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<mods:title id="3F506A3CF7F42939BA0F0F561837C282">Revision of the genus Odocnemis Allard, 1876 (Coleoptera: Tenebrionidae: Helopini) from Turkey, the Caucasus and Iran with observations on feeding habits</mods:title>
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<taxonomicName id="4C0435BDF8014923FF1EFA403E8CFA7F" authority="Faldermann, 1837" authorityName="Faldermann" authorityYear="1837" box="[151,646,1414,1440]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="gloriosa" status="comb. nov.">
<emphasis id="B970922CF8014923FF1EFA403D8DFA7F" bold="true" box="[151,391,1414,1440]" italics="true" pageId="82" pageNumber="83">Odocnemis gloriosa</emphasis>
(
<bibRefCitation id="EF9533CFF8014923FE1FFA403E74FA7F" author="Faldermann" box="[406,638,1414,1440]" pageId="82" pageNumber="83" refString="Faldermann, F. (1837) Fauna Entomologica Trans-Caucasica. Coleoptera. Pars II. Auguste Semen, Moscou, 433 pp, 15 pls." type="book" year="1837">Faldermann, 1837</bibRefCitation>
)
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,
<taxonomicNameLabel id="A2432F57F8014923FD1DFA403EF5FA7F" box="[660,767,1414,1440]" pageId="82" pageNumber="83">comb. n.</taxonomicNameLabel>
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<figureCitation id="133F52BBF8014923FF1EFA6F3CF8FA1E" box="[151,242,1449,1474]" captionStart="FIGURE 39. O" captionStartId="72.[151,250,1157,1179]" captionTargetBox="[156,1432,585,1146]" captionTargetId="figure@72.[150,1437,577,1150]" captionTargetPageId="72" captionText="FIGURE 39. O. subtuberculigera thracica subsp. n. and O. gloriosa. A) Pronotum of O. t. thracica, male. B) The same, female. C) Protibia of O. gloriosa. Scale bars— 1 mm." httpUri="https://zenodo.org/record/192259/files/figure.png" pageId="82" pageNumber="83">Figs. 39</figureCitation>
C, 48G,H, 53
</paragraph>
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<emphasis id="B970922CF8014923FF1EFA373CFFF9D8" box="[151,245,1521,1543]" italics="true" pageId="82" pageNumber="83">gloriosus</emphasis>
<bibRefCitation id="EF9533CFF8014923FF72FA373DBCF9D8" author="Faldermann" box="[251,438,1521,1543]" pageId="82" pageNumber="83" refString="Faldermann, F. (1837) Fauna Entomologica Trans-Caucasica. Coleoptera. Pars II. Auguste Semen, Moscou, 433 pp, 15 pls." type="book" year="1837">Faldermann, 1837</bibRefCitation>
: 67, tab. I. f. 2 (
<taxonomicName id="4C0435BDF8014923FDDDFA373E96F9D8" box="[596,668,1521,1543]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FDDDFA373E96F9D8" box="[596,668,1521,1543]" italics="true" pageId="82" pageNumber="83">Helops</emphasis>
</taxonomicName>
);
<bibRefCitation id="EF9533CFF8014923FD38FA373F6CF9D8" author="Faldermann" box="[689,870,1521,1543]" pageId="82" pageNumber="83" refString="Faldermann, F. (1838) Fauna Entomologica Trans-Caucasica. Coleoptera. Pars III. Auguste Semen, Moscou, 338 pp." type="book" year="1838">Faldermann 1838</bibRefCitation>
: 1 (
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<emphasis id="B970922CF8014923FC03FA373FD8F9D8" box="[906,978,1521,1543]" italics="true" pageId="82" pageNumber="83">Helops</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FBE4FA3738E0F9D8" author="Allard" box="[1133,1258,1521,1543]" pageId="82" pageNumber="83" refString="Allard, E. (1876) Revision des Helopines vrais de Lacordaire. L'Abeille, Journal d'Entomologie, 14, 1 - 80." type="journal article" year="1876">Allard 1876</bibRefCitation>
: 16 (
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<emphasis id="B970922CF8014923FA92FA343977F9D8" box="[1307,1405,1522,1543]" italics="true" pageId="82" pageNumber="83">Stenomax</emphasis>
</taxonomicName>
, in key);
<bibRefCitation id="EF9533CFF8014923FE8DF9D63D8FF9F9" author="Allard" box="[260,389,1552,1574]" pageId="82" pageNumber="83" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">Allard 1877</bibRefCitation>
: 34, 134 (
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<emphasis id="B970922CF8014923FE7AF9D73E5EF9F9" box="[499,596,1553,1574]" italics="true" pageId="82" pageNumber="83">Stenomax</emphasis>
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, in key, redescription);
<bibRefCitation id="EF9533CFF8014923FCD8F9D63852F9F9" author="Schneider" box="[849,1112,1552,1574]" pageId="82" pageNumber="83" refString="Schneider, O. &amp; Leder, H. (1878) Beitrage zur Kenntniss der kaukasischen Kaferfauna. Ferhandlungen des Naturforschendes Fereines in Brunn, 16, 3 - 262." type="journal article" year="1878">Schneider &amp; Leder 1878</bibRefCitation>
: 243 (
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<emphasis id="B970922CF8014923FB14F9D738F5F9F9" box="[1181,1279,1553,1574]" italics="true" pageId="82" pageNumber="83">Stenomax</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FF4EF9E83D9FF99B" author="Heyden" box="[199,405,1582,1604]" pageId="82" pageNumber="83" refString="Heyden, L. von, Reitter, E. &amp; Weise, J. (1883) Catalogus Coleopterorum Europae et Caucasi. Editio tertia. Libraria Nicolai, Berlin, 228 pp." type="book" year="1883">
Heyden
<emphasis id="B970922CF8014923FE96F9E93D5CF99B" box="[287,342,1582,1604]" italics="true" pageId="82" pageNumber="83">et al.</emphasis>
1883
</bibRefCitation>
: 136 (
<taxonomicName id="4C0435BDF8014923FE52F9E93E37F99B" box="[475,573,1583,1604]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FE52F9E93E37F99B" box="[475,573,1583,1604]" italics="true" pageId="82" pageNumber="83">Stenomax</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FD6DF9E83F6FF99B" author="Radde" box="[740,869,1582,1604]" pageId="82" pageNumber="83" refString="Radde, G. I. (1899) Collections of Caucasian Museum. Fol. 1. Zoology. Typography Office of civilian supreme commander in the Caucasus, Tiflis, 520 pp." type="book" year="1899">Radde 1899</bibRefCitation>
: 381 (
<taxonomicName id="4C0435BDF8014923FC23F9E83FF8F99B" box="[938,1010,1582,1604]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FC23F9E83FF8F99B" box="[938,1010,1582,1604]" italics="true" pageId="82" pageNumber="83">Helops</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FB13F9E83963F99B" author="Heyden" box="[1178,1385,1582,1604]" pageId="82" pageNumber="83" refString="Heyden, L. von, Reitter, E. &amp; Weise, J. (1891) Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. E. Reitter, Modling, 420 pp." type="book" year="1891">
Heyden
<emphasis id="B970922CF8014923FB7AF9E93923F99B" box="[1267,1321,1582,1604]" italics="true" pageId="82" pageNumber="83">et al.</emphasis>
1891
</bibRefCitation>
: 252 (
<taxonomicName id="4C0435BDF8014923FF46F98B3D9BF9BC" authority="(Stenomax)" baseAuthorityName="Stenomax" box="[207,401,1613,1635]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FF46F98B3D1DF9BC" box="[207,279,1613,1635]" italics="true" pageId="82" pageNumber="83">Helops</emphasis>
(
<emphasis id="B970922CF8014923FEAFF9883D82F9BC" box="[294,392,1614,1635]" italics="true" pageId="82" pageNumber="83">Stenomax</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FDA4F98B3EB2F9BC" author="Seidlitz" box="[557,696,1613,1635]" pageId="82" pageNumber="83" refString="Seidlitz, G. (1896) Tenebrionidae. In: Kiesenwetter H. von, Seidlitz G. von, Naturgeschichte der Insecten Deutschlands. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800" type="book chapter" year="1896">Seidlitz 1896</bibRefCitation>
: 729;
<bibRefCitation id="EF9533CFF8014923FD7CF98B3F73F9BC" author="Gebien" box="[757,889,1613,1635]" pageId="82" pageNumber="83" refString="Gebien, H. (1911) Pars. 28. Tenebrionidae. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus auspiciis et auxilio. Fol. 18. Junk, W., Berlin, pp. 355 - 585" type="journal article" year="1911">Gebien 1911</bibRefCitation>
: 543 (
<taxonomicName id="4C0435BDF8014923FC31F98B3871F9BC" authority="(Stenomax)" baseAuthorityName="Stenomax" box="[952,1147,1613,1635]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FC31F98B380AF9BC" box="[952,1024,1613,1635]" italics="true" pageId="82" pageNumber="83">Helops</emphasis>
(
<emphasis id="B970922CF8014923FB86F988387BF9BC" box="[1039,1137,1614,1635]" italics="true" pageId="82" pageNumber="83">Stenomax</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FA9EF98B3992F9BC" author="Reitter" box="[1303,1432,1613,1635]" pageId="82" pageNumber="83" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44, 113 - 171. http: // dx. doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter 1922</bibRefCitation>
: 138 (
<taxonomicName id="4C0435BDF8014923FF75F9AA3DF0F95D" authority="(Omaleis)" baseAuthorityName="Omaleis" box="[252,506,1644,1666]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FF75F9AA3D86F95D" box="[252,396,1644,1666]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
(
<emphasis id="B970922CF8014923FE15F9AA3DFAF95D" box="[412,496,1644,1666]" italics="true" pageId="82" pageNumber="83">Omaleis</emphasis>
)
</taxonomicName>
, in key);
<bibRefCitation id="EF9533CFF8014923FDD2F9AA3EE2F95D" author="Winkler" box="[603,744,1644,1666]" pageId="82" pageNumber="83" refString="Winkler, A. (1928 - 1929) Pars. 8, 9: D. Heteromera. In: Winkler, A. (Ed.) Catalogus Coleopterorum regionis Palaearctica. Verlag von Albert Winkler, Wien, pp. 881 - 1136." type="book chapter" year="1928">
Winkler
<date id="FFBA68FEF8014923FD3DF9AA3EE2F95D" box="[692,744,1644,1666]" pageId="82" pageNumber="83" value="1928" valueMax="1929">1928</date>
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–1929: 1018 (
<taxonomicName id="4C0435BDF8014923FCFFF9AA387EF95D" authority="(Omaleis)" baseAuthorityName="Omaleis" box="[886,1140,1644,1666]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FCFFF9AA380CF95D" box="[886,1030,1644,1666]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
(
<emphasis id="B970922CF8014923FB9FF9AA3860F95D" box="[1046,1130,1644,1666]" italics="true" pageId="82" pageNumber="83">Omaleis</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FA9BF9AA3992F95D" author="Gebien" box="[1298,1432,1644,1666]" pageId="82" pageNumber="83" refString="Gebien, H. (1943) Katalog der Tenebrioniden. Teil. 3. Mitteilungen der Munchner Entomologischen Gesellschaft, 33, 399 - 430 (778 - 809)." type="journal article" year="1943">Gebien 1943</bibRefCitation>
: 428 (807) (
<taxonomicName id="4C0435BDF8014923FEB4F94C3E37F97F" authority="(Omaleis)" baseAuthorityName="Omaleis" box="[317,573,1674,1696]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FEB4F94C3DC7F97F" box="[317,461,1674,1696]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
(
<emphasis id="B970922CF8014923FE57F94C3E38F97F" box="[478,562,1674,1696]" italics="true" pageId="82" pageNumber="83">Omaleis</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FD56F94C3FE5F97F" author="Iablokoff-Khnzorian" box="[735,1007,1674,1696]" pageId="82" pageNumber="83" refString="Iablokoff-Khnzorian, S. M. (1964) Bemmerkungen uber einige Reiiter'sche Typen aus dem Ungarischen Naturwissenschaftlichen Museum (Coleoptera). Folia entomologica Hungarica, 17, 293 - 315." type="journal article" year="1964">Iablokoff-Khnzorian 1964</bibRefCitation>
: 306, abb. 1h (
<taxonomicName id="4C0435BDF8014923FB05F94C38DEF97F" box="[1164,1236,1674,1696]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FB05F94C38DEF97F" box="[1164,1236,1674,1696]" italics="true" pageId="82" pageNumber="83">Helops</emphasis>
</taxonomicName>
, faunistic records);
<bibRefCitation id="EF9533CFF8014923FF4EF96F3DB4F960" author="Dzhambazishvili" box="[199,446,1705,1727]" pageId="82" pageNumber="83" refString="Dzhambazishvili, M. Ya. (1983) Vertical zonal distribution of darkling beetles of Lagodekhi Nature Reserve. In: Fauna and ecology of Invertebrates of Georgia. Mitsniereba, Tbilisi, pp. 96 - 103. [in Russian]" type="book chapter" year="1983">Dzhambazishvili 1983</bibRefCitation>
: 96–103 (
<taxonomicName id="4C0435BDF8014923FDC8F96F3EDBF960" box="[577,721,1705,1727]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FDC8F96F3EDBF960" box="[577,721,1705,1727]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
</taxonomicName>
, faunistic records);
<bibRefCitation id="EF9533CFF8014923FC43F96F3963F960" author="Medvedev" box="[970,1385,1705,1727]" pageId="82" pageNumber="83" refString="Medvedev, G. S. &amp; Abdurakhmanov, G. M. (1988) Check-list of beetles of the family Tenebrionidae of South West of Palaearctic. In: Abdurakhmanov, G. M. Eastern Caucasus through the eyes of an entomologist. Dagestanian book Publishing House, Makhachkala, pp. 71 - 99. [in Russian]" type="book chapter" year="1988">Medvedev &amp; Abdurakhmanov 1988</bibRefCitation>
: 98 (
<taxonomicName id="4C0435BDF8014923FF46F90E3D55F901" box="[207,351,1736,1758]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FF46F90E3D55F901" box="[207,351,1736,1758]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FE75F90E3F64F901" author="Abdurakhmanov" box="[508,878,1736,1758]" pageId="82" pageNumber="83" refString="Abdurakhmanov, G. M. &amp; Medvedev, G. S. (1994) Catalogue of darkling beetles of the Caucasus. Daghestan State Pedagogical Institute, Makhachkala. 212 pp. [in Russian]" type="book" year="1994">Abdurakhmanov &amp; Medvedev 1994</bibRefCitation>
: 192 (
<taxonomicName id="4C0435BDF8014923FC25F90E3836F901" box="[940,1084,1736,1758]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FC25F90E3836F901" box="[940,1084,1736,1758]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FB50F90E3D2AF923" author="Abdurakhmanov" pageId="82" pageNumber="83" refString="Abdurakhmanov, G. M., Ismailov, Sh. I. &amp; Lobanov, A. L. (1995) A new approach to the problem of objective zoogeographical districting. Part 5. Computer base of dates TENKAF about fauna of darkling beetles of the Caucasus. Dagestanian State Pedagogical Institute, Makhachkala. 325 pp. [in Russian]" type="book" year="1995">
Abdurakhmanov
<emphasis id="B970922CF8014923FA03F90F3CE8F923" italics="true" pageId="82" pageNumber="83">et al.</emphasis>
1995
</bibRefCitation>
: 212 (
<taxonomicName id="4C0435BDF8014923FEECF9203DFFF923" box="[357,501,1766,1788]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FEECF9203DFFF923" box="[357,501,1766,1788]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
</taxonomicName>
, in check-list);
<bibRefCitation id="EF9533CFF8014923FD14F9203F57F923" author="Nabozhenko" box="[669,861,1766,1788]" pageId="82" pageNumber="83" refString="Nabozhenko, M. V. (1999) Landscape-ecological characteristics and common regularities of the darkling beetles (Coleoptera, Tenebrionidae) fauna distribution in south Russia. Izvesyia Kharkovskogo entomologicheskogo obshchestva, 7, 40 - 54. [in Russian]" type="journal article" year="1999">Nabozhenko 1999</bibRefCitation>
: 52 (
<taxonomicName id="4C0435BDF8014923FC1DF920382EF923" box="[916,1060,1766,1788]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FC1DF920382EF923" box="[916,1060,1766,1788]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
</taxonomicName>
, bionomics);
<bibRefCitation id="EF9533CFF8014923FB3FF920397DF923" author="Nabozhenko" box="[1206,1399,1766,1788]" pageId="82" pageNumber="83" refString="Nabozhenko, M. V. (2000) New data about the distribution of darkling beetles of the tribe Helopini (Coleoptera, Tenebrionidae). Fauna Stavropol'ya, 10, 50 - 56." type="journal article" year="2000">Nabozhenko 2000</bibRefCitation>
: 53 (
<taxonomicName id="4C0435BDF8014923FF46F8C33D55F8C4" box="[207,351,1797,1819]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FF46F8C33D55F8C4" box="[207,351,1797,1819]" italics="true" pageId="82" pageNumber="83">Cylindronotus</emphasis>
</taxonomicName>
, faunistic records);
<bibRefCitation id="EF9533CFF8014923FDB2F8C33F23F8C4" author="Dzhambazishvili" box="[571,809,1797,1819]" pageId="82" pageNumber="83" refString="Dzhambazishvili, M. Ya. (2000) Catalogue of darkling beetles (Coleoptera, Tenebrionidae) of Georgia. Trudy Instituta Zoologii, Tbilisi, 20, 185 - 193. [in Russian]" type="journal article" year="2000">Dzhambazishvili 2000</bibRefCitation>
: 191 (
<taxonomicName id="4C0435BDF8014923FCFBF8C3383BF8C4" box="[882,1073,1797,1819]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="cloriosus">
<emphasis id="B970922CF8014923FCFBF8C3383BF8C4" box="[882,1073,1797,1819]" italics="true" pageId="82" pageNumber="83">Nalassus cloriosus</emphasis>
</taxonomicName>
, incorrect spelling, in catalogue);
<bibRefCitation id="EF9533CFF8014923FF4EF8E23D89F8E5" author="Nabozhenko" box="[199,387,1828,1850]" pageId="82" pageNumber="83" refString="Nabozhenko, M. V. (2001) On the classification of the Tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomological Review, 81, 909 - 942." type="journal article" year="2001">Nabozhenko 2001</bibRefCitation>
: 656, 662 (
<taxonomicName id="4C0435BDF8014923FE7FF8E23EE4F8E5" authority="(Helopondrus)" baseAuthorityName="Helopondrus" box="[502,750,1828,1850]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FE7FF8E23E5BF8E5" box="[502,593,1828,1850]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FDE9F8E23EEEF8E5" box="[608,740,1828,1850]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, redescription in key);
<bibRefCitation id="EF9533CFF8014923FC53F8E23993F8E5" author="Abdurakhmanov" box="[986,1433,1828,1850]" pageId="82" pageNumber="83" refString="Abdurakhmanov, G. M. &amp; Abdulmuslimova, K. M. (2002) Composition, morphoecological structure and zoogeographic peculiarities of tenebrionid populations in the Caucasus. Russian entomological journal, 11, 41 - 48." type="journal article" year="2002">Abdurakhmanov &amp; Abdulmuslimova, 2002</bibRefCitation>
: 45 (
<taxonomicName id="4C0435BDF8014923FF66F8843D40F887" box="[239,330,1858,1880]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FF66F8843D40F887" box="[239,330,1858,1880]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
</taxonomicName>
, faunistic records);
<bibRefCitation id="EF9533CFF8014923FD9CF8843ED8F887" author="Nabozhenko" box="[533,722,1858,1880]" pageId="82" pageNumber="83" refString="Nabozhenko, M. V. (2004) Darkling-beetles of the tribe Helopini (Coleoptera, Tenebrionidae) in the collection of E. E. Ballion (Part 1. Subtribe Cylindronotina). Izvestiya Muzeinogo Fonda im. A. A. Braunera, 1 (3 - 4), 1 - 6. [in Russian]" type="journal article" year="2004">Nabozhenko 2004</bibRefCitation>
: 4
<taxonomicName id="4C0435BDF8014923FD67F8843FECF887" authority="(Helopondrus)" baseAuthorityName="Helopondrus" box="[750,998,1858,1880]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FD67F8843F43F887" box="[750,841,1858,1880]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FCD1F8843FD6F887" box="[856,988,1858,1880]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, faunistic records); Nabozhenko 2005b: 39 (
<taxonomicName id="4C0435BDF8014923FF46F8A73DD8F8A8" authority="(Helopondrus)" baseAuthorityName="Helopondrus" box="[207,466,1889,1911]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FF46F8A73D20F8A8" box="[207,298,1889,1911]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FECDF8A73DC2F8A8" box="[324,456,1889,1911]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, bionomics);
<bibRefCitation id="EF9533CFF8014923FDF8F8A73F96F8A8" author="Nabozhenko" box="[625,924,1889,1911]" pageId="82" pageNumber="83" refString="Nabozhenko, M. V. &amp; Lobl, I. (2008) Tribe Helopini. In: Lobl, I. &amp; Smetana, A. (Eds.) Catalogue of Palearctic Coleoptera. Fol. 5. Tenebrionoidea. Stenstrup, Apollo books, pp. 241 - 257." type="journal article" year="2008">Nabozhenko &amp; Löbl 2008</bibRefCitation>
: 242 (
<taxonomicName id="4C0435BDF8014923FC66F8A738F8F8A8" authority="(Helopondrus)" baseAuthorityName="Helopondrus" box="[1007,1266,1889,1911]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FC66F8A73841F8A8" box="[1007,1099,1889,1911]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FBEDF8A738E2F8A8" box="[1124,1256,1889,1911]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8014923FF4EF8463E51F849" author="Abdurakhmanov" box="[199,603,1920,1942]" pageId="82" pageNumber="83" refString="Abdurakhmanov, G. M. &amp; Nabozhenko, M. V. (2009) Relict and endemic elements in the fauna of tenebrionid beetles (Coleoptera, Tenebrionidae) of the Big Caucasus. Yug Rossii: ekologia, razvitie, 2, 6 - 14. [in Russian]" type="journal article" year="2009">Abdurakhmanov &amp; Nabozhenko 2009</bibRefCitation>
: 12 (
<taxonomicName id="4C0435BDF8014923FD1CF8463F98F849" authority="(Helopondrus)" baseAuthorityName="Helopondrus" box="[661,914,1920,1942]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FD1CF8463EFBF849" box="[661,753,1920,1942]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FC8DF8463F82F849" box="[772,904,1920,1942]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, distribution, bionomics); Keskin &amp; Nabozhenko 2010: 24 (
<taxonomicName id="4C0435BDF8014923FEB7F8583E34F86B" authority="(Helopondrus)" baseAuthorityName="Helopondrus" box="[318,574,1950,1972]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FEB7F8583D90F86B" box="[318,410,1950,1972]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FE26F8583E39F86B" box="[431,563,1950,1972]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, faunistic records);
<bibRefCitation id="EF9533CFF8014923FC94F85838B3F86B" author="Abdurakhmanov" box="[797,1209,1950,1972]" pageId="82" pageNumber="83" refString="Abdurakhmanov, G. M. &amp; Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov &amp; Nabozhenko 2011</bibRefCitation>
: 142, 293 (
<taxonomicName id="4C0435BDF8014923FAC9F8583D57F80C" authority="(Helopondrus)" baseAuthorityName="Helopondrus" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FAC9F8583996F86B" box="[1344,1436,1950,1972]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
(
<emphasis id="B970922CF8014923FF46F87B3D59F80C" box="[207,339,1981,2003]" italics="true" pageId="82" pageNumber="83">Helopondrus</emphasis>
)
</taxonomicName>
, in key, in catalogue);
<bibRefCitation id="EF9533CFF8014923FDCBF87B3F54F80C" author="Nabozhenko" box="[578,862,1981,2003]" pageId="82" pageNumber="83" refString="Nabozhenko, M. V., Lebedeva, N. V., Nabozhenko, S. V. &amp; Lebedev, V. D. (2016) The taxocene of lichen-feeding darkling beetles (Coleoptera, Tenebrionidae: Helopini) in a forest-steppe ecotone. Entomological Review, 96, 101 - 113. http: // dx. doi. org / 10.1134 / S 0013873816010115" type="journal article" year="2016">Nabozhenko &amp; Kolov 2016</bibRefCitation>
: 110 (
<taxonomicName id="4C0435BDF8014923FC15F87B3FF2F80C" box="[924,1016,1981,2003]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8014923FC15F87B3FF2F80C" box="[924,1016,1981,2003]" italics="true" pageId="82" pageNumber="83">Nalassus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBB4E3EF8004922FF1EFF5F3E86FF13" blockId="83.[151,1436,152,204]" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FF1EFF5F3CF6FF71" box="[151,252,153,174]" italics="true" pageId="83" pageNumber="84">impressus</emphasis>
<bibRefCitation id="EF9533CFF8004922FE88FF5F3D8DFF71" author="Kuster" box="[257,391,152,174]" pageId="83" pageNumber="84" refString="Kuster, H. C. (1850) Die Kafer Europa's. Nach der Natur Beschrieben. Heft 21. Bauer und Raspe, Nurnberg. 4 + 100 p., 2 pls." type="book" year="1850">Küster, 1850</bibRefCitation>
: 21, 51 (
<taxonomicName id="4C0435BDF8004922FE56FF5E3E2DFF71" box="[479,551,152,174]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FE56FF5E3E2DFF71" box="[479,551,152,174]" italics="true" pageId="83" pageNumber="84">Helops</emphasis>
</taxonomicName>
, description); Baudi di Selve 1876b: 275 (
<taxonomicName id="4C0435BDF8004922FC51FF5E382AFF71" box="[984,1056,152,174]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FC51FF5E382AFF71" box="[984,1056,152,174]" italics="true" pageId="83" pageNumber="84">Helops</emphasis>
</taxonomicName>
, redescription); Baudi di Selve 1877: 93 (
<taxonomicName id="4C0435BDF8004922FF66FF703D3DFF13" box="[239,311,182,204]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FF66FF703D3DFF13" box="[239,311,182,204]" italics="true" pageId="83" pageNumber="84">Helops</emphasis>
</taxonomicName>
, faunistic records, redescription).
</paragraph>
<paragraph id="8BBB4E3EF8004922FF1EFF3238DEFE5B" blockId="83.[151,1436,244,388]" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FF1EFF323CEEFED5" box="[151,228,244,266]" italics="true" pageId="83" pageNumber="84">pulcher</emphasis>
<bibRefCitation id="EF9533CFF8004922FF64FF323D78FED5" author="Allard" box="[237,370,244,266]" pageId="83" pageNumber="84" refString="Allard, E. (1877) Revision des Helopides vrais. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 13 - 268." type="journal article" year="1877">Allard, 1877</bibRefCitation>
: 256 (
<taxonomicName id="4C0435BDF8004922FE3FFF323E12FED6" box="[438,536,244,265]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FE3FFF323E12FED6" box="[438,536,244,265]" italics="true" pageId="83" pageNumber="84">Stenomax</emphasis>
</taxonomicName>
, pulcher);
<bibRefCitation id="EF9533CFF8004922FD03FF323F5FFED5" author="Heyden" box="[650,853,244,266]" pageId="83" pageNumber="84" refString="Heyden, L. von, Reitter, E. &amp; Weise, J. (1883) Catalogus Coleopterorum Europae et Caucasi. Editio tertia. Libraria Nicolai, Berlin, 228 pp." type="book" year="1883">
Heyden
<emphasis id="B970922CF8004922FD6BFF333F1DFED5" box="[738,791,244,266]" italics="true" pageId="83" pageNumber="84">et al.</emphasis>
1883
</bibRefCitation>
: 136 (
<taxonomicName id="4C0435BDF8004922FC10FF323FF1FED6" box="[921,1019,244,265]" class="Insecta" family="Tenebrionidae" genus="Stenomax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FC10FF323FF1FED6" box="[921,1019,244,265]" italics="true" pageId="83" pageNumber="84">Stenomax</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8004922FB16FF323960FED5" author="Heyden" box="[1183,1386,244,266]" pageId="83" pageNumber="84" refString="Heyden, L. von, Reitter, E. &amp; Weise, J. (1891) Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. E. Reitter, Modling, 420 pp." type="book" year="1891">
Heyden
<emphasis id="B970922CF8004922FB7FFF333926FED5" box="[1270,1324,244,266]" italics="true" pageId="83" pageNumber="84">et al.</emphasis>
1891
</bibRefCitation>
: 252 (
<taxonomicName id="4C0435BDF8004922FF46FED43D9BFEF7" authority="(Stenomax)" baseAuthorityName="Stenomax" box="[207,401,274,296]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FF46FED43D1DFEF7" box="[207,279,274,296]" italics="true" pageId="83" pageNumber="84">Helops</emphasis>
(
<emphasis id="B970922CF8004922FEACFED53D8CFEF7" box="[293,390,275,296]" italics="true" pageId="83" pageNumber="84">Stenomax</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8004922FDA2FED43EBFFEF7" author="Seidlitz" box="[555,693,274,296]" pageId="83" pageNumber="84" refString="Seidlitz, G. (1896) Tenebrionidae. In: Kiesenwetter H. von, Seidlitz G. von, Naturgeschichte der Insecten Deutschlands. Erste Abteilung Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, pp. 609 - 800" type="book chapter" year="1896">Seidlitz 1896</bibRefCitation>
: 729 (
<taxonomicName id="4C0435BDF8004922FD7AFED43FBFFEF7" authority="(Stenomax)" baseAuthorityName="Stenomax" box="[755,949,274,296]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FD7AFED43F31FEF7" box="[755,827,274,296]" italics="true" pageId="83" pageNumber="84">Helops</emphasis>
(
<emphasis id="B970922CF8004922FCC0FED53FA1FEF7" box="[841,939,275,296]" italics="true" pageId="83" pageNumber="84">Stenomax</emphasis>
)
</taxonomicName>
, in key);
<bibRefCitation id="EF9533CFF8004922FB9AFED43892FEF7" author="Gebien" box="[1043,1176,274,296]" pageId="83" pageNumber="84" refString="Gebien, H. (1911) Pars. 28. Tenebrionidae. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus auspiciis et auxilio. Fol. 18. Junk, W., Berlin, pp. 355 - 585" type="journal article" year="1911">Gebien 1911</bibRefCitation>
: 551 (
<taxonomicName id="4C0435BDF8004922FB5CFED4399DFEF7" authority="(Stenomax)" baseAuthorityName="Stenomax" box="[1237,1431,274,296]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FB5CFED43917FEF7" box="[1237,1309,274,296]" italics="true" pageId="83" pageNumber="84">Helops</emphasis>
(
<emphasis id="B970922CF8004922FAA5FED53984FEF7" box="[1324,1422,275,296]" italics="true" pageId="83" pageNumber="84">Stenomax</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8004922FED0FEF73DD0FE98" author="Reitter" box="[345,474,305,327]" pageId="83" pageNumber="84" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44, 113 - 171. http: // dx. doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter 1922</bibRefCitation>
: 138 (
<taxonomicName id="4C0435BDF8004922FD93FEF73F1DFE98" authority="(Omaleis)" baseAuthorityName="Omaleis" box="[538,791,305,327]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FD93FEF73EA0FE98" box="[538,682,305,327]" italics="true" pageId="83" pageNumber="84">Cylindronotus</emphasis>
(
<emphasis id="B970922CF8004922FD30FEF73F07FE98" box="[697,781,305,327]" italics="true" pageId="83" pageNumber="84">Omaleis</emphasis>
)
</taxonomicName>
, in note);
<bibRefCitation id="EF9533CFF8004922FCF6FEF73806FE98" author="Winkler" box="[895,1036,305,327]" pageId="83" pageNumber="84" refString="Winkler, A. (1928 - 1929) Pars. 8, 9: D. Heteromera. In: Winkler, A. (Ed.) Catalogus Coleopterorum regionis Palaearctica. Verlag von Albert Winkler, Wien, pp. 881 - 1136." type="book chapter" year="1928">
Winkler
<date id="FFBA68FEF8004922FC51FEF73806FE98" box="[984,1036,305,327]" pageId="83" pageNumber="84" value="1928" valueMax="1929">1928</date>
</bibRefCitation>
–1929: 1018 (
<taxonomicName id="4C0435BDF8004922FB13FEF7399DFE98" authority="(Omaleis)" baseAuthorityName="Omaleis" box="[1178,1431,305,327]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FB13FEF73920FE98" box="[1178,1322,305,327]" italics="true" pageId="83" pageNumber="84">Cylindronotus</emphasis>
(
<emphasis id="B970922CF8004922FAB0FEF73987FE98" box="[1337,1421,305,327]" italics="true" pageId="83" pageNumber="84">Omaleis</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8004922FED2FE963DEBFEB9" author="Gebien" box="[347,481,336,358]" pageId="83" pageNumber="84" refString="Gebien, H. (1943) Katalog der Tenebrioniden. Teil. 3. Mitteilungen der Munchner Entomologischen Gesellschaft, 33, 399 - 430 (778 - 809)." type="journal article" year="1943">Gebien 1943</bibRefCitation>
: 428 (807) (
<taxonomicName id="4C0435BDF8004922FDEBFE963F6AFEB9" authority="(Omaleis)" baseAuthorityName="Omaleis" box="[610,864,336,358]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FDEBFE963EF8FEB9" box="[610,754,336,358]" italics="true" pageId="83" pageNumber="84">Cylindronotus</emphasis>
(
<emphasis id="B970922CF8004922FC8BFE963F5CFEB9" box="[770,854,336,358]" italics="true" pageId="83" pageNumber="84">Omaleis</emphasis>
)
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8004922FB88FE963972FEB9" author="Medvedev" box="[1025,1400,336,358]" pageId="83" pageNumber="84" refString="Medvedev, G. S. &amp; Abdurakhmanov, G. M. (1988) Check-list of beetles of the family Tenebrionidae of South West of Palaearctic. In: Abdurakhmanov, G. M. Eastern Caucasus through the eyes of an entomologist. Dagestanian book Publishing House, Makhachkala, pp. 71 - 99. [in Russian]" type="book chapter" year="1988">Medvedev &amp; Abdurakhmanov 1988</bibRefCitation>
: 98 (
<taxonomicName id="4C0435BDF8004922FF46FEA83D55FE5B" box="[207,351,366,388]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FF46FEA83D55FE5B" box="[207,351,366,388]" italics="true" pageId="83" pageNumber="84">Cylindronotus</emphasis>
</taxonomicName>
, in catalogue);
<bibRefCitation id="EF9533CFF8004922FE75FEA83F64FE5B" author="Abdurakhmanov" box="[508,878,366,388]" pageId="83" pageNumber="84" refString="Abdurakhmanov, G. M. &amp; Medvedev, G. S. (1994) Catalogue of darkling beetles of the Caucasus. Daghestan State Pedagogical Institute, Makhachkala. 212 pp. [in Russian]" type="book" year="1994">Abdurakhmanov &amp; Medvedev 1994</bibRefCitation>
: 194 (
<taxonomicName id="4C0435BDF8004922FC24FEA83837FE5B" box="[941,1085,366,388]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FC24FEA83837FE5B" box="[941,1085,366,388]" italics="true" pageId="83" pageNumber="84">Cylindronotus</emphasis>
</taxonomicName>
, in catalogue).
</paragraph>
</subSubSection>
<subSubSection id="C31E1DB5F8004922FF1EFE773E20FAF5" pageId="83" pageNumber="84" type="materials_examined">
<paragraph id="8BBB4E3EF8004922FF1EFE773D87FD19" blockId="83.[151,1437,433,1322]" pageId="83" pageNumber="84">
<materialsCitation id="3B6C4463F8004922FF1EFE773E64FE15" ID-GBIF-Occurrence="1413110243" box="[151,622,433,458]" country="Russia" location="Krasnodar krai" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Krasnodar krai">
<emphasis id="B970922CF8004922FF1EFE773D6EFE17" bold="true" box="[151,356,433,458]" pageId="83" pageNumber="84">
Material.
<collectingCountry id="F3130EAEF8004922FE87FE773D6EFE17" box="[270,356,433,458]" name="Russia" pageId="83" pageNumber="84">RUSSIA</collectingCountry>
</emphasis>
:
<collectingRegion id="49C080DCF8004922FEFDFE773E2AFE15" box="[372,544,433,458]" country="Russia" name="Krasnodar" pageId="83" pageNumber="84">Krasnodar krai</collectingRegion>
: Sochi
</materialsCitation>
;
<materialsCitation id="3B6C4463F8004922FDF3FE773D03FE31" ID-GBIF-Occurrence="1413110231" country="Russia" location="Kabardino-Balkaria" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Kabardino-Balkaria">
<collectingRegion id="49C080DCF8004922FDF3FE773F52FE15" box="[634,856,433,458]" country="Russia" name="Kabardino-Balkariya" pageId="83" pageNumber="84">Kabardino-Balkaria</collectingRegion>
(invasion:
<bibRefCitation id="EF9533CFF8004922FC51FE77393BFE15" author="Nabozhenko" box="[984,1329,433,458]" pageId="83" pageNumber="84" refString="Nabozhenko, M. V., Lebedeva, N. V., Nabozhenko, S. V. &amp; Lebedev, V. D. (2016) The taxocene of lichen-feeding darkling beetles (Coleoptera, Tenebrionidae: Helopini) in a forest-steppe ecotone. Entomological Review, 96, 101 - 113. http: // dx. doi. org / 10.1134 / S 0013873816010115" type="journal article" year="2016">Nabozhenko &amp; Kolov (2016))</bibRefCitation>
: Nalchik, Nart-Bala
</materialsCitation>
.
<materialsCitation id="3B6C4463F8004922FE9FFE133F81FD85" ID-GBIF-Occurrence="1413110232" country="Georgia" location="Shaori Lake" municipality="South Ossetia" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Abkhazia">
<collectingCountry id="F3130EAEF8004922FE9FFE133D8DFE33" box="[278,391,469,494]" name="Georgia" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FE9FFE133D8DFE33" bold="true" box="[278,391,469,494]" pageId="83" pageNumber="84">GEORGIA</emphasis>
</collectingCountry>
: (including
<collectingMunicipality id="6BDFD444F8004922FD9CFE133EB9FE31" box="[533,691,469,494]" pageId="83" pageNumber="84">South Ossetia</collectingMunicipality>
and
<collectingRegion id="49C080DCF8004922FD67FE133F55FE31" box="[750,863,469,494]" country="Georgia" name="Abkhazia" pageId="83" pageNumber="84">Abkhazia</collectingRegion>
): Adigeni, Atskuri, Babaneuri, Bakuriani, Betania Monastery, Bobokvati, Borjomi, Chiatura, Chikhareshi, Djo-djo, Eldari, Kazbegi, Kechkobi, Kiketi, Kintrishi Reserve, Kobuleti, Kojori, Kutaisi, Lagodekhi, Lentekhi, Manglisi, Martkopi, Mtskheta, Natanebi, Ochamchira,
<location id="8EDB18E5F8004922FF1EFD873D2FFD85" LSID="urn:lsid:plazi:treatment:03ADFF28F8014922FF1EFA403E20FAF5:8EDB18E5F8004922FF1EFD873D2FFD85" box="[151,293,577,602]" country="Georgia" municipality="South Ossetia" name="Shaori Lake" pageId="83" pageNumber="84" stateProvince="Abkhazia">Shaori Lake</location>
, Sioni (
<collectingRegion id="49C080DCF8004922FE09FD873EB8FD85" box="[384,690,577,602]" country="Georgia" name="Mtskheta-Mtianeti" pageId="83" pageNumber="84">
Mtskheta-Mtianeti
<collectingRegion id="49C080DCF8004922FDD4FD873EB8FD85" box="[605,690,577,602]" country="Iraq" pageId="83" pageNumber="84">Region</collectingRegion>
</collectingRegion>
), Sukhum, Surami
</materialsCitation>
,
<materialsCitation id="3B6C4463F8004922FC11FD873E0FFDA1" ID-GBIF-Occurrence="1413110237" country="Georgia" location="Tbilisi" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Tbilisi">
<collectingRegion id="49C080DCF8004922FC11FD873FEEFD85" box="[920,996,577,602]" country="Georgia" name="Tbilisi" pageId="83" pageNumber="84">Tbilisi</collectingRegion>
, Tianeti, Tkibuli, Tsagveri, Tskhinval, Tskhneti, Utsera, Zemo-Avchala
</materialsCitation>
.
<materialsCitation id="3B6C4463F8004922FD86FDA338A9FDA1" ID-GBIF-Occurrence="1413110240" box="[527,1187,613,638]" country="Armenia" location="Tavush Province" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Tavush Province">
<collectingCountry id="F3130EAEF8004922FD86FDA33E8AFDA3" box="[527,640,613,638]" name="Armenia" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FD86FDA33E8AFDA3" bold="true" box="[527,640,613,638]" pageId="83" pageNumber="84">ARMENIA</emphasis>
</collectingCountry>
: Norashen (
<collectingRegion id="49C080DCF8004922FC81FDA33FC3FDA1" box="[776,969,613,638]" country="Armenia" name="Tavush" pageId="83" pageNumber="84">Tavush Province</collectingRegion>
), Kozman, Gyumri
</materialsCitation>
.
<materialsCitation id="3B6C4463F8004922FB24FDA33EF5FD7E" ID-GBIF-Occurrence="1413110245" country="Azerbaijan" location="Mount Bazarduzu" pageId="83" pageNumber="84" specimenCount="1">
<collectingCountry id="F3130EAEF8004922FB24FDA33948FDA3" box="[1197,1346,613,638]" name="Azerbaijan" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FB24FDA33948FDA3" bold="true" box="[1197,1346,613,638]" pageId="83" pageNumber="84">AZERBAIJAN</emphasis>
</collectingCountry>
:
<location id="8EDB18E5F8004922FAD8FDA03D05FD7D" LSID="urn:lsid:plazi:treatment:03ADFF28F8014922FF1EFA403E20FAF5:8EDB18E5F8004922FAD8FDA03D05FD7D" country="Azerbaijan" name="Mount Bazarduzu" pageId="83" pageNumber="84">Mount Bazardüzü</location>
(Bumga Canyon), Zakatataly State Reserve
</materialsCitation>
,
<materialsCitation id="3B6C4463F8004922FC82FD4F380FFD7E" ID-GBIF-Occurrence="1413110235" box="[779,1029,649,674]" country="Azerbaijan" location="Ismailli" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Ismailli">
<collectingRegion id="49C080DCF8004922FC82FD4F3F6BFD7D" box="[779,865,649,674]" country="Azerbaijan" name="Ismayilli" pageId="83" pageNumber="84">Ismailli</collectingRegion>
State Reserve
</materialsCitation>
.
<materialsCitation id="3B6C4463F8004922FB86FD4F3D87FD19" ID-GBIF-Occurrence="1413110252" country="Turkey" location="Artvin Province" pageId="83" pageNumber="84" specimenCount="1" stateProvince="Artvin Province">
<collectingCountry id="F3130EAEF8004922FB86FD4F387AFD7F" box="[1039,1136,649,674]" name="Turkey" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FB86FD4F387AFD7F" bold="true" box="[1039,1136,649,674]" pageId="83" pageNumber="84">TURKEY</emphasis>
</collectingCountry>
:
<collectingRegion id="49C080DCF8004922FBF6FD4F393DFD7D" box="[1151,1335,649,674]" country="Turkey" name="Artvin" pageId="83" pageNumber="84">Artvin Province</collectingRegion>
: Karagöl Sahara National Park.
</materialsCitation>
</paragraph>
<paragraph id="8BBB4E3EF8004922FF4EFD173D62FB8D" blockId="83.[151,1437,433,1322]" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FF4EFD173D1BFD35" bold="true" box="[199,273,721,746]" pageId="83" pageNumber="84">Notes.</emphasis>
This species was included in the subgenus
<taxonomicName id="4C0435BDF8004922FD73FD173F86FD35" box="[762,908,721,746]" class="Insecta" family="Tenebrionidae" genus="Helopondrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FD73FD173F86FD35" box="[762,908,721,746]" italics="true" pageId="83" pageNumber="84">Helopondrus</emphasis>
</taxonomicName>
of the genus
<taxonomicName id="4C0435BDF8004922FBADFD173883FD35" box="[1060,1161,721,746]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FBADFD173883FD35" box="[1060,1161,721,746]" italics="true" pageId="83" pageNumber="84">Nalassus</emphasis>
</taxonomicName>
based on the absence of teeth on inner side of male tibiae, absence of tubercles or granules on elytra, structure of epipleura and dorsoventrally flattened parameres (
<bibRefCitation id="EF9533CFF8004922FE71FCDF3EC5FCED" author="Nabozhenko" box="[504,719,793,818]" pageId="83" pageNumber="84" refString="Nabozhenko, M. V. (2001) On the classification of the Tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomological Review, 81, 909 - 942." type="journal article" year="2001">Nabozhenko 2001</bibRefCitation>
). Later we studied additional material and found granules on inner side of tibia of males from some populations. This character is variable even among specimens of one population. Genitalia and terminalia of male and genital tubes of female of the species also belong to the ‘cylindrinotoid’
<typeStatus id="54BFF09CF8004922FED8FC403D8BFC41" box="[337,385,902,926]" pageId="83" pageNumber="84">type</typeStatus>
(Nabozhenko 2005). This species has a keel-shaped 8th elytral interval, which is connected with epipleural carina and 1st elytral interval apically and, therefore, is similar in this respect to species of the
<taxonomicName id="4C0435BDF8004922FF1EFC083D22FC39" box="[151,296,973,998]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="recticollis">
<emphasis id="B970922CF8004922FF1EFC083D22FC39" box="[151,296,973,998]" italics="true" pageId="83" pageNumber="84">O. recticollis</emphasis>
</taxonomicName>
species-group. Two other characters, namely presence of small reduced wings and epipleura not reaching elytral apex, distinguish this species from the
<taxonomicName id="4C0435BDF8004922FC8CFC343F9DFBD5" box="[773,919,1009,1034]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="recticollis">
<emphasis id="B970922CF8004922FC8CFC343F9DFBD5" box="[773,919,1009,1034]" italics="true" pageId="83" pageNumber="84">O. recticollis</emphasis>
</taxonomicName>
species-group. Based on these considerations we assign this species to the genus
<taxonomicName id="4C0435BDF8004922FDCCFBD33EC9FBF1" box="[581,707,1045,1070]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B970922CF8004922FDCCFBD33EC9FBF1" box="[581,707,1045,1070]" italics="true" pageId="83" pageNumber="84">Odocnemis</emphasis>
</taxonomicName>
and, therefore, a new combination is established:
<taxonomicName id="4C0435BDF8004922FA97FBD33CFEFB8D" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="gloriosa" status="comb. nov.">
<emphasis id="B970922CF8004922FA97FBD33CFEFB8D" italics="true" pageId="83" pageNumber="84">Odocnemis gloriosa</emphasis>
</taxonomicName>
,
<emphasis id="B970922CF8004922FE8BFBFF3D62FB8D" bold="true" box="[258,360,1081,1106]" pageId="83" pageNumber="84">
<taxonomicNameLabel id="A2432F57F8004922FE8BFBFF3D62FB8D" box="[258,360,1081,1106]" pageId="83" pageNumber="84">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BBB4E3EF8004922FF4EFB9B3D63FB3D" blockId="83.[151,1437,433,1322]" pageId="83" pageNumber="84">
Material, synonymy, figures of genitalia and habitus of this species were published in our previous works (
<bibRefCitation id="EF9533CFF8004922FF17FB473D7BFB45" author="Nabozhenko" box="[158,369,1153,1178]" pageId="83" pageNumber="84" refString="Nabozhenko, M. V. (2001) On the classification of the Tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomological Review, 81, 909 - 942." type="journal article" year="2001">Nabozhenko 2001</bibRefCitation>
; Keskin &amp; Nabozhenko 2010;
<bibRefCitation id="EF9533CFF8004922FD5DFB473883FB45" author="Abdurakhmanov" box="[724,1161,1153,1178]" pageId="83" pageNumber="84" refString="Abdurakhmanov, G. M. &amp; Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov &amp; Nabozhenko 2011</bibRefCitation>
). Here only localities of this widespread species are given from studied specimens deposited in
<collectionCode id="ED15D6FBF8004922FC57FB603805FB62" box="[990,1039,1190,1213]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="83" pageNumber="84">ZIN</collectionCode>
, ZM
<collectionCode id="ED15D6FBF8004922FBDDFB60389EFB61" box="[1108,1172,1190,1214]" country="USA" httpUri="http://grbio.org/cool/53w1-8r3k" name="Michigan State University Museum" pageId="83" pageNumber="84">MSU</collectionCode>
,
<collectionCode id="ED15D6FBF8004922FB2AFB6038F8FB61" box="[1187,1266,1190,1214]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="83" pageNumber="84">MPSU</collectionCode>
,
<collectionCode id="ED15D6FBF8004922FA8BFB603950FB62" box="[1282,1370,1190,1213]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="83" pageNumber="84">HNHM</collectionCode>
,
<collectionCode id="ED15D6FBF8004922FAE3FB603992FB61" box="[1386,1432,1190,1214]" pageId="83" pageNumber="84">IZG</collectionCode>
,
<collectionCode id="ED15D6FBF8004922FF1EFB0C3CEFFB3E" box="[151,229,1226,1249]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="83" pageNumber="84">ZDEU</collectionCode>
,
<collectionCode id="ED15D6FBF8004922FF66FB0C3D25FB3D" box="[239,303,1226,1250]" pageId="83" pageNumber="84">ONU</collectionCode>
, CN.
</paragraph>
<paragraph id="8BBB4E3EF8004922FF4EFB2B3E20FAF5" blockId="83.[151,1437,433,1322]" pageId="83" pageNumber="84">
<emphasis id="B970922CF8004922FF4EFB2B3D49FAD9" bold="true" box="[199,323,1261,1286]" pageId="83" pageNumber="84">Bionomics</emphasis>
. The species inhabits forest landscapes; it feeds on lichens on
<taxonomicName id="4C0435BDF8004922FB8BFB28386AFADA" box="[1026,1120,1262,1285]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="B970922CF8004922FB8BFB28386AFADA" box="[1026,1120,1262,1285]" italics="true" pageId="83" pageNumber="84">Quercus</emphasis>
</taxonomicName>
spp.,
<taxonomicName id="4C0435BDF8004922FB2AFB2B38F0FAD9" box="[1187,1274,1261,1286]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="B970922CF8004922FB2AFB2B38F0FAD9" box="[1187,1274,1261,1286]" italics="true" pageId="83" pageNumber="84">Juglans</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0435BDF8004922FA80FB2B394DFAD9" box="[1289,1351,1261,1286]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="B970922CF8004922FA80FB2B394DFAD9" box="[1289,1351,1261,1286]" italics="true" pageId="83" pageNumber="84">Abies</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0435BDF8004922FADCFB28399FFADA" box="[1365,1429,1262,1285]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="B970922CF8004922FADCFB28399FFADA" box="[1365,1429,1262,1285]" italics="true" pageId="83" pageNumber="84">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0435BDF8004922FF1EFAD43CD7FAF6" box="[151,221,1298,1321]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="B970922CF8004922FF1EFAD43CD7FAF6" box="[151,221,1298,1321]" italics="true" pageId="83" pageNumber="84">Fagus</emphasis>
</taxonomicName>
, active at night (20:00–2:00).
</paragraph>
</subSubSection>
</treatment>
</document>
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