber="417">
<taxonomicNameLabel id="A241EA7A8D62FFBBF8C0F83FA051F86F" box="[766,822,1939,1963]" pageId="16" pageNumber="417" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
5762, tergalii IIII and VVII; 6365, tergalii I, IV and VII of other specimens (6465, holotype); 66, claw.
</paragraph>
</caption>
<caption id="DF79DB9B8D63FFBAFAA9F8D2A027F87F" ID-DOI="http://doi.org/10.5281/zenodo.4397893" ID-Zenodo-Dep="4397893" httpUri="https://zenodo.org/record/4397893/files/figure.png" pageId="17" pageNumber="418" startId="17.[151,264,1918,1942]" targetBox="[151,1436,181,1895]" targetPageId="17">
<paragraph id="8BB98B138D63FFBAFAA9F8D2A027F87F" blockId="17.[151,1436,1918,1979]" pageId="17" pageNumber="418">
<emphasis id="B97257018D63FFBAFAA9F8D2A232F852" bold="true" box="[151,341,1918,1942]" pageId="17" pageNumber="418">FIGURES 6770.</emphasis>
Male
<taxonomicName id="4C06F0908D63FFBAFBA6F8D3A1B5F852" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[408,722,1919,1943]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D63FFBAFBA6F8D3A166F853" box="[408,513,1919,1943]" italics="true" pageId="17" pageNumber="418">Procloeon</emphasis>
(
<emphasis id="B97257018D63FFBAF82EF8D3A1EAF853" box="[528,653,1919,1943]" italics="true" pageId="17" pageNumber="418">Oculogaster</emphasis>
)
<emphasis id="B97257018D63FFBAF8A3F8D3A1B5F852" box="[669,722,1919,1942]" italics="true" pageId="17" pageNumber="418">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D63FFBAF8E4F8D3A073F853" bold="true" box="[730,788,1919,1943]" pageId="17" pageNumber="418">
<taxonomicNameLabel id="A241EA7A8D63FFBAF8E4F8D3A073F853" box="[730,788,1919,1943]" pageId="17" pageNumber="418" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype).
</paragraph>
</caption>
<paragraph id="8BB98B138D60FFB9FAF9FF3BA7D1FD6D" blockId="18.[151,1437,150,681]" pageId="18" pageNumber="419">
<emphasis id="B97257018D60FFB9FAF9FF3BA231FF75" bold="true" box="[199,342,150,177]" italics="true" pageId="18" pageNumber="419">Imago, male</emphasis>
(
<figureCitation id="133D97968D60FFB9FB5AFF3BA282FF76" box="[356,485,151,178]" captionStart="FIGURES 6770" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-16@17.[151,1436,181,1895]" captionTargetPageId="17" captionText="FIGURES 6770. Male Procloeon (Oculogaster) niger sp. n. 6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397893" httpUri="https://zenodo.org/record/4397893/files/figure.png" pageId="18" pageNumber="419">Figs 6768</figureCitation>
). Head ocher with brown. Antenna pale ocher, joining of scapus and pedicellus brown. Turbinate eyes widened apically; facetted surface and dorsal part of stem light yellow, ventral part of stem brown. Thorax brown, with sterna lighter yellowish, pleura with longitudinal white and darker brown bands. Legs light yellow. In
<typeStatus id="54BD35B18D60FFB9FAD3FEAFA237FED9" box="[237,336,259,285]" pageId="18" pageNumber="419" type="paratype">paratype</typeStatus>
, ratio femur / tibia / tarsal segments (mm) on fore leg 1.10: 1.55: 0.05: 0.72: 0.60: 0.30: 0.17; on middle and hind leg 0.92: 0.88: 0.30: 0.13: 0.05: 0.15. Middle and hind legs normally with 2 apical spines: on 1st+2nd and on primary 3rd tarsomeres (as in
<figureCitation id="133D97968D60FFB9F8A1FEE7A197FEA1" box="[671,752,331,357]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="18" pageNumber="419">Fig. 77</figureCitation>
) (among specimens examined, only in
<typeStatus id="54BD35B18D60FFB9FE95FEE7A66AFEA1" box="[1195,1293,331,357]" pageId="18" pageNumber="419" type="holotype">holotype</typeStatus>
one of these spines lacking on some legs). Wing with membrane colorless, veins ocher; subcostal vein proximad of costal brace colored with brown. Pterostigma with one oblique cross vein; besides it, costal field with very slender cross vein belonging to middle cord of cross veins. Marginal intercalaries present in most of interspaces, both behind convex and concave longitudinal veins. Hind wing absent. Abdominal terga mostly brown; terga IIVI with lateral areas dark brown, median area lighter, with pair of light spots adjacent to anterior margin, light submedian sigilla and unpaired light spot adjacent to posterior margin; tergum VII dark brown with pair of light spots adjacent to anterior margin and light submedian sigilla; terga
<collectionCode id="ED1713D68D60FFB9F853FDEBA1D8FDA5" box="[621,703,583,609]" pageId="18" pageNumber="419">VIIIX</collectionCode>
dark brown with light submedian sigilla. Abdominal sterna light ocher, with pair of longitudinal lateral brown stripes. Gonostyli pale ocher, penis brown. Median projection of penis semicircular (
<figureCitation id="133D97968D60FFB9FB0FFD23A2E3FD6D" box="[305,388,655,681]" captionStart="FIGURES 7172" captionStartId="18.[151,264,1960,1984]" captionTargetBox="[170,1428,712,1926]" captionTargetId="figure-305@18.[151,1436,705,1936]" captionTargetPageId="18" captionText="FIGURES 7172. Male Procloeon (Oculogaster) niger sp. n. 71, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 72, their subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.4397895" httpUri="https://zenodo.org/record/4397895/files/figure.png" pageId="18" pageNumber="419">Fig. 71</figureCitation>
). Cerci whitish, with brown apex of each segment or every 2nd segment.
</paragraph>
<caption id="DF79DB9B8D60FFB9FAA9F804A01CF820" ID-DOI="http://doi.org/10.5281/zenodo.4397895" ID-Zenodo-Dep="4397895" httpUri="https://zenodo.org/record/4397895/files/figure.png" pageId="18" pageNumber="419" startId="18.[151,264,1960,1984]" targetBox="[170,1428,712,1926]" targetPageId="18">
<paragraph id="8BB98B138D60FFB9FAA9F804A01CF820" blockId="18.[151,1436,1960,2020]" pageId="18" pageNumber="419">
<emphasis id="B97257018D60FFB9FAA9F804A234F804" bold="true" box="[151,339,1960,1984]" pageId="18" pageNumber="419">FIGURES 7172.</emphasis>
Male
<taxonomicName id="4C06F0908D60FFB9FBAFF804A1A0F804" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[401,711,1960,1984]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D60FFB9FBAFF804A29DF804" box="[401,506,1960,1984]" italics="true" pageId="18" pageNumber="419">Procloeon</emphasis>
(
<emphasis id="B97257018D60FFB9F839F804A1E3F804" box="[519,644,1960,1984]" italics="true" pageId="18" pageNumber="419">Oculogaster</emphasis>
)
<emphasis id="B97257018D60FFB9F8ACF805A1A0F804" box="[658,711,1961,1984]" italics="true" pageId="18" pageNumber="419">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D60FFB9F8F2F804A063F804" bold="true" box="[716,772,1960,1984]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="A241EA7A8D60FFB9F8F2F804A063F804" box="[716,772,1960,1984]" pageId="18" pageNumber="419" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
71, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 72, their subimaginal exuviae.
</paragraph>
</caption>
<paragraph id="8BB98B138D61FFB8FAF9FF3BA389FE31" blockId="19.[151,1437,150,969]" pageId="19" pageNumber="420">
<emphasis id="B97257018D61FFB8FAF9FF3BA208FF75" bold="true" box="[199,367,150,177]" italics="true" pageId="19" pageNumber="420">Imago, female</emphasis>
(
<figureCitation id="133D97968D61FFB8FB40FF3BA165FF76" box="[382,514,151,178]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Figs 7376</figureCitation>
). Head moderately wide, distance between eyes subequal to eye length; eyes only slightly elevated above head surface. Head ocher, with reddish brown and white stripes. Pronotum ocher, with small brown spots. Mesonotum ocher, bordered with reddish-brown and white stripes. Thoracic pleura ocher with wide white and brown longitudinal bands. Sterna and legs light yellow. Fore leg without apical spine on 2nd tarsomere, with single apical spine on primary 3rd tarsomere (
<figureCitation id="133D97968D61FFB8F8E7FE8BA048FE86" box="[729,815,295,322]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Fig. 76</figureCitation>
); middle and hind legs each with 2 spines (
<figureCitation id="133D97968D61FFB8FF27FE8BA60CFE85" box="[1305,1387,295,321]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Fig. 77</figureCitation>
) (as in male see above). Wings as in male (see above). Abdominal terga with peculiar contrasting ornament gradually changing from tergum II to tergum IX, including dark reddish-brown lateral areas, medial ocher area and paired white spots; terga
<collectionCode id="ED1713D68D61FFB8FB5EFE3FA2DBFE69" box="[352,444,403,429]" pageId="19" pageNumber="420">VIIIIX</collectionCode>
with median reddish-brownish stripe; tergum X reddish-brownish (
<figureCitation id="133D97968D61FFB8FE94FE3FA641FE69" box="[1194,1318,402,429]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Fig. 73, 75</figureCitation>
). Abdominal sterna ocher, with pair of longitudinal lateral reddish-brown stripes (
<figureCitation id="133D97968D61FFB8F982FE1BA777FE15" box="[956,1040,439,465]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Fig. 74</figureCitation>
). Cerci as in male (see above) (
<figureCitation id="133D97968D61FFB8FF51FE1BA385FE31" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Fig. 7374</figureCitation>
).
</paragraph>
<paragraph id="8BB98B138D61FFB8FAF9FE53A1E2FDDE" blockId="19.[151,1437,150,969]" box="[199,645,511,538]" pageId="19" pageNumber="420">
<emphasis id="B97257018D61FFB8FAF9FE53A39BFDDD" bold="true" box="[199,252,511,537]" italics="true" pageId="19" pageNumber="420">Egg.</emphasis>
Without rigid chorion (viviparity).
</paragraph>
<paragraph id="8BB98B138D61FFB8FAF9FD8EA01AFDF9" blockId="19.[151,1437,150,969]" box="[199,893,546,573]" pageId="19" pageNumber="420">
<emphasis id="B97257018D61FFB8FAF9FD8EA229FDF8" bold="true" box="[199,334,546,572]" pageId="19" pageNumber="420">Dimension.</emphasis>
Fore wing length of male
<quantity id="4CFE26F68D61FFB8F849FD8EA1B1FDF9" box="[631,726,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="19" pageNumber="420" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, female
<quantity id="4CFE26F68D61FFB8F90BFD8FA01EFDF9" box="[821,889,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="420" unit="mm" value="6.0">6 mm</quantity>
.
</paragraph>
<paragraph id="8BB98B138D61FFB8FAF9FDEBA61AFD09" blockId="19.[151,1437,150,969]" pageId="19" pageNumber="420">
<emphasis id="B97257018D61FFB8FAF9FDEBA21AFDA4" bold="true" box="[199,381,582,609]" pageId="19" pageNumber="420">Larval habitat.</emphasis>
Larvae were collected in a small stream running in dark forest; just below the waterfall, this stream represent a chain of pools bordered by rocks, among which the first pool (into which the waterfall falls) is large, and downstream pools are smaller; all larvae of
<taxonomicName id="4C06F0908D61FFB8F8D5FD23A014FD6C" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[747,883,655,681]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8F8D5FD23A067FD6C" box="[747,768,655,680]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8F930FD23A04EFD6D" box="[782,809,655,681]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8F906FD23A014FD6C" box="[824,883,655,680]" italics="true" pageId="19" pageNumber="420">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D61FFB8F947FD23A0D0FD6D" bold="true" box="[889,951,655,681]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A241EA7A8D61FFB8F947FD23A0D0FD6D" box="[889,951,655,681]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected in a single pool, which is the 3rd from the waterfall; in spite of intensive collecting, in other places of this stream such larvae were not found.
</paragraph>
<paragraph id="8BB98B138D61FFB8FAF9FD7AA18FFC0D" blockId="19.[151,1437,150,969]" pageId="19" pageNumber="420">
<emphasis id="B97257018D61FFB8FAF9FD7AA205FD34" bold="true" box="[199,354,726,752]" pageId="19" pageNumber="420">Comparison.</emphasis>
Larva of the new species
<taxonomicName id="4C06F0908D61FFB8F8B6FD7BA073FD34" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[648,788,727,753]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8F8B6FD7BA1F9FD34" box="[648,670,727,752]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8F893FD7BA1AEFD35" box="[685,713,727,753]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8F8E7FD7BA073FD34" box="[729,788,727,752]" italics="true" pageId="19" pageNumber="420">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D61FFB8F922FD7BA03CFD35" bold="true" box="[796,859,727,753]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A241EA7A8D61FFB8F922FD7BA03CFD35" box="[796,859,727,753]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C06F0908D61FFB8F9CEFD7BA7C3FD35" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1008,1188,727,753]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8F9CEFD7BA762FD34" box="[1008,1029,727,752]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8FE2BFD7BA757FD35" box="[1045,1072,727,753]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8FE7FFD7BA7C3FD35" box="[1089,1188,727,753]" italics="true" pageId="19" pageNumber="420">barnardi</emphasis>
</taxonomicName>
<emphasis id="B97257018D61FFB8FE95FD7BA78CFD35" bold="true" box="[1195,1259,727,753]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A241EA7A8D61FFB8FE95FD7BA78CFD35" box="[1195,1259,727,753]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C06F0908D61FFB8FF1FFD7BA270FCD1" authorityName=": Kluge" authorityYear="2016" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8FF1FFD7BA650FD34" box="[1313,1335,727,752]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8FF78FD7BA605FD35" box="[1350,1378,727,753]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8FF4CFD7BA270FCD1" italics="true" pageId="19" pageNumber="420">cylindroculum</emphasis>
</taxonomicName>
by the uniform brown coloration of abdominal terga, longer rows of denticles on claws and by the presence of lateral spines on abdominal segment IX only. Thin terminal portions of cerci lacking swimming setae and spines, in
<taxonomicName id="4C06F0908D61FFB8FB39FCEFA2F6FC98" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[263,401,835,861]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8FB39FCEFA27BFC98" box="[263,284,835,860]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8FB15FCEFA221FC99" box="[299,326,835,861]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8FB68FCEFA2F6FC98" box="[342,401,835,860]" italics="true" pageId="19" pageNumber="420">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D61FFB8FBA6FCEFA2B0FC99" bold="true" box="[408,471,835,861]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A241EA7A8D61FFB8FBA6FCEFA2B0FC99" box="[408,471,835,861]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are usually longer than in other species (
<figureCitation id="133D97968D61FFB8F9A3FCEFA096FC99" box="[925,1009,835,861]" captionStart="FIGURES 4348" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="19" pageNumber="420">Fig. 43</figureCitation>
). Imago and subimago of
<taxonomicName id="4C06F0908D61FFB8FF2CFCEFA6FBFC98" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1298,1436,835,861]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8FF2CFCEFA640FC98" box="[1298,1319,835,860]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8FF08FCEFA636FC99" box="[1334,1361,835,861]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8FF5FFCEFA6FBFC98" box="[1377,1436,835,860]" italics="true" pageId="19" pageNumber="420">niger</emphasis>
</taxonomicName>
<emphasis id="B97257018D61FFB8FAA9FCCBA3BFFC45" bold="true" box="[151,216,871,897]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A241EA7A8D61FFB8FAA9FCCBA3BFFC45" box="[151,216,871,897]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C06F0908D61FFB8FB51FCCBA142FC45" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[367,549,871,897]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8FB51FCCBA2E2FC44" box="[367,389,871,896]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8FBABFCCBA2D6FC45" box="[405,433,871,897]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8FBFCFCCBA142FC45" box="[450,549,871,897]" italics="true" pageId="19" pageNumber="420">barnardi</emphasis>
</taxonomicName>
<emphasis id="B97257018D61FFB8F813FCCBA109FC45" bold="true" box="[557,622,871,897]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A241EA7A8D61FFB8F813FCCBA109FC45" box="[557,622,871,897]" pageId="19" pageNumber="420" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C06F0908D61FFB8F898FCCBA0FDFC45" authorityName=": Kluge" authorityYear="2016" box="[678,922,871,897]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="B97257018D61FFB8F898FCCBA1DBFC44" box="[678,700,871,896]" italics="true" pageId="19" pageNumber="420">P.</emphasis>
(
<emphasis id="B97257018D61FFB8F8F2FCCBA180FC45" box="[716,743,871,897]" italics="true" pageId="19" pageNumber="420">O.</emphasis>
)
<emphasis id="B97257018D61FFB8F8C7FCCBA0FDFC45" box="[761,922,871,897]" italics="true" pageId="19" pageNumber="420">cylindroculum</emphasis>
</taxonomicName>
by the presence of 2 apical spines on middle and hind tarsi (
<figureCitation id="133D97968D61FFB8FB03FC27A2F6FC61" box="[317,401,907,933]" captionStart="FIGURES 7377" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="19" pageNumber="420">Fig. 77</figureCitation>
). Winged stages of male and female differ from other species by more extensive and uniform brown hypodermal pigmentation of abdominal terga.
</paragraph>
</subSubSection>
</treatment>
</document>